taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
D06EF633FFC6743AFF231D3A9B427B42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC6743AFF231D3A9B427B42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC6743AFF231D3A9B427B42.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC6743BFF231BC698087CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC6743BFF231BC698087CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC6743BFF231BC698087CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231D1498427B57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231D1498427B57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231D1498427B57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231D1498427B57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231BB59B4279D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231BB59B4279D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231BB59B4279D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC7743BFF231BB59B4279D2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743CFF231E9D99AA7852.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743CFF231E9D99AA7852.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743CFF231E9D99AA7852.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743CFF231E9D99AA7852.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743DFF2318B298467CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743DFF2318B298467CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743DFF2318B298467CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743DFF2318B298467CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC0743DFF2318B298467CF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142304/files/figure.png	https://doi.org/10.5281/zenodo.21142304	FIGURE 1. Celonites maussi Fateryga, sp. nov., ♀, holotype. A, habitus in dorsal view; B, habitus in lateral view; C, terga III–VI in dorsal view; D, head in frontal view; E, labels; F, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 1. Celonites maussi Fateryga, sp. nov., ♀, holotype. A, habitus in dorsal view; B, habitus in lateral view; C, terga III–VI in dorsal view; D, head in frontal view; E, labels; F, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142306/files/figure.png	https://doi.org/10.5281/zenodo.21142306	FIGURE 2. Celonites maussi Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, terga III–VII in dorsal view. Scale bars 1 mm.	FIGURE 2. Celonites maussi Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, terga III–VII in dorsal view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC17430FF231D1498427CBC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCC7433FF231CDC98427D2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142310/files/figure.png	https://doi.org/10.5281/zenodo.21142310	FIGURE 3. Celonites mediasiaticus Fateryga, sp. nov. A, B, D, E, ♀, holotype; C, F–I, ♀, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, labels; E, sternum VI in ventral view; F, terga III–VI in dorsal view; G, head in lateral view; H, left tegula and part of scutum in dorsal view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 3. Celonites mediasiaticus Fateryga, sp. nov. A, B, D, E, ♀, holotype; C, F–I, ♀, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, labels; E, sternum VI in ventral view; F, terga III–VI in dorsal view; G, head in lateral view; H, left tegula and part of scutum in dorsal view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCC7433FF231CDC98427D2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142312/files/figure.png	https://doi.org/10.5281/zenodo.21142312	FIGURE 4. Celonites mediasiaticus Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in lateral view; D, head in frontal view; E, terga III–VII in dorsal view. Scale bars 1 mm.	FIGURE 4. Celonites mediasiaticus Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in lateral view; D, head in frontal view; E, terga III–VII in dorsal view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCC7433FF231CDC98427D2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCC7433FF231CDC98427D2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCC7433FF231CDC98427D2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCF7433FF231D549B567862.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCF7434FF2318A99E6D7AD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCF7434FF2318A99E6D7AD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCF7434FF2318A99E6D7AD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCF7434FF2318A99E6D7AD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC87437FF231B309F177FCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC87437FF231B309F177FCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC87437FF231B309F177FCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFC87437FF231B309F177FCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7437FF231C4C98427A77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7437FF231C4C98427A77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7437FF231C4C98427A77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7437FF231C4C98427A77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7437FF231C4C98427A77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7429FF231A959F127DD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7429FF231A959F127DD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7429FF231A959F127DD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFCB7429FF231A959F127DD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD5742AFF231A349ADF7CC8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD5742AFF231A349ADF7CC8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD5742AFF231A349ADF7CC8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD5742AFF231A349ADF7CC8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD5742AFF231A349ADF7CC8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142322/files/figure.png	https://doi.org/10.5281/zenodo.21142322	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	FIGURE 7. Celonites spp. A, I, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, H, C. montanus Mocsáry, 1906, ♀, Kazakhstan; D, K, C. sibiricus Gusenleitner, 2007, ♀, Russia; E, L, C. octoannulatus (Kuznetzov, 1923), ♀, Tajikistan; F, C. tristiculus Kostylev, 1935, ♀, Kazakhstan; G, C. tristiculus, ♂, Kazakhstan; J, C. hystrix Kostylev, 1940, ♀, paralectotype; M, C. kozlovi Kostylev, 1935, ♀, Russia. A–G, head in frontal view; H–M, head in lateral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142304/files/figure.png	https://doi.org/10.5281/zenodo.21142304	FIGURE 1. Celonites maussi Fateryga, sp. nov., ♀, holotype. A, habitus in dorsal view; B, habitus in lateral view; C, terga III–VI in dorsal view; D, head in frontal view; E, labels; F, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 1. Celonites maussi Fateryga, sp. nov., ♀, holotype. A, habitus in dorsal view; B, habitus in lateral view; C, terga III–VI in dorsal view; D, head in frontal view; E, labels; F, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142310/files/figure.png	https://doi.org/10.5281/zenodo.21142310	FIGURE 3. Celonites mediasiaticus Fateryga, sp. nov. A, B, D, E, ♀, holotype; C, F–I, ♀, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, labels; E, sternum VI in ventral view; F, terga III–VI in dorsal view; G, head in lateral view; H, left tegula and part of scutum in dorsal view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 3. Celonites mediasiaticus Fateryga, sp. nov. A, B, D, E, ♀, holotype; C, F–I, ♀, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, labels; E, sternum VI in ventral view; F, terga III–VI in dorsal view; G, head in lateral view; H, left tegula and part of scutum in dorsal view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142326/files/figure.png	https://doi.org/10.5281/zenodo.21142326	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	FIGURE 8. Celonites spp. A, C. osseus Morawitz, 1888, ♀, Armenia; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. hystrix Kostylev, 1940, ♂, Uzbekistan; D, H, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; E, C. modestus Kostylev,1935, ♀, Tajikistan;F, C. octoannulatus(Kuznetzov, 1923), ♀, Tajikistan; G, C. kurzenkoi, ♀, paratype;I, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–D, left tegula and part of scutum in dorsal view; E, F, posterior part of scutum, tegulae, and scutellum in dorsal view; G, H, right part of propodeum in dorso-posterior view; I, left part of propodeum in dorso-posterior view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142328/files/figure.png	https://doi.org/10.5281/zenodo.21142328	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	FIGURE 9. Celonites spp. A, C. crenulatus Morawitz, 1888, ♀, Turkmenistan; B, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, ♀, holotype; C, C. biinterruptus Kostylev, 1940, ♀, paralectotype; D, C. osseus Morawitz, 1888, ♂, Iran; E, C. crenulatus, ♂, Turkmenistan; F, C. montanus Mocsáry, 1906, ♂, Kazakhstan; G, C. kozlovi Kostylev, 1935, ♂, Russia; H, C. sibiricus Gusenleitner, 2007, ♀, Russia; I, C. foveolatus Kostylev, 1935, ♀, Turkmenistan; J, C. tristiculus Kostylev, 1935, ♀, Kazakhstan. A–C, terga II–VI in dorsal view; D–G, terga IV–VII in dorsal view; H–J, sternum VI in ventral view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142314/files/figure.png	https://doi.org/10.5281/zenodo.21142314	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 5. Male sternum VII+VIII in ventral view. A, Celonites osseus Morawitz, 1888; B, C. crenulatus Morawitz, 1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142306/files/figure.png	https://doi.org/10.5281/zenodo.21142306	FIGURE 2. Celonites maussi Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, terga III–VII in dorsal view. Scale bars 1 mm.	FIGURE 2. Celonites maussi Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in frontal view; D, terga III–VII in dorsal view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142312/files/figure.png	https://doi.org/10.5281/zenodo.21142312	FIGURE 4. Celonites mediasiaticus Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in lateral view; D, head in frontal view; E, terga III–VII in dorsal view. Scale bars 1 mm.	FIGURE 4. Celonites mediasiaticus Fateryga, sp. nov., ♂, paratype. A, habitus in dorsal view; B, habitus in lateral view; C, head in lateral view; D, head in frontal view; E, terga III–VII in dorsal view. Scale bars 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
D06EF633FFD7742DFF23183F9F947992.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21142320/files/figure.png	https://doi.org/10.5281/zenodo.21142320	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	FIGURE 6. Malegenitalia in dorsal view.A, Celonitesosseus Morawitz, 1888; B, C. crenulatus Morawitz,1888; C, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026; D, C. hystrix Kostylev, 1940; E, C. montanus Mocsáry, 1906; F, C. sibiricus Gusenleitner, 2007; G, C. octoannulatus (Kuznetzov, 1923); H, C. biinterruptus Kostylev, 1940; I, C. mediasiaticus Fateryga, sp. nov.; J, C. kozlovi Kostylev, 1935; K, C. foveolatus Kostylev, 1935; L, C. maussi Fateryga, sp. nov.; M, C. tristiculus Kostylev, 1935. Scale bar 1 mm.	2026-06-09	Fateryga, Alexander V.		Zenodo	biologists	Fateryga, Alexander V.			
