identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D14E87C9FF857F5EFF676A26FAA00BA1.text	D14E87C9FF857F5EFF676A26FAA00BA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cosmos harkerae Art. Castro & Aaron Rodr. A - B. Different 2025	<div><p>Cosmos harkerae Art. Castro &amp; Aarón Rodr., sp. nov. (Figures 1–2)</p><p>Cosmos harkerae resembles C. longipetiolatus and C. sherffii, but differs from them by its fusiform to subglobose tuberous roots 0.5–2.3 × 2–4.2 cm, petioles 0.6–5 cm long, opposite or alternate lateral pinnae segments of the leaves with the basal lobes often reclinate and pointing backward and resembling stipels, disk florets protected by the first paleae series and showing overall a tubular appearance, and light brown to reddish brown cypsela.</p><p>Type:— MEXICO. Guerrero: Olinalá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.849045&amp;materialsCitation.latitude=17.95574" title="Search Plazi for locations around (long -98.849045/lat 17.95574)">Xixila, km 19.5 de la carretera Papalutla-Xixila, cuenca del río Atoyac-Alto Balsas</a>, 1640 m, 17.955740ºN, - 98.849043ºW, 31 August 2017 (fl, fr), A. Castro-Castro, S.C. Saucedo-Heredia &amp; J. Esteban 4305 (holotype CIIDIR!; isotypes, FCME!, HUMO!, IBUG!, IEB!, MEXU!, OAX!, QMEX!, TEX!, UAGC!, XAL!) .</p><p>Perennial herbs, 50–112 cm, from a rhizome and tuberous roots, the roots 0.5–2.3 × 2–4.2 cm, fusiform to subglobose. Stems erect, simple, 19–53 cm, terete, sulcate, tomentose, sometimes glabrate on upper parts or old stems, trichomes, simple, multicellular. Leaves 4–8 pairs per stem, opposite, petiolate, membranaceous, lanceolate to deltate, rarely ovate in outline, 1.5–18 × 1–19 cm, 1–2 pinnatifid, 3–5(–7)-pinnate, tapering, acute to rounded, acuminate in the apex, margin antrorse spinulose-ciliate, somewhat thickened, lamina glabrous or appressed-hispid especially beneath and along the mid vein; petiole 0.6–5 cm, narrowly winged, channelled; central pinna 1–13-lobed, lanceolate or ovate even elliptic in outline, sometimes linear, the lobes entire or with one short secondary lobe, acuminate, 1.3–13 × 0.2–10 cm; lateral pinnae 1–9-lobed, lanceolate to elliptic, rarely linear in outline, the lobes entire or with one short secondary lobe, acuminate, opposite or alternate, 1–9.7 × 0.2–6.3 cm, the basal lobes often reclinate and pointing backward, resembling stipels. Capitulescence terminal or axillary, of 1–2-capitula, peduncles 17–45 cm, slender, glabrous. Capitula radiate, heterogamous, subhemispheric, 4–8 cm wide across the extended rays. Involucre 1.1–2.2 cm wide, 0.5–1.2 cm high, cup-shaped. Phyllaries biseriate; outer series 7–8, green, 4.3–10 × 0.8–2.8 mm, ensiform to oblong, ascending to inclined, glabrous, shiny, with 3–5 resiniferous ducts on the abaxial face, apices acute; inner series 6–8, translucent, pale yellow to light green, glabrous, conspicuously striped, 5.4–11 × 1.2–3 mm, ensiform to oblong, apex acute. Receptacle convex, paleaceous, 1.4–3 mm high, 4–6.8 mm wide. Paleae translucent yellow, 9.8–14.8 × 0.9–2.5 mm, persistent and resembling the inner phyllaries. Ray florets 8, uniseriate, sterile, limb obovate to elliptic, pale lilac, apex obtuse, entire to irregularly toothed, 1.9–3.9 × 0.5–1.6 cm, 3 ribbed outside, glabrous. Disk florets 34–46, hermaphrodite, 6.9–8.1 mm, 1.3–2 mm diameter of extended lobes, yellow, protected by the first paleae series, showing a tubular appearance; tube 1.5–3 × 0.6–0.7 mm, glabrous; throat 5.4–5.5 × 0.7–1.1 mm, glabrous; lobes 5, 0.8–1 mm long, erect to slightly involute, triangular, acute, margins ciliate, slightly revolute. Anthers 5, 1.8–2.3 mm long, apical appendages ovate, cylinder brown; pollen grain yellow; filaments 1.5–2 mm long, pubescent throughout. Style 5–5.8 mm long; stigma 0.9–1.3 mm long, lobes yellow, apices acute, penicillate, papillate beneath and along margins. Cypsela 6.9–19.1 × 0.8–2 mm, light brown to reddish brown, fusiform, tetrangular, with deep furrows between angles, carpopodium annular; pappus awns 2, 0.8–3.5 mm long, each with 6–14 retrorse barbs, often loosening.</p><p>Distribution, habitat and phenology: — Cosmos harkerae has been collected in northwestern Guerrero, at the transition zone between the Balsas Basin and the SMS, biogeographical provinces (Figure 3). It grows on northwestfacing slopes in oak forest, with deep, clayed and reddish soil. It is associated with Bessera elegans Schult. f., Begonia monophylla Pav. ex A. DC., Brongniartia sp., Cosmos purpureus (DC.) Benth. &amp; Hook. f. ex Hemsl., Euphorbia campestris S. Geuns, E. sphaerorhiza Benth., Eysenhardtia polystachya (Ortega) Sarg., Hypenia violacea Mart. Gord. &amp; S. Valencia, Mammillaria albilanata Backeb., Mesosphaerum suaveolens (L.) Kuntze, Mimosa sp., Pinguicula heterophylla Benth., Polianthes geminiflora (Lex.) Rose, Sessilanthera citrina Cruden, Vachellia pennatula (Schltdl. &amp; Cham.) Seigler &amp; Ebinger, Vitex mollis Kunth, and Zornia sp. The plants grow at elevations between 1290 and 1810 m. Cosmos harkerae flowers in August and early September, while fruiting occurs from September to December.</p><p>Etymology: —This is the second species of Cosmos named after a female botanist. The name C. steenisiae Veldkamp in Veldkamp &amp; Kreffer (1991: 479) was dedicated to Maria Johanna van Steenis-Kruseman. Now, we honor Professor Mollie F. Harker of the Institute of Botany at the Universidad de Guadalajara. Mollie advanced the knowledge of Mexican Asteraceae, and with dedication she trained a new generation of botanists.</p><p>Taxonomic relationships: — Cosmos harkerae is a geophyte growing from a short rhizome with tuberous roots, and therefore it belongs to C. sect. Discopoda . It is characterized by a combination of characters which include fusiform to subglobose tuberous roots, long petioles 0.6–5 cm long, opposite or alternate lateral pinnae segments of the leaves with the basal lobes often reclinate and pointing backward and resembling stipels, disk florets protected by the first paleae series and showing a tubular appearance, and light brown to reddish brown cypselae. Morphologically, C. harkerae shares similarities with C. longipetiolatus and C. sherffii . Table 1 and Figure 4 show a morphological comparison among these species. Furthermore, the key presented in the following lines allows the identification of the taxa within C. sect. Discopoda .</p><p>Based on the biogeographic regionalization by Morrone et al. (2017), all three species grow in the SMS (Figure 3). Cosmos longipetiolatus is endemic to the western end of the mountain range in the state of Jalisco, in a transition zone between the pine and oak forest and the cloud forest. Cosmos sherffii inhabits the Pinus - Quercus -Juniperus forest at the eastern portion of the SMS, in the state of Oaxaca. In contrast, C. harkerae is found in the central region of the SMS, on the slopes facing the Balsas Basin in the state of Guerrero. Here, a tropical Quercus forest develops (Figure 3). The TMVB and the SMS limit the Balsas Basin. Temperate Pinus and Quercus forest dominate the first two provinces, above the elevation of 1000 m. In contrast, the tropical dry forest covers the Balsas Basin. Morrone (2017) suggested a relationship among the SMS, the TMVB and the Balsas Basin along the contact zone.</p><p>Conservation assessment: — Cosmos harkerae has been collected in five locations close to each other and probably all form just one population with a broader geographic range (Figure 3). In all the sites explored, we found numerous plants. Based on the criteria of the IUCN (2024), it is proposed to include this species in the Critically Endangered category (CR/B1a), taking into consideration the values calculated for the EOO (10.716 km 2) and Endangered (EN/ B2a) considering the value of the AOO (20 km 2), both based on 2 × 2 km cells.</p><p>Most likely, Cosmos harkerae has a wider geographical range. A morphologically similar plant was identified on the citizen science platform iNaturalistMx. The observation was made in the municipality of Madero, Michoacán, almost 280 km west in a straight line from the type locality. The area falls in the transition zone to the Balsas Basin but now with the TMVB (Figure 3). It was not possible to visit the site, and we did not find herbarium specimens in the consulted herbaria.</p><p>Additional material examined (paratypes): — MEXICO. Guerrero: Olinalá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.847855&amp;materialsCitation.latitude=17.995115" title="Search Plazi for locations around (long -98.847855/lat 17.995115)">Papalutla, 14 km al SE de Papalutla, crucero Olinalá-Xixila-Papalutla</a>, 1290 m, 17.995115ºN, - 98.847852ºW, 7 September 1993 (fl), M. Luna-Flores 692 (FCME) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.88437&amp;materialsCitation.latitude=18.01704" title="Search Plazi for locations around (long -98.88437/lat 18.01704)">6.6 km al SE de Papalutla</a>, 1550 m, 18.017040ºN, - 98.884366ºW, 1 December 2006 (fr), O. Hinojosa-Espinosa 251 (FCME) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.853165&amp;materialsCitation.latitude=17.953444" title="Search Plazi for locations around (long -98.853165/lat 17.953444)">Xixila, crucero Xixila-Papalutla-Olinalá</a>, 1400 m, 17.953444ºN, - 98.853165ºW, 17 August 2007 (fl), O. Hinojosa-Espinosa 137 (FCME) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.853165&amp;materialsCitation.latitude=17.953444" title="Search Plazi for locations around (long -98.853165/lat 17.953444)">crucero Olinalá-Xixila-Papalutla</a>, 1400 m, 17.953444ºN, - 98.853165ºW, 21 August 1992 (fl), J. Calónico S. 232 (FCME) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.85027&amp;materialsCitation.latitude=17.94998" title="Search Plazi for locations around (long -98.85027/lat 17.94998)">km 21 de la carretera Papalutla-Xixila, laderas cercanas a la comunidad de Xixila</a>, 1810 m, 17.949980ºN, - 98.850276ºW, 31 August 2017 (fl, fr), A. Castro-Castro, S.C. Saucedo-Heredia &amp; J. Esteban 4311 (CIIDIR) .</p><p>Observation in iNaturalistMx: — MEXICO. Michoacán: Madero, southeast Las Mesas, 1965 m, 19.374373ºN, - 101.020338ºW, 8 September 2016 (fl), Acatecatl (https://mexico.inaturalist.org/observations/5038567).</p></div>	https://treatment.plazi.org/id/D14E87C9FF857F5EFF676A26FAA00BA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Castro-Castro, Arturo;Rodríguez, Aarón;Neri-Mondragón, Alberto Yahir;Siordia-González, Nadia Elizabeth	Castro-Castro, Arturo, Rodríguez, Aarón, Neri-Mondragón, Alberto Yahir, Siordia-González, Nadia Elizabeth (2025): A new species of Cosmos section Discopoda (Coreopsideae, Asteraceae) from Guerrero, Mexico. Phytotaxa 725 (1): 91-100, DOI: 10.11646/phytotaxa.725.1.8, URL: https://doi.org/10.11646/phytotaxa.725.1.8
D14E87C9FF827F50FF676AD4FA1D0C83.text	D14E87C9FF827F50FF676AD4FA1D0C83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cosmos (sect. Discopoda)	<div><p>Key to the species of Cosmos section Discopoda</p><p>1. Plants semiaquatic, from a rhizome with fibrous roots; leaves pinnatisect, segments simple or elliptic-lobed; outer phyllaries elliptic-obovate, obtuse; ray florets white; cypsela exaristate; from western Jalisco, Mexico .................................... C. mcvaughii</p><p>– Plants terrestrial, from a short rhizome with tuberous roots; leaves simple to 1–3-pinatisect, segments simple to 1–2-pinatisect; outer phyllaries deltoid, lanceolate, ensiform, trullate or penicilliform, acuminate; ray florets purple, violet or white; cypsela usually awned; ample distribution .................................................................................................................................................... 2</p><p>2. Plants scapose to subscapose; capitula 1(3–5); petioles 1.5–9 (15) cm, frequently longer than leaves ........................................... 3</p><p>– Plants leafy caulescent; capitula 3–6(–9); petioles 0.5–2(–4) cm, shorter than the leaves or completely sessile ............................ 7</p><p>3. External phyllaries 1.0–2.5 × 0.3–0.6 cm; from South America (Argentina, Bolivia and Peru) ......................... C. peucedanifolius</p><p>– External phyllaries 0.5–1.0 × 0.1–0.3 cm; from Mexico .................................................................................................................. 4</p><p>4. Disk and ray florets dark purple; from Guanajuato, Hidalgo, Querétaro and San Luis Potosí ............................ C. atrosanguineus</p><p>– Disk florets yellow, ray florets white, violet to light purple ............................................................................................................. 5</p><p>5. Leaves bipinnate (rarely tripinnate); primary pinnae 3–5(–7), on unwinged rachis, the larger ones petiolulate and again divided; secondary segments 1.5–5 mm wide; petioles 1.5–7(–9) cm, often as long as the blade; capitula prevailingly solitary (rarely 2–3) on terminal peduncles 15–35(–45) cm; external phyllaries 6–8, linear-lanceolate; ray florets 5–8, white; pappus awns lacking; from western Jalisco, Mexico ................................................................................................................................................... C. deficiens</p><p>– Leaves simple, pinnatifid or pinnate; pinnae 3–5, on partially winged rachis, 0.5–2.5 cm wide; petioles 3–5 cm, generally shorter than the blades (when longer, the terminal segment is dentate and twice as large as the lateral pinnae); capitula 1–5 on terminal peduncles 10–45 cm; external phyllaries 8–10, linear-oblanceolate; ray florets 8, violet to purple (sometimes nearly white); pappus awns 2–4(–6); central and southern Mexico .................................................................................................................................... 6</p><p>6. Leaves simple or pinnatifid; all pinnae similar, entire; capitula 3–7 cm wide across the extended rays; external phyllaries 8, oblonglanceolate to oblanceolate; ample distribution in Mexico to Guatemala .................................................................. C. diversifolius</p><p>– Leaves pinnatifid; terminal segment larger than the laterals, denticulate; capitula 2.5–4 cm wide across the extended rays; external phyllaries 8–10, linear-lanceolate; from Taxco, Guerrero, Mexico ............................................................................... C. schaffneri</p><p>7. Disk florets yellow, rarely with purple lobes .................................................................................................................................... 8</p><p>– Disk florets dark purple, occasionally yellow with purple lobes ................................................................................................... 19</p><p>8. Ray florets reflexed to reclinate; leaves pinnatisect ......................................................................................................................... 9</p><p>– Ray florets spreading; leaves simple, lobed, pinnatifid or pinnatisect ........................................................................................... 10</p><p>9. Leaves 2.5–7.5 cm long, 1.5–5 cm wide, 2–3 pinnatisect; petioles 0.7–2 cm; capitula 3–18, 2–4 cm wide across the reflexed rays; external phyllaries 8, 5–9 mm long, 1–2 mm wide; ray florets 7–10, dark purple to reddish; limb 1.2–1.6 cm long; disk florets 18–32; cypsela 1–1.3 cm long, 1–2 mm wide; from Juxtlahuaca, Oaxaca, Mexico ............................................ C. juxtlahuacensis</p><p>– Leaves 4–11 cm long, 4–9 cm wide, bipinnatisect, sessile or with winged petioles 1.5–2.5 cm; capitula solitary, 4.5–7.5 cm wide across the reflexed to reclinate rays; external phyllaries 6–8, 1–2.5 cm long, 5–7 mm wide; ray florets 8–10, dark red; limb 2–2.5 cm long; disk florets 51–80; cypsela 1.5–2.5 cm long, 2–3 mm wide; from San María Lachixio, Oaxaca; Mexico ...... C. nelsonii</p><p>10. Leaves sessile or with petioles 3 mm; ray florets light to dark purple or violet; disk florets yellow ............................................. 11</p><p>– Leaves petiolate; petioles 1–4(–9) cm; ray florets white, violet or dark purple; disk florets various............................................. 13</p><p>11. Leaves tomentose on both faces; ray florets violet; disk florets 50–80; external phyllaries (5–)7–8, 5–9 mm long, 3–4 mm wide, with 5 conspicuous resiniferous ducts on the abaxial face; from Guachinango, Jalisco, Mexico ................... C. pseudoperfoliatus</p><p>– Leaves glabrous or only pubescent on the central vein of the adaxial face; ray florets violet or dark purple; disk florets 10–30; external phyllaries 8, 5–7 mm long, 1–1.5 mm wide, with 3 conspicuous resiniferous ducts on the abaxial face ........................ 12</p><p>12. Lobes of the cauline leaves usually undivided, oblong to lanceolate, 5–8 mm wide; ray florets white, violet or dark purple .......... ............................................................................................................................................................................................. C. sessilis</p><p>– Lobes of the cauline leaves often divided, linear to lanceolate, 1–3 mm wide; ray florets light to purple ...................... C. stellatus</p><p>13. Petioles 0.3–1.5(–2) cm; leaves simple, pinnatisect or bipinnatisect; segments linear, 0.4–3.5 cm long, 1–3(–5) mm wide; ray florets white or violet ...................................................................................................................................................................... 14</p><p>– Petioles (1–)2–6(–9) cm; leaves bipinnatisect to tripinnatisect; segments linear-lanceolate to lanceolate or oblanceolate, 0.3–1.5 cm long, 3–6 mm wide; ray florets white, violet or dark purple .................................................................................................... 15</p><p>14. Capitula 3–6 cm wide across extended rays; peduncles 15–30 cm; disk florets 25–30 .................................................... C. palmeri</p><p>– Capitula 5–7.5 cm wide across extended rays; peduncles 30–40 cm; disk florets 35–50 ................................................ C. pringlei</p><p>15. Leaves pinnate or bipinnatifid; petioles 1.5–9 cm .......................................................................................................................... 16</p><p>– Leaves simple or pinnatifid, decurrent; sessile or with petioles up to 2.5 cm ................................................................................ 18</p><p>16. Roots fusiform to subglobose; leaves with opposite or alternate lateral pinnae segments, with the basal lobes often reclinate and pointing backward and resembling stipels; outer phyllaries ensiform to oblong; disk florets protected by the first paleae series and showing overall a tubular appearance ............................................................................................................................. C. harkerae</p><p>– Roots fusiform; leaves with opposite lateral pinnae segments, without basal lobes resembling stipels; outer phyllaries linear lanceolate; disk florets not protected by the first paleae series and somewhat loose ..................................................................... 17</p><p>17. Leaves 10–23 cm long; petioles 3–9 cm; terminal segments 0.3–1 cm wide; capitula 3–6, peduncles 7–16(–27) cm; from Ayutla, Jalisco, Mexico ...................................................................................................................................................... C. longipetiolatus</p><p>– Leaves 5–7(–10) cm long; petioles 1.5–3 cm; terminal segments, 1–2 mm wide; capitula 1–2; peduncles 15–25 cm; from Santiago Tenango, Santiago Textitlán and Zimatlán, Oaxaca, Mexico ............................................................................................. C. sherffii</p><p>18. Stems puberulent; ray florets violet; leaves ferruginous below, light green above; external phyllaries 6–8; disk florets 21–30, yellow; from southern Estado de México, Mexico ........................................................................................................ C. modestus</p><p>– Stems glabrous; ray florets violet, light or dark purple; leaves concolour; external phyllaries 8; disk florets 25–45, yellow or with purple lobes; from Durango, Jalisco, Nayarit and Zacatecas, Mexico .......................................................................... C. montanus</p><p>19. External paleae tubular enveloping and isolating disk florets ........................................................................................................ 20</p><p>– External paleae not tubular nor enveloping disk florets ................................................................................................................. 21</p><p>20. Plants scapose to subscapose; leaves simple or bipinnatisect; pinnae ovate to lanceolate, entire, rounded; peduncle 30–60 cm; cypsela glabrous to strigillose above, pappus awns 2–4; from Sierra Madre Oriental in Guanajuato, Hidalgo, Querétaro and San Luis Potosí, Mexico .............................................................................................................................................. C. atrosanguineus</p><p>– Plants leafy caulescent; leaves pinnatisect; pinnae lanceolate, dentate, cuneate; peduncles 15–27 cm; cypsela glabrous, pappus awns 2; from Sierra Madre del Sur in Coahuayutla, Guerrero, Mexico ......................................................................... C. mattfeldii</p><p>21. Ray florets violet ............................................................................................................................................................................ 22</p><p>– Ray florets purple to dark purple .................................................................................................................................................... 23</p><p>22. Internodes (2–) 5–15 cm; leaves bipinnatisect; pinnae rhomboid-ovate or deltoid-ovate, glabrescent on both faces, terminal pinnae frequently exceeding the laterals; segment margins 4–5 dentate below; from Sultepec, Estado de México, Mexico ....... C. nitidus</p><p>– Internodes 2.5–6.5(–11) cm; leaves pinnatisect; pinnae ovate-lanceolate, hirsute on both faces, terminal pinnae almost equal to the laterals; segment margins entire or 1–4 dentate above; from Estado de México, Guerrero and Morelos, Mexico ...... C. purpureus</p><p>23. External phyllaries ovate to deltoid; from Durango, Mexico .......................................................................................... C. concolor</p><p>– External phyllaries linear-lanceolate; ample distribution in Mexico .............................................................................................. 24</p><p>24. Leaves pinnatisect; segments (3–)5(–7), linear, glabrous, 4–14 cm long, 2–5 mm wide; petioles 2–3 cm; external phyllaries 7–8(– 10); from central and western Jalisco, Mexico ......................................................................................................... C. ramirezianus</p><p>– Leaves simple, lobed, pinnatisect or ternate; leaflets 3–5, ovate to lanceolate, pubescent, 2–4 cm long, 1.5–3 cm wide; petioles 0.3–2.5 cm; external phyllaries 8; ample distribution .................................................................................................................... 25</p><p>25. Leaves simple, lobed or pinnatisect; pinnae 3–5; petioles 3–5 mm; internal phyllaries purple; ray florets light to dark purple; disk florets 20–50, yellow with purple lobes; ample distribution in Mexico .................................................................... C. scabiosoides</p><p>– Leaves ternate; petioles 0.8–2.5 cm; internal phyllaries translucent with a purple apex; ray florets dark purple; disk florets 20–35, yellow with purple lobes or sometimes purple; from Atenguillo and Mascota, Jalisco, Mexico ................................. C. jaliscensis</p></div>	https://treatment.plazi.org/id/D14E87C9FF827F50FF676AD4FA1D0C83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Castro-Castro, Arturo;Rodríguez, Aarón;Neri-Mondragón, Alberto Yahir;Siordia-González, Nadia Elizabeth	Castro-Castro, Arturo, Rodríguez, Aarón, Neri-Mondragón, Alberto Yahir, Siordia-González, Nadia Elizabeth (2025): A new species of Cosmos section Discopoda (Coreopsideae, Asteraceae) from Guerrero, Mexico. Phytotaxa 725 (1): 91-100, DOI: 10.11646/phytotaxa.725.1.8, URL: https://doi.org/10.11646/phytotaxa.725.1.8
