identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D28ED20EE61F593CA35934109B63BC81.text	D28ED20EE61F593CA35934109B63BC81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luffy Hu & Solodovnikov 2026	<div><p>Genus Luffy gen. nov.</p><p>Figs 1, 2, 3, 5 A, 6 A</p><p>Type species:</p><p>Luffy schillhammeri sp. nov.</p><p>Included species:</p><p>Luffy nika sp. nov., L. schillhammeri sp. nov.</p><p>Diagnosis.</p><p>The genus can be diagnosed by narrow, not expanded protarsi in both sexes (Fig. 2 C, D) and all antennomeres longer than wide (Figs 1 A, 1 B, 3 A, 3 B), both traits being unique synapomorphies within Staphylinina . In addition to these characters, the new genus can be further distinguished by the combination of the presence of semimembranous extension on the labrum (Fig. 5 A) and the presence of DRT 1, DRT 2, and BET on the left mandible (Fig. 6 A).</p><p>Description.</p><p>Medium-sized Staphylinina (TL: 16.4–20.5 mm; FL: 8.4–10.3 mm).</p><p>Head. Head slightly transverse (HL / HW = 0.8). Tempora not expanded in both sexes. Eyes large and convex, slightly shorter than tempora (EL / TL: 0.7–0.8). Head densely punctate on both dorsal and ventral faces, punctures rounded to irregular. Labrum moderately large, trapezoidal, with distinct semimembranous margin (Fig. 5 A); anterior portion with two macrosetae. Gular sutures not separated. Antennae very long, all antennomeres longer than wide, with dense pubescence. Mentum with two pairs of macrosetae located in the anterior portion; two pairs of thinner setae located at the anterior margin. Labial palps long; last segment setose, longer than the first two segments (Fig. 2 A). Maxillary palps long; first segment short; second segment as long as the third and last segments combined; last segment with very minute setae, almost unobservable even at 50 × magnification (Fig. 2 B). Mandibles very long and slender, both left and right mandibles with DRT 1, DRT 2 and BET, DRT 1, and DRT 2 forming a larger bidentate tooth; BET close to the base of mandible on both sides of mandible, located above the prostheca (Fig. 6 A). Prostheca single-lobed. Head chaetotaxy: FCP present; TM close to posterior margin of eyes; 1 POP present; PFM missing.</p><p>Thorax. Pronotum pyriform, longer than wide (PL / PW: 1.32–1.41), densely punctate; impunctate midline almost invisible. Anterior angles rounded. Superior and inferior marginal lines join each other approximately one-third from the anterior margins; pronotal hypomera asetose. Pronotum chaetotaxy: PAMM 1 present, PALM absent, PMM present, located at approximately one-third of pronotal length from the anterior margin. Mesoventrite ridge absent. Elytra and hind wings fully developed, elytra with a distinct transverse curved stripe of pale setae in both species. Elytral chaetotaxy: EDM 1, 2, 3 present; PsP 2 absent. Protibiae slender, without modification, with 2–3 spines on lateral face; meso-and metatibiae straight, not arcuate, mesotibiae with numerous spines; metatibiae with 2–3 spines on lateral face. Protarsi not expanded; underside densely setose, but setae unmodified (Fig. 2 C, D).</p><p>Abdomen. All segments densely covered with long setae. Tergites III – V with distinct accessory ridges, with paler setal patches besides the outer margin of accessory ridges; tergites VI and VII with paler setae in anterior margin. Tergite VII with a distinct pale apical seam of palisade setae. Abdominal segment VII longer than other segments. Abdominal segment chaetotaxy: MTM and PTM present; segments VII and VIII with numerous macrosetae.</p><p>Male. Sternite VIII with a moderately deep, subtriangular emargination at midpoint of posterior margin. Abdominal segment IX clavate (Fig. 2 G). Tergite X triangular (Fig. 2 E). Aedeagus strongly asymmetrical; paramere slightly bent to the left side in parameral view; peg setae present in the anterior portion but not strongly pigmented.</p><p>Female. The only known female of L. schillhammeri sp. nov. is slightly larger than males of both species. Abdominal segment IX clavate, slightly wider than in male (Fig. 2 H). Tergite X triangular (Fig. 2 F), fully sclerotized, without a membranous part.</p><p>Etymology.</p><p>The generic name Luffy is the given name of the main character of the famous manga “ One Piece ”, Monkey D. Luffy, who has the ability to extend his body parts. It refers to the elongated antennae, palps, and mandibles of the new genus. The gender is masculine.</p></div>	https://treatment.plazi.org/id/D28ED20EE61F593CA35934109B63BC81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hu, Fang-Shuo;Solodovnikov, Alexey	Hu, Fang-Shuo, Solodovnikov, Alexey (2026): Luffy gen. nov., a new genus of Staphylinina (Coleoptera, Staphylinidae, Staphylininae), remarkable for understanding the Eucibdelus lineage. ZooKeys 1281: 247-264, DOI: 10.3897/zookeys.1281.198593
961DFFD617395FFF802402D6F7C59CB8.text	961DFFD617395FFF802402D6F7C59CB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luffy nika Hu & Solodovnikov 2026	<div><p>Luffy nika sp. nov.</p><p>Fig. 3</p><p>Material examined.</p><p>Holotype: • male, Laos, NW., Kiolom, 1050 m alt., 36 km NW from L N. Louang Namtha, 11. V. 2004, Y. Shibata leg. / NSMT -I-C Shibata C No. SBC 006913 (NSMT) .</p><p>Paratype: • 1 male, same data as the holotype, but NSMT -I-C Shibata C No. SBC 006914 (NSMT) .</p><p>Diagnosis and comparison.</p><p>Luffy nika sp. nov. is very similar to L. schillhammeri sp. nov. It can be distinguished from L. schillhammeri sp. nov. by differences in pronotal shape: L. nika sp. nov. has a slightly longer pronotum (PL / PW = 1.41), distinctly narrowing basad from the anterior third of its length (Fig. 3 A), whereas L. schillhammeri sp. nov. has a slightly shorter and more gradually narrowing pronotum (PL / PW = 1.32) (Fig. 1 A). More reliable identification of the two species requires examination of the aedeagus: L. nika sp. nov. has a deeper apical emargination of the median lobe (Fig. 3 D), a thinner paramere (Fig. 3 D, E) with a rounded apex (Fig. 3 E), and a more symmetrical transverse area of its articulation to the median lobe (Fig. 3 E); whereas L. schillhammeri sp. nov. has a shallower apical emargination of the median lobe (Fig. 1 D), and a wider paramere (Fig. 1 D, E) with a subtriangular apex (Fig. 1 E), and a distinct asymmetrical area of its articulation to the median lobe (Fig. 1 E).</p><p>Description.</p><p>Measurements (given in mm, average provided in brackets): Male, WL: 16.39–17.36 (16.88); FL: 8.64–8.95 (8.8); EL: 0.8 (0.8); EW: 0.4 (0.4); HL: 2.5 (average 2.5); HW: 2.7 (2.7); TL: 1 (1); PL: 3.1 (3.1); PW: 2.2 (2.2).</p><p>Habitus (Fig. 3 A, B). Body black, without metallic lustre. Head, pronotum and elytra densely punctate with long black setae; punctures mostly rounded, more irregular on vertex area.</p><p>Head slightly transverse, without impunctate midline. Eyes large and protruding. Antennae rufotestaceous to black, first five antennomeres darker than others; all antennomeres distinctly longer than wide. Head with patches of white setae on tempora, with sparse white setae on disc.</p><p>Pronotum pyriform, longer than wide, distinctly narrowing basad from one-third of its length; densely punctate, with rounded punctures; with small patches of white setae at midlength; without impunctate midline. Elytra and scutellum densely covered with black setae; elytra with a distinct, transverse, curved stripe of white setae. Legs covered with dense white setae.</p><p>Abdomen tergites black; tergites III – V each with accessory basal ridges. Tergites III – VII densely covered with long, black setae; tergites III – V each with paired patches of white setae antero-laterally, extending to paratergites; tergites VI – VII with transverse stripe of white setae across anterior margin; tergite VIII with mixture of white and black setae. Sternites III – VIII black; sternites III – IV densely covered with white setae; sternite V with white setae on anterior and posterior margins, with brown setae in the middle portion; sternites VI – VIII with white setae in the anterior portion and brown setae in the posterior portion.</p><p>Male sternite VIII with a moderately deep, subtriangular emargination at midpoint of posterior margin. Aedeagus slender, asymmetrical; median lobe parallel-sided in basal portion in parameral view, apex with a shallow emargination (Fig. 3 C – F). Paramere thin, slightly twisted to the left, apex rounded; underside with rather sparse and not strongly pigmented peg setae forming an apical cluster, with a few additional ones sparse in anterior portion (Fig. 3 C – F). Underside of paramere with a distinct, narrow, transverse symmetrical area of its articulation to the median lobe (Fig. 3 F).</p><p>Female unknown.</p><p>Etymology.</p><p>The specific epithet nika is derived from the Devil Fruit of the main character, Luffy, in the famous manga One Piece, in reference to the genus name. It also alludes to the pale hairs of the new species, reminiscent of Luffy’s “ Nika ” form, in which his clothing and hair turn white. It is a noun in apposition.</p><p>Ecology.</p><p>Nothing is known about the habitat or collecting circumstances for L. nika sp. nov.</p><p>Distribution.</p><p>The species is known from the type locality in Louang Namtha in northwestern Laos.</p><p>Remarks.</p><p>The terminal segment of the maxillary and labial palps in both specimens of L. nika sp. nov. bears a protruding membranous structure, although it is less pronounced in one specimen. We are uncertain whether this represents a species-specific character or an artefact resulting from specimen preparation. Therefore, this character was not included in the diagnosis, pending examination of additional material in the future.</p></div>	https://treatment.plazi.org/id/961DFFD617395FFF802402D6F7C59CB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hu, Fang-Shuo;Solodovnikov, Alexey	Hu, Fang-Shuo, Solodovnikov, Alexey (2026): Luffy gen. nov., a new genus of Staphylinina (Coleoptera, Staphylinidae, Staphylininae), remarkable for understanding the Eucibdelus lineage. ZooKeys 1281: 247-264, DOI: 10.3897/zookeys.1281.198593
7FEA42F8FB6E5FE1A137058F2CAF2EA8.text	7FEA42F8FB6E5FE1A137058F2CAF2EA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luffy schillhammeri Hu & Solodovnikov 2026	<div><p>Luffy schillhammeri sp. nov.</p><p>Figs 1, 2</p><p>Material examined.</p><p>Holotype: • male, China: Yunnan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.1951&amp;materialsCitation.latitude=21.9266" title="Search Plazi for locations around (long 101.1951/lat 21.9266)">CCCC Manzhang</a> (曼掌), 27. III. 2018. leg. Y.-T. Wang / Mengla County, 21.9266, 101.1951, alt. 650 m, FIT, primary and secondary broadleaf forests (TARI) .</p><p>Paratype: • 1 female, China: Yunnan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.6312&amp;materialsCitation.latitude=21.4066" title="Search Plazi for locations around (long 101.6312/lat 21.4066)">CCCC Manfeng</a> (曼粉), 12. V. 2016. leg. Y.-T. Wang / Mengla County, 21.4066, 101.6312, alt. 750 m, light trap, primary and secondary broadleaf forests / Project: Revision of Ocypus group DNA Voucher specimen: FS 363 (TARI) .</p><p>Diagnosis and comparison.</p><p>Luffy schillhammeri sp. nov. is very similar to L. nika sp. nov. It can be distinguished from L. nika sp. nov. by differences in pronotal shape: L. schillhammeri sp. nov. has a slightly shorter pronotum (PL / PW = 1.32), gradually narrowing basad from the anterior third of its length (Fig. 1 A), whereas L. nika sp. nov. has a slightly longer pronotum (PL / PW = 1.41), more distinctly narrowing basad (Fig. 3 A). More reliable identification of these species requires examination of the aedeagus: L. schillhammeri sp. nov. has a shallower emargination of the median lobe (Fig. 1 D), a wider paramere (Fig. 1 D, E), a subtriangular parameral apex (Fig. 1 D, E), and (on the underside) a small asymmetrical area of its articulation to the median lobe (Fig. 1 E); whereas L. nika sp. nov. has a deeper apical emargination of the median lobe (Fig. 3 D), a thinner paramere (Fig. 3 D, E), a rounded parameral apex (Fig. 3 E), and (on the underside) a transverse, more symmetrical area of its articulation to the median lobe (Fig. 3 E).</p><p>Description.</p><p>Measurements (given in mm): Male, WL: 16.79; FL: 8.4; EL: 0.7; EW: 0.3; HL: 2.4; HW: 2.6; TL: 1; PL: 2.9; PW: 2.2. Female, WL: 20.48; FL: 10.25; EL: 0.8; EW: 0.4; HL: 2.7; HW: 2.9; TL: 1.1; PL: 3.3; PW: 2.5.</p><p>Habitus (Figs 1, 2). Body black, without metallic lustre. Head, pronotum and elytra densely punctate with long black setae, punctures mostly rounded, more irregular on vertex area.</p><p>Head slightly transverse, without impunctate midline. Eyes large and protruding. Antennae rufotestaceous to black, first five antennomeres darker than others; all antennomeres distinctly longer than wide. Head with patches of white setae on tempora, with sparse white setae on disc.</p><p>Pronotum pyriform, longer than wide, gradually narrowing basad from one-third of pronotal length; densely punctate, with rounded punctures; with small patches of white setae at midlength of lateral margin; without impunctate midline. Elytra and scutellum densely covered with black setae; elytra with a distinct, transverse, curved stripe of white setae. Legs covered with dense white setae.</p><p>Abdomen tergites black; tergites III – V each with accessory basal ridges. Tergites III – VII densely covered with long, black setae; tergites III – V each with paired patches of white setae antero-laterally, extending to paratergites; tergites VI – VII with transverse stripe of white setae across anterior margin; tergite VIII with mixture of white and black setae. Sternites III – VIII black; sternites III – IV densely covered with white setae; sternite V with white setae on anterior and posterior margins, with brown setae in the middle portion; sternites VI – VIII with white setae on anterior portion and brown setae in the posterior portion.</p><p>Male sternite VIII with a moderately deep, subtriangular emargination at midpoint of posterior margin. Aedeagus slender, asymmetrical; median lobe parallel-sided in basal portion in parameral view, apex with a shallow emargination (Fig. 1 C – F). Paramere moderately wide, slightly twisted to the left, apex subtriangular; underside with rather sparse and not strongly pigmented peg setae forming an apical cluster, with a few additional ones sparse in anterior portion (Fig. 1 C – F). Underside of paramere with a distinct asymmetrical area of articulation to the median lobe (Fig. 1 E).</p><p>Female slightly larger and more robust than male, but otherwise no marked sexual dimorphism.</p><p>Etymology.</p><p>The species is named in honour of Harald Schillhammer (Vienna, Austria), in recognition of his decades of contributions to the taxonomy of Staphylininae .</p><p>Distribution.</p><p>The species is known from the type locality in Mengla County, southwestern Yunnan Province, China.</p><p>Ecology.</p><p>Luffy schillhammeri sp. nov. was collected in low-altitude primary and secondary broadleaf forests (Fig. 4) by flight-intercept and light traps.</p></div>	https://treatment.plazi.org/id/7FEA42F8FB6E5FE1A137058F2CAF2EA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hu, Fang-Shuo;Solodovnikov, Alexey	Hu, Fang-Shuo, Solodovnikov, Alexey (2026): Luffy gen. nov., a new genus of Staphylinina (Coleoptera, Staphylinidae, Staphylininae), remarkable for understanding the Eucibdelus lineage. ZooKeys 1281: 247-264, DOI: 10.3897/zookeys.1281.198593
