taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795501	https://doi.org/10.3897/dez.73.210225.figure1	Figure 1. Habitus of male Lucanus apollon sp. nov. A, B, E – H. Paratypes; C, D. Holotype. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	Figure 1. Habitus of male Lucanus apollon sp. nov. A, B, E – H. Paratypes; C, D. Holotype. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795502	https://doi.org/10.3897/dez.73.210225.figure2	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795503	https://doi.org/10.3897/dez.73.210225.figure3	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795504	https://doi.org/10.3897/dez.73.210225.figure4	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795505	https://doi.org/10.3897/dez.73.210225.figure5	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795506	https://doi.org/10.3897/dez.73.210225.figure6	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
94E9AECB7C305B30A04745F890BBFBAA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795507	https://doi.org/10.3897/dez.73.210225.figure7	Figure 7. Habitat of Lucanus apollon sp. nov. in Pisia, Corinthia, Greece. A. Forest habitat; B. Dirt road through the forest on steep slopes with a streetlight; C. Characteristic vegetation of the habitat, including Quercus coccifera, Quercus ilex, and Pinus halepensis; D. Living male of Lucanus apollon sp. nov. at the type locality.	Figure 7. Habitat of Lucanus apollon sp. nov. in Pisia, Corinthia, Greece. A. Forest habitat; B. Dirt road through the forest on steep slopes with a streetlight; C. Characteristic vegetation of the habitat, including Quercus coccifera, Quercus ilex, and Pinus halepensis; D. Living male of Lucanus apollon sp. nov. at the type locality.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
0E53B3D80BAD5D4CB1919F0E8DA2F206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795502	https://doi.org/10.3897/dez.73.210225.figure2	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
0E53B3D80BAD5D4CB1919F0E8DA2F206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795503	https://doi.org/10.3897/dez.73.210225.figure3	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
0E53B3D80BAD5D4CB1919F0E8DA2F206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795504	https://doi.org/10.3897/dez.73.210225.figure4	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
0E53B3D80BAD5D4CB1919F0E8DA2F206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795505	https://doi.org/10.3897/dez.73.210225.figure5	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
0E53B3D80BAD5D4CB1919F0E8DA2F206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795506	https://doi.org/10.3897/dez.73.210225.figure6	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
479690ED6BE5581A826E3F6DDA399C5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795502	https://doi.org/10.3897/dez.73.210225.figure2	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
479690ED6BE5581A826E3F6DDA399C5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795503	https://doi.org/10.3897/dez.73.210225.figure3	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
479690ED6BE5581A826E3F6DDA399C5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795504	https://doi.org/10.3897/dez.73.210225.figure4	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
479690ED6BE5581A826E3F6DDA399C5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795505	https://doi.org/10.3897/dez.73.210225.figure5	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
479690ED6BE5581A826E3F6DDA399C5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795506	https://doi.org/10.3897/dez.73.210225.figure6	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
30FBE17526C556F4812D2E228DF3A55C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795502	https://doi.org/10.3897/dez.73.210225.figure2	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	Figure 2. Habitus of male Lucanus species. A, B. L. apollon sp. nov., holotype; C, D. L. ibericus; E, F. L. busignyi; G, H. L. macrophyllus. A, C, E, G. Dorsal view; B, D, F, H. Ventral view. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
30FBE17526C556F4812D2E228DF3A55C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795503	https://doi.org/10.3897/dez.73.210225.figure3	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	Figure 3. Head details of male Lucanus species in dorsal view. A, L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 10.0 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
30FBE17526C556F4812D2E228DF3A55C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795504	https://doi.org/10.3897/dez.73.210225.figure4	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	Figure 4. Male abdominal ventrite VIII (A, E, I, M), abdominal tergite VIII (B, F, J, N) and abdominal segment IX (C, D, G, H, K, L, O, P) of Lucanus species. A – D, L. apollon sp. nov., holotype; E – H, L. ibericus; I – L, L. busignyi; M – P, L. macrophyllus. B, C, F, G, J, K, N, O. Dorsal view; A, D, E, H, I, L, M, P. Ventral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
30FBE17526C556F4812D2E228DF3A55C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795505	https://doi.org/10.3897/dez.73.210225.figure5	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	Figure 5. Aedeagus of Lucanus species in ventral view. A. L. apollon sp. nov., holotype; B. L. ibericus; C. L. busignyi; D. L. macrophyllus. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
30FBE17526C556F4812D2E228DF3A55C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1795506	https://doi.org/10.3897/dez.73.210225.figure6	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	Figure 6. Aedeagus of Lucanus species, excluding flagellum. A – C. L. apollon sp. nov., holotype; D – F. L. ibericus; G – I. L. busignyi; J – L. L. macrophyllus. B, E, H, K. Dorsal view; A, D, G, J. Ventral view; C, F, I, L. Lateral view. Scale bar: 2.5 mm.	2026-10-01	Kristl, Jonas;Flocon, Gaëtan		Zenodo	biologists	Kristl, Jonas;Flocon, Gaëtan			
