taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
642CA9675DFB5BD7ADE89349A82AE2DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715969	https://doi.org/10.3897/asp.84.e174388.figure3	Figure 3. Cephalonomia hyalinipennis lectotype ♀ (A – E) and paralectotype ♀ (F – K). A, F Face view of head; B, G dorsal view; C side view; D, H ocellar view; E anterior view of head showing mandible; I, J gaster; K: wings.	Figure 3. Cephalonomia hyalinipennis lectotype ♀ (A – E) and paralectotype ♀ (F – K). A, F Face view of head; B, G dorsal view; C side view; D, H ocellar view; E anterior view of head showing mandible; I, J gaster; K: wings.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
642CA9675DFB5BD7ADE89349A82AE2DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715983	https://doi.org/10.3897/asp.84.e174388.figure17	Figure 17. Ventral view of C. hyalinipennis lectotype (above) and C. soconuscensis holotype (below). Sculpture of the second gastral sternum (S 3) can be used to differentiate the two species.	Figure 17. Ventral view of C. hyalinipennis lectotype (above) and C. soconuscensis holotype (below). Sculpture of the second gastral sternum (S 3) can be used to differentiate the two species.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
642CA9675DFB5BD7ADE89349A82AE2DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715981	https://doi.org/10.3897/asp.84.e174388.figure15	Figure 15. Morphometric ratios among Cephalonomia spp. nr. hyalinipennis (females): the five species found in Hawai'i; C. soconuscensis; C. nr. hyalinipennis from near the C. hyalinipennis type locality in Florida; and the C. hyalinipennis lectotype and paralectotype. Acronyms for morphometric ratios explained in section 2.4. and shown in Fig. 1.	Figure 15. Morphometric ratios among Cephalonomia spp. nr. hyalinipennis (females): the five species found in Hawai'i; C. soconuscensis; C. nr. hyalinipennis from near the C. hyalinipennis type locality in Florida; and the C. hyalinipennis lectotype and paralectotype. Acronyms for morphometric ratios explained in section 2.4. and shown in Fig. 1.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
642CA9675DFB5BD7ADE89349A82AE2DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715982	https://doi.org/10.3897/asp.84.e174388.figure16	Figure 16. Morphometric ratios among Cephalonomia spp. nr. hyalinipennis (males): the four species found in Hawai'i for which males are known; C. soconuscensis; and C. nr. hyalinipennis from near the C. hyalinipennis type locality in Florida. Acronyms for morphometric ratios explained in section 2.4. and shown in Fig. 1.	Figure 16. Morphometric ratios among Cephalonomia spp. nr. hyalinipennis (males): the four species found in Hawai'i for which males are known; C. soconuscensis; and C. nr. hyalinipennis from near the C. hyalinipennis type locality in Florida. Acronyms for morphometric ratios explained in section 2.4. and shown in Fig. 1.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
65991BAE03B15A0AB53AA88B4B533A15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715976	https://doi.org/10.3897/asp.84.e174388.figure10	Figure 10. Cephalonomia kala sp. nov. holotype ♀. A: Side view; B: Dorsal view; C: Face view of head; D: Ocellar view; E: Mandibles.	Figure 10. Cephalonomia kala sp. nov. holotype ♀. A: Side view; B: Dorsal view; C: Face view of head; D: Ocellar view; E: Mandibles.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
65991BAE03B15A0AB53AA88B4B533A15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715984	https://doi.org/10.3897/asp.84.e174388.figure18	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715967	https://doi.org/10.3897/asp.84.e174388.figure1	Figure 1. Morphometric measurements. Acronyms explained in section 2.4. A Cephalonomia liula ♀ side view (paratype); B Cephalonomia soconuscensis ♀ dorsal view; C Cephalonomia lynnea ♀ face view of head (paratype); D Cephalonomia lynnea ocellar view ♀ (paratype); E Cephalonomia soconuscensis ♀ wings.	Figure 1. Morphometric measurements. Acronyms explained in section 2.4. A Cephalonomia liula ♀ side view (paratype); B Cephalonomia soconuscensis ♀ dorsal view; C Cephalonomia lynnea ♀ face view of head (paratype); D Cephalonomia lynnea ocellar view ♀ (paratype); E Cephalonomia soconuscensis ♀ wings.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715974	https://doi.org/10.3897/asp.84.e174388.figure8	Figure 8. Cephalonomia liula sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H ocellar view. Note that the apex of the gaster in (A) appears differently in this and other DNA extracted individuals; usually appears as in Figs 9 and 18 in fresh specimens.	Figure 8. Cephalonomia liula sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H ocellar view. Note that the apex of the gaster in (A) appears differently in this and other DNA extracted individuals; usually appears as in Figs 9 and 18 in fresh specimens.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715975	https://doi.org/10.3897/asp.84.e174388.figure9	Figure 9. Cephalonomia liula sp. nov. paratype ♀ (A – E) and paratype ♂ (F – I). A, F Side view; B, G dorsal view; C, H face view of head; D, I wings; E anterior view of head showing mandibles.	Figure 9. Cephalonomia liula sp. nov. paratype ♀ (A – E) and paratype ♂ (F – I). A, F Side view; B, G dorsal view; C, H face view of head; D, I wings; E anterior view of head showing mandibles.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715980	https://doi.org/10.3897/asp.84.e174388.figure14	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715984	https://doi.org/10.3897/asp.84.e174388.figure18	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715985	https://doi.org/10.3897/asp.84.e174388.figure19	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
14091B0008DF5247AD368C7AD207198B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715986	https://doi.org/10.3897/asp.84.e174388.figure20	Figure 20. Host associations of Cephalonomia lynnea (A – C) and Cephalonomia liula (D – F). All photographs except C are of naturally occurring situations, taken while peeling bark from branches found in forests. A: Hypothenemus eruditus larva with a Cephalonomia lynnea larva developing ectoparasitically on it, the adult wasp in the tunnel system possibly the mother of the ectoparasitic larva, guarding it; B: Cephalonomia lynnea pupa having developed on the desiccated H. eruditus larva next to it, the parasitoid removed from its pupal cocoon; C: Cephalonomia lynnea larva developing on a H. hampei larva, its mother having stung and parasitized the beetle larva in a phloem sandwich observation chamber, after switching out H. eruditus larvae in host wood with H. hampei larvae; D: Two Leptophloeus sp. larvae, each with a Cephalonomia liula egg attached; E: Two Cephalonomia liula cocoons, the parasitoids having developed on the dessicated Leptophloeus sp. larva adjacent and stuck to them; F: Leptophloeus sp. larva consuming the inside of a Cephalonomia liula cocoon, the parasitoid having developed on another Leptophloeus sp. individual.	Figure 20. Host associations of Cephalonomia lynnea (A – C) and Cephalonomia liula (D – F). All photographs except C are of naturally occurring situations, taken while peeling bark from branches found in forests. A: Hypothenemus eruditus larva with a Cephalonomia lynnea larva developing ectoparasitically on it, the adult wasp in the tunnel system possibly the mother of the ectoparasitic larva, guarding it; B: Cephalonomia lynnea pupa having developed on the desiccated H. eruditus larva next to it, the parasitoid removed from its pupal cocoon; C: Cephalonomia lynnea larva developing on a H. hampei larva, its mother having stung and parasitized the beetle larva in a phloem sandwich observation chamber, after switching out H. eruditus larvae in host wood with H. hampei larvae; D: Two Leptophloeus sp. larvae, each with a Cephalonomia liula egg attached; E: Two Cephalonomia liula cocoons, the parasitoids having developed on the dessicated Leptophloeus sp. larva adjacent and stuck to them; F: Leptophloeus sp. larva consuming the inside of a Cephalonomia liula cocoon, the parasitoid having developed on another Leptophloeus sp. individual.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
16A3BB077A7954FAAACFD5DB643F5F85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715972	https://doi.org/10.3897/asp.84.e174388.figure6	Figure 6. Cephalonomia luiji sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H ocellar view. Note that the apex of the gaster in (A) appears differently in this and other DNA extracted individuals; usually appearing as in Figs 7 and 18 in fresh specimens.	Figure 6. Cephalonomia luiji sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H ocellar view. Note that the apex of the gaster in (A) appears differently in this and other DNA extracted individuals; usually appearing as in Figs 7 and 18 in fresh specimens.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
16A3BB077A7954FAAACFD5DB643F5F85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715973	https://doi.org/10.3897/asp.84.e174388.figure7	Figure 7. Cephalonomia luiji sp. nov. paratype ♀ (A – D) and paratype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H wings.	Figure 7. Cephalonomia luiji sp. nov. paratype ♀ (A – D) and paratype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H wings.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
16A3BB077A7954FAAACFD5DB643F5F85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715980	https://doi.org/10.3897/asp.84.e174388.figure14	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
16A3BB077A7954FAAACFD5DB643F5F85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715984	https://doi.org/10.3897/asp.84.e174388.figure18	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
16A3BB077A7954FAAACFD5DB643F5F85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715985	https://doi.org/10.3897/asp.84.e174388.figure19	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
1C4AF36C8E935B04BD3DE01AAAF80B14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715967	https://doi.org/10.3897/asp.84.e174388.figure1	Figure 1. Morphometric measurements. Acronyms explained in section 2.4. A Cephalonomia liula ♀ side view (paratype); B Cephalonomia soconuscensis ♀ dorsal view; C Cephalonomia lynnea ♀ face view of head (paratype); D Cephalonomia lynnea ocellar view ♀ (paratype); E Cephalonomia soconuscensis ♀ wings.	Figure 1. Morphometric measurements. Acronyms explained in section 2.4. A Cephalonomia liula ♀ side view (paratype); B Cephalonomia soconuscensis ♀ dorsal view; C Cephalonomia lynnea ♀ face view of head (paratype); D Cephalonomia lynnea ocellar view ♀ (paratype); E Cephalonomia soconuscensis ♀ wings.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
1C4AF36C8E935B04BD3DE01AAAF80B14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715970	https://doi.org/10.3897/asp.84.e174388.figure4	Figure 4. Cephalonomia lynnea sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H ocellar view.	Figure 4. Cephalonomia lynnea sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G face view of head; D, H ocellar view.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
1C4AF36C8E935B04BD3DE01AAAF80B14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715971	https://doi.org/10.3897/asp.84.e174388.figure5	Figure 5. Cephalonomia lynnea sp. nov. paratype ♀ (A – E) and paratype ♂ (F – J). A, F Side view; B, G dorsal view; C, H face view of head; D, I wings; E, J view of head showing mandible.	Figure 5. Cephalonomia lynnea sp. nov. paratype ♀ (A – E) and paratype ♂ (F – J). A, F Side view; B, G dorsal view; C, H face view of head; D, I wings; E, J view of head showing mandible.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
1C4AF36C8E935B04BD3DE01AAAF80B14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715980	https://doi.org/10.3897/asp.84.e174388.figure14	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
1C4AF36C8E935B04BD3DE01AAAF80B14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715984	https://doi.org/10.3897/asp.84.e174388.figure18	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
1C4AF36C8E935B04BD3DE01AAAF80B14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715985	https://doi.org/10.3897/asp.84.e174388.figure19	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
9ABE23C863E450C2804973D55A78E7A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715967	https://doi.org/10.3897/asp.84.e174388.figure1	Figure 1. Morphometric measurements. Acronyms explained in section 2.4. A Cephalonomia liula ♀ side view (paratype); B Cephalonomia soconuscensis ♀ dorsal view; C Cephalonomia lynnea ♀ face view of head (paratype); D Cephalonomia lynnea ocellar view ♀ (paratype); E Cephalonomia soconuscensis ♀ wings.	Figure 1. Morphometric measurements. Acronyms explained in section 2.4. A Cephalonomia liula ♀ side view (paratype); B Cephalonomia soconuscensis ♀ dorsal view; C Cephalonomia lynnea ♀ face view of head (paratype); D Cephalonomia lynnea ocellar view ♀ (paratype); E Cephalonomia soconuscensis ♀ wings.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
9ABE23C863E450C2804973D55A78E7A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715978	https://doi.org/10.3897/asp.84.e174388.figure12	Figure 12. Cephalonomia soconuscensis sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G: face view of head; D, H ocellar view.	Figure 12. Cephalonomia soconuscensis sp. nov. holotype ♀ (A – D) and allotype ♂ (E – H). A, E Side view; B, F dorsal view; C, G: face view of head; D, H ocellar view.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
9ABE23C863E450C2804973D55A78E7A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715979	https://doi.org/10.3897/asp.84.e174388.figure13	Figure 13. Cephalonomia soconuscensis sp. nov. paratype ♀ (A – E) and paratype ♂ (F – I). A, F Side view; B, G dorsal view; C, H face view of head; D view of head showing mandible; E, I wings.	Figure 13. Cephalonomia soconuscensis sp. nov. paratype ♀ (A – E) and paratype ♂ (F – I). A, F Side view; B, G dorsal view; C, H face view of head; D view of head showing mandible; E, I wings.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
9ABE23C863E450C2804973D55A78E7A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715980	https://doi.org/10.3897/asp.84.e174388.figure14	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	Figure 14. Genitalia (♂). A Cephalonomia lynnea sp. nov.; B Cephalonomia luiji sp. nov.; C Cephalonomia liula sp. nov.; D Cephalonomia soconuscensis sp. nov.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
9ABE23C863E450C2804973D55A78E7A8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715983	https://doi.org/10.3897/asp.84.e174388.figure17	Figure 17. Ventral view of C. hyalinipennis lectotype (above) and C. soconuscensis holotype (below). Sculpture of the second gastral sternum (S 3) can be used to differentiate the two species.	Figure 17. Ventral view of C. hyalinipennis lectotype (above) and C. soconuscensis holotype (below). Sculpture of the second gastral sternum (S 3) can be used to differentiate the two species.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
7F9360F21A8C5B4CA5BA83C29DB903A0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715977	https://doi.org/10.3897/asp.84.e174388.figure11	Figure 11. Cephalonomia zander sp. nov. holotype ♀ (A – E) and allotype ♂ (F – I). A, F: Side view; B, G: Dorsal view; C, H: Face view of head; D, I: Ocellar view.	Figure 11. Cephalonomia zander sp. nov. holotype ♀ (A – E) and allotype ♂ (F – I). A, F: Side view; B, G: Dorsal view; C, H: Face view of head; D, I: Ocellar view.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
7F9360F21A8C5B4CA5BA83C29DB903A0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715984	https://doi.org/10.3897/asp.84.e174388.figure18	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	Figure 18. Cephalonomia spp. females near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), C. liula (G – I), C. kala (J – L), C. zander (M – O).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
7F9360F21A8C5B4CA5BA83C29DB903A0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715985	https://doi.org/10.3897/asp.84.e174388.figure19	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	Figure 19. Cephalonomia spp. males near C. hyalinipennis found in Hawai'i. C. lynnea (A – C), C. luiji (D – F), and C. liula (G – I), C. zander (J – L) (males are unknown for C. kala).	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
7F9360F21A8C5B4CA5BA83C29DB903A0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715987	https://doi.org/10.3897/asp.84.e174388.figure21	Figure 21. Behavior of C. lynnea in phloem sandwich observation chambers. A, B: Stinging; C – F: Chewing for host feeding or malaxation, mandibles moving in opposite phase as a tile nipper at the beginning of a chewing episode (E), and in phase as grinding for a prolonged period subsequently (F). The larvae in A, C – F are H. eruditus; in B is H. hampei.	Figure 21. Behavior of C. lynnea in phloem sandwich observation chambers. A, B: Stinging; C – F: Chewing for host feeding or malaxation, mandibles moving in opposite phase as a tile nipper at the beginning of a chewing episode (E), and in phase as grinding for a prolonged period subsequently (F). The larvae in A, C – F are H. eruditus; in B is H. hampei.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
7F9360F21A8C5B4CA5BA83C29DB903A0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1715988	https://doi.org/10.3897/asp.84.e174388.figure22	Figure 22. Development of C. lynnea on H. eruditus larva (A – H), and C. liula on Leptophloeus sp. larva (I – L). Time elapsed after presence of an egg was first observed on the host are noted in the photographs. The parasitized beetles were found in natural situations upon opening wood, and the host with the ectoparasitoid was moved into a hole in the wood substrate and covered with a piece of glass slide cover to create an enclosed environment.	Figure 22. Development of C. lynnea on H. eruditus larva (A – H), and C. liula on Leptophloeus sp. larva (I – L). Time elapsed after presence of an egg was first observed on the host are noted in the photographs. The parasitized beetles were found in natural situations upon opening wood, and the host with the ectoparasitoid was moved into a hole in the wood substrate and covered with a piece of glass slide cover to create an enclosed environment.	2026-07-14	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.		Zenodo	biologists	Honsberger, David;Pérez-Lachaud, Gabriela;Magnacca, Karl N.;Lorenzo-Elarco, Jacob Hauʻoli;Kelly, Sandor L.;Wright, Mark G.;Gariepy, Tara D.			
