identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D06EF633FFC5743AFF2319389B817CE6.text	D06EF633FFC5743AFF2319389B817CE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites Latreille 1802	<div><p>Genus Celonites Latreille, 1802</p><p>Celonites Latreille, 1802: 368, genus. Type species: Masaris apiformis Fabricius, 1793 (= Vespa abbreviata Villers, 1789), by monotypy.</p><p>Paracelonites Kostylev, 1935: 108, subgenus of Celonites Latreille, 1802 . Unavailable under Art. 13.3 of ICZN (1999), no type species designated.</p><p>Eucelonites Kostylev, 1935: 112, subgenus of Celonites Latreille, 1802 . Unavailable under Art. 13.3 of ICZN (1999), no type species designated.</p><p>Eucelonites Richards, 1962: 215, subgenus of Celonites Latreille, 1802 . Type species: Celonites cyprius de Saussure, 1854, by original designation.</p><p>Diagnosis. Antennal scape less than 1.5 times longer than the pedicel; antennal club robust and roundly separated from the preceding flagellomeres; forewing can be folded longitudinally in rest and in death; metasoma with sharply produced lateral margins of terga forming more or less developed teeth; metasomal sterna rather flat; imagines able to roll up in the same manner as cuckoo wasps ( Chrysididae) (Richards 1962; Gess &amp; Gess 2010; Fateryga 2025c).</p><p>Number of species and distribution. The genus has a disjunct distribution with 20 species occurring in the semi-arid to arid areas in the southwest of the Afrotropical region (Gess &amp; Gess 2010, although they claimed to report 21 species, their list contained 20 valid species and one synonym) and 46 species in the Palaearctic and adjacent dry areas in the north of the Afrotropical region (according to the list of Carpenter 2001, combined with newly described or synonymized species by Gusenleitner 2002, 2007, 2012, 2018; Mauss 2013; Mauss et al. 2016, 2022a; Mauss &amp; Prosi 2018; Fateryga &amp; Fadeev 2023; Fateryga 2025b; Fateryga et al. 2025; Mauss et al. 2026; present contribution).</p></div>	https://treatment.plazi.org/id/D06EF633FFC5743AFF2319389B817CE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC6743AFF231D3A9B427B42.text	D06EF633FFC6743AFF231D3A9B427B42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites biinterruptus Kostylev 1940	<div><p>Celonites biinterruptus Kostylev, 1940</p><p>(Figs 5H, 6H, 9C)</p><p>Celonites modestus biinterruptus Kostylev, 1940: 138, ♀ ♂ (type locality: “Ruidacht, 40 km. de Stalinabad” [Tajikistan]), lectotype (Fateryga 2025b: 211), ♀, ZISP (examined).</p><p>Celonites biinterruptus: Fateryga 2025b: 211.</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 211)): ♀, labeled “РУЙдашт [ Ruydasht] / 22-VII 37 / В. ГУссакОвск. [V. Gussakovskij] // Celonites ♀ / modestus / biinterruptus Typ. / G Kostylev det. 1938 // к. ГУссакОвскОгО [collection of Gussakovskij] // Lectotype ♀ Celonites / modestus biinterruptus / Kostylev, 1940 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000319 [pale yellow label]” [ZISP]. PARALECTOTYPES: TAJIKISTAN: Ruydasht, 22.VII.1937, 1 ♀, 1 ♂, leg. V. Gussakovskij [1 ♀, ZMMU; 1 ♂, ZISP]. ADDITIONAL MATERIAL: TAJIKISTAN: Kondara Canyon, 25.VI.1937, 1 ♀, leg. V. Gussakovskij [ZISP]; ibid., 26.VII.1937, 1 ♀, leg. V. Gussakovskij [ZISP].</p><p>Distribution. Tajikistan.</p><p>Trophic relationships. Popov (1948) reported this species collected at flowers of Anchusa azurea Mill. ( Boraginaceae).</p></div>	https://treatment.plazi.org/id/D06EF633FFC6743AFF231D3A9B427B42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC6743BFF231BC698087CF4.text	D06EF633FFC6743BFF231BC698087CF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites crenulatus Morawitz 1888	<div><p>Celonites crenulatus Morawitz, 1888</p><p>(Figs 5B, 6B, 9A, E)</p><p>Celonites crenulatus Morawitz, 1888: 267, ♀ (type locality: “in territorio transcaspico (Kiltitschinar)” [Turkmenistan]), lectotype (Fateryga 2025b: 203), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 203)): ♀, labeled “[golden disc] // Kiltitschi- / nar. ♀. // Frans-Caspi G. / Turcmenien / E.König . // Celonites / crenulatus / F. Morawitz. ♀. // к. Ф МОРавица [collection of F. Morawitz] // Lectotypus ♀ / Celonites crenulatus / Morawitz, 1888 / des. Fateryga, 2021 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000320 [pale yellow label]” [ZISP]. ADDITIONAL MATERIAL: TURKMENISTAN: Tash-Kepri [ Daşköpri], 5.V.1923, 1♀ [ZMMU] ; Ashgabat, 10.VI.1928, 1♀,leg. V.Gussakovskij [ZISP]; ibid., 10.VII.1928, 1 ♀, leg. V. Gussakovskij [ZISP] ; Uch-Adzhi [ Bagtyýarlyk], 1–3.V.1929, 7 ♀, 5 ♂, leg. A. Shestakov [ZISP] ; Imam-Baba, 285 m, 1932, 29 ♀, 12 ♂, leg. A. Shestakov [ZISP] ; ibid., 19.V.1932, 1 ♀, leg. Kuzjakin [ZMMU] ; Karabata, 7.VI.1932, 1 ♀, leg. Kuzjakin [ZMMU] ; Repetek, 1.VI.1937, 7 ♂, leg. G. Kostylev [ZMMU] ; ibid., 9.VI.1937, 1 ♂, leg. G. Kostylev [ZMMU]; ibid., 30.V.1976, 1 ♀, leg. N. Kurzenko [FSCV]; Ashgabat, sands, 7.VII.1937, 2 ♀, 2 ♂, leg. Ushinskiy [ZMMU]; ibid., 8.VII.1937, 1 ♂, leg. Ushinskiy [ZMMU] ; 20 km W Kerki [ Kerkichi], sands, 20.VI.1971, 2 ♀, leg. V. Zaythsev [ZISP] ; 9 km NEE Anau, 30.VIII.1973, 1 ♂, leg. V. Zaythsev [ZISP] ; Akhcha-Kuyma, NW Kazandzik [ Bereket], 4.VI.1976, 1 ♀, 1 ♂, leg. N. Kurzenko [1 ♀, FSCV; 1 ♂, CVMM]; ibid., 6.VI.1976, 1 ♀, leg. N. Kurzenko [CVMM]; 25 km E Ashgabat, 20.VI.1976, 8 ♀, 5 ♂, leg. N. Kurzenko [7 ♀, 4 ♂, FSCV; 1 ♀, 1 ♂, CVMM]; Vicinity of Ashgabat, sands, 21.VI.1976, 7 ♀, 3 ♂, leg. N. Kurzenko [6 ♀, 2 ♂, FSCV; 1 ♀, 1 ♂, CVMM]. TAJIKISTAN: Ayvadzh, mouth of Kofarnihon River, 26.VI.1934, 1 ♀, leg. V. Gussakovskij [ZMMU] ; ibid., 16–17.VI.1936, 1 ♀, leg. V. Gussakovskij [ZISP] ; Mt. Koy-Pyaz-Tau near Kobadiyan [ Qubodiyon], 14.VII.1934, 2 ♀, 1 ♂, leg. V. Gussakovskij [ZISP] ; Sands 5 km S <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=68.166664&amp;materialsCitation.latitude=37.133335" title="Search Plazi for locations around (long 68.166664/lat 37.133335)">Kurdjalalkum</a>, 37°08′N, 68°10′E, 6.V.2002, 1 ♀, 1 ♂, leg. S. Zonstein [CSZT] ; Tigrovaya Balka Nature Reserve, 37°16ʹ51ʺN, 68°20ʹ53ʺE, 330 m, 14.VI.2025, 1 ♂, leg. M. Proshchalykin [CAFK]. UZBEKISTAN: Karakalpakstan, Baday-Tugay Nature Reserve, 24.V.1977, 1 ♀, leg. Zlobin [ZISP]. COUNTRY UNCERTAIN: [cannot be read], 1 ♀ [ZMMU] ; Middle Asia ?, 2 ♀ [ZISP].</p><p>Additional literature records. TURKMENISTAN: Kurtly, near Ashgabat (Mauss et al. 2026).</p><p>Distribution. Iran, Turkmenistan, Tajikistan, Uzbekistan.</p><p>Trophic relationships. Probably narrowly oligolectic (sensu Müller &amp; Kuhlmann 2008) on the genus Heliotropium ( Boraginaceae); the pollen from the rectum of a male of this species consisted of 100% pollen of this plant genus (Mauss et al. 2026).</p></div>	https://treatment.plazi.org/id/D06EF633FFC6743BFF231BC698087CF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC7743BFF231D1498427B57.text	D06EF633FFC7743BFF231D1498427B57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites foveolatus Kostylev 1935	<div><p>Celonites foveolatus Kostylev, 1935</p><p>(Figs 5K, 6K, 8D, H, 9I)</p><p>Celonites foveolatus Kostylev, 1935: 112, ♀ ♂ (type locality: “ЗакаспИЙскИЕ пУстынИ” [Turkmenistan, Transcaspian deserts]), lectotype (Fateryga 2025b: 204), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 204)): ♀, labeled “[golden disc] // УчЬ—АдЖИ [ Uch-Adzhi] / 1-3. V. 1929 / А. ШЕстакОва [A. Shestakova] // Celonites ♀ / foveolatus n. sp. / G. Kostylev det. 1929 // Lectotype ♀ / Celonites foveolatus / Kostylev, 1935 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000321 [pale yellow label]” [ZISP]. PARALECTOTYPES: TURKMENISTAN: Uch-Adzhi, 1–3.V.1929, 13 ♀, 1 ♂, leg. A. Shestakov [13 ♀, ZISP; 1 ♂, ZMMU]; ibid., 1–3.V.1929, 11 ♀, 1 ♂, leg. A. Shestakova [9 ♀, 1 ♂, ZISP; 2 ♀, ZMMU]. ADDITIONAL MATERIAL: TURKMENISTAN: Bamy, 15.IV, 1 ♀ [ZISP]; Bathyz Nature Reserve, Kyzyl-Zhar [Gyzyljar] Place, 6–7.V.1989, 1 ♀, leg. Volkovich [ZISP]; ibid., 13.V.1990, 1 ♀, leg. A. Lelej [FSCV]; ibid., 13.V.1990, 1 ♀, leg. N. Kurzenko [FSCV]; Repetek, 4–10.V.1990, 2 ♀, 1 ♂, leg. V. Kazenas [FSCV]; 50 km NEE Tejen, Ju-Ju-Klu, 23.V.1985, 1 ♂, leg. V. Makarkin [FSCV].</p><p>Distribution. Turkmenistan.</p><p>Trophic relationships. Unknown.</p></div>	https://treatment.plazi.org/id/D06EF633FFC7743BFF231D1498427B57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC7743BFF231BB59B4279D2.text	D06EF633FFC7743BFF231BB59B4279D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites hystrix Kostylev 1940	<div><p>Celonites hystrix Kostylev, 1940</p><p>(Figs 5D, 6D, 7J, 8C)</p><p>Celonites hystrix Kostylev, 1940: 138, ♀ ♂ (type locality: “Kondara, Chaîne Ghissar, 30 km vers le N de Stalinabad (autrefois Duchambé), 1 100 m au-d. du niv. de la mer” [Tajikistan]), lectotype (Fateryga 2025b: 208), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 208)): ♀, labeled “УЩ. КОндаРа [ Kondara Canyon] / 25-VI 37 / В. ГУссакОвск. [V. Gussakovskij] // Celonites / hystryx Typus ♀ / G Kostylev det. 1938 // к. ГУссакОвскОгО [collection of Gussakovskij] // Lectotype ♀ / Celonites hystryx / Kostylev, 1940 / des. Fateryga, 2024 [red label] //Zoological Institute/St.Petersburg/ INS_HYM_0000322 [pale yellow label]”[ZISP]. PARALECTOTYPES: TAJIKISTAN: Kondara Canyon, 25.VI.1937, 1 ♀, leg. V. Gussakovskij [ZMMU]; ibid., 26.VII.1937, 1 ♀, 1 ♂, leg. V. Gussakovskij [ZISP]. ADDITIONAL MATERIAL: UZBEKISTAN: Qashqadaryo Province, Tashkurgan, 2.VII.1942, 1 ♂, leg. K. Arnoldi [ZISP].</p><p>Distribution. Tajikistan, * Uzbekistan.</p><p>Trophic relationships. Popov (1948) reported this species collected in Tajikistan at flowers of Anchusa azurea ( Boraginaceae).</p></div>	https://treatment.plazi.org/id/D06EF633FFC7743BFF231BB59B4279D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC0743CFF231E9D99AA7852.text	D06EF633FFC0743CFF231E9D99AA7852.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites kozlovi Kostylev 1935	<div><p>Celonites kozlovi Kostylev, 1935</p><p>(Figs 5J, 6J, 7M, 9G)</p><p>Celonites kozlovi Kostylev, 1935: 110, [♀] (type locality: “ПУстынЯ ГОБИ (МОнгОЛИЯ)” [Mongolia, Gobi Desert]), lectotype (Fateryga 2020: 22), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Fateryga (2020: 22)): ♀, labeled “[golden disc] // ЦЕнт. ГОБИ, кЛ. / УЛанъ-БУЛак [Central Gobi, Ulan-Bulak Spring] / 6–8.VII.1909 / П. КОЗЛОвъ. М.-С. Эк. [P. Kozlov, Mongolia– Sichuan expedition] // Celonites ♀ / kozlovi, n. sp. / G. Kostylev det. 1930 // Lectotypus ♀ / Celonites kozlovi / Kostylev, 1935 / des. Fateryga, 2019 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000323 [pale yellow label]” [ZISP]. ADDITIONAL MATERIAL: RUSSIA. Altai Republic: 2 km SE Chagan-Uzun, Balkhash River valley, 9.VII.2016, 3 ♀, leg. M. Proshchalykin, V. Loktionov [FSCV] ; 5 km SE Chagan-Uzun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.42&amp;materialsCitation.latitude=50.07361" title="Search Plazi for locations around (long 88.42/lat 50.07361)">Tydtuyaryk River</a> valley, 50°04′25″N, 88°25′12″E, on Dracocephalum peregrinum, 8.VII.2019, 2 ♀, leg. A. Fateryga [CAFK] ; ibid., on Ziziphora clinopodioides, 8.VII.2019, 2 ♀, leg. A. Fateryga [CAFK] ; ibid., on Aster alpinus, 8.VII.2019, 1 ♀, leg. A. Fateryga [CAFK] ; ibid., on Youngia tenuifolia subsp. altaica, 8.VII.2019, 1 ♀, leg. A. Fateryga [CAFK] ; ibid., on Ziziphora clinopodioides, 8.VII.2019, 2 ♀, leg. A. Fateryga [CVMM] ; ibid., 8.VII.2019, 2 ♀, leg. V. Dorofeev [ASUB]; ibid., 8–9.VII.2019, 1 ♀, leg. Yu. Danilov [ISEN]; ibid., on Dracocephalum peregrinum, 9.VII.2019, 1 ♀, leg. A. Fateryga [CVMM] ; ibid., 9.VII.2019, 1 ♀, leg. V. Dorofeev [ASUB]; ibid., 12.VI.2022, 1 ♀, leg. A. Fateryga [ZISP]; ibid., on Klasea marginata, 12.VI.2022, 1 ♂, leg. A. Fateryga [CAFK] ; ibid., 12.VI.2022, 5 ♀, leg. M. Proshchalykin [CAFK]; ibid., 18.VI.2022, 5 ♀, leg. M. Proshchalykin [CAFK]; ibid., 21.VI.2022, 2 ♀, leg. A. Fateryga [CVMM]; ibid., 21.VI.2022, 12 ♀, 1 ♂, leg. M. Proshchalykin [3 ♀, IBUM; 9 ♀, 1 ♂, CAFK]; ibid., 22.VI.2022, 2 ♀, leg. M. Proshchalykin [CAFK]; “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.3125&amp;materialsCitation.latitude=50.06389" title="Search Plazi for locations around (long 88.3125/lat 50.06389)">Mars</a> ”, 50°03′50″N, 88°18′45″E, 17.VI.2022, 2 ♀, leg. M. Proshchalykin [CAFK] ; ibid., 22.VI.2022, 2 ♀, leg. M. Proshchalykin [CAFK]; ibid., 23.VI.2022, 2 ♀, leg. M. Proshchalykin [CAFK]; ibid., 25.VI.2022, 1 ♂, leg. A. Fateryga [ZISP]; ibid., 25.VI.2022, 1 ♀, leg. M. Proshchalykin [AMNH]; ibid., 25.VI.2022, 1 ♀, leg. M. Proshchalykin [CAFK]; 5 km NE Kokorya, 49°57′00″N, 89°04′19″E, on Caragana bungei, 26.VI.2022, 1 ♀, leg. A. Fateryga [CAFK] .</p><p>Additional literature records. MONGOLIA: Khovd Province, Conocharajch-gol ( Choit-dalai-nuur) (Gusenleitner 1991). RUSSIA. Altai Republic: 3 km NE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=89.04583&amp;materialsCitation.latitude=49.941944" title="Search Plazi for locations around (long 89.04583/lat 49.941944)">Kokorya</a>, 49°56′31″N, 89°02′45″E (Fateryga et al. 2023).</p><p>Distribution. Russia (Altai Republic), Mongolia.</p><p>Trophic relationships. Celonites kozlovi is a polylectic species in the narrow sense (sensu Müller &amp; Kuhlmann 2008), collecting pollen from flowers of plants belonging to at least five families (with the predomination of Asteraceae and Lamiaceae) and using diverse methods for both pollen and nectar uptake. Females consume pollen directly from the anthers with the mouthparts and the foretarsi. In addition, this species can be a secondary nectar robber (sensu Inouye 1980) that was never observed in other species of Masarinae . Morphological and behavioral adaptations of C. kozlovi for pollen uptake from flowers were described in detail by Fateryga (2020) and Fateryga et al. (2023). Flower-visiting records (all in the Altai Republic of Russia): Aster alpinus L., Crepidiastrum tenuifolium (Willd.) Sennikov (≡ Youngia tenuifolia subsp. altaica Babc. &amp; Stebbins), Klasea marginata (Tausch) Kitag., Pseudopodospermum pubescens (DC.) Zaika, Sukhor. &amp; N. Kilian, Saussurea pricei N.D. Simpson, and Taraxacum sp. (all Asteraceae); Sisymbrium polymorphum (Murray) Roth ( Brassicaceae); Gypsophila patrinii Ser. ( Caryophyllaceae); Convolvulus ammannii Desr. ( Convolvulaceae); Caragana bungei Ledeb. ( Fabaceae); Dracocephalum peregrinum L. and Ziziphora clinopodioides Lam. (both Lamiaceae); Sibbaldianthe bifurca (L.) Kurtto &amp; T. Erikss. ( Rosaceae) (Fateryga 2020; Fateryga et al. 2023).</p></div>	https://treatment.plazi.org/id/D06EF633FFC0743CFF231E9D99AA7852	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC0743DFF2318B298467CF4.text	D06EF633FFC0743DFF2318B298467CF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites kurzenkoi Mauss, Fateryga & Carpenter 2026	<div><p>Celonites kurzenkoi Mauss, Fateryga &amp; Carpenter, 2026</p><p>(Figs 5C, 6C, 7B, 8B, G, 9B)</p><p>Celonites osseus: Richards 1962: 226, ♀.</p><p>Celonites kurzenkoi Mauss, Fateryga &amp; Carpenter, 2026: 8, figs 1B, D–G, 2B, D, F, 3B, C, E, G, 4B, D, 5B, D, F, 6C, D, 7C, D, 8C, ♀ ♂ (type locality: “ТУРкмЕнИЯ АхчакУЙма NW КаЗандЖИка [Turkmenistan, Akhcha-Kuyma, NW Gazandjyk (currently Bereket)]”), holotype, ♀, ZISP (examined).</p><p>Material examined. HOLOTYPE: ♀, labeled “ТУРкмЕнИЯ / АхчакУЙма / NW КаЗандЖИка [Turkmenistan, Akhcha-Kuyma, NW Gazandjyk (currently Bereket)] / 6.VI.1976 Н. КУРЗЕнкО [N. Kurzenko] // Holotypus ♀ / Celonites kurzenkoi Mauss, Fateryga / &amp; Carpenter [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000372 [pale yellow label]” [ZISP] . PARATYPES: TURKMENISTAN: Ashgabat, 10.VII.1928, 1 ♂, leg. V. Gussakovskij [ZISP]; ibid., 18.IX.1937, 1 ♀, leg. Ushinskiy [ZISP]; Imam-Baba, 285 m, 1932, 1 ♀, leg. A. Shestakov [ZISP]; Repetek, 9.VI.1937, 1 ♀, leg. G. Kostylev [ZMMU]; ibid., 30.V.1976, 1 ♂, leg. N. Kurzenko [CVMM]; Vicinity of Jebel, Mollakara, 19.V.1969, 1 ♂, leg. V. Zaythsev [ZISP] ; Tash-Kepri [Daşköpri], 20.V.1976, 1 ♂, leg. N. Kurzenko [AMNH]; Akhcha-Kuyma, NW Kazandzik [ Bereket], 3.VI.1976, 5 ♀, leg. N. Kurzenko [1 ♀, AMNH ; 3 ♀, FSCV; 1 ♀, CVMM]; ibid., 6.VI.1976, 2 ♀, 1 ♂, leg. N. Kurzenko [AMNH]; ibid., 4.VI.1976, 1 ♀, leg. N. Kurzenko [FSCV]; ibid., 6.VI.1976, 5 ♀, leg. N. Kurzenko [FSCV]; 25 km E Ashgabat, 20.VI.1976, 2 ♀, 1 ♂, leg. N. Kurzenko [1 ♀, FSCV ; 1 ♀, CVMM; 1 ♂, AMNH]; Vicinity of Ashgabat, sands, 21.VI.1976, 1 ♀, leg. N. Kurzenko [FSCV] ; Akhcha-Kuyma, 30 km W Kazandzik [ Bereket], 30.V.1985, 1 ♀, leg. A. Lelej [FSCV] ; 30 km W Kazandzik [Bereket], Akhcha-Kuyma, 4.VI.1988, 2 ♀, leg. N. Kurzenko [FSCV, OLML] ; ibid., 5.VI.1988, 2 ♂, leg. N. Kurzenko [FSCV, OLML] .</p><p>Distribution. Turkmenistan.</p><p>Trophic relationships. Probably narrowly oligolectic (sensu Müller &amp; Kuhlmann 2008) on the genus Heliotropium ( Boraginaceae); pollen samples from the rectum of two males of this species consisted of 100% pollen of this plant genus (Mauss et al. 2026).</p></div>	https://treatment.plazi.org/id/D06EF633FFC0743DFF2318B298467CF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC17430FF231D1498427CBC.text	D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites maussi Fateryga 2026	<div><p>Celonites maussi Fateryga, sp. nov.</p><p>(Figs 1, 2, 5L, 6L)</p><p>Diagnosis. The species is closely related to Celonites tristiculus Kostylev, 1935 and can be distinguished from it by a regularly and sparsely punctate frons, with dull and distinctly reticulately microsculptured interspaces exceeding the puncture diameter in both sexes (Figs 1D, 2C) (versus a densely rugosely punctate frons, with rather shining, less distinctly microsculptured interspaces less than the puncture diameter in C. tristiculus, Fig. 7F, G), sternum VI of the female with moderately coarse macropunctures and reticulately micropunctured interspaces between them (Fig. 1F) (versus with much coarser macropunctures and smoothened interspaces between them in C. tristiculus, Fig. 9J), apical margin of the male sternum VIII without distinct short acute processes (Fig. 5L) (versus with two distinct short acute processes in C. tristiculus, Fig. 5M); harpide with a distinctly produced postero-lateral lobe, and volsella with a straight posterior margin (Fig. 6L) (versus harpide with the postero-lateral lobe not so produced and volsella with a roundly produced posterior margin in C. tristiculus, Fig. 6M).</p><p>Description. Female. Body length (from head to apical margin of tergum II) 5 mm (total body length about 6.5 mm); forewing length 5 mm. Head in frontal view about 1.2× as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus about 1.3× as wide as long; its apical emargination shallow, about 0.2× as deep as wide, taking about 0.35 of clypeal width, apical teeth rounded. Distance between lateral ocellus and occiput about 0.8× as long as distance between lateral ocelli and about as distance between lateral ocellus and compound eye. Antenna with articles 8–12 forming ventrally flattened club about 2.4× as long as wide in dorsal view. Width of gena in front of ocular sinus about equal to diameter of lateral ocellus. Preoccipital carina weak but distinct along nearly whole outer margin of compound eye except uppermost dorsal surface. Pronotum with anterior side nearly vertical and slightly concave, roundly angled to dorsal surface. Pronotal humerus broadly rounded in dorsal view; in lateral view, pronotal carina distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, moderately convex, bluntly angled posteriorly. Axilla of scutellum with obsolete lateral projection towards tegula. Tegula without depression in front of axilla; outer side of tegula distinctly shallowly emarginate. Epipleural and episternal sulci indistinct. Ventral mesepisternum unevenly rounded ventrally. Epicnemial carina distinct. Mesepimeron produced posteriorly as small tubercle. Dorsal carinae of propodeum forming sharp lamellar tooth on medial end, adjacent to metanotum. Outline of emargination between lateral lamella and postero-lateral process of propodeum small, comma-like, its medially directed apex hardly enlarged. Terga I–V medially with straight, not crenulate margins, laterally with small tooth. Tergum VI bluntly angulate apically, laterally with rounded tooth. Sternum VI postero-medially with small acute process. Basitarsus of foreleg about 3.4× as long as broad in dorsal view.</p><p>Clypeus regularly punctate with sparse and very shallow punctures; interspaces exceeding puncture diameter, dull, strongly reticulately microsculptured. Punctation on frons generally similar to that on clypeus but macropunctures evidently deeper. Punctation on vertex and gena much denser, interspaces less than puncture diameter, also evidently reticulately microsculptured. Pronotum, scutum, and scutellum densely punctate, punctures larger than those anywhere on head, interspaces less than puncture diameter, dull, with reticulate microsculpture. Tegula dull, mostly microsculptured in anterior half, with few indistinct punctures, and deeply punctate in posterior half, interspaces between punctures reaching puncture diameter. Dorsal and ventral mesepisterna and mesepimeron generally sculptured similarly to pronotum, scutum, and scutellum but some punctures slightly coarser. Dorsal surface of metanotum densely punctate with punctures smaller than those on pronotum, scutum, and scutellum, interspaces much less than puncture diameter, dull, reticulately microsculptured; posterior surface of metanotum without distinct macropunctures, only with microsculpture. Metapleuron and lateral surface of propodeum dull, longitudinally wrinkled, without distinct punctures. Posterior surface of propodeum (except lateral lamella) densely punctate, punctures forming longitudinal rows, interspaces less than puncture diameter, dull, with reticulate microsculpture; lateral lamella regularly punctate with coarse macropunctures, interspaces less than puncture diameter, with less distinct microsculpture. Metasomal terga deeply punctate, about as coarse as pronotum, scutum, and scutellum, interspaces less than puncture diameter on disc and reaching puncture diameter on depression, dull, with weak microsculpture. Sternum I with dull reticulate microsculpture only. Sterna II–V with scattered fine punctures, interspaces much exceeding puncture diameter, rather dull, with strong reticulate microsculpture. Sternum VI with moderately coarse punctures, interspaces reaching puncture diameter, rather dull, with distinct reticulate microsculpture.</p><p>Mandible, labrum, and clypeus covered with thin pale setae, reaching in length diameter of median ocellus. Frons and pronotum with very short setae, reaching at most 1/3 of diameter of median ocellus. Vertex between ocelli with few setae reaching in length 1/2 of diameter of median ocellus. Compound eye with very short dust-like setae. Scutum, scutellum, and terga without noticeable setae. Sterna mostly with short appressed setae, especially at apical margins. Coxae and femora with setae as long as those on clypeus. Tibia and tarsus of foreleg with stiff setae as long as diameter of median ocellus, those setae on mid and hind legs much shorter; pollen-collecting brush of stiff setae on inner surface of basitarsus of foreleg not modified. Spur of foretibia slightly curved and rather blunt apically.</p><p>Basal color black. Following parts whitish-yellow: median spot on pronotum, short and narrow band along posterior side of pronotum near tegula, tegula (except antero-central part), lateral side of lateral lamella of propodeum, apical band on tergum I extended anteriorly at lateral sides, apical bands on terga II–III, apical bands on medial parts of terga IV–V. Whitish-yellow color on body slightly outlined by brown. Mandible, labrum, antero-central part of tegula, and tarsi rather brown. Ventral side of antennal club ferruginous. Small basal parts of tibiae pure yellow. Wings strongly fuscous; veins and pterostigma brown.</p><p>Male. Body length (from head to apical margin of tergum II) 4.5 mm (total body length about 6 mm); forewing length 4.5 mm. Structure resembles that in female except the following: head in frontal view about 1.3× as wide as long; distance between lateral ocellus and occiput about 1.65× as distance between lateral ocellus and compound eye; antennal club larger than that in female; ventral side of antennal club with three oval-shaped tyloids on articles 9, 10, and 11; tyloid of article 10 taking about 2/3 of length of article; tyloid of article 9 smaller than that of article 10 and larger than that of article 11; tergum VI and sternum VI as those of preceding segment; tergum VII roundly truncate apically, laterally with tooth; sternum VIII apically truncate, without processes. Genitalia as in Fig. 6L; harpide with distinctly produced postero-lateral lobe; volsella with straight posterior margin; aedeagus rather broad, slightly narrowing towards apex.</p><p>Sculpture and setation as in female except that clypeus more shining, with less strong reticulate microsculpture on interspaces between macropunctures. Tergum VII punctate coarsely than preceding terga, similarly to lateral lamella of propodeum. Sternum VI sculptured as previous sterna; sternum VII with band of scattered coarse punctures on distal half. Spur of foretibia less curved and dorsally with large transparent lamella. Coloration resembles that in female but tergum VI medially with whitish-yellow apical spot and tibiae rather brown than black.</p><p>Material examined. HOLOTYPE: ♀, labeled: “КаРатаУ [Karatau] / 30-V 30. // Celonites ♀ / tristiculus / karatauicus m / G. Kostylev det. 1933 // Holotypus ♀ / Celonites maussi / Fateryga [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000373 [pale yellow label]” [ZISP] . PARATYPE: KAZAKHSTAN. Turkestan Province: Karatau, Karasay Place, 22.V.1932, 1 ♂, leg. Pravdin [ZISP] .</p><p>Etymology. It is a pleasure to name this species after my colleague Volker Mauss (Staatliches Museum für Naturkunde, Stuttgart, Germany), the leading expert in the taxonomy and the biology of the Palaearctic Masarinae, in recognizing his support of my study of the pollen wasps.</p><p>Distribution. Kazakhstan (Turkestan Province).</p><p>Trophic relationships. Unknown.</p></div>	https://treatment.plazi.org/id/D06EF633FFC17430FF231D1498427CBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFCC7433FF231CDC98427D2C.text	D06EF633FFCC7433FF231CDC98427D2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites mediasiaticus Fateryga 2026	<div><p>Celonites mediasiaticus Fateryga, sp. nov.</p><p>(Figs 3, 4, 5I, 6I)</p><p>Diagnosis. The species is closely related to Celonites biinterruptus and can be distinguished from it by complete yellow apical bands on first three to four metasomal terga and apical yellow spots medially on the remaining terga except the last one in both sexes (Figs 3B, F, 4B, E) (versus twice-interrupted whitish apical bands outlined by ferruginous on most terga except the last one in C. biinterruptus, Fig. 9C), a longer sternum VII+VIII of the male (Fig. 5I) (versus a shorter sternum VII+VIII in C. biinterruptus, Fig. 5H), a broader harpide, rather parallel-sided aedeagus, and volsella with a less produced posterior margin (Fig. 6I) (versus a narrower harpide, aedeagus narrowing towards the apex, and volsella with a strongly produced posterior margin in C. biinterruptus, Fig. 6H).</p><p>Description. Female. Body length (from head to apical margin of tergum II) 5.5 mm (total body length about 7 mm); forewing length 5.5 mm. Head in frontal view about 1.2× as wide as long (from top of median ocellus to apical margin of clypeus). Clypeus about 1.3× as wide as long; its apical emargination shallow, about 0.2× as deep as wide, taking about 0.45 of clypeal width, apical teeth rounded. Distance between lateral ocellus and occiput about 0.85× as long as distance between lateral ocelli and about as distance between lateral ocellus and compound eye. Antenna with articles 8–12 forming ventrally flattened club about 2.3× as long as wide in dorsal view. Width of gena in front of ocular sinus about equal to diameter of lateral ocellus. Preoccipital carina distinct, developed along nearly whole outer margin of compound eye except uppermost dorsal surface. Pronotum with anterior side nearly vertical, roundly angled to dorsal surface. Pronotal humerus broadly rounded in dorsal view; in lateral view, pronotal carina distinctly separating antero-ventral area of pronotum from its dorsal area at antero-lateral sides. Scutellum separated from mesoscutum by transverse suture, moderately convex, bluntly angled posteriorly.Axilla of scutellum with very weak to obsolete lateral projection towards tegula. Tegula without depression in front of axilla; outer side of tegula distinctly shallowly emarginate. Epipleural and episternal sulci distinct. Ventral mesepisternum unevenly rounded ventrally. Epicnemial carina weakly developed. Mesepimeron produced posteriorly as small tubercle. Dorsal carinae of propodeum weakly developed, lamellar tooth on its medial end (adjacent to metanotum) obsolete. Outline of emargination between lateral lamella and postero-lateral process of propodeum small, comma-like, its medially directed apex hardly enlarged. Terga I–V medially with straight, not crenulate margins, laterally with small tooth. Tergum VI bluntly angulate apically, laterally with blunt to rounded tooth. Sternum VI postero-medially without distinct process. Basitarsus of foreleg about 2.9× as long as broad in dorsal view.</p><p>Clypeus irregularly punctate with sparse and very shallow punctures; interspaces reaching puncture diameter, rather dull, reticulately microsculptured. Frons regularly punctate with punctures deeper than those on clypeus, interspaces exceeding puncture diameter, dull, strongly reticulately microsculptured. Punctation on vertex and gena much denser than that on frons, interspaces less than puncture diameter, also evidently reticulately microsculptured. Pronotum, scutum, and scutellum densely punctate, punctures larger than those anywhere on head, interspaces much less than puncture diameter, dull, with visible reticulate microsculpture. Tegula dull, mostly microsculptured in anterior 2/5, with few indistinct punctures, and deeply punctate in posterior 3/5, interspaces between punctures reaching puncture diameter. Dorsal and ventral mesepisterna and mesepimeron generally sculptured similarly to pronotum, scutum, and scutellum or very slightly coarser. Dorsal surface of metanotum coarsely punctate similarly to pronotum, scutum, and scutellum, interspaces much less than puncture diameter; posterior surface of metanotum without distinct macropunctures, only with microsculpture. Metapleuron and lateral surface of propodeum dull, longitudinally wrinkled, without any punctures. Posterior surface of propodeum (except lateral lamella) densely punctate, punctures fusing to longitudinal rows, interspaces less than puncture diameter, edge-like; lateral lamella regularly punctate with coarse macropunctures, interspaces less than puncture diameter, with weakly developed microsculpture. Metasomal terga deeply punctate, about as coarse as pronotum, scutum, and scutellum, interspaces less than puncture diameter on disc and reaching puncture diameter on depression, dull, with weak microsculpture.</p><p>Sternum I with dull reticulate microsculpture only. Sterna II–V with scattered fine punctures, interspaces much exceeding puncture diameter, rather dull, with strong reticulate microsculpture. Sternum VI with moderately coarse macropunctures, interspaces less than puncture diameter, rather dull, with distinct reticulate microsculpture and few scattered micropunctures.</p><p>Mandible, labrum, and clypeus covered with thin pale setae, reaching in length diameter of median ocellus. Frons and pronotum with shorter setae, reaching approximately 2/3 of diameter of median ocellus. Vertex between ocelli with few setae reaching in length diameter of median ocellus. Compound eye with very short dust-like setae. Scutum, scutellum, and terga without noticeable setae. Sterna mostly with short appressed setae, especially at apical margins. Coxae and femora with setae as long as 1/3 of diameter of median ocellus. Tibia and tarsus of foreleg with stiff setae as long as diameter of median ocellus, those setae on mid and hind legs much shorter; pollen-collecting brush of stiff setae on inner surface of basitarsus of foreleg not modified. Spur of foretibia slightly curved and equipped with pointed spike at apical end, directed at blunt angle from spur body.</p><p>Basal color black. Following parts yellow: median spot on pronotum, spot on antero-lateral side of pronotum, tegula (except small antero-central part), lateral side of lateral lamella of propodeum, apical bands on terga I–III extended anteriorly at center and lateral sides, apical spots medially on terga IV–V. Distal half of mandible and ventral side of antennal club ferruginous. Small antero-central part of tegula and tarsi rather brown. Tops of femora and whole tibiae brownish-yellow. Wings strongly fuscous; veins and pterostigma brown.</p><p>Male. Body length (from head to apical margin of tergum II) 5 mm (total body length about 6.5 mm); forewing length 5 mm. Structure resembles that in female except the following: head in frontal view about 1.3× as wide as long; distance between lateral ocellus and occiput about 1.45× as distance between lateral ocellus and compound eye; antennal club larger than that in female; ventral side of antennal club with three large elongate tyloids on articles 9, 10, and 11; tyloid of article 10 taking nearly whole length of article; tyloid of article 9 smaller than that of article 10 and larger than that of article 11; tergum VI and sternum VI as those of preceding segment; tergum VII roundly angulate apically, laterally with strong tooth; sternum VIII apically emarginate, without processes. Genitalia as in Fig. 6I; harpide broad; volsella with slightly produced posterior margin; aedeagus rather parallel-sided.</p><p>Sculpture and setation as in female except that clypeus slightly more shining, with less distinct reticulate microsculpture on interspaces between macropunctures. Sternum VI sculptured as previous sterna; sternum VII with band of scattered coarse punctures on distal half. Setae on frons longer, about as diameter of median ocellus. Spur of foretibia less curved and dorsally with large transparent lamella. Coloration resembles that in female but tergum VI medially with yellow apical spot and tibiae rather yellow than brownish-yellow; in two specimens, apical spot on tergum IV replaced by complete apical band.</p><p>Material examined. HOLOTYPE: ♀, labeled “КАРЖАНТАУ [ Karzhantau] К [Kazakhstan?] / СУБАЛьпиЙСКиЙ ЛУГ [Sub-alpine meadow] / 2-6 ОЗ. СУсИнган [Susingen Lake] / VII 1939 ОБУХОВА [ Obukhova] // Celonites / octoannulatus / Kuzn. / G. Kostylev / det. 19.. // Holotypus ♀ / Celonites mediasiaticus / Fateryga [red label]” [ZMMU] . PARATYPES: UZBEKISTAN: Ak-Tash, 25.V.1921, 1 ♂, leg. N. Kuznetsov [ZMMU] . KAZAKHSTAN. Turkestan Province: Karatau, Zhylagan-Ata Place, 1.VI.1931, 1 ♂, leg. Pogodina [ZISP]; Karatau, Karasay Place, 6.VI.1932, 1 ♂, leg. Pravdin [ZISP]; Karzhantau, sub-alpine meadow, left of Susingen Lake, 28.VII.1934, 1 ♀, leg. Obukhova [ZMMU]; Chimkent Province, 5 km NE Darbaza Station, steppe, 24.IV.1988, 1 ♀, leg. N. Vinokurov [FSCV] .</p><p>Etymology. The specific name is an adjective meaning “Middle Asian” as a reference to the distribution of this species in Middle Asia.</p><p>Distribution. Uzbekistan, Kazakhstan (Turkestan Province).</p><p>Trophic relationships. Unknown.</p></div>	https://treatment.plazi.org/id/D06EF633FFCC7433FF231CDC98427D2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFCF7433FF231D549B567862.text	D06EF633FFCF7433FF231D549B567862.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites modestus Kostylev 1935	<div><p>Celonites modestus Kostylev, 1935</p><p>(Fig. 8E)</p><p>Celonites modestus Kostylev, 1935: 110, [♀] (type locality: “ПамИР, ХРЕБЕт ПЕтРа ВЕЛИкОгО” [Tajikistan, Pamir, Peter the Great Range]), lectotype (Fateryga 2025b: 210), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Fateryga (2025b: 210)): ♀, labeled “[golden disc] // ХР. ПЕтРа ВЕЛИкагО / ДОЛИна Р. КаРа-ШУРы / Р. ГУРсы-Ташъ [Peter the Great Range, Kara-Shura River valley, Gursy-Tash River] / 29.VII.1911. А. ГОЛЬБЕкъ [A. Golbek] // Celonites ♀ / modestus, n. sp / G. Kostylev det. 1932 // Lectotype ♀ / Celonites modestus / Kostylev, 1935 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000324 [pale yellow label]” [ZISP]. PARALECTOTYPE: TAJIKISTAN: Peter the Great Range, Kara-Shura River valley, Yashil-Kul Lake, 6.VIII.1911, leg. A. Golbek [ZISP]. ADDITIONAL MATERIAL: TAJIKISTAN: Pamir, Murgab River valley, 3 km below mouth of Western Pshart River, Chottukay Palce, 3325 m, on Myricaria squamosa, Lonicera, Dracocephalum, 29.VI.1958, 7 ♀, leg. Gorodkov [ZISP].</p><p>Distribution. Tajikistan.</p><p>Trophic relationships. A series of females of this species was probably collected at flowers of Dracocephalum sp. ( Lamiaceae). Despite three plant species were indicated simultaneously on the labels, two others are hardly probable, while species of the genus Dracocephalum are a known pollen source for two other more or less closely related species of the genus Celonites (Fateryga 2020; Fateryga et al. 2023).</p><p>Remarks. The species is closely related to Celonites octoannulatus (Kuznetzov, 1923) . To verify the taxonomic independence of C. modestus, males of this species should be collected and compared with those of C. octoannulatus .</p></div>	https://treatment.plazi.org/id/D06EF633FFCF7433FF231D549B567862	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFCF7434FF2318A99E6D7AD0.text	D06EF633FFCF7434FF2318A99E6D7AD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites montanus Mocsary 1906	<div><p>Celonites montanus Mocsáry, 1906</p><p>(Figs 5E, 6E, 7C, H, 9F)</p><p>Celonites montanus Mocsáry, 1906: 198, ♂ (type locality: “Turkestan: Alai montes” [Kyrgyzstan]), lectotype (Fateryga &amp; Fadeev 2023: 568), ♂, HNHM (photographs examined).</p><p>Celonites kostylevi Panfilov, 1961: 165, ♀ ♂ (type locality: “КИРгИЗИЯ: ЗападнОЕ пОБЕРЕЖЬЕ ОЗ. Иссык-КУЛЬ, УРОчИЩЕ Ак-УЛЕн” [Kyrgyzstan, west coast of Issyk-Kul Lake, Ak-Ulen Place]), lectotype (Fateryga &amp; Fadeev 2023: 568), ♀, ZMMU (examined), synonymized by Fateryga &amp; Fadeev 2023: 565.</p><p>Celonites haemorrhoidalis Gusenleitner, 2012: 323, figs 11–13, ♀ (type locality: “Kyrghyzstan: Alai Mt. R. Katta-Karskoir [Katta-Karakol], 2550 m ” [Kyrgyzstan]), holotype, ♀, OLML (photographs examined), synonymized by Fateryga &amp; Fadeev 2023: 565.</p><p>Material examined. LECTOTYPE OF CELONITES MONTANUS (designated by Fateryga &amp; Fadeev (2023: 568)): ♂, labeled “ Alai mont. / 1905 Korb // Celonites / montanus / Mocs. // [blank, red label] // id nr. 130126 / HNHM / Hym. coll. [blue label]” [HNHM]. LECTOTYPE OF CELONITES KOSTYLEVI (designated by Fateryga &amp; Fadeev (2023: 568)): ♀, labeled “ ИссЫК-КУЛЬ / АК-УЛен [Issyk-Kul, Ak-Ulen] 1.VII.1953 / Д. ПанфиЛОв [D. Panfilov] // Holotypus ♀ 1961 / Celonites (s. str.) / kostylevi Panfilov [red label] // Lectotypus ♀ / Celonites kostylevi / Panfilov, 1961 / des. Fateryga, 2022 [red label]” [ZMMU]. HOLOTYPE OF CELONITES HAEMORRHOIDALIS: ♀, labeled “Kyrghyzstan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.36667&amp;materialsCitation.latitude=39.866665" title="Search Plazi for locations around (long 73.36667/lat 39.866665)">Alai Mt. R.</a> / Katta-Karakol r., 2550m / 39°52’N 73°22’E / Makogonova, 15.07.2000 // ♀ // Celonites / haemorrhoidalis nov. spec. ♀ / J.Gusenleitner,det.2011 / Holotypus // Biologiezentrum Linz, / Austria (LI) / ex Coll. S. Zonstein / Eingang 2001 // HOLOTYPUS [red label]” [OLML: ZOBODAT 2025a]. PARALECTOTYPES OF CELONITES KOSTYLEVI: KYRGYZSTAN: Issyk-Kul, Ak-Ulen, 28.VI.1953, 3 ♀, leg. D. Panfilov [ZMMU]; ibid., 29.VI.1953, 1 ♀, leg. D. Panfilov [ZMMU]; ibid., 1.VII.1953, 6 ♀, 3 ♂, leg. D. Panfilov [ZMMU]; ibid., 2.VII.1953, 2 ♀, leg. D. Panfilov [ZMMU]. ADDITIONAL MATERIAL: KYRGYZSTAN: Issyk-Kul, Ak-Ulen, 5.VII.1954, 1 ♀, leg. D. Panfilov [ZMMU]; Alai Mountain Range, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.36667&amp;materialsCitation.latitude=39.866665" title="Search Plazi for locations around (long 73.36667/lat 39.866665)">Katta-Karakol River</a>, 39°52′N, 73°22′E, 2550 m, 15.VII.2000, 22 ♀, leg. I. Makogonova [19 ♀, CSZT; 3 ♀, ZISP]; Kasgharia, vicinity of Irkeshtam [Erkeshtam], left bank of Kyzyl-Suu, Karavan-Kul Canyon, on Otostegia, 17.VII.2003, 3 ♀, leg. V. Dubatolov, D. Milko [ISEN]. KAZAKHSTAN. Almaty Province: 3 km NE Aktogay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=78.880005&amp;materialsCitation.latitude=43.243057" title="Search Plazi for locations around (long 78.880005/lat 43.243057)">Charyn River</a> canyon, 43°14′35″N, 78°52′48″E, 1160 m, 25.V.2016, on Lagochilus, 14 ♀, 2 ♂, leg. K. Fadeev [2 ♀, 1 ♂, CAFK; 12 ♀, 1 ♂, ZISP]; 14 km NE Aktogay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=78.991104&amp;materialsCitation.latitude=43.295555" title="Search Plazi for locations around (long 78.991104/lat 43.295555)">Charyn River</a> canyon, 43°17′44″N, 78°59′28″E, 1006 m, 25.V.2016, 1 ♀, 2 ♂, leg. K. Fadeev [1 ♀, 1 ♂, CAFK; 1 ♂, ZISP]; Charyn National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=79.0811&amp;materialsCitation.latitude=43.3496" title="Search Plazi for locations around (long 79.0811/lat 43.3496)">Canyon</a>, 43.3496°N, 79.0811°E, on Convolvulus fruticosus, 13.V.2024, 1 ♂, leg. A. Fateryga [CAFK]; ibid., on Lagochilus diacanthophyllus, 14.V.2024, 2 ♂, leg. A. Fateryga [CAFK]; ibid., on Lagochilus diacanthophyllus, 16.V.2024, 4 ♀, leg. A. Fateryga [3 ♀, CAFK; 1 ♀, CVMM]; 6 km NNW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=78.6225&amp;materialsCitation.latitude=43.5017" title="Search Plazi for locations around (long 78.6225/lat 43.5017)">Kokpek</a>, 43.5017°N, 78.6225°E, 19.V.2024, 2 ♀, leg. A. Fateryga [CAFK, CVMM]. Jetisu Province: Altyn-Emel National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.8626&amp;materialsCitation.latitude=43.9163" title="Search Plazi for locations around (long 77.8626/lat 43.9163)">Taygak</a>, 43.9163°N, 77.8626°E, 7.V.2024, 2 ♂, leg. A. Fateryga [CVMM].</p><p>Distribution. Kyrgyzstan, Kazakhstan (Almaty and Jetisu provinces).</p><p>Trophic relationships. Celonites montanus is broadly oligolectic (sensu Müller &amp; Kuhlmann 2008) on Lamiaceae . Females take up pollen with pollen-collecting setae on the frontal surface of the head, by rubbing it over the nototribic anthers; rubbing movements are alternated with pollen brushing from the head with the foretarsi, from which the pollen is finally consumed with the mouthparts (Fateryga 2025a). Flower-visiting records: Convolvulus fruticosus Pall. ( Convolvulaceae) in Kazakhstan; Lagochilus diacanthophyllus (Pall.) Benth., L. platyacanthus Rupr., and Ziziphora tenuior L. (all Lamiaceae) in Kazakhstan; Moluccella olgae (Regel) Ryding (≡ Otostegia olgae (Regel) Korsh., Lamiaceae) in Kyrgyzstan (Fateryga &amp; Fadeev 2023; Fateryga 2025a).</p></div>	https://treatment.plazi.org/id/D06EF633FFCF7434FF2318A99E6D7AD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFC87437FF231B309F177FCC.text	D06EF633FFC87437FF231B309F177FCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites octoannulatus (Kuznetzov 1923)	<div><p>Celonites octoannulatus (Kuznetzov, 1923)</p><p>(Figs 5G, 6G, 7E, L, 8F)</p><p>Saryara octoannulata Kuznetzov, 1923: 127, ♀ ♂ (type locality: “Montes Ac-Tasch, distr. Taschkent, Turkestan. Alt 7500ʹ” [Uzbekistan]), lectotype (Fateryga 2025b: 211), ♀, ZMMU (examined).</p><p>Celonites octoannulatus: Kostylev 1935: 109.</p><p>Celonites octoannulatus hissaricus Kostylev, 1940: 137, ♀ ♂ (type locality: “Ruidacht, 40 km vers le N de Stalinabad (autrefois Duchambé). 3000 m. au-dessus du niv. de la m” [Tajikistan]), lectotype (Fateryga 2025b: 211), ♀, ZISP (examined), synonymized by Fateryga 2025b: 215.</p><p>Material examined. LECTOTYPE OF SARYARA OCTOANNULATA (designated by Fateryga (2025b: 211)): ♀, labeled “[golden disc] // Ак-Таш [ Ak-Tash] / 29/VI-22. 7000ʹ // Celonites / 8-annulatus / N Kuzn // ТИп [type] ♀ / Saryara 8- / annulata Kuz / = Celonites 8- / annulata / G Kostylev / 1941, 23 VI [Kostylev’s signature] // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Lectotype ♀ / Saryara octoannulata / Kuznetzov, 1923 / des. Fateryga, 2024 [red label]” [ZMMU]. LECTOTYPE OF CELONITES OCTOANNULATUS HISSARICUS (designated by Fateryga (2025b: 211)): ♀, labeled “РУЙдашт [Ruydasht] / 22-VII 37 / В. ГУссакОвск. [V. Gussakovskij] // C. octoannulatus / hissaricus Typ. ♀ / G Kostylev det. 1938 // к. ГУссакОвскОгО [collection of Gussakovskij] // Lectotype ♀ Celonites / octoannulatus hissaricus / Kostylev, 1940 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000325 [pale yellow label]” [ZISP]. PARALECTOTYPES OF SARYARA OCTOANNULATA: UZBEKISTAN: Ak-Tash, 7000ʹ, 29.VI.1922, 1 ♀, 1 ♂ [ZMMU]. PARALECTOTYPE OF CELONITES OCTOANNULATUS HISSARICUS: TAJIKISTAN: Ruydasht, 22.VII.1937, 1 ♂, leg. V. Gussakovskij [ZISP]. ADDITIONAL MATERIAL: TAJIKISTAN: Kvak, 17.VIII.1937, 1 ♀, leg. V. Gussakovskij [ZMMU] ; Ruydasht Place, 3000 m, 4.IX.1937, 2 ♀, leg. V. Gussakovskij [ZISP]; ibid., 20.VII.1938, 11 ♀, leg. V. Gussakovskij [ZISP]; Kok-Kul Place, 3000 m, 20–22.VIII.1940, 10 ♀, leg. V. Gussakovskij [ZISP] ; Shuroobod, 37°51ʹ07ʺN, 70°05ʹ22ʺE, 2290 m, 25–26.VI.2025, 1 ♀, leg. M. Proshchalykin [CAFK]. KYRGYZSTAN: Transalai Mountain Range, E part, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.833336&amp;materialsCitation.latitude=39.65" title="Search Plazi for locations around (long 73.833336/lat 39.65)">Kaltabulak River</a> gorge, 39°39′N, 73°50′E, 2950 m, 15.VII.1998, 1 ♀, 1 ♂, leg. I. Makogonova [CSZT] ; Kasgharia, lower reaches of Archalsu River (right inflow of Koksu River), 20.VII.2003, 1 ♀, leg. V. Dubatolov, E. Nikolayeva, V. Zinchenko, R. Dudko [ISEN].</p><p>Distribution. Tajikistan, Uzbekistan,?* Kyrgyzstan.</p><p>Trophic relationships. Popov (1948) reported this species collected in Tajikistan at flowers of Anchusa azurea ( Boraginaceae).</p><p>Remarks. Identification of the specimens from Kyrgyzstan is preliminary and requires further confirmation. They differ from specimens from both Tajikistan and Uzbekistan by completely black dorsal mesepisternum and scutellum (like in C. modestus). At the same time, specimens from Kyrgyzstan have completely orange antenna, similarly to those from Uzbekistan, while specimens of both C. octoannulatus and C. modestus from Tajikistan have the antenna always significantly darkened dorsally. Males of C. octoannulatus from Kyrgyzstan, as well as males of C. modestus should be collected and compared with those of C. octoannulatus from Tajikistan and Uzbekistan. The available male from Kyrgyzstan (from CSZT) is lacking the genitalia. When I received the material, female genitalia were glued to a piece of cartoon and pinned under this male specimen, while the female from the same series was without genitalia. It is obvious that the genitalia had been extracted by somebody from both specimens from CSZT but the male’s ones were lost and the female’s ones were pinned under the male specimen by a mistake.</p></div>	https://treatment.plazi.org/id/D06EF633FFC87437FF231B309F177FCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFCB7437FF231C4C98427A77.text	D06EF633FFCB7437FF231C4C98427A77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites osseus Morawitz 1888	<div><p>Celonites osseus Morawitz, 1888</p><p>(Figs 5A, 6A, 7A, I, 8A, 9D)</p><p>Celonites osseus Morawitz, 1888: 268, ♀ (type locality: “in territorio transcaspico (Tschikischljar)” [Turkmenistan]), lectotype (Mauss et al. 2022a: 113), ♀, ZISP (examined).</p><p>Material examined. LECTOTYPE (designated by Mauss et al. (2022a: 113)): ♀, labeled “[golden disc] // Tschikisch-/ ljar. Pom. [Pomeranzev] // к. Ф МОРавица [collection of F. Morawitz] // Celonites / osseus ♀. / F. Moraw. // Lectotypus ♀ / Celonites osseus / Morawitz / des. Fateryga, 2018 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000326 [pale yellow label]” [ZISP] . ADDITIONAL MATERIAL: TURKMENISTAN: Krasnovodsk [Türkmenbaşy], 13.VI.1928, 1 ♀, leg. V. Gussakovskij [ZISP] ; ibid., 16.VI.1928, 1 ♂, leg. V. Gussakovskij [ZISP] .</p><p>Additional literature records. TURKMENISTAN: Jebel, 25 km NW Balkanabat (Mauss et al. 2022a).</p><p>Distribution. Armenia, Azerbaijan (Nakhchivan AR), Iran, Turkmenistan.</p><p>Trophic relationships. Unknown.</p></div>	https://treatment.plazi.org/id/D06EF633FFCB7437FF231C4C98427A77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFCB7429FF231A959F127DD4.text	D06EF633FFCB7429FF231A959F127DD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites sibiricus Gusenleitner 2007	<div><p>Celonites sibiricus Gusenleitner, 2007</p><p>(Figs 5F, 6F, 7D, K, 9H)</p><p>Celonites sibiricus Gusenleitner, 2007: 133, figs 1, 2, ♀ ♂ (type locality: “ Russia, Sibiria, Altaya, Cevero-Chuyskiy [ Siberia, Altai, Severo-Chuyskiy Range], 50°05’ 87°46’, 2500m ”), holotype, ♂, in the collection of J.T. Smit, Duiven, The Netherlands (not examined).</p><p>Material examined. PARATYPE OF CELONITES SIBIRICUS: RUSSIA. Altai Republic: Kosh-Agach District, Aktru, S of Kuray, 50°05ʹN, 87°45ʹE, 2100 m, Malaise trap, 3–6.VII.2006, 1 ♂, leg. J. Smit, Th. Zeegers [OLML: ZOBODAT 2025b] . ADDITIONAL MATERIAL: RUSSIA. Altai Republic: Kosh-Agach District, middle raches of Tara River, 22.VII.2003, leg. L. Petrozhitskaya [ISEN]; Kosh-Agach District, upper reaches of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.7&amp;materialsCitation.latitude=50.1" title="Search Plazi for locations around (long 87.7/lat 50.1)">Aktru Glacier</a>, 50.1°N, 87.7°E, 2519 m, 6.VII.2006, 1 ♀, leg. A. Barkalov [ISEN]; Kosh-Agach District, lower reaches of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.74&amp;materialsCitation.latitude=50.07" title="Search Plazi for locations around (long 87.74/lat 50.07)">Aktru Glacier</a>, 50.07°N, 87.74°E, 2600 m, 14–15.VII.2008, 1 ♀, leg. A. Barkalov [ISEN]; 15 km SE of Kuray, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.11778&amp;materialsCitation.latitude=50.18611" title="Search Plazi for locations around (long 88.11778/lat 50.18611)">Chuya River</a>, 50°11′10″N, 88°07′04″E, 1600 m, 5.VII.2016, 1 ♀, leg. M. Proshchalykin, V. Loktionov [FSCV] ; ibid., 15.VI.2022, 2 ♂, leg. A. Fateryga [CAFK]; ibid., 15.VI.2022, 1 ♀, leg. M. Proshchalykin [CAFK]; ibid., 17.VI.2022, 6 ♀, leg. M. Proshchalykin [2 ♀, IBUM; 4 ♀, CAFK]; ibid., 21.VI.2022, 2 ♀, leg. M. Proshchalykin [AMNH, CAFK]; 2 km SE Chagan-Uzun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.42&amp;materialsCitation.latitude=50.07361" title="Search Plazi for locations around (long 88.42/lat 50.07361)">Balkhash River valley</a>, 9.VII.2016, 7 ♀, leg. M. Proshchalykin, V. Loktionov [FSCV]; 5 km SE Chagan-Uzun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.42&amp;materialsCitation.latitude=50.07361" title="Search Plazi for locations around (long 88.42/lat 50.07361)">Tydtuyaryk River valley</a>, 50°04′25″N, 88°25′12″E, 11.VII.2016, 5 ♀, leg. M. Proshchalykin, V. Loktionov [FSCV] ; ibid., 8.VII.2019, 4 ♂, leg. A. Fateryga [2 ♂, CAFK; 2 ♂, CVMM]; ibid., on Dracocephalum peregrinum, 8.VII.2019, 4 ♀, leg. A. Fateryga [3 ♀, CAFK; 1 ♀, CVMM]; ibid., 8–9.VII.2019, 4 ♀, leg. Yu. Danilov [ISEN]; ibid., 9.VII.2019, 3 ♀, leg. V. Dorofeev [ASUB]; ibid., 9.VII.2019, 1 ♂, leg. A. Fateryga [ZISP]; ibid., on Dracocephalum peregrinum, 9.VII.2019, 1 ♀, leg. A. Fateryga [ZISP] ; ibid., 10.VII.2019, 1 ♂, leg. A. Fateryga [CAFK]; ibid., 14.VI.2022, 1 ♀, leg. M. Proshchalykin [CAFK]; ibid., 18.VI.2022, 1 ♀, leg. M. Proshchalykin [CAFK]; ibid., 21.VI.2022, 2 ♀, leg. A. Fateryga [CVMM]; ibid., 21.VI.2022, 3 ♀, leg. M. Proshchalykin [CAFK]; ibid., 26.VI.2022, 1 ♂, leg. M. Proshchalykin [CAFK]; 12 km SE Aktash, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.270836&amp;materialsCitation.latitude=50.35861" title="Search Plazi for locations around (long 87.270836/lat 50.35861)">Chuya River valley</a>, 13.VII.2016, 1 ♀, leg. M. Proshchalykin, V. Loktionov [FSCV]; 24 km NWW Aktash, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.270836&amp;materialsCitation.latitude=50.35861" title="Search Plazi for locations around (long 87.270836/lat 50.35861)">Chuya River valley</a>, 50°21′31″N, 87°16′15″E, 6.VII.2019, 3 ♂, leg. A. Fateryga [2 ♂, IBUM; 1 ♂, CAFK]; ibid., on Dracocephalum nutans, 6.VII.2019, 3 ♀, 1 ♂, leg. A. Fateryga [2 ♀, CAFK; 1 ♀, 1 ♂, CVMM]; ibid., 6.VII.2019, 1 ♂, leg. V. Dorofeev [ASUB]; ibid., 7.VII.2019, 1 ♀, leg. V. Dorofeev [ASUB]; ibid., 7.VII.2019, 1 ♂, leg. A. Fateryga [CAFK]; ibid., on Dracocephalum peregrinum, 7.VII.2019, 2 ♀, leg. A. Fateryga [CAFK, CVMM]; “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.3125&amp;materialsCitation.latitude=50.06389" title="Search Plazi for locations around (long 88.3125/lat 50.06389)">Mars</a> ”, 50°03′50″N, 88°18′45″E, 17.VI.2022, 1 ♂, leg. A. Fateryga [CAFK] ; ibid., on Taraxacum sp., 17.VI.2022, 1 ♀, leg. A. Fateryga [CAFK] ; ibid., on Sisymbrium polymorphum, 17.VI.2022, 1 ♂, leg. A. Fateryga [CAFK] ; ibid., 17.VI.2022, 1 ♂, leg. M. Proshchalykin [AMNH]; ibid., on Dracocephalum peregrinum, 23.VI.2022, 1 ♂, leg. A. Fateryga [CAFK]; 5 km NE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=89.071945&amp;materialsCitation.latitude=49.95" title="Search Plazi for locations around (long 89.071945/lat 49.95)">Kokorya</a>, 49°57′00″N, 89°04′19″E, 24.VI.2022, 1 ♂, leg. A. Fateryga [CAFK] ; ibid., on Gypsophila patrinii, 26.VI.2022, 1 ♂, leg. M. Proshchalykin [CAFK] . KAZAKHSTAN. East Kazakhstan Province: Central Tarbagatay, 80 km S Aksuat, 13.VII.1986, 9 ♀, 1 ♂, leg. V. Kazenas [FSCV] .</p><p>Additional literature records. RUSSIA. Altai Republic: 3 km NE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=89.04583&amp;materialsCitation.latitude=49.941944" title="Search Plazi for locations around (long 89.04583/lat 49.941944)">Kokorya</a>, 49°56′31″N, 89°02′45″E (Fateryga et al. 2023).</p><p>Distribution. Russia (Altai Republic), Kazakhstan (East Kazakhstan Province).</p><p>Trophic relationships. Celonites sibiricus is broadly oligolectic(sensu Müller&amp;Kuhlmann2008),predominantly collecting pollen from flowers of Lamiaceae . Morphological and behavioral adaptations of C. sibiricus for pollen uptake from flowers were described in detail by Fateryga (2020) and Fateryga et al. (2023). Females take up pollen with pollen-collecting setae on the frontal surface of the head, by rubbing it over the nototribic anthers; rubbing movements are alternated with pollen brushing from the head with the foretarsi, from which the pollen is finally consumed with the mouthparts. Flower-visiting records (all in the Altai Republic of Russia): Taraxacum sp. ( Asteraceae); Sisymbrium polymorphum ( Brassicaceae); Dianthus chinensis L. and Gypsophila patrinii (both Caryophyllaceae); Geranium sibiricum L. ( Geraniaceae); Dracocephalum peregrinum, D. nutans L., and Ziziphora clinopodioides (all Lamiaceae); Potentilla sericea L. ( Rosaceae) (Fateryga 2020; Fateryga et al. 2023).</p></div>	https://treatment.plazi.org/id/D06EF633FFCB7429FF231A959F127DD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFD5742AFF231A349ADF7CC8.text	D06EF633FFD5742AFF231A349ADF7CC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites tristiculus Kostylev 1935	<div><p>Celonites tristiculus Kostylev, 1935</p><p>(Figs 5M, 6M, 7F, G, 8I, 9J)</p><p>Celonites tristiculus Kostylev, 1935: 111, ♀ (type locality: “ЧаРкУ (ФЕРгана)” [Uzbekistan, Fergana, Charku]), holotype, ♀, ZMMU (examined).</p><p>Celonites tristiculus karatauicus Kostylev, 1935: 111, ♀ ♂ (type locality: “КаРатаУ, ОкР. ДЖУЛЕка” [Kazakhstan, Karatau, vicinity of Dzhulek]), lectotype (Fateryga 2025b: 217), ♀, ZISP (examined), synonymized by Fateryga 2025b: 219.</p><p>Material examined. HOLOTYPE OF CELONITES TRISTICULUS: ♀, labeled “ЧаРкУ / КОканд. У. [Charku, Kokand Uezd] / 12/VII 1920 г. // Celonites ♀ Typ / tristiculus, n. sp / G. Kostylev det. 1932 // МОсКва / ЗООЛОГичесКий / МУЗей МГУ [Moscow, Zoological Museum MSU] // Holotype ♀ / Celonites tristiculus / Kostylev, 1935 [red label]” [ZMMU]. LECTOTYPE OF CELONITES TRISTICULUS KARATAUICUS (designated by Fateryga (2025b: 217)): ♀, labeled “[golden disc] // Karataugeb. / bei Djulek / Balamurun / Koshantschikow. / 23/V 1913 [on underside] // к. ВОЛЬмана [collection of Wollmann] // Celonites tristiculus / karatauicus n sp ssp / G. Kostylev det 1932 // Lectotype ♀ Celonites / tristiculus karatauicus / Kostylev, 1935 / des. Fateryga, 2024 [red label] // Zoological Institute / St. Petersburg / INS_HYM_0000328 [pale yellow label]” [ZISP]. PARALECTOTYPES OF CELONITES TRISTICULUS KARATAUICUS: KAZAKHSTAN. Kyzylorda Province: Karataugebiet, bei Djulek, Balamurun, 9.V.1913, 1 ♂, leg. Koshantschikow [ZISP] ; ibid., 16.V.1913, 3 ♀, leg. Koshantschikow [ZISP]; ibid., 18.V.1913, 4 ♀, leg. Koshantschikow [ZISP]; ibid., 21.V.1913, 2 ♀, leg. Koshantschikow [ZISP]; ibid., 23.V.1913, 9 ♀, leg. Koshantschikow [ZISP]; ibid., 24.V.1913, 1 ♀, leg. Koshantschikow [ZISP]; ibid., 25.V.1913, 2 ♀, leg. Koshantschikow [ZISP]; ibid., 26.V.1913, 1 ♀, leg. Koshantschikow [ZISP]; ibid., 28.V.1913, 1 ♀, leg. Koshantschikow [ZISP]; ibid., 3.VI.1913, 1 ♀, leg. Koshantschikow [ZISP]. ADDITIONAL MATERIAL: KAZAKHSTAN. Abai Province: 30 km SW Semipalatinsk [Semey], Mukur River, 4.VII.1958, 1 ♀, leg. D. Panfilov [ZMMU]. Almaty Province: Middle reaches of Charyn River, 7.VI.1968, 2 ♀, leg. V. Kazanas [FSCV]. Jetisu Province: Vicinity of Kopaly, 20.V.1938, 1 ♀, leg. B. Tsvetkov [ZMMU] ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.8626&amp;materialsCitation.latitude=43.9582" title="Search Plazi for locations around (long 77.8626/lat 43.9582)">Altyn-Emel National Park</a>, Shylbyr, 43.9582°N, 78.1869°E, 5.V.2024, 1 ♀, 2 ♂, leg. A. Fateryga [1 ♂, CAFK ; 1 ♀, 1 ♂, ZISP]; ibid., on Allium caricifolium, 5.V.2024, 1 ♀, leg. M. Proshchalykin [CAFK] ; ibid., on Nepeta micrantha, 5.V.2024, 1 ♀, leg. M. Proshchalykin [CAFK] ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.8626&amp;materialsCitation.latitude=43.9163" title="Search Plazi for locations around (long 77.8626/lat 43.9163)">Altyn-Emel National Park</a>, Taygak, 43.9163°N, 77.8626°E, 6.V.2024, 1 ♂, leg. A. Fateryga [CAFK] ; ibid., on Ziziphora tenuior, 6.V.2024, 1 ♂, leg. A. Fateryga [CAFK] ; ibid., 7.V.2024, 1 ♀, 1 ♂, leg. A. Fateryga [CVMM] .</p><p>Additional observations on iNaturalist. KAZAKHSTAN. Almaty Province: Iliysk District, 44.0844°N, 77.0096°E, on Orobanche amoena, 7.V.2025, 1 ♀, L. Abramova (https://www.inaturalist.org/ observations/280174094) .</p><p>Distribution. Uzbekistan, Kazakhstan ( Abai, Almaty, Jetisu, and Kyzylorda provinces).</p><p>Trophic relationships. Celonites tristiculus is probably polylectic or at most eclectically oligolectic. Females collect pollen (but not nectar) from flowers of Allium caricifolium Kar. &amp; Kir. ( Amaryllidaceae), consuming it directly from the anthers with the mouthparts and the foretarsi. At the same time, they collect nectar mainly from flowers of Ziziphora tenuior ( Lamiaceae), which are hardly used for pollen uptake. Flower-visiting records (all in Kazakhstan): Allium caricifolium ( Amaryllidaceae); Nepeta micrantha Bunge and Ziziphora tenuior (both Lamiaceae) (Fateryga 2025a); Orobanche amoena C.A. Mey. ( Orobanchaceae) (new record). The record of this species at O. amoena is apparently the first documented report of a pollen wasp visiting flowers of holoparasitic plants in the family Orobanchaceae . However, it was not documented whether C. tristiculus collected pollen or nectar (or both) at this plant. Although a few flowering individuals of O. amoena were present at the locality where the foraging behavior of C. tristiculus was studied (Fateryga 2025a), I did not observe this wasp species visiting them.</p></div>	https://treatment.plazi.org/id/D06EF633FFD5742AFF231A349ADF7CC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
D06EF633FFD7742DFF23183F9F947992.text	D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Celonites Latreille 1802	<div><p>Key to the species of Celonites Latreille, 1802 from Central Asia</p><p>1. Females......................................................................................... 2</p><p>- Males (unknown in C. modestus).................................................................... 15</p><p>2. Head in frontal view not wider than long, clypeus and frons with cell-like sculpture (Fig. 7C); setae on frons about two times longer than diameter of median ocellus (Fig. 7H); body black with orange pattern, without yellow or whitish markings............................................................................ C. montanus Mocsáry, 1906</p><p>- Head in frontal view at least 1.2× as wide as long, clypeus and frons with different sculpture, never cell-like (Figs 1D, 3C, 7A, B, D–G); setae on frons at most as long as diameter of median ocellus (Fig. 7K, L) or shorter (Figs 3G, 7I, J, M); body usually with yellow or whitish markings (except C. kozlovi)................................................ 3</p><p>3. Gena very narrow, much narrower than diameter of lateral ocellus (Fig. 7I); head 1.4–1.6× as wide as long, with at least some yellow markings (Fig. 7A, B); axilla of scutellum with distinct lateral projection lying on tegula (Fig. 8A, B); outline of emargination between lateral lamella and postero-lateral process of propodeum apically enlarged (Fig. 8G)........ 4</p><p>- Width of gena in front of ocular sinus about as diameter of lateral ocellus or sometimes wider (Figs 3G, 7J–M); head 1.2–1.3× as wide as long, usually without pale markings or at most with very small spot on clypeus (Figs 1D, 3C, 7D, E, F); axilla of scutellum usually with less distinct or even obsolete lateral projection lying on tegula (Figs 3H, 8D–F) (except C. hystrix, Fig. 8C); outline of emargination between lateral lamella and postero-lateral process of propodeum usually not so enlarged apically (Figs 3I, 8I) (except C. foveolatus, Fig. 8H)............................................ 6</p><p>4. Apical margins of terga II–V strongly crenulate (Fig. 9A).......................... C. crenulatus Morawitz, 1888</p><p>- Apical margins of terga II–V at most weakly crenulate (Fig. 9B)............................................ 5</p><p>5. Scutum shining, with distinct smoothened interspaces between punctures (Fig. 7B)....................................................................................... C. kurzenkoi Mauss, Fateryga &amp; Carpenter, 2026</p><p>- Scutum duller, with very narrow, edge-like interspaces between punctures (Fig. 7A)......... C. osseus Morawitz, 1888</p><p>6. Disc of clypeus shallowly concave; mesosoma and metasomal terga very coarsely punctate, scutum shining, with distinct smoothened interspaces between punctures (Fig. 8C); axilla of scutellum with distinct lateral projection lying on tegula (Fig. 8C); apical margins of terga II–V strongly crenulate (see Fig. 9A) and deflected upwards....... C. hystrix Kostylev, 1940</p><p>- Clypeus uniformly convex; mesosoma and metasomal terga less coarsely punctate, scutum duller, with either reticulately microsculptured or very narrow, edge-like interspaces between punctures (Figs 3H, 8D–F); axilla of scutellum with less distinct or even obsolete lateral projection lying on tegula (Figs 3H, 8D–F); apical margins of terga II–V usually not crenulate or at most weakly crenulate (Figs 1C, 3F, 9C) and never deflected upwards............................ 7</p><p>7. Outline of emargination between lateral lamella and postero-lateral process of propodeum apically enlarged (Fig. 8H); scutum with distinct interspaces between punctures (Fig. 8D); sternum VI shining, with smooth interspaces between macropunctures (Fig. 9I)..................................................... C. foveolatus Kostylev, 1935</p><p>- Outline of emargination between lateral lamella and postero-lateral process of propodeum not so enlarged apically (Figs 3I, 8I); scutum with narrower interspaces between punctures (Figs 3H, 8E, F); sternum VI usually duller, with more or less reticulately microsculptured interspaces between macropunctures (Fig. 1F) (except C. tristiculus, Fig. 8J), sometimes also with additional micropunctures (Figs 3E, 9H)........................................................... 8</p><p>8. Frons covered with setae approximately as long as diameter of median ocellus, forming pollen brush (Fig. 7K, L); sternum VI with reticulately microsculptured interspaces and numerous micropunctures between macropunctures (Fig. 9H).... 9</p><p>- Frons covered mostly with setae less in length than diameter of median ocellus, not forming pollen brush (Fig 3G, 7M); sternum VI with reticulately microsculptured or smoothened interspaces and with few or without micropunctures between macropunctures (Figs 1F, 3E, 8J).................................................................... 11</p><p>9. Setae on frons stiff and spatula-like apically (Fig. 7K); clypeus and frons regularly sparsely punctate, interspaces exceeding puncture diameter, dull, distinctly reticulately microsculptured (Fig. 7D).............. C. sibiricus Gusenleitner, 2007</p><p>- Setae on frons thin and pointed apically (Fig. 7K); clypeus and frons densely rugosely punctate, interspaces less than puncture diameter, rather shining, less distinctly microsculptured (Fig. 7E)................................... 10</p><p>10. Punctation on scutellum somewhat smoothened (Fig. 8E); scutellum and dorsal mesepisternum always black; flagellum of antenna always dark dorsally.................................................. C. modestus Kostylev, 1935</p><p>- Punctation on scutellum more distinct (Fig. 8F); scutellum and dorsal mesepisternum usually with yellow markings; flagellum of antenna sometimes completely orange.......................... C. octoannulatus (Kuznetzov, 1923)</p><p>11. Setae on frons about as long as 2/3 of diameter of median ocellus (Fig. 3G); sternum VI with few micropunctures on reticulately microsculptured interspaces between macropunctures (Fig. 3E)................................... 12</p><p>- Setae on frons about as long as 1/3 of diameter of median ocellus (Fig. 8M); sternum VI usually without micropunctures on reticulately microsculptured or smoothened interspaces between macropunctures (Figs 1F, 9J) (except C. kozlovi)..... 13</p><p>12. Metasomal terga I–V with twice-interrupted whitish apical bands, whitish color outlined by ferruginous (Fig. 9C)............................................................................. C. biinterruptus Kostylev, 1940</p><p>- Metasomal terga I–III with complete yellow apical bands, terga IV–V with apical yellow spots medially (Fig. 3B, F)....................................................................... C. mediasiaticus Fateryga, sp. nov.</p><p>13. Body black with orange pattern, without yellow or whitish markings..................... C. kozlovi Kostylev, 1935</p><p>- Body black with whitish-yellow pattern, whitish-yellow color often outlined by ferruginous..................... 14</p><p>14. Frons regularly sparsely punctate, interspaces exceeding puncture diameter, dull, distinctly reticulately microsculptured (Fig. 1D); sternum VI with less coarse macropunctures and reticulately micropunctured interspaces between them (Fig. 1F)....................................................................... C. maussi Fateryga, sp. nov.</p><p>- Frons densely rugosely punctate, interspaces less than puncture diameter, rather shining, less distinctly microsculptured (Fig. 7F); sternum VI with coarser macropunctures and smoothened interspaces between them (Fig. 9J)....................................................................................... C. tristiculus Kostylev, 1935</p><p>15. Sternum VIII either acutely produced, running into two tapering lobes at posterior end with deep median incision between them (Fig. 5A), or roundly produced apically (Fig. 5B, C); gena very narrow, much narrower than diameter of lateral ocellus; head 1.4–1.6× as wide as long, frons with yellow markings; axilla of scutellum with distinct lateral projection lying on tegula (Fig. 8A, B); outline of emargination between lateral lamella and postero-lateral process of propodeum apically enlarged (see Fig. 8G)............................................................................. 16</p><p>- Sternum VIII broadly emarginate to truncate apically, with two short acute processes at center (Fig. 5D, F, J, M) or without them (Fig. 5E, G–I, K, L); width of gena in front of ocular sinus about as diameter of lateral ocellus or sometimes wider; head about 1.3× as wide as long, frons and sometimes clypeus black (Figs 2C, 4D, 7G); axilla of scutellum usually with less distinct or even obsolete lateral projection lying on tegula (Figs 3H, 8D–F) (except C. hystrix, Fig. 8C); outline of emargination between lateral lamella and postero-lateral process of propodeum usually not so enlarged apically (see Figs 3I, 8I) (except C. foveolatus, see Fig. 8H)............................................................. 18</p><p>16. Sternum VIII acutely produced, running into two tapering lobes at posterior end with deep median incision between them (Fig. 5A); lateral margin of harpide slightly emarginate (Fig. 6A); tergum VII trilobed, median lobe rounded apically (Fig. 9D); scutum duller, with very narrow, edge-like interspaces between punctures (Fig. 8A)..... C. osseus Morawitz, 1888</p><p>- Sternum VIII roundly produced apically (Fig. 5B, C); lateral margin of harpide different, not emarginate (Fig. 6B, C); tergum VII with median lobe truncate (Fig. 9F) to even emarginate apically; scutum shining, with distinct smoothened interspaces between punctures (Fig. 8B).............................................................. 17</p><p>17. Harpide truncate apically, volsella with straight oblique postero-lateral margin (Fig. 6B); apical margins of terga II–VI strongly crenulate (see Fig. 9A); tergum VII with median lobe truncate (Fig. 9F)....... C. crenulatus Morawitz, 1888</p><p>- Harpide rounded apically, volsella with distinct more or less parallel-sided posterior process (Fig. 6C); apical margins of terga II–VI at most weakly crenulate (see Fig. 9B); tergum VII rather quadrilobed due to more or less deeply emarginate median lobe............................................... C. kurzenkoi Mauss, Fateryga &amp; Carpenter, 2026</p><p>18. Mesosoma and metasoma black with orange pattern, without yellow or whitish markings; clypeus with at least small orange-yellow spot............................................................................... 19</p><p>- Mesosoma and metasoma with at least some yellow or whitish markings, whitish color often outlined by ferruginous; coloration of clypeus variable....................................................................... 20</p><p>19. Setae on frons about two times longer than diameter of median ocellus (see Fig. 7H); tergum VII strongly quadrilobed due to deeply emarginate median lobe (Fig. 9F); apical margin of sternum VIII without short acute processes at center (Fig. 5E); genitalia strongly broadened posteriorly, with harpide directed postero-laterally and terminated by small triangular lobe (Fig. 6E).................................................................. C. montanus Mocsáry, 1906</p><p>- Setae on frons shorter than diameter of median ocellus (see Fig. 7M); tergum VII trilobed, with median lobe at most truncate (Fig. 9G); apical margin of sternum VIII with two short acute processes at center (Fig. 5J); genitalia hardly broadened posteriorly, with harpide directed posteriorly, rounded postero-medially, and with small triangular lobe postero-laterally (Fig. 6J).............................................................. C. kozlovi Kostylev, 1935</p><p>20. Clypeus with at least small yellow or whitish spot....................................................... 21</p><p>- Clypeus completely black (Figs 2C, 4D, 7G)........................................................... 23</p><p>21. Setae on frons shorter than diameter of median ocellus (see Fig. 7I, J, M); outline of emargination between lateral lamella and postero-lateral process of propodeum enlarged apically (see Fig. 8H); apical margin of harpide with lateral lobe shorter than medial rounded end, volsella somewhat produced medially, with posterior margin rounded (Fig. 6K)..................................................................................... C. foveolatus Kostylev, 1935</p><p>- Setae on frons about as long as diameter of median ocellus (see Fig. 7K, L) or even somewhat longer; outline of emargination between lateral lamella and postero-lateral process of propodeum not so enlarged apically (see Figs 3I, 8I); apical margin of harpide with lateral lobe not shorter than medial end, volsella not produced medially, either triangularly produced posteriorly (Fig. 6F) or with straight posterior margin (Fig. 6G)............................................ 22</p><p>22. Sternum VII+VIII longer, its apical margin with two short fused acute processes (Fig. 5F); genitalia longer, harpide terminated by small triangular lobe, volsella triangularly produced posteriorly (Fig. 6F)................................................................................................... C. sibiricus Gusenleitner, 2007</p><p>- Sternum VII+VIII shorter, its apical margin without short acute processes (Fig. 5G); genitalia shorter, harpide terminated by two equal lobes separated by deep rounded emargination, volsella with straight posterior margin (Fig. 6G)............................................................................ C. octoannulatus (Kuznetzov, 1923)</p><p>23. Disc of clypeus shallowly concave; mesosoma and metasomal terga very coarsely punctate, scutum shining, with distinct smoothened interspaces between punctures (Fig. 8C); axilla of scutellum with distinct lateral projection lying on tegula (Fig. 8C); apical margins of terga II–VI strongly crenulate (see Fig. 9A) and deflected upwards; sternum VII+VIII long (Fig. 5D); genitalia long, harpide nearly rounded apically, with indistinct postero-lateral lobe, volsella strongly produced posteriorly (Fig. 6D)........................................................... C. hystrix Kostylev, 1940</p><p>- Clypeus uniformly convex; mesosoma and metasomal terga less coarsely punctate, scutum duller, with very narrow, edge-like interspaces between punctures (Fig. 3H); axilla of scutellum with less distinct or even obsolete lateral projection lying on tegula (Fig. 3H); apical margins of terga II–V usually not crenulate or at most weakly crenulate (Figs 2D, 4D) and never deflected upwards; sternum VII+VIII shorter (Fig. 5H, I, L, M); genitalia shorter, harpide emarginate apically, often with distinct postero-lateral lobe, volsella less produced or not produced posteriorly (Fig. 6H, I, L, M)................. 24</p><p>24. Setae on frons reaching in length diameter of median ocellus (Fig. 4C); apical margin of sternum VIII distinctly emarginate (Fig. 5H, I); genitalia somewhat longer, volsellae situated at about their half-length, aedeagus narrow (Fig. 6H, I).... 25</p><p>- Setae on frons shorter than diameter of median ocellus (see Fig. 7M); apical margin of sternum VIII rather truncate (Fig. 5L, M); genitalia shorter, volsellae situated strongly posterior to their half-length, aedeagus broader (Fig. 6L, M)..... 26</p><p>25. Metasomal terga I–VI with twice-interrupted whitish apical bands, whitish color outlined by ferruginous (see Fig. 9C); sternum VII+VIII shorter (Fig. 5H); harpide narrower, aedeagus narrowing towards apex, volsella with strongly produced posterior margin (Fig. 6H)................................................. C. biinterruptus Kostylev, 1940</p><p>- Metasomal terga I–III or I–IV with complete yellow apical bands, terga IV–VI or V–VI with apical yellow spots medially (Fig. 4B, E); sternum VII+VIII longer (Fig. 5I); harpide broader, aedeagus rather parallel-sided, volsella with less produced posterior margin (Fig. 6I)............................................... C. mediasiaticus Fateryga, sp. nov.</p><p>26. Frons regularly sparsely punctate, interspaces exceeding puncture diameter, dull, distinctly reticulately microsculptured (Fig. 2C); apical margin of sternum VIII without distinct short acute processes (Fig. 5L); harpide with distinctly produced postero-lateral lobe, volsella with straight posterior margin (Fig. 6L)................... C. maussi Fateryga, sp. nov.</p><p>- Frons densely rugosely punctate, interspaces less than puncture diameter, rather shining, less distinctly microsculptured (Fig. 7G); apical margin of sternum VIII with two distinct short acute processes (Fig. 5M); harpide with postero-lateral lobe not so produced, volsella with roundly produced posterior margin (Fig. 6M)............. C. tristiculus Kostylev, 1935</p></div>	https://treatment.plazi.org/id/D06EF633FFD7742DFF23183F9F947992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Fateryga, Alexander V.	Fateryga, Alexander V. (2026): The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802. Zootaxa 5828 (2): 277-302, DOI: 10.11646/zootaxa.5828.2.5, URL: https://doi.org/10.11646/zootaxa.5828.2.5
