identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
47E49F0055AC55919C4D63F614B35828.text	47E49F0055AC55919C4D63F614B35828.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus attarum (Bernhauer 1912) Guzman Sarmiento & Żyła 2026	<div><p>Lypeticus attarum (Bernhauer, 1912) comb. nov.</p><p>Fig. 3 A – E</p><p>= Medon (Leiporaphes) attarum Bernhauer, 1912: 37 (original description);</p><p>= Leiporaphes attarum Blackwelder, 1939: 119 (as separate genus); Blackwelder 1944: 118 (checklist); Blackwelder 1952: 213 (list of generic names).</p><p>Material examined.</p><p>lectotype (designated here); Argentina • ♂; “ RLA Argentina Prov. Buenos Aires 7. VIII 1911 C. Bruch \\ 7. [red paper] \\ [B. atta hypiris Carhue.] [red label] \\ [attarum Brh. Typus] [yellow label] \\ Chicago NHMus M. Bernhauer Collection ”; FMNH . paralectotype; Argentina • ♂; “ RLA Argentina Prov. Buenos Aires 7. VIII 1911 C. Bruch \\ 7. [red label] \\ [attarum Brh. Cotypus] [yellow label] \\ Chicago NHMus M. Bernhauer Collection ”; FMNH .</p><p>Diagnosis.</p><p>Lypeticus attarum is distinguished from other species by the aedeagus with median lobe subquadrate (elongate in all species except for L. nori sp. nov.) and widened ventral process (narrowed in L. nori sp. nov.). From L. nori sp. nov., it also differs in the lack of the additional sclerotization at the ventral process apex (present in L. nori sp. nov.). Additionally, ridge behind antennal insertion is short, not extending beyond anterior margin of eye.</p><p>Redescription.</p><p>Measurements (n = 1, Lectotype), FBL (1.30), H (0.39, 0.27, 0.43), A (0.6), NKW (0.13), P (0.38, 0.37, 0.20), E (0.47, 0.41, 0.46).</p><p>Coloration. Head, thorax, and abdomen dark brown; legs paler, brownish (Fig. 3 A).</p><p>Head. Anterior margin 1.5 times longer than posterior margin, slightly concave, carinate. Gena strongly protruding; ridge behind antennal insertion reaching anterior margin of eye. Gular sutures separated, widening posteriorly. Antennomeres 3–8 quadrate and subequal; antennomeres 9–10 slightly wider than preceding ones; antennomere 11 obtusely pointed. Labrum with deep, distinctly V-shaped median emargination; lateral emarginations narrow, situated below level of median emargination. Neck 1 / 4 width of posterior margin of head; in ventral view, median portion of postoccipital suture reaching posterior margin of head.</p><p>Thorax. Basisternal transverse carina well developed, darkened; pronotosternal suture present as sulcus. Marginal carina of mesocoxal acetabulum incomplete.</p><p>Legs. Procoxa 2.5 times longer than wide. Mesotarsomere 1 as long as mesotarsomeres 2 and 3 combined; mesotarsomere 2 longer than 3; mesotarsomere 4 as long as 3; mesotarsomere 5 as long as 1. Metatarsomere 1 as long as metatarsomeres 2 and 3 combined; metatarsomere 2 longer than 3; metatarsomere 3 longer than 4; metatarsomere 5 as long as 1.</p><p>Aedeagus. Median lobe subquadrate; ventral process broad; apical margin straight, with deep median emargination flanked by rounded lobes. Internal sac with two slender, apically tapering sclerites projecting beyond median emargination (Fig. 3 B – E).</p><p>Female. Unknown.</p><p>Distribution.</p><p>The species is only known from the type locality in Argentina, in the province of Buenos Aires.</p><p>Notes.</p><p>Bernhauer (1912) did not designate a holotype and presumably based the original description on more than one specimen, as indicated by the presence of two syntypes bearing identical collection data in the FMNH collection. To ensure nomenclatural stability and facilitate future identification, we hereby designate the male specimen labelled “ attarum Brh. Typus ” as the lectotype. The second male specimen from the same type series, labelled “ attarum Brh. Cotypus, ” is a paralectotype.</p><p>According to the original description, L. attarum was collected from a nest of Acromyrmex hystrix . However, it was not specified whether the specimens were collected from within the nest or its immediate surroundings.</p></div>	https://treatment.plazi.org/id/47E49F0055AC55919C4D63F614B35828	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
258FF375CDC659548587C05752176A5F.text	258FF375CDC659548587C05752176A5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus celatus Sharp 1886	<div><p>Lypeticus aff. celatus Sharp, (1886)</p><p>Material examined.</p><p>Costa Rica • ♀; ” Südabhg. Barba [most likely Volcán Barva], 1200 m, 23. m. vii. 35 (Neverm.), Costa Rica [Hand writing] \\ Field Mus. Nat. Hist. 1966, A. Biering Colln., Acc. Z-13812 ”; FMNH .</p><p>Mexico • ♀; “ Orizaba, Veracruz, MEX. VII: 9: 41 \\ Elevation 4000 ft. \\ Col. By H. S. Dybas ”; AMNH .</p></div>	https://treatment.plazi.org/id/258FF375CDC659548587C05752176A5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
1C185ACA20F45E078DA86DA4D1402DC1.text	1C185ACA20F45E078DA86DA4D1402DC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus celatus Sharp 1886	<div><p>Lypeticus celatus Sharp, 1886</p><p>Fig. 4 A – E</p><p>Lypeticus celatus Sharp, 1886: 556 (original description); Blackwelder 1944: 122 (checklist)</p><p>Material examined.</p><p>holotype; Guatemala • ♂; “ Type [red circular label] \\ “ El Reposo, Guatemala. Champion. \\ B. C. A. Col. I. 2. Lypeticus celatus, Sharp. \\ Sharp Coll. 1995, – 313. \\ NHMUK 013580423 [QR code] ”; NHMUK . additional specimens; Costa Rica • 11 ♂, 7 ♀; San Jose province 6 ♂ • “ COSTA RICA, F. NEVERMANN, 2. T. 27 \\ SAN JOSE, 1000–1200 \\ anges [indet. handwriting] \\ Lipethicus celatus Shp. [hand writing] \\ Hamburg Mus Tilsch [indet. handwriting] \\ Chicago NHMus, M. Bernhauer Collection ” • ” San José, 1936, Costa Rica [hand writing] \\ Field Mus. Nat. Hist. 1966, A. Bierig Colln., Acc. Z-13812 \\ Lypeticus celatus ”; • ” San José, 1937, Costa Rica [hand writing] \\ Field Mus. Nat. Hist. 1966, A. Bierig Colln., Acc. Z-13812 ”; • “ Sta Ana, 17. VIII. 39, Costa Rica \\ Field Mus. Nat. Hist., 1966, A. Bierig Colln., Acc. Z-13812 ”; • “ Sta Ana, 3. VI. 39, Costa Rica \\ Field Mus. Nat. Hist., 1966, A. Bierig Colln., Acc. Z-13812 ” ; “ Aserré, 29. vii. 45, Costa Rica [Hand writing] \\ Field Mus. Nat. Hist. 1966, A. Biering Colln., Acc. Z–13812 ” ; 1 ♀ • ” San José, 1937, Costa Rica [hand writing] \\ Field Mus. Nat. Hist. 1966, A. Bierig Colln., Acc. Z-13812 ”; Alajuela province 3 ♂ • “ Atenas, 5–8. V. 1939, Costa Rica \\ Field Mus. Nat. Hist., 1966, A. Bierig Colln., Acc. Z-13812 ” ; • ” La Caja: 8 kil. w., San Jose, C. R. Schmidt 1931 \\ Field Mus. Nat. Hist. 1966, A. Bierig Colln., Acc. Z-13812 ” ; • “ Naranjo, 2. VII. 1939, Costa Rica [hand writing] \\ Field Mus. Nat. Hist. 1966, A. Bierig Colln., Acc. Z-13812 ”; 4 ♀ • La Caja: 8 kil. w., San Jose, C. R. Schmidt 1931 \\ Field Mus. Nat. Hist. 1966, A. Bierig Colln., Acc. Z-13812 “; 1 ♀ • ” La Caja, 1940, Costa Rica [Hand writing] \\ Field Mus. Nat. Hist. 1966, A. Biering Colln., Acc. Z–13812 ” ; 1 ♀ • ” La Caja, 1931, Costa Rica [Hand writing] \\ Field Mus. Nat. Hist. 1966, A. Biering Colln., Acc. Z–13812 ” ; Guatemala • 1 ♂ and 1 ♀; “ Finca El Zapote, Zapote, Escuintla, VII: 8: 48 GUAT Elev. 2400 ft. [731.52 m], leg. R. D. MITCHELL \\ under bark ”; FMNH .</p><p>Diagnosis.</p><p>L. celatus closely resembles L. mundus but can be differentiated by the ridge behind antenna insertion extending to the middle of the eyes (extending to the posterior margin of the eye (Fig. 5 A) in L. mundus). Additionally, in L. celatus, the transversal carina of the basisternum of the prosternum is only weakly outlined, and pronotosternal suture is developed as a sulcus. In contrast, in L. mundus, the transversal carina of the basisternum is well-developed and darker, while pronotosternal suture is absent.</p><p>Redescription.</p><p>Measurements (n = 1, Holotype), FBL (1.38), H (0.38, 0.2, 0.40), A (0.62), NKW (0.13), GL (0.05), P (0.35, 0.35, 0.25), E (0.50, 0.4, 0.4).</p><p>Coloration. Head, prothorax and abdomen brown; legs and elytra paler, yellowish (Fig. 4 A).</p><p>Head. Anterior margin straight, approximately half as long as posterior margin. Ridge behind antennal insertion reaching middle of eye. Gular sutures parallel. Antennomeres 4–6 longer than wide; antennomeres 7–9 quadrate; antennomere 11 triangular, with straight base and rounded apex. Labral lobes with apices curved outward. Neck 1 / 4 width of posterior margin of head; in ventral view, median portion of postoccipital suture reaching half of neck length.</p><p>Thorax. Basisternal transverse carina weakly developed; pronotosternal suture present as shallow sulcus. Marginal carina of mesocoxal acetabulum incomplete.</p><p>Legs. Mesotarsomere 1 longer than 2; mesotarsomere 2 longer than 3; mesotarsomeres 3 and 4 subequal; mesotarsomere 5 as long as 1. Metatarsomere 1 as long as metatarsomeres 3 and 4 combined; metatarsomere 2 longer than 3; metatarsomeres 3 and 4 subequal; metatarsomere 5 as long as 2.</p><p>Aedeagus. Median lobe elongate; ventral process asymmetrical, with median emargination flanked by inflated, inward-directed lobes. Internal sac with two centrally positioned sclerites curved laterally (Fig. 4 B – E).</p><p>Female. Similar to male, except basisternal transverse carina well developed; marginal carina of mesocoxal acetabulum complete, but weak medially.</p><p>Distribution.</p><p>This species is known from Central America, namely from Guatemala, where it was originally described, and Costa Rica, based on the additional material examined in this study. It has been collected on sandy stream banks (Sharp 1886). This represents the first record of the genus from Costa Rica.</p><p>Notes.</p><p>In the original description, David Sharp (1886) stated that the only available specimen was “ in poor condition ” and probably male, based on external morphology. Because the species was described from a single specimen, it is the holotype by monotypy. We dissected the holotype and provided the first description, illustrations, and photographs of the male genitalia.</p></div>	https://treatment.plazi.org/id/1C185ACA20F45E078DA86DA4D1402DC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
1366C5D0297B5FD3BFCB3733F927F61E.text	1366C5D0297B5FD3BFCB3733F927F61E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus mundus (Sharp 1876)	<div><p>Lypeticus mundus (Sharp, 1876)</p><p>Fig. 5 A – E</p><p>Lithocharis munda: Sharp, 1876: 266 (original description); Sharp 1886: 556 (moved to Lypeticus); Blackwelder 1944: 122 (checklist).</p><p>Material examined.</p><p>syntypes; Brazil • 6 ♀; “ [on case while beetle enclosed] Lithocharis munda amazons Type D. S. \\ TYPE [white-red circular label] \\ Tapajos [green oval label] \\ S. America: Brazil. \\ Sharp Coll 1905–313. \\ [printed barcode NHMUK 014532892 ” ; “ [on case while beetle enclosed] Lithocharis munda amazons. Ind. Typ. \\ Tapajos [green oval label] \\ SYNTYPE [white-blue circular label] \\ S. America: Brazil. \\ Sharp Coll 1905-313. ”; ” SYNTYPE [white-blue circular label] \\ Sharp Coll. 1905. – 313. \\ NHMUK 013580424 [QR code] ” ; 3 ♀ with the same label “ Tapajos [green oval label] \\ ” S. America: Brazil. \\ Sharp Coll 1905-313. \\ Lithocharis munda Ind. typ. D. S. ”; NHMUK .</p><p>Diagnosis.</p><p>L. mundus resembles L. celatus but can be distinguished by the ridge behind antenna insertion extending to the posterior margin of the eye (extending to the middle of the eye in L. celatus). Additionally, in L. mundus, the transversal carina on the basisternum of the prosternum is well-developed and darker, and the pronotosternal suture is absent. In contrast, in L. celatus, the transversal carina of the basisternum of the prosternum is only weakly outlined, and the pronotosternal suture forms a depressed area or sulcus.</p><p>Redescription.</p><p>Measurements (n = 1; Syntype), FBL (1.33), H (0.38, 0.2, 0.40), A (0.60), NKW (0.13), GL (0.02), P (0.36, 0.34, 0.28), E (0.50, 0.42, 0.33).</p><p>Coloration. Body reddish; legs and elytra yellowish (Fig. 5 A).</p><p>Head. Anterior margin straight, approximately twice as long as posterior margin. Ridge behind antennal insertion reaching posterior margin of eye. Gular sutures parallel; surface with distinct, coarse micropunctures. Antennomeres 4–8 quadrate and subequal; antennomeres 9–10 slightly wider than preceding ones; antennomere 11 obtusely pointed. Labrum with broad, shallow, U-shaped median emargination. Neck 1 / 4 width of posterior margin of head; in ventral view, median portion of postoccipital suture reaching posterior margin of head.</p><p>Thorax. Basisternal transverse carina well developed and darkened (Fig. 5 B, C); pronotosternal suture absent. Marginal carina of mesocoxal acetabulum complete.</p><p>Legs. Mesotarsomere 1 as long as mesotarsomeres 2 and 3 combined; mesotarsomere 2 longer than 3; mesotarsomeres 3 and 4 subequal; mesotarsomere 5 as long as 1. Metatarsomere 1 as long as metatarsomeres 2 and 3 combined; metatarsomere 2 longer than 3; metatarsomere 3 longer than 4; metatarsomere 4 approximately one-third as long as 3; metatarsomere 5 as long as 1.</p><p>Male. Unknown.</p><p>Distribution.</p><p>This species is known from the Amazon forest in Brazil and was collected in the Tapajos River basin.</p><p>Notes.</p><p>Sharp (1876) did not designate a holotype for Lithocharis munda and instead annotated six specimens in the original series with the handwritten note “ Ind. Typ. ” (individuum typicum). All specimens bearing this annotation are therefore part of the original type material and constitute syntypes under ICZN Article 73.2. The presence or absence of later curatorial labels indicating “ SYNTYPE ” has no nomenclatural effect. Accordingly, all specimens labelled “ Ind. Typ. ” and originating from Sharp’s material are treated herein as syntypes. Because all syntypes are females and provide no additional diagnostic advantage, we refrain from designating a lectotype.</p></div>	https://treatment.plazi.org/id/1366C5D0297B5FD3BFCB3733F927F61E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
E1E338AEF049533C934E937EE0BF068B.text	E1E338AEF049533C934E937EE0BF068B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus nori Guzman Sarmiento & Żyła 2026	<div><p>Lypeticus nori sp. nov.</p><p>Fig. 6 A – E</p><p>Material examined.</p><p>holotype; Bolívia • ♂; “ BOL / Nov. 2013 / 17 BOLIVIA 2013, SANTA CRUZ dep., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.579166&amp;materialsCitation.latitude=-18.324722" title="Search Plazi for locations around (long -59.579166/lat -18.324722)">Santiago de Chiquitos env.</a>, 630 m, 6. xii. 2013; S18°19’29’’ W59°34’45’’; sifting litter on stream bank in the forest, Winkler app. Extr.; P. Baňař lgt. \\ Collection P. BAŇAŘ, Moravian Museum Brno; [red printed label] holotype Lypeticus nori sp. nov. Guzman &amp; Żyła ”; UASC ; paratypes; Bolívia • 3 ♀; “ BOL / Nov. 2013 / 10 BOLIVIA 2013, SANTA CRUZ dep., W Amboro NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.80111&amp;materialsCitation.latitude=-18.098612" title="Search Plazi for locations around (long -63.80111/lat -18.098612)">Barientos</a>, 1806 m, 29. xi.; sifting rotten wood S18°5’55’’ W63°48’04’’, Winkler app. Extr.; P. Baňař lgt. \\ Collection P. BAŇAŘ, Moravian Museum Brno; [yellow printed label] Lypeticus nori sp. nov. Guzman &amp; Żyła ”; 1 MMBC, 2 ZMH .</p><p>Diagnosis.</p><p>L. nori sp. nov. closely resembles L. plaumanni sp. nov. but can be differentiated by the following characters: the median lobe of the aedeagus is subquadrate (elongate in L. plaumanni), ventral process lacks the distinct apical lobes (lobes present in L. plaumanni sp. nov.), and presence of the additional sclerotization at the ventral process apex.</p><p>Description.</p><p>Measurements (n = 1, Holotype), FBL (1.55), H (0.47, 0.25, 0.47), A (0.8), NKW (0.15), GL (0.05), P (0.48, 0.40, 0.37), E (0.54, 0.43, 0.53).</p><p>Coloration. Head, thorax, and abdomen brownish; legs and elytra yellowish (Fig. 6 A). Body larger than in congeners.</p><p>Head. Anterior margin approximately half as long as posterior margin. Ridge behind antennal insertion reaching middle of eye. Gular sutures parallel. Antennomeres 7–10 wider than long; antennomere 11 triangular, with straight base and rounded apex. Median emargination of labrum deeper than lateral emarginations. Neck 1 / 3 width of posterior margin of head; in ventral view, median portion of postoccipital suture reaching half of neck length.</p><p>Thorax. Basisternal transverse carina developed; pronotosternal suture present. Marginal carina of mesocoxal acetabulum complete.</p><p>Legs. Mesotarsomere 1 as long as mesotarsomeres 2 and 3 combined; mesotarsomere 2 longer than 3; mesotarsomere 3 longer than 4; mesotarsomere 4 approximately three-fourths as long as 3; mesotarsomere 5 as long as 1. Metatarsomere 1 as long as metatarsomeres 2 and 3 combined; metatarsomere 2 longer than metatarsomeres 3 and 4 combined; metatarsomeres 3 and 4 subequal; metatarsomere 5 as long as 2.</p><p>Aedeagus. Median lobe elongate; ventral process narrow, with shallow median emargination; margins of emargination strongly sclerotized. Internal sac with two slender, slightly curved sclerites projecting beyond median emargination (Fig. 6 B – E).</p><p>Female. Similar to male; abdomen as described for genus.</p><p>Etymology.</p><p>The specific epithet nori is named after Nori, a fictional dwarf from “ The Hobbit ” by J. R. R. Tolkien, referring to the generally small body size of the genus Lypeticus .</p><p>Distribution.</p><p>This species has been recorded from Santiago de Chiquitos in the Serranía de Santiago and the foothills of Amboró National Park and Integrated Management Natural Area, both in the Santa Cruz Department of Bolivia. This represents the first record of the genus from Bolivia.</p></div>	https://treatment.plazi.org/id/E1E338AEF049533C934E937EE0BF068B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
B0BE095728E055F2A53BE068674006C3.text	B0BE095728E055F2A53BE068674006C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus paraguayensis Guzman Sarmiento & Żyła 2026	<div><p>Lypeticus paraguayensis sp. nov.</p><p>Fig. 7 A – E</p><p>Material examined.</p><p>holotype; Paraguay • ♂; “ PARAGUAY: Itapua prov. 10 km s Santa Maria 25. x. 1982, tamisage forét, bois mort feuilles mortes # 15 \\ leg. F. Baud, C. Dlouhy, M. - C. Durette-Desset, V. Manhert, J. - L. Perret, C. &amp; P. Vaucher \\ [red printed label] HOLOTYPE Lypeticus paraguayensis sp. nov. Guzman &amp; Żyła ”; MHNG ; paratypes Paraguay • 4 ♂; [same labels as holotype] \\ [yellow printed label] PARATYPE Lypeticus paraguayensis sp. nov. Guzman &amp; Żyła ” ; 1 ♂; “ PARAGUAY: Mission zoologique du Museum d’Histoire Naturelle de Genève, x-xi. 1979 \\ [yellow printed label] PARATYPE Lypeticus paraguayensis sp. nov. Guzman &amp; Żyła ” ; 1 ♂; “ PARAG: Central prov. Villa del Maestro, San Lorenzo, Asuncion 05. x. 1979 # 4 \\ tamisage feuilles mortes leg. F. Baud, C. Dlouhy, V. Manhert # 4 J. - L. Perret, Cl. Vaucher \\ [yellow printed label] PARATYPE Lypeticus paraguayensis sp. nov. Guzman &amp; Żyła ” ; 1 ♂; “ PARAGUAY: Itapua prov. Salto Tembey, 4 km amont des chutes, 01. xi. 1982 tamisage forét, bois mort feuilles mortes # 20 \\ leg. F. Baud, C. Dlouhy, M. - C. Durette-Desset, V. Manhert, J. - L. Perret, C. &amp; P. Vaucher \\ [yellow printed label] PARATYPE Lypeticus paraguayensis sp. nov. Guzman &amp; Żyła ”; MHNG .</p><p>Diagnosis.</p><p>Distinguished from all congeners by male sternite VIII with three emarginations (lack of emargination in all other species known from males), posterior margin appearing wavy. Aedeagus with internal sac sclerites robust, apically pointed, not protruding beyond median emargination. Additionally, it differs from Lypeticus plaumanni sp. nov. by the presence of ridge behind antennal insertion (absent in L. plaumanni sp. nov.), from the other species by the ridge reaching posterior margin of eye (shorter in other species), and from Lypeticus mundus by the presence of pronotosternal suture in form of a sulcus (suture absent in L. mundus).</p><p>Description.</p><p>Measurements (n = 1, Holotype), FBL (1.1), H (0.33, 0.22, 0.32), A (0.5), GL (0.05), P (0.3, 0.3, 0.2), E (0.4, 0.3, 0.3).</p><p>Coloration. Head, thorax, and abdomen brownish; appendages lighter (Fig. 7 A). Body smaller than in congeners.</p><p>Head. Anterior margin approximately half as long as posterior margin. Ridge behind antennal insertion reaching posterior margin of eye. Gular sutures reaching posterior margin of head. Antennomeres 4–8 longer than wide; antennomeres 9–10 wider than long; antennomere 11 triangular, with straight base and rounded apex. Median emargination of labrum V-shaped. Neck 1 / 3 width of posterior margin of head; in ventral view, median portion of postoccipital suture reaching half of neck length.</p><p>Thorax. Pronotum slightly longer than wide. Basisternal transverse carina present, weaker medially; pronotosternal suture present as sulcus. Marginal carina of mesocoxal acetabulum complete.</p><p>Legs. Mesotarsomere 1 as long as mesotarsomeres 2 and 3 combined; mesotarsomere 2 longer than 3; mesotarsomere 3 longer than 4; mesotarsomere 4 slightly shorter than 3; mesotarsomere 5 shorter than 1. Metatarsomere 1 as long as metatarsomeres 2 and 3 combined; metatarsomere 2 longer than metatarsomeres 3 and 4 combined; metatarsomere 3 longer than 4; metatarsomere 5 as long as 2.</p><p>Abdomen, male. Sternite VIII with three emarginations; posterior margin consequently appearing wavy; emarginations extending approximately 1 / 6 length of sternite.</p><p>Aedeagus. Median lobe elongate; ventral process broad, slightly curved in lateral view, with deep median emargination flanked by broad, rounded lobes. Internal sac with two robust, apically pointed sclerites not protruding beyond median emargination (Fig. 7 B – E).</p><p>Female. Similar to male; abdomen as described for genus.</p><p>Etymology.</p><p>The specific epithet paraguayensis is derived from the name of Paraguay and refers to the geographic origin of the type series.</p><p>Distribution.</p><p>Known from Paraguay, based on material from Itapúa Department (10 km south of Santa María; Salto Tembey, upstream of the falls) and Central Department (San Lorenzo, Villa del Maestro, near Asunción). It is the first record of the genus from Paraguay.</p></div>	https://treatment.plazi.org/id/B0BE095728E055F2A53BE068674006C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
FDD02FFCC1BF5363B226401895DA5FFB.text	FDD02FFCC1BF5363B226401895DA5FFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus plaumanni Guzman Sarmiento & Żyła 2026	<div><p>Lypeticus plaumanni sp. nov.</p><p>Fig. 8 A – E</p><p>Material examined.</p><p>Holotype; brazil • ♂; “ Brasilien, Nova Teutonia, F. PLAUMANN \\ Lypeticus mundus Sharp. [hand written] \\ ex coll. Scheerpeltz [Blue label] \\ mundus Sharp \\ [red printed label] HOLOTYPE Lypeticus plaumanni sp. nov. Guzman &amp; Żyła ”; NMW ; paratype; Brazil • ♀; ” BRASIL: Sta. Catharina; Nova Teutonia, X: XII: 1952, leg. F. Plaumann \\ Leiporaphes / Lypeticus [hand writing]; [yellow printed label] PARATYPE Lypeticus plaumanni sp. nov. Guzman &amp; Żyła ”; AMNH .</p><p>Diagnosis.</p><p>L. plaumanni sp. nov. closely resembles L. celatus but can be differentiated by the absence of the carinate edge of the antenna insertions, whereas in L. celatus it is present. Additionally, in L. plaumanni sp. nov., ventral process of aedeagus is widened apically (same width as basally in L. celatus).</p><p>Description.</p><p>Measurements (n = 1, Holotype), FBL (1.35), H (0.40, 0.18, 0.35), A (0.64), NKW (0.11), GL (0.04), P (0.36, 0.33, 0.27), E (0.5, 0.4, 0.44).</p><p>Coloration. Head, thorax, and abdomen brownish; legs and elytra yellowish (Fig. 8 A).</p><p>Head. Anterior margin approximately half as long as posterior margin. Ridge behind antennal insertion absent. Gular sutures parallel. Antennomeres 4–8 longer than wide; antennomeres 9–10 wider than long; antennomere 11 triangular, with straight base and rounded apex. Median emargination of labrum V-shaped. Neck 1 / 3 width of posterior margin of head; in ventral view, median portion of postoccipital suture reaching half of neck length.</p><p>Thorax. Pronotum slightly longer than wide. Basisternal transverse carina faint; pronotosternal suture present as sulcus. Marginal carina of mesocoxal acetabulum incomplete.</p><p>Legs. Mesotarsomere 1 as long as mesotarsomeres 2 and 3 combined; mesotarsomere 2 longer than 3; mesotarsomere 3 longer than 4; mesotarsomere 4 approximately three-fourths as long as 3; mesotarsomere 5 as long as 1. Metatarsomere 1 as long as metatarsomeres 2 and 3 combined; metatarsomere 2 longer than metatarsomeres 3 and 4 combined; metatarsomeres 3 and 4 subequal; metatarsomere 5 as long as 2.</p><p>Aedeagus. Median lobe elongate; ventral process broad, with deep median emargination flanked by flat, asymmetrical lobes. Internal sac with two dissimilar sclerites projecting beyond median emargination, one with obtuse apex, the other with pointed apex (Fig. 8 B – E).</p><p>Female. Similar to male; abdomen as described for genus.</p><p>Etymology.</p><p>The specific epithet plaumanni honors the Brazilian entomologist Fritz Plaumann, in recognition of his significant contributions to Brazilian entomology and for collecting the type specimens.</p><p>Distribution.</p><p>Known only from southern Brazil, based on material from Nova Teutônia, Santa Catarina.</p><p>Note.</p><p>This species is described from material previously identified as L. mundus . Comparison of the external morphology with the type series of L. mundus, together with comparison of the male genitalia and other male characters with those of all other Lypeticus species, demonstrates that this material represents a distinct species.</p><p>Additional examined material.</p><p>We identified the specimens based on the diagnoses presented in this study. The specimens from Costa Rica and Mexico exhibit intermediate characters between L. celatus and L. mundus . Based on their geographic origin, we tentatively assign them to L. celatus . This constitutes the first confirmed record of the genus from Mexico.</p></div>	https://treatment.plazi.org/id/FDD02FFCC1BF5363B226401895DA5FFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
800923AA64005C478D20DD575DF0C17C.text	800923AA64005C478D20DD575DF0C17C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lypeticus Sharp 1886	<div><p>Genus: Lypeticus Sharp, 1886</p><p>Figs 1, 2, 3, 4, 5, 6, 7, 8</p><p>Lypeticus Sharp, 1886: 556 (original description); Blackwelder 1944: 122 (checklist); Blackwelder 1952: 228 (list of generic names); Navarrete-Heredia et al. 2002: 268 (characters in key).</p><p>= Medon (Leiporaphes) Bernhauer, 1912: 37 (original description in the rank of a subgenus).</p><p>= Leiporaphes Bernhauer: Blackwelder, 1939: 119 (elevated to genus rank), syn. nov.</p><p>Type species.</p><p>Medon (Leiporaphes) attarum Bernhauer, 1912, by original designation.</p><p>Synonymy and taxonomic remarks.</p><p>In the original description, Bernhauer (1912) established Leiporaphes attarum (Bernhauer, 1912) based particularly on the apparent absence of the gular suture, without comparing the new taxon with Lypeticus . Examination of the type specimen of Leiporaphes attarum, however, revealed clearly separated gular sutures. Furthermore, examination of the type material of Leiporaphes and Lypeticus revealed no characters supporting their separation at the generic level. The shared longitudinal wavy microsculpture behind the eyes, together with the remaining external and male genitalic characters, indicates that Leiporaphes is congeneric with Lypeticus . We therefore place Leiporaphes Bernhauer, 1912 in synonymy with Lypeticus Sharp, 1886, and propose the new combination Lypeticus attarum (Bernhauer, 1912), comb. nov.</p><p>Type species.</p><p>Lithocharis munda Sharp 1876, by original designation.</p><p>Diagnosis.</p><p>Lypeticus is distinct from other genera within Paederinae by head sculpture with longitudinal wavy microsculpture on both sides behind eyes (Figs 1 A, 2 A, 2 B, arrow) and small body size (forebody length at most 1.5 mm). Additionally, it is recognizable by the following combination of features: eyes with setae between ommatidia; antennomeres bead-like; labrum with anterior margin slightly trimarginated (Fig. 2 C, arrow), with two distinctive lobes and wide emargination between; mandibles bidentate; maxillary palpomere 3 fusiform, maxillary palpomere 4 acicular, small (Fig. 2 D, arrow); posterior margin of head emarginated (Fig. 2 A, arrow); neck narrow; anterior angles of pronotum obtuse; metatarsi slender.</p><p>Redescription.</p><p>Coloration brownish to reddish; legs and elytra paler.</p><p>Body small; forebody length 1.10–1.55; surface with micropunctures, most distinct on head.</p><p>Head. Head capsule trapezoidal; anterior margin shorter than posterior, straight; membrane between head and labrum slightly exposed. Eyes protruding, situated in anterior third of head; interommatidial setae present. Gena extending beyond antennal insertion in dorsal view; lateral margin of head with ridge behind antennal insertion (Fig. 1 A, anir), reaching middle of eye or extending beyond; three thick preorbital setae present; longitudinal, wavy microsculpture present behind eyes (Fig. 1 A, wvm). Posterior margin deeply emarginate medially, rounded laterally. Epicranium with denser micropunctures between eyes, sparser behind eyes (Figs 1 A, right side, 2 A, B). Antennae longer than head, but shorter than head and pronotum combined; antennomere 1 as long as antennomeres 2 and 3 combined, wider, with emarginate apex; antennomere 2 shorter than or subequal to antennomere 3, with narrower base and wider apex; antennomeres 4–9 longer than wide or quadrate; antennomere 10 wider than long; antennomere 11 longer than wide. Mandibles asymmetrical; each with two teeth; anterior tooth longer on right mandible, shorter on left. Labrum slightly trimarginate, with broad V-shaped median emargination between two lobes, and two narrow lateral emarginations on each side (Figs 1 A, 2 C). Maxillary palpomere 1 small, curved, glabrous; palpomere 2 longer than palpomere 1, as long as palpomere 3, wider, with setae, and emarginate apically; palpomere 3 broader apically than palpomere 2; palpomere 4 small, glabrous, slender, elongate (Fig. 2 D, arrow). Gula with glabrous median strip between gular sutures; sutures parallel, widening posteriorly in L. attarum (Fig. 1 A, left side, dashed line); surface with micropunctures. Labial palpomere 1 short and narrow; palpomere 2 widest and elongate; palpomere 3 shorter and narrower than palpomere 2. Ligula bilobed. Neck narrow; in ventral view, median portion of postoccipital suture curving anteriorly (Fig. 1 A, pos), reaching at least half of neck length (e. g. L. mundus, Fig. 1 A, dashed line).</p><p>Thorax. Pronotum trapezoidal, narrower than elytra, slightly punctate; anterior margin wider than posterior; anterior angles slightly obtuse, tapering obliquely towards slender neck; posterior margin rounded; posterior angles rounded.</p><p>Prosternum smooth; pronotosternal suture absent or present as sulcus (Fig. 1 B, pss); superior marginal line deflexed (Fig. 1 B, sml); basisternum with faint or well-developed transverse carina (Fig. 1 B, tcbs); longitudinal carina absent; prosternal carina well developed; furcasternum with pointed process reaching apex of postcoxal process; longitudinal furcasternal carina darker (Fig. 1 B, lcfs).</p><p>Mesoventrite nearly as long as wide; anepisterna elevated, laterally adjacent to external prepectal sclerotisation, and separated from posterior mesoventrite by strong posterior anepisternal ridges (Fig. 1 C, par). Posterior anepisternal ridge divided posteriorly into two short branches, reaching lateral angles of external central sclerotisation of prepectus (Fig. 1 C, ecsp) anteriorly, and transversely divided by anterior anepisternal ridges (Fig. 1 C, taar). Anterior anepisternal ridges reaching anterior mesoventral margin, with median posteriorly directed branch connected to posterior anepisternal ridge. External central prepectal sclerotisation (ecsp) elevated, trapezoidal, extending through upper third of plate, posteriorly produced into paired secondary prepectal ridges (Fig. 1 C, spr), directed posterolaterally and reaching posterior anepisternal ridge, forming two rounded-rectangular windows. Posteromedian ventrite nearly as long as wide, with central oval invagination (Fig. 1 C, vmoi). Paired longitudinal secondary ridges (Fig. 1 C, plsr) well developed. Internal prepectal structure present as thin lamina aligned with anterior mesoventral margin. Furcal arms long, slender, apically tapering, nearly reaching prepectal lamina. Mesofurcasternum with longitudinal intercoxal carina. Mesobasisternum with shallow, elongate depression. Metaventrite fused with mesoventrite; marginal carina of mesocoxal acetabulum well developed, emarginate medially or not meeting medially (Fig. 1 C, rbcr); metasternal intercoxal process (metakatepisternal process) bilobed. Elytra slightly longer than pronotum, with dense micropunctures; epipleural ridge absent; apical angles rounded; costal margin curved and longer than anal margin. Scutellum with one well-developed ridge. Hind wings present; veins MP 4 and CuA fused; vein MP 3 absent.</p><p>Legs. Procoxa approximately three times longer than wide; protibia with two longitudinal combs, inner comb longer; outer margin with three or four dark, spine-like setae; protarsomeres 1–4 quadrate and subequal in length, fully divided in dorsal view (Fig. 2 E, F, arrow); protarsomere 5 as long as protarsomeres 1 and 2 combined. Mesocoxa triangular, wider than long; mesofemur as long as mesotibia; mesotibial ctenidium present on inner and outer margins. Mesotarsomeres variable in length among species. Metacoxa as long as wide; metafemur longer than metatibia; metatibial ctenidium present on inner margin; metatarsomeres slender, variable in length among species.</p><p>Abdomen. Abdomen widening posteriorly to segment VII, then narrowing apically; segment VII as long as the two preceding segments combined, anterior margin wider than posterior. Two pairs of paratergites present on segments III – VII. Tergites III – VI transversely impressed basally; posterior margins of segments III – VI straight. Sternite III with median longitudinal keel, nearly reaching posterior margin. Posterior margin of tergite VII slightly convex, without emargination. Tergite IX wrapping ventromedially; posterior margin broadly U-shaped, occupied by tergite X; middorsal base fused.</p><p>Abdomen, males. Posterior margin of sternite VII straight, without emargination. Posterior margin of tergite VIII slightly convex, without emargination (except in L. paraguayensis). Posterior margin of sternite VIII with shallow, broad, indistinct emargination, similar among species. Tergite IX with shallow U-shaped emargination; middorsal base fused medially. Sternite IX narrow, elongate; posterior margin straight. Tergite X triangular.</p><p>Aedeagus. Median lobe elongate or subquadrate; basal portion rounded. Ventral process variable in shape, symmetrical or asymmetrical; apical margin with median emargination of variable depth, flanked by paired lobes differing in shape and development. Internal sac bearing two sclerites, varying in shape (slender to robust, symmetrical or asymmetrical, apices obtuse to pointed), and degree of protrusion beyond median emargination; surface finely scaled.</p><p>Abdomen, females. Posterior margin of sternite VII straight, without emargination. Posterior margin of tergite VIII slightly convex, without emargination. Posterior margin of sternite VIII straight. Tergite IX with middorsal base fused close to anterior margin. Gonocoxal plate divided into two long, moderately broad lateral plates.</p><p>Systematic position.</p><p>The genus is currently classified within the subtribe Medonina of the tribe Lathrobiini . Its placement in Lathrobiini is supported by the presence of a minute, acicular maxillary palpomere 4, one of the diagnostic characters of the tribe. In contrast, its assignment to Medonina reflects the current traditional classification, as the subtribe has been shown to be non-monophyletic (e. g. Żyła et al. 2021). A comprehensive phylogenomic reassessment of the classification of Paederinae is currently underway and will be presented elsewhere. Pending that revision, we retain Lypeticus in Medonina for the purposes of the present study.</p><p>Distribution.</p><p>Lypeticus stat. rev. is distributed throughout the Neotropical region, occurring in Argentina, Bolivia, Brazil, Costa Rica, Guatemala, Mexico, Guatemala, and Paraguay.</p><p>Habitat.</p><p>The original description of L. celatus indicates that the species was collected on sandy stream banks. Although the original description of L. mundus provides only the locality (the Tapajós basin in Brazil), it likely occupies a similar habitat. Lypeticus attarum was originally reported from a nest of Atta hystrix, now considered a synonym of Acromyrmex hystrix (Latreille, 1802), although it remains unclear whether the specimens were collected inside the nest or in its immediate surroundings. The Bolivian specimens were collected by sifting decomposing wood, whereas those from Paraguay were obtained by sifting dead wood and leaf litter in forest habitats. Navarrete-Heredia et al. (2006) also reported Lypeticus from a pine – oak forest.</p></div>	https://treatment.plazi.org/id/800923AA64005C478D20DD575DF0C17C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Guzman Sarmiento, Yoan Camilo;Żyła, Dagmara	Guzman Sarmiento, Yoan Camilo, Żyła, Dagmara (2026): Revised concept of the paederine rove beetle genus Lypeticus Sharp, 1886 with the genus Leiporaphes Bernhauer, 1912 as a synonym and three species new to science (Coleoptera, Staphylinidae). Evolutionary Systematics 10 (2): 327-343, DOI: 10.3897/evolsyst.10.206974
