taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
D28ED20EE61F593CA35934109B63BC81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670283	https://doi.org/10.3897/zookeys.1281.198593.figure1	Figure 1. Luffy schillhammeri sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	Figure 1. Luffy schillhammeri sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
D28ED20EE61F593CA35934109B63BC81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670284	https://doi.org/10.3897/zookeys.1281.198593.figure2	Figure 2. Luffy schillhammeri sp. nov., labium, maxillary palps, protarsi, abdominal segments IX – X. A. Labium; B. Maxillary palps; C. Protarsi, dorsal view; D. Protarsi, ventral view; E. Tergite X, male; F. Tergite X, female; G. Abdominal segment IX, male; H. Abdominal segment IX and gonocoxite, female. Scale bars: 1 mm.	Figure 2. Luffy schillhammeri sp. nov., labium, maxillary palps, protarsi, abdominal segments IX – X. A. Labium; B. Maxillary palps; C. Protarsi, dorsal view; D. Protarsi, ventral view; E. Tergite X, male; F. Tergite X, female; G. Abdominal segment IX, male; H. Abdominal segment IX and gonocoxite, female. Scale bars: 1 mm.	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
D28ED20EE61F593CA35934109B63BC81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670285	https://doi.org/10.3897/zookeys.1281.198593.figure3	Figure 3. Luffy nika sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	Figure 3. Luffy nika sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
D28ED20EE61F593CA35934109B63BC81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670287	https://doi.org/10.3897/zookeys.1281.198593.figure5	Figure 5. Diversity of labrum morphology of Ocypus - group and Staphylinus. A. Luffy; B. Agelosus; C. Cyanocypus; D. Protocypus; E. Ocypus; F. Eucibdelus; G. Guillaumius; H. Menoedius; I. Parapalaestrinus; J. Paraphytolinus; K. Rhynchocheilus; L. Rhyncocheilus; M. Trichocosmetes; N. Staphylinus.	Figure 5. Diversity of labrum morphology of Ocypus - group and Staphylinus. A. Luffy; B. Agelosus; C. Cyanocypus; D. Protocypus; E. Ocypus; F. Eucibdelus; G. Guillaumius; H. Menoedius; I. Parapalaestrinus; J. Paraphytolinus; K. Rhynchocheilus; L. Rhyncocheilus; M. Trichocosmetes; N. Staphylinus.	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
D28ED20EE61F593CA35934109B63BC81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670288	https://doi.org/10.3897/zookeys.1281.198593.figure6	Figure 6. Mandibular morphology of Luffy and Eucibdelus lineage. A. Luffy; B. Menoedius; C. Parapalaestrinus; D. Eucibdelus; E. Rhynchocheilus (Photos A by the authors; photos B-E by Q. - H. Zhao). Abbreviations: Abbreviations: BET: basal extra tooth; DR: dorsal ridge; DRT: dorsal ridge teeth; VR: ventral ridge; VRT: ventral ridge teeth.	Figure 6. Mandibular morphology of Luffy and Eucibdelus lineage. A. Luffy; B. Menoedius; C. Parapalaestrinus; D. Eucibdelus; E. Rhynchocheilus (Photos A by the authors; photos B-E by Q. - H. Zhao). Abbreviations: Abbreviations: BET: basal extra tooth; DR: dorsal ridge; DRT: dorsal ridge teeth; VR: ventral ridge; VRT: ventral ridge teeth.	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
961DFFD617395FFF802402D6F7C59CB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670285	https://doi.org/10.3897/zookeys.1281.198593.figure3	Figure 3. Luffy nika sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	Figure 3. Luffy nika sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
961DFFD617395FFF802402D6F7C59CB8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670283	https://doi.org/10.3897/zookeys.1281.198593.figure1	Figure 1. Luffy schillhammeri sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	Figure 1. Luffy schillhammeri sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
7FEA42F8FB6E5FE1A137058F2CAF2EA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670283	https://doi.org/10.3897/zookeys.1281.198593.figure1	Figure 1. Luffy schillhammeri sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	Figure 1. Luffy schillhammeri sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
7FEA42F8FB6E5FE1A137058F2CAF2EA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670284	https://doi.org/10.3897/zookeys.1281.198593.figure2	Figure 2. Luffy schillhammeri sp. nov., labium, maxillary palps, protarsi, abdominal segments IX – X. A. Labium; B. Maxillary palps; C. Protarsi, dorsal view; D. Protarsi, ventral view; E. Tergite X, male; F. Tergite X, female; G. Abdominal segment IX, male; H. Abdominal segment IX and gonocoxite, female. Scale bars: 1 mm.	Figure 2. Luffy schillhammeri sp. nov., labium, maxillary palps, protarsi, abdominal segments IX – X. A. Labium; B. Maxillary palps; C. Protarsi, dorsal view; D. Protarsi, ventral view; E. Tergite X, male; F. Tergite X, female; G. Abdominal segment IX, male; H. Abdominal segment IX and gonocoxite, female. Scale bars: 1 mm.	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
7FEA42F8FB6E5FE1A137058F2CAF2EA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670285	https://doi.org/10.3897/zookeys.1281.198593.figure3	Figure 3. Luffy nika sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	Figure 3. Luffy nika sp. nov., habitus and aedeagus. A. Habitus, dorsal view; B. Habitus, ventral view; C, F. Aedeagus, lateral view; D. Aedeagus, parameral view; E. Paramere, underside. The arrow indicates the articulation on paramere. The black dots on the paramere strengthen the position of the peg setae. Scale bars: 3 mm (A, B); 1 mm (C – F).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
7FEA42F8FB6E5FE1A137058F2CAF2EA8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1670286	https://doi.org/10.3897/zookeys.1281.198593.figure4	Figure 4. Habitats of Luffy schillhammeri sp. nov. in Yunnan, China. A. Manzhang (曼掌); B. Manfeng (曼粉) (Photos by Y. - T. Wang).	Figure 4. Habitats of Luffy schillhammeri sp. nov. in Yunnan, China. A. Manzhang (曼掌); B. Manfeng (曼粉) (Photos by Y. - T. Wang).	2026-06-05	Hu, Fang-Shuo;Solodovnikov, Alexey		Zenodo	biologists	Hu, Fang-Shuo;Solodovnikov, Alexey			
