taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
0C557C5CFADC5DE3AB35D35EDC02F30F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675866	https://doi.org/10.3897/dez.73.170965.figure1	Figure 1. Habitus, dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A, B. Ceratocombus ishigaki sp. nov., holotype, male; C, D. Same, paratype, female; E, F. C. meridianus sp. nov., holotype, male; G, H. Same, paratype, female; I, J. C. trimaculatus sp. nov., holotype, male; K, L. Same, paratype, female. Scale bars: 0.5 mm.	Figure 1. Habitus, dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A, B. Ceratocombus ishigaki sp. nov., holotype, male; C, D. Same, paratype, female; E, F. C. meridianus sp. nov., holotype, male; G, H. Same, paratype, female; I, J. C. trimaculatus sp. nov., holotype, male; K, L. Same, paratype, female. Scale bars: 0.5 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
0C557C5CFADC5DE3AB35D35EDC02F30F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675868	https://doi.org/10.3897/dez.73.170965.figure3	Figure 3. Head and pronotum (A – C) and right forewing (setae omitted) (D – F). A, D. Ceratocombus ishigaki sp. nov., male; B, E. C. meridianus sp. nov., male; C, F. C. trimaculatus sp. nov., male (C) and female (F). Abbreviations: An 1 – first anal vein; An 2 – second anal vein; cf – costal fracture; C + Sc – costa + subcosta; Cu – cubitus; cu – cubital cell; M – media; m – medial cell; R – radius; r – radial cell. Scale bars: 0.2 mm.	Figure 3. Head and pronotum (A – C) and right forewing (setae omitted) (D – F). A, D. Ceratocombus ishigaki sp. nov., male; B, E. C. meridianus sp. nov., male; C, F. C. trimaculatus sp. nov., male (C) and female (F). Abbreviations: An 1 – first anal vein; An 2 – second anal vein; cf – costal fracture; C + Sc – costa + subcosta; Cu – cubitus; cu – cubital cell; M – media; m – medial cell; R – radius; r – radial cell. Scale bars: 0.2 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
0C557C5CFADC5DE3AB35D35EDC02F30F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675869	https://doi.org/10.3897/dez.73.170965.figure4	Figure 4. Female right leg, outer view. A – C. Fore (A), mid- (B) and hind (C) legs of Ceratocombus ishigaki sp. nov.; D – F. Fore (D), mid- (E) and hind (F) legs of C. meridianus sp. nov.; G – I. Fore (G), mid- (H) and hind (I) legs of C. trimaculatus sp. nov. Scale bars: 0.2 mm.	Figure 4. Female right leg, outer view. A – C. Fore (A), mid- (B) and hind (C) legs of Ceratocombus ishigaki sp. nov.; D – F. Fore (D), mid- (E) and hind (F) legs of C. meridianus sp. nov.; G – I. Fore (G), mid- (H) and hind (I) legs of C. trimaculatus sp. nov. Scale bars: 0.2 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
0C557C5CFADC5DE3AB35D35EDC02F30F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675870	https://doi.org/10.3897/dez.73.170965.figure5	Figure 5. Male (A – H) and female (I, J) genitalia of Ceratocombus ishigaki sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views; C left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	Figure 5. Male (A – H) and female (I, J) genitalia of Ceratocombus ishigaki sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views; C left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
0C557C5CFADC5DE3AB35D35EDC02F30F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675873	https://doi.org/10.3897/dez.73.170965.figure8	Figure 8. Living individuals and habitats. A. Ceratocombus ishigaki sp. nov., male; B. C. meridianus sp. nov., female; C. C. trimaculatus sp. nov., female; D. Habitat of C. ishigaki, Kabira, Ishigaki-jima Island; E. Habitat of C. meridianus, Sumiyô-chô, Amami-Ôshima Island; F. Habitat of C. trimaculatus, Shiramizu, Ishigaki-jima Island.	Figure 8. Living individuals and habitats. A. Ceratocombus ishigaki sp. nov., male; B. C. meridianus sp. nov., female; C. C. trimaculatus sp. nov., female; D. Habitat of C. ishigaki, Kabira, Ishigaki-jima Island; E. Habitat of C. meridianus, Sumiyô-chô, Amami-Ôshima Island; F. Habitat of C. trimaculatus, Shiramizu, Ishigaki-jima Island.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
8875B0CA2B2B5872A507C08B44F7AA33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675867	https://doi.org/10.3897/dez.73.170965.figure2	Figure 2. Lectotype of Ceratocombus japonicus Poppius, 1910 (A) and C. plebejus Poppius, 1910 (B), dorsal view. Photographed by HNHM, published with permission.	Figure 2. Lectotype of Ceratocombus japonicus Poppius, 1910 (A) and C. plebejus Poppius, 1910 (B), dorsal view. Photographed by HNHM, published with permission.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
150B1880D9955BC0A09320698A9765A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675866	https://doi.org/10.3897/dez.73.170965.figure1	Figure 1. Habitus, dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A, B. Ceratocombus ishigaki sp. nov., holotype, male; C, D. Same, paratype, female; E, F. C. meridianus sp. nov., holotype, male; G, H. Same, paratype, female; I, J. C. trimaculatus sp. nov., holotype, male; K, L. Same, paratype, female. Scale bars: 0.5 mm.	Figure 1. Habitus, dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A, B. Ceratocombus ishigaki sp. nov., holotype, male; C, D. Same, paratype, female; E, F. C. meridianus sp. nov., holotype, male; G, H. Same, paratype, female; I, J. C. trimaculatus sp. nov., holotype, male; K, L. Same, paratype, female. Scale bars: 0.5 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
150B1880D9955BC0A09320698A9765A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675868	https://doi.org/10.3897/dez.73.170965.figure3	Figure 3. Head and pronotum (A – C) and right forewing (setae omitted) (D – F). A, D. Ceratocombus ishigaki sp. nov., male; B, E. C. meridianus sp. nov., male; C, F. C. trimaculatus sp. nov., male (C) and female (F). Abbreviations: An 1 – first anal vein; An 2 – second anal vein; cf – costal fracture; C + Sc – costa + subcosta; Cu – cubitus; cu – cubital cell; M – media; m – medial cell; R – radius; r – radial cell. Scale bars: 0.2 mm.	Figure 3. Head and pronotum (A – C) and right forewing (setae omitted) (D – F). A, D. Ceratocombus ishigaki sp. nov., male; B, E. C. meridianus sp. nov., male; C, F. C. trimaculatus sp. nov., male (C) and female (F). Abbreviations: An 1 – first anal vein; An 2 – second anal vein; cf – costal fracture; C + Sc – costa + subcosta; Cu – cubitus; cu – cubital cell; M – media; m – medial cell; R – radius; r – radial cell. Scale bars: 0.2 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
150B1880D9955BC0A09320698A9765A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675869	https://doi.org/10.3897/dez.73.170965.figure4	Figure 4. Female right leg, outer view. A – C. Fore (A), mid- (B) and hind (C) legs of Ceratocombus ishigaki sp. nov.; D – F. Fore (D), mid- (E) and hind (F) legs of C. meridianus sp. nov.; G – I. Fore (G), mid- (H) and hind (I) legs of C. trimaculatus sp. nov. Scale bars: 0.2 mm.	Figure 4. Female right leg, outer view. A – C. Fore (A), mid- (B) and hind (C) legs of Ceratocombus ishigaki sp. nov.; D – F. Fore (D), mid- (E) and hind (F) legs of C. meridianus sp. nov.; G – I. Fore (G), mid- (H) and hind (I) legs of C. trimaculatus sp. nov. Scale bars: 0.2 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
150B1880D9955BC0A09320698A9765A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675871	https://doi.org/10.3897/dez.73.170965.figure6	Figure 6. Male (A – H) and female (I, J) genitalia of Ceratocombus meridianus sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views (one of the two pairs of long, stout setae near the caudal portion is missing); C. Left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, left lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	Figure 6. Male (A – H) and female (I, J) genitalia of Ceratocombus meridianus sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views (one of the two pairs of long, stout setae near the caudal portion is missing); C. Left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, left lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
150B1880D9955BC0A09320698A9765A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675873	https://doi.org/10.3897/dez.73.170965.figure8	Figure 8. Living individuals and habitats. A. Ceratocombus ishigaki sp. nov., male; B. C. meridianus sp. nov., female; C. C. trimaculatus sp. nov., female; D. Habitat of C. ishigaki, Kabira, Ishigaki-jima Island; E. Habitat of C. meridianus, Sumiyô-chô, Amami-Ôshima Island; F. Habitat of C. trimaculatus, Shiramizu, Ishigaki-jima Island.	Figure 8. Living individuals and habitats. A. Ceratocombus ishigaki sp. nov., male; B. C. meridianus sp. nov., female; C. C. trimaculatus sp. nov., female; D. Habitat of C. ishigaki, Kabira, Ishigaki-jima Island; E. Habitat of C. meridianus, Sumiyô-chô, Amami-Ôshima Island; F. Habitat of C. trimaculatus, Shiramizu, Ishigaki-jima Island.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
150B1880D9955BC0A09320698A9765A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675867	https://doi.org/10.3897/dez.73.170965.figure2	Figure 2. Lectotype of Ceratocombus japonicus Poppius, 1910 (A) and C. plebejus Poppius, 1910 (B), dorsal view. Photographed by HNHM, published with permission.	Figure 2. Lectotype of Ceratocombus japonicus Poppius, 1910 (A) and C. plebejus Poppius, 1910 (B), dorsal view. Photographed by HNHM, published with permission.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
74AA14FB09E455A3A323D1258BFF4D81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675867	https://doi.org/10.3897/dez.73.170965.figure2	Figure 2. Lectotype of Ceratocombus japonicus Poppius, 1910 (A) and C. plebejus Poppius, 1910 (B), dorsal view. Photographed by HNHM, published with permission.	Figure 2. Lectotype of Ceratocombus japonicus Poppius, 1910 (A) and C. plebejus Poppius, 1910 (B), dorsal view. Photographed by HNHM, published with permission.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
6308FC3EF89E571FB7A7120AD5DAFB33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675866	https://doi.org/10.3897/dez.73.170965.figure1	Figure 1. Habitus, dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A, B. Ceratocombus ishigaki sp. nov., holotype, male; C, D. Same, paratype, female; E, F. C. meridianus sp. nov., holotype, male; G, H. Same, paratype, female; I, J. C. trimaculatus sp. nov., holotype, male; K, L. Same, paratype, female. Scale bars: 0.5 mm.	Figure 1. Habitus, dorsal (A, C, E, G, I, K) and lateral (B, D, F, H, J, L) views. A, B. Ceratocombus ishigaki sp. nov., holotype, male; C, D. Same, paratype, female; E, F. C. meridianus sp. nov., holotype, male; G, H. Same, paratype, female; I, J. C. trimaculatus sp. nov., holotype, male; K, L. Same, paratype, female. Scale bars: 0.5 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
6308FC3EF89E571FB7A7120AD5DAFB33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675868	https://doi.org/10.3897/dez.73.170965.figure3	Figure 3. Head and pronotum (A – C) and right forewing (setae omitted) (D – F). A, D. Ceratocombus ishigaki sp. nov., male; B, E. C. meridianus sp. nov., male; C, F. C. trimaculatus sp. nov., male (C) and female (F). Abbreviations: An 1 – first anal vein; An 2 – second anal vein; cf – costal fracture; C + Sc – costa + subcosta; Cu – cubitus; cu – cubital cell; M – media; m – medial cell; R – radius; r – radial cell. Scale bars: 0.2 mm.	Figure 3. Head and pronotum (A – C) and right forewing (setae omitted) (D – F). A, D. Ceratocombus ishigaki sp. nov., male; B, E. C. meridianus sp. nov., male; C, F. C. trimaculatus sp. nov., male (C) and female (F). Abbreviations: An 1 – first anal vein; An 2 – second anal vein; cf – costal fracture; C + Sc – costa + subcosta; Cu – cubitus; cu – cubital cell; M – media; m – medial cell; R – radius; r – radial cell. Scale bars: 0.2 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
6308FC3EF89E571FB7A7120AD5DAFB33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675869	https://doi.org/10.3897/dez.73.170965.figure4	Figure 4. Female right leg, outer view. A – C. Fore (A), mid- (B) and hind (C) legs of Ceratocombus ishigaki sp. nov.; D – F. Fore (D), mid- (E) and hind (F) legs of C. meridianus sp. nov.; G – I. Fore (G), mid- (H) and hind (I) legs of C. trimaculatus sp. nov. Scale bars: 0.2 mm.	Figure 4. Female right leg, outer view. A – C. Fore (A), mid- (B) and hind (C) legs of Ceratocombus ishigaki sp. nov.; D – F. Fore (D), mid- (E) and hind (F) legs of C. meridianus sp. nov.; G – I. Fore (G), mid- (H) and hind (I) legs of C. trimaculatus sp. nov. Scale bars: 0.2 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
6308FC3EF89E571FB7A7120AD5DAFB33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675872	https://doi.org/10.3897/dez.73.170965.figure7	Figure 7. Male (A – H) and female (I, J) genitalia of Ceratocombus trimaculatus sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views; C. Left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	Figure 7. Male (A – H) and female (I, J) genitalia of Ceratocombus trimaculatus sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views; C. Left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
6308FC3EF89E571FB7A7120AD5DAFB33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675873	https://doi.org/10.3897/dez.73.170965.figure8	Figure 8. Living individuals and habitats. A. Ceratocombus ishigaki sp. nov., male; B. C. meridianus sp. nov., female; C. C. trimaculatus sp. nov., female; D. Habitat of C. ishigaki, Kabira, Ishigaki-jima Island; E. Habitat of C. meridianus, Sumiyô-chô, Amami-Ôshima Island; F. Habitat of C. trimaculatus, Shiramizu, Ishigaki-jima Island.	Figure 8. Living individuals and habitats. A. Ceratocombus ishigaki sp. nov., male; B. C. meridianus sp. nov., female; C. C. trimaculatus sp. nov., female; D. Habitat of C. ishigaki, Kabira, Ishigaki-jima Island; E. Habitat of C. meridianus, Sumiyô-chô, Amami-Ôshima Island; F. Habitat of C. trimaculatus, Shiramizu, Ishigaki-jima Island.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
6308FC3EF89E571FB7A7120AD5DAFB33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1675870	https://doi.org/10.3897/dez.73.170965.figure5	Figure 5. Male (A – H) and female (I, J) genitalia of Ceratocombus ishigaki sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views; C left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	Figure 5. Male (A – H) and female (I, J) genitalia of Ceratocombus ishigaki sp. nov. A, B. Genital segments, dorsal (A) and ventral (B) views; C left laterotergite 8, lateral view; D – F. Left laterotergite 9, dorsal (D), anterior (E) and lateral (F) views; G, H. Left paramere, two different views; I. Anterior part of first valvula, lateral view; J. Spermatheca. Abbreviations: mt 8 – mediotergite 8; lt 8 – laterotergite 8; lt 9 – laterotergite 9; lp – left paramere; pp – preparameral process of pygophore; py – pygophore; rp – right paramere; s 8 – sternum 8. Scale bars: 0.05 mm.	2026-06-10	Yamada, Kazutaka		Zenodo	biologists	Yamada, Kazutaka			
