taxonID	type	description	language	source
D1130013FFA21E77FF7BFC13FB9AE244.taxon	materials_examined	Type species. Orthocis aterrima Casey, 1898 (= Cis punctatus Mellie, 1848)	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA21E74FF7BFBB6FC60E46A.taxon	description	(Fig. 1)	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA21E74FF7BFBB6FC60E46A.taxon	materials_examined	Type material. Holotype: NHMD 153507 with label ' Ciidae, C. V. Hennigsen / 1 - 10 - 1964 ', Danish amber, late Eocene.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA21E74FF7BFBB6FC60E46A.taxon	etymology	Etymology. The epithet of this new species is dedicated to our friend and colleague A. V. Kompantsev (1952 – 2022) who studied Ciidae.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA21E74FF7BFBB6FC60E46A.taxon	description	Description. Body black-brownish, oblong-oval, convex dorsally, glabrous. Clypeus without teeth. Forehead convex, wide, finely punctate. Eyes well protruded, coarsely faceted, 0.7 times as long as wide. Antennae clavate, extend beyond middle of pronotum, ten-segmented. Antennomeres first and second suboval. First antennomere about 1.4 times as long as wide at apex. Second antennomere about 1.2 times as long as wide at apex, about 0.8 times as long as and of same width as antennomere one. Third antennomere long-oval, about 2.5 times as long as wide at apex, slightly longer than and 0.5 times as wide as antennomere two. Fourth antennomere oval, about 1.6 times as long as wide at apex, about 0.6 times as long as and about as wide as antennomere three. Antennomeres four-nine suboval, transverse. Fifth antennomere about 0.9 times as long as wide at apex, about 0.7 times as long as and 1.3 times as wide as antennomere four. Sixth antennomere about 0.6 times as long as wide at apex, slightly shorter than and about 1.2 times as wide as antennomere five. Seventh antennomere about 0.5 times as long as wide at apex, about of same length as and slightly wider than antennomere six. Antennal club consists of antennomeres eight – ten. Eighth antennomere about 0.6 times as long as wide at apex, about 1.6 times as long as and 1.5 times as wide as antennomere seven. Ninth antennomere about 0.6 times as long as wide at apex, about 1.3 times as long as and 1.3 times as wide as antennomere eight. Tenth antennomere cordate, about 1.2 times as long as wide at base, about twice as long as and of same width as antennomere nine. Pronotum almost trapezoid, of same length as wide across apex, about 0.9 times as long as wide across middle and across base. Lateral margins narrow, slightly explanate, visible for their entire lengths from above, lacking bristles. Disc with large and small punctures. Scutellum small, subpentagonal. Elytra oval, with lateral margins subparallel at basal three-fourths, then converging and forming moderately acute apex, widest at middle, 1.8 times as long as wide across base, about 1.6 times as long as wide across middle and across apical fourth, about 2.2 times as long as pronotum, with indistinct humeri. Elytral striae absent. Metaventrite weakly convex, about 2.8 times as long as length of mesocoxal cavity, sparsely punctate. Metepisternum narrow, about 11.3 times as long as its median width, with row of punctures. Abdomen convex. First ventrite about 1.7 times as long as metacoxal cavity. Second ventrite 0.6 times as long as first ventrite. Third ventrite about 0.8 times as long as second ventrite. Fourth ventrite about 1.1 times as long as third ventrite. Fifth ventrite subequal to ventrite four. Legs long. Femora clavate. Protibiae untoothed on the outer edge. Apex of protibiae simply rounded. Tarsi long, four-segmented. Last tarsomere very long. Body length: 1.1 mm. Comparison. The new species differs from European Orthocis linearis (J. Sahlberg, 1901) in its glabrous body, smaller body size and shorter antennomere 10.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA11E72FF7BF8D8FB98E2BE.taxon	description	(Fig. 2 - 3)	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA11E72FF7BF8D8FB98E2BE.taxon	materials_examined	Type material. Holotype: NHMD 153508 with label ' Ciidae, C. V. Hennigsen / 14 - 11 - 1959 ', Danish amber, late Eocene.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA11E72FF7BF8D8FB98E2BE.taxon	etymology	Etymology. The epithet of this new species is dedicated to outstanding hymenopterist Lars Vilhelmsen, keeper of NHMD amber collection.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA11E72FF7BF8D8FB98E2BE.taxon	description	Description. Body black, oblong-oval, convex dorsally, with bristles. Clypeus without teeth. Forehead convex, wide, finely punctate. Eyes well protruded, coarsely faceted, about 0.7 times as long as wide. Antennae clavate, extend beyond middle of pronotum, ten-segmented. Antennomeres first and second suboval. Third antennomere long-oval. Fourth antennomere oval, about 0.8 times as long as antennomere three. Antennomeres four-nine suboval, transverse. Antennal club consists of antennomeres eight – ten. Tenth antennomere about 1.5 times as long as antennomere nine. Pronotum almost trapezoid, of same length as wide across apex, about 0.8 times as long as wide across middle and about 0.7 times as long as wide across base. Lateral margins narrow, slightly explanate, visible for their entire lengths from above, lacking bristles. Disc with large and small punctures. Scutellum small, subpentagonal. Elytra oval, coarsely punctate, with lateral margins subparallel at basal three-fourths, then converging and forming moderately acute apex, widest at middle, about 2.2 times as long as wide across base, about 1.5 times as long as wide across middle, about 2.4 times as long as wide across apical fourth, about 3.3 times as long as pronotum, with indistinct humeri. Elytral striae absent. Metaventrite weakly convex, about 2.5 times as long as length of mesocoxal cavity, sparsely punctate. Metepisternum narrow, with row of punctures. Abdomen convex. Legs long. Femora clavate. Protibiae untoothed on the outer edge. Apex of protibiae simply rounded. Tarsi long, four-segmented. Last tarsomere very long. Body length: 1.3 mm. Comparison. The new species differs from Orthocis kompantsevi sp. nov. in having a bristly body, a densely punctate pronotum and elytra, and longer antennomere nine.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA71E71FF7BFAD2FC28E362.taxon	description	(Fig. 4)	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA71E71FF7BFAD2FC28E362.taxon	materials_examined	Type material. Holotype: NHMD 153503 with label ' Ciidae, A. Hennigsen / 9 - 9 - 1974 ', Danish amber, late Eocene. Syninclusion — Chthonioidea (Pseudoscorpiones).	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA71E71FF7BFAD2FC28E362.taxon	etymology	Etymology. The epithet of this new species is dedicated to Sven Gisle Larsson from Zoological Museum, University of Copenhagen (now NHMD), who published a first comprehensive overview of the European amber biota.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA71E71FF7BFAD2FC28E362.taxon	description	Description. Body black, oblong-oval, convex dorsally, glabrous. Clypeus without teeth. Forehead convex, wide. Eyes well protruded, coarsely faceted. Antennae clavate, probably ten-segmented. Antennomeres first and second oval. Third antennomere long-oval. Antennomeres four-nine suboval. Antennal club appears to consist of antennomeres eight – ten. Tenth antennomere cordate. Pronotum almost trapezoid, of same length as wide across apex and about 0.7 times as long as wide across base. Lateral margins narrow, slightly explanate, lacking bristles. Disc with densely subequal punctures. Scutellum small, subpentagonal. Elytra oval, densely and coarsely punctate, with lateral margins subparallel at basal three-fourths, widest at middle, about 1.6 times as long as wide across base, about 1.5 times as long as wide across middle, about 1.6 times as long as wide across apical fourth, about 1.6 times as long as pronotum, with indistinct humeri. Elytral striae absent. Metaventrite weakly convex, about 2.9 times as long as length of mesocoxal cavity, sparsely punctate. Metepisternum narrow, about 12.5 times as long as its median width, with row of punctures. Abdomen convex. First ventrite about 1.7 times as long as metacoxal cavity. Second ventrite about 0.5 times as long as first ventrite. Third ventrite about 0.9 times as long as second ventrite. Fourth ventrite about 0.8 times as long as third ventrite. Fifth ventrite subequal to ventrite four. Legs long. Femora clavate. Protibiae untoothed on the outer edge. Apex of protibiae simply rounded. Tarsi long, four-segmented. Last tarsomere very long. Body length: 1.8 mm. Comparison. The new species differs from Orthocis kompantsevi Legalov et Perkovsky, sp. nov. in its larger body size, simple punctate pronotum and coarser and more densely punctate elytra.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
D1130013FFA71E71FF7BFAD2FC28E362.taxon	discussion	Remark. Chthonioidea from fossil resins were epigean or endogean, so amber trap was situated on the base of the tree trunk (Turbanov et al. 2024; Turbanov, pers. com. 2024); spread wing indicates that beetle rested on the trunk. Discussion Goldsmith C. V. Hennigsen from Thisted, northwestern Jutland (sometimes abbreviation " C. V. " handwritten on the labels was mentioned in publicatons as " G. V. " or even " B. V. "), contributed not only two thirds of the Danish amber aphids, housed in NHMD until 1967 (Heie 1967), but a lion share of other Danish amber inclusions as well (e. g., Simutnik et al. 2020; Simutnik & Perkovsky 2023; Shavrin et al. 2023; Legalov et al. 2024 a; Lyubarsky et al. 2024 a, 2024 b; Perkovsky et al. 2024) including 60 % of ciid imagines. Mycophagous beetles from Danish amber were nearly unstudied in details (Larsson 1978) until the recent descriptions of two cryptophagids from the East African subgenus Neomicrambe (Lyubarsky et al. 2024 a, 2024 b), unknown from Baltic amber and a new mycetophagid genus (Legalov et al. 2024 b). Species of the Orthocis are known to feed and breed almost exclusively in Auricularia and Exidia basidioms (Lopes-Andrade 2010), and they are the only members of the Auricularia host-use group (Orledge & Reinolds 2005), so finding Orthocis in Danish amber is a first proxy for Auriculariaceae in the fossil record. Acknowledgements We thank Lars Vilhelmsen (NHMD) for the loan of the specimen and useful advices, Ilya S. Turbanov (Institute for Biology of Inland Waters, Yaroslavl Region, Russia) for Chthonioidea determination and anonymous reviewers for improving the overall quality of the manuscript. EEP’s research was supported by a “ Support and development of the amber collection of the Statens Naturhistoriske Museum (continuation) ” grant from Dr. Bøje Benzons Støttefond.	en	Legalov, Andrei A., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2024): Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber. Ecologica Montenegrina 80: 78-85, DOI: 10.37828/em.2024.80.7, URL: https://doi.org/10.37828/em.2024.80.7
