identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
409E777A864855539CCF56B0EFBA13D9.text	409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ilyoplax kagga Prema & Kakati & Ng 2026	<div><p>Ilyoplax kagga sp. nov.</p><p>Figs 1, 2 A, C, E, G, I, 3, 4 A, C, E, G, 5, 6, 7 A, C, E, G – I, 8 A – E, 9, 10, 11</p><p>Material examined.</p><p>Holotype: India • ♂ (4.2 × 2.5 mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.381065&amp;materialsCitation.latitude=14.522905" title="Search Plazi for locations around (long 74.381065/lat 14.522905)">Aghanashini River Estuary</a>, upper mudflat areas in mangroves with soft-muddy substrate and oyster bed, off Karnataka, west coast of India; 14°31'22.46"N, 74°22'51.82"E; M. Prema leg.; 17 Dec. 2020; CASAU CR 1048 . Paratypes: India • 9 ♂♂ (5.5 × 3.5 mm, 5.3 × 3.5 mm, 5.2 × 3.3 mm, 5.0 × 3.2 mm, 4.9 × 3.6 mm, 4.8 × 3.3 mm, 4.7 × 3.2 mm, 4.6 × 3.2 mm, 4.6 × 3.1 mm), 11 ♀♀ (5.1 × 3.4 mm, 5.0 × 3.1 mm, 4.8 × 3.1 mm, 4.9 × 3.4 mm, 4.9 × 3.3 mm, 4.7 × 3.1 mm, 4.5 × 3.0 mm, 4.5 × 2.9 mm, 4.5 × 2.9 mm, 4.3 × 2.5 mm, 4.1 × 2.7 mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.376625&amp;materialsCitation.latitude=14.520641" title="Search Plazi for locations around (long 74.376625/lat 14.520641)">Aghanashini River Estuary</a>, upper mudflat areas in mangroves of soft-muddy, and oyster bed habitat, off Karnataka, west coast of India; 14°31'14.31"N, 74°22'35.83"E; M. Prema leg.; 22 Oct. 2025; CASAU CR 1045 • 2 ♂♂ (4.0 × 2.5 mm, 3.6 × 2.6 mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.381065&amp;materialsCitation.latitude=14.522905" title="Search Plazi for locations around (long 74.381065/lat 14.522905)">Aghanashini River Estuary</a>, Kumta, Karnataka; 14°31'22.46"N, 74°22'51.82"E; M. Prema leg.; 12 Dec. 2022; CASAU CR 1046 • 1 ♂ (5.5 × 3.1 mm), 2 ♀♀ (5.0 × 3.5 mm, 4.5 × 2.9 mm); same collection data as previous; CASAU CR 1047 a • 1 ♂ (4.5 × 3.0 mm), 2 juv. ♀♀ (4.5 × 3.0 mm, 4.2 × 2.9 mm); same collection data as previous; CASAU CR 1047 b • 6 ♂♂ (with subequal chelae, 6.2 × 3.8 mm, 6.0 × 3.9 mm, 5.5 × 3.7 mm, 4.8 × 3.3 mm, 4.6 × 3.2 mm, 4.5 × 3.1 mm); same collection data as previous; CASAU CR 1047 c • 10 ♂♂ (6.1 × 4.0 mm, 5.8 × 3.9 mm, 5.8 × 3.7 mm, 5.3 × 3.5 mm, 5.2 × 3.6 mm, 5.2 × 3.4 mm, 5.0 × 3.3 mm, 4.6 × 3.5 mm, 4.6 × 3.3 mm, 4.4 × 3.0 mm), 19 ♂♂ (5.0 × 3.2 mm, 4.7 × 3.9 mm, 4.7 × 2.8 mm, 4.7 × 2.7 mm, 4.6 × 3.3 mm, 4.5 × 3.1 mm, 4.5 × 3.0 mm, 4.5 × 3.0 mm, 4.4 × 3.0 mm, 4.4 × 2.9 mm, 4.4 × 2.9 mm, 4.4 × 2.8 mm, 4.4 × 2.8 mm, 4.4 × 2.7 mm, 4.2 × 3.1 mm, 4.0 × 2.9 mm, 4.0 × 2.7 mm, 3.7 × 2.7 mm, 3.4 × 2.1 mm), 15 ♀♀ (5.4 × 3.4 mm, 5.1 × 3.2 mm, 5.0 × 3.5 mm, 4.9 × 3.3 mm, 4.9 × 3.2 mm, 4.8 × 3.3 mm, 4.8 × 3.0 mm, 4.7 × 3.0 mm, 4.7 × 2.9 mm, 4.6 × 3.1 mm, 4.6 × 3.0 mm, 4.6 × 2.8 mm, 4.4 × 2.5 mm, 4.3 × 3.1 mm, 4.1 × 2.3 mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.37728&amp;materialsCitation.latitude=14.5191965" title="Search Plazi for locations around (long 74.37728/lat 14.5191965)">Aghanashini River Estuary</a>, upper mudflat areas in mangroves with soft-muddy substrates and oyster beds, off Karnataka, west coast of India; 14°31'09.11"N, 74°22'38.20"E; M. Prema leg.; 22 Oct. 2025; CASAU CR 1047 d .</p><p>Additional material examined.</p><p>India • 4 ♂♂ (5.5 × 3.7 mm, 4.9 × 3.0 mm, 4.7 × 3.0 mm, 4.6 × 3.1 mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.143394&amp;materialsCitation.latitude=14.843216" title="Search Plazi for locations around (long 74.143394/lat 14.843216)">Kali River, near Kali Matha Temple</a>, soft-muddy areas in mangroves, Karwar, off Karnataka, west coast of India; 14°50'35.58"N, 74°08'36.22"E; M. Prema leg.; 07 Dec. 2022; CASAU CR 1049 . Specimens not suitable to be type material.</p><p>Comparative material examined.</p><p>Ilyoplax gangetica (Kemp, 1919) . Holotype: India • ♂ (5.7 × 3.9 mm); Kaikal, Maree, near junction of Matla and Bidyadhari rivers, Gangetic Delta, West Bengal; S. Kemp leg.; 01 Dec. 1916; ZSIK 9794 / 10. Paratype: India • 1 ♀ (5.5 × 3.6 mm); Matla Riverbank, opposite Port Canning, Canning Town, near Kolkata, West Bengal; B. Prashad leg.; 10 Mar. 1918; ZSIK 9808 / 10. Non-type: Thailand • 1 ♂ (5.6 × 4.5 mm); Ranong, mud, KR (S) ’; P. Clark leg.; 11 Nov. 2001; ZRC 2001.2339.</p><p>Ilyoplax stapletoni (De Man, 1908) Non-types: India • 5 ♂♂ (5.2 × 3.7 mm, 5.2 × 3.7 mm, 5.0 × 3.2 mm, 4.2 × 3.3 mm, 4.1 × 3.0 mm), 2 ♀♀ (3.4 × 2.4 mm, 3.2 × 2.4 mm); Diamond Harbour River side, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.19375&amp;materialsCitation.latitude=22.17861" title="Search Plazi for locations around (long 88.19375/lat 22.17861)">Hooghly River</a>; 22°10'43.0"N, 88°11'37.5"E; M. Prema leg.; 17 Jan. 2024; ZRC 2026.1330. Bangladesh • 3 ♂♂ (6.9 × 4.7 mm, 5.5 × 3.7 mm, 5.3 × 3.6 mm), 3 ♀♀ (6.3 × 4.5 mm, 5.6 × 4.5 mm, 4.8 × 3.6 mm), 1 ovigerous ♀ (5.8 × 4.6 mm); “ Jhala Kati, Backergunji dist. Bengal ” [Jhalakathi, Jhalakathi District, Barisal Division]; H. E. Stapleton leg.; Jul. 1908; ZSIK 5851 / 10 Syntypes: • 4 ♂♂ (8.4 × 5.4 mm, 7.9 × 5.3 mm, 6.6 × 4.9 mm, 6.0 × 4.9 mm), 1 ♀ (6.7 × 4.7 mm); Bangladesh and India • Kanaigunji, Backergunji district, banks of Passur River, Khulna, Bangladesh, and banks of Hugli River, near Calcutta ( Sibpur, Shalimar, Budge-Budge, and Takta Ghat, West Bengal, India); Jul. 1908; ZSIK 5137 / 10.</p><p>Ilyoplax frater (Kemp, 1919) Syn-types: Pakistan • 2 ♂♂ (7.6 × 5.2 mm, G 1 detached; 5.8 × 3.8 mm), 1 ♀ (4.7 × 3.8 mm); Karachi; collector and date not known; ZSIK 9861 / 10. Non-types: Kuwait • 1 ♂ (5.5 × 3.5 mm); station SDG 57, Kuwait; S. De Grave leg.; 2014 / 2015; ZRC 2021.0495.</p><p>Ilyoplax stevensi (Kemp, 1919) Syn-types: Pakistan • 1 ♂ (7.6 × 4.8 mm), 2 females (7.3 × 4.9 mm, 4.8 × 3.5 mm), 1 ovigerous ♀ (6.5 × 4.9 mm), 6 ♂♂ (5.7 × 3.7 mm, 4.7 × 3.2 mm, 4.7 × 2.6 mm, 4.5 × 2.6 mm, 3.8 × 2.3 mm, 3.5 × 2.6 mm), 2 ♀♀ (6.0 × 3.6 mm, 4.7 × 2.9 mm); Karachi; collector and date not known; ZSIK 9796 / 10. Non-types: Kuwait • 2 ovigerous ♀♀ (9.0 × 5.7 mm, 5.6 × 4.5 mm); station SDG 56, Kuwait; S. De Grave leg.; 2014 / 2015; ZRC 2021.0494.</p><p>Ilyoplax delsmani De Man, 1926 Non-types: Singapore • 1 ♂ (4.5 × 3.7 mm); Sungai Buloh Mangrove, Singapore; P. K. L. Ng leg.; 10 Jan. 1992; ZRC 1992.7927 – 7935.</p><p>Diagnosis.</p><p>Carapace rectangular, smooth; suborbital margin subovate, lined with evenly spaced small, rounded granules; stridulatory ridge with 22–24 subconical tubercles, inner half of ridge with 5–7 fungiform tubercles, last tubercle being largest, with cap of tubercle fully corneous; epistome triangular. Cheliped massive, carpus short, inner carpal spine blunt, palm broad and lower margin lined with 2 rows of granules gradually increased size towards dactylus; upper margin of dactylus granulated; P 2 – P 4 of similar length, relatively broad, merus length ca. 2.5 times width, tympanum present; carpus and propodus of P 2 and P 3 with dense patch of plumose setae in male. Female carapace rectangular, suborbital margin subovate, lined with evenly spaced small, conical granules; stridulatory ridge with 22 subconical tubercles, densely arranged, inner half ridge with 4–6 small fungiform tubercles; chelipeds small, subequal; vulvae on sternite 6, comma-shaped, vulvar cover absent.</p><p>Description.</p><p>Carapace (Figs 1 A, 1 C, 2 A, 7 A) transversely rectangular, smooth, ca. 1.5–1.6 times wider than long, maximum length medially; regions poorly defined; microscopically granular, with low ridges and transverse setae. Front broad, ca. one-quarter distance between external orbital angles, deflexed, lateral margins converging anteriorly, frontal margin bilobed with gently convex margins when viewed dorsally, median concavity shallow (Fig. 1 C). Gastrocardiac groove defined; cardiac region swollen, well defined by clear ridge with anterior margin deeply curved medially, straight, sloping posteriorly. Branchial regions with short transverse ridges or crests, with short stellate setae. Posterior carapace margin straight, broadly rounded laterally, with broad rim; supraorbital margin smooth, sinuous, sloping obliquely; external orbital angle forming acutely triangular, sharp tooth, separated from remainder of margin by distinct concavity; anterolateral margin minutely denticulated (Fig. 2 A, G), first anterolateral tooth low, without trace of second tooth or lobe; lateral margins slightly diverging posteriorly. Eyestalks elongated, peduncle thick, short; cornea bulbous (Figs 1, 2 C, 3 A, C, E).</p><p>Male: Suborbital margin subovate, lined with evenly spaced small rounded granules; stridulatory ridge immediately below infraorbital margin long, starting outside lateral edge of orbit, appressed against margin medially, before bifurcating around median part of suborbital region (Fig. 3 A, B, D, G); outer (lower) half of main stridulatory ridge with 22–24 subconical tubercles, each with corneous tip, gradually becoming larger medially, inner half of ridge with 5–7 fungiform tubercles, last one being largest, with cap of tubercle entirely corneous (Figs 2 C, 2 E, 3 D, 3 G, 3 H); upper part of bifurcated stridulatory ridge shorter than lower ridge, extending to near base of ocular peduncle, lined with subconical tubercles without corneous tip (Fig. 3 D, G); additional short ridge of stridulatory granules below and subparallel to outer part of main stridulatory ridge, with 25 rounded tubercles, each with corneous tip, outer tubercles largest, closely packed, becoming smaller and more spaced out inwards (Fig. 3 H). Female: Similar to male, except with additional short ridge of stridulatory granules below outer part of main stridulatory ridge with 30 rounded tubercles and curving downwards along inner part (Fig. 3 A, B, D).</p><p>Suborbital, subhepatic, sub-branchial and pterygostomial regions covered with evenly spaced short stellate setae (Fig. 3 A – H); external orbital tooth lined with long papposerrate (Fig. 3 I); row of long pappose setae on anterolateral tooth (Figs 2 G, 3 B, 3 G – I).</p><p>Maxilliped 3 (Fig. 2 I) moderately bulging or convex, completely closing buccal cavity; merus longer than ischium, surface with evenly spaced obliquely stellate setae; antero-internal angle of ischium triangular, apically blunt, rounded, produced along edge of merus; ischium anterior margin deeply curved with line of plumose setae densely arranged, surface with few stellate setae, granules absence.</p><p>Chelipeds (Fig. 4 A, C) stout, heterochelous; merus trihedral, inner and outer margins granular; carpus short, rhomboidal, armed with short inner tooth. Major chela of adult male holotype distinctly swollen, palm broader, slenderer, margin lined with granules gradually increased size towards dactylus; dactylus upper margin granulated, with broad gap when fingers closed, cutting margin with large, rounded teeth on proximal half and small evenly sized teeth on distal half; pollex cutting margin with small, evenly sized teeth along length (Fig. 5 A, B); minor chela similar in form to major chela but proportionately more slender (Figs 1 B, 4 A). Female chelipeds (Figs 5 C, 5 D, 7 C) small, subequal; carpus rounded, inner carpal spine short; palm with 1 row of rounded granules, tip of dactylus and pollex spatulate.</p><p>P 2 – P 4 length subequal, relatively broad, P 5 shortest. P 2 – P 4 merus length ca. 2.5 times width, about as long as carpus and propodus combined, P 2 – P 5 merus margins denticulated inner surface tympanum present (Fig. 5 F); dactylus slender, smooth, longitudinal grooves with row of simple setae, tapering to point, shorter than propodus (Figs 5 E, 6 E – H, 7 A); carpus and propodus of P 2 and P 3 with dense patch of setae in male (Figs 4 A, 5 E, 5 G), female with less dense bed of setae only on carpus and part of propodus of P 3, absent on P 2 (Figs 1 C, 1 D, 7 A).</p><p>Gastric mill: median tooth plate (Fig. 6 A, B) broad, smooth surface, pentagonal, surface slightly convex laterally, convex medially, simple structure, consisting of 1 tooth, each side of lateral margin with 3 curved, pointed, simple spines. Lateral tooth plate (Fig. 6 C – I) consisting of 8 teeth, including larger tooth with cup-like structure; each tooth with row of short, bristle-like setae anteriorly and tuft of long, serrulate setae on lateral edge (Fig. 6 C, D); lateral tooth palm-like with 7 closely appressed branches, each branch densely covered with scaled macrosetae (Fig. 6 C, E, F); anteriorly with cusp-like fang structure comprised of triangular plate, each with long, curved, tuft of serrate setae on edges (Fig. 6 G – I).</p><p>Male thoracic sternum wide; male sternopleonal cavity reaching to base of maxilliped 3 (Fig. 2 I); male telson (Fig. 4 G) elongate, apically rounded, slightly shorter than pleonal somite 6; pleonal somite 5 much shorter than pleonal somite 6, markedly constricted at base; pleonal somites 3 and 4 similar, wider at base forming triangular margin laterally; pleonal somites 1 and 2 similar in width and subparallel to posterior margin of carapace (Fig. 4 G). Female pleon relatively narrow, not fully covering sternum, with short, semicircular telson; female sternum wide, sparsely distributed plumose setae (Fig. 7 E, G, H); female sternopleonal cavity broad, smooth, vulvae located medially on sternite 6, divergent towards median region, simple, comma-shaped, vulvar cover absent (Fig. 7 I).</p><p>G 1 long, slender, sinuous, proximal third gently bent at angle; distal part prominently elongate, triangular, gently curved, bent obliquely outwards at ca. 90 ° from longitudinal axis of G 1 distal portion; subdistal part prominently swollen just before bend, rounded (Figs 8 A – D, 9 A – D). G 2 short with broad base and slender distal part (Fig. 8 E).</p><p>Variations.</p><p>Carapace in larger males have the ridges lined with more stellate setae and with tufts of short setae clearly visible in the frontal view, with surface more densely packed with minute granules; supra orbital margin microscopically granulated in young males more prominent in larger specimens; chelipeds subequal in few males of both large and small size; palm broader, surface smooth, equal in size, with the lower margin lined with granules that are gradually increased size towards dactylus, and the upper margins of palm and dactylus denticulated; some males have the margin of the dactylus and adjacent part granulated. In both males and females, the number of tubercles on the stridulatory ridge after the bifurcating inner (upper) part varies between 5–8.</p><p>Etymology.</p><p>Ilyoplax kagga sp. nov. is named in recognition of an endemic rice “ Kagga ”, a highly salt-tolerant variety that is cultivated traditionally and only in the type locality, Aghanashini village. The name also honours the many generations of ethnic farmers who have preserved and cultivated this endemic rice variety through traditional practices. The specific epithet is used as a noun in apposition.</p><p>Live colour.</p><p>Male dark brown dorsally; reddish on inner side of ambulatory legs, P 2 and P 3 with tuft of soft, plumose setae densely arranged on outer surface of carpus and propodus is pale-whitish in colour (Fig. 10 A); outer surface of the chela pale brownish-green palm when freshly frozen (Fig. 1 B) and more white with brown spotted pattern in life (Fig. 10). Female carapace greenish brown when freshly frozen (Fig. 1 C) and reddish in life (Fig. 10 B); chelipeds pale white with blackish network pattern (Fig. 1 C, D).</p><p>Habitat.</p><p>The new species was collected from the upper tidal regions of the mangroves and oyster beds. It lives in the burrows of muddy-sandy bottom in the upper limit of intertidal areas. This species is often difficult to collect because of its small body size, cryptic colouration and burrowing behaviour (Figs 10 E – G, 11 C, 11 D), often remaining concealed in narrow burrows and emerging only briefly under suitable tidal and environmental conditions for feeding (Fig. 10 G, H). We found this species in large numbers only in Aghanashini River Estuary (Figs 10, 11), with much fewer individuals in the Islands of Kali River Estuary at Karwar.</p><p>Distribution.</p><p>Ilyoplax kagga sp. nov. is currently known from the coastal villages of Aghanashini near Kumta and Kali River Estuary near Karwar on the west coast of India, Karnataka, Arabian Sea. The species might also occur in the Mandavi River Estuary, Goa, on the west coast of India.</p><p>Remarks.</p><p>The genus Ilyoplax Stimpson, 1858, contains 28 recognised species, of which five species are known from the Indian subcontinent (Ng et al. 2008; Davie and Naruse 2010; Trivedi et al. 2015, 2018; Poore and Ahyong 2023). In general appearance, Ilyoplax kagga sp. nov. is most similar to I. gangetica (Kemp, 1919) but can be easily distinguished by the following characters: (1) carapace rectangular, smooth, ca. 1.5–1.6 times wider than long, with the upper orbital margin smooth and the lateral parts minutely granulated (Figs 1 A, 1 C, 2 A, 7 A) (vs. carapace subrectangular, ca. 1.2–1.4 times as wide as long, with the upper orbital as well as lateral margins microscopically beaded / minutely granulated in I. gangetica; Figs 2 B, 4 F, 7 B); (2) external orbital angle more acute and sharp, followed by a relatively broader and deeper concavity, with the first anterolateral tooth low and no second tooth visible (Figs 2 G, 3 E) (vs. external orbital angle more obtuse, wider, followed by a narrow and deep concavity, with the first tooth low but broad and separated from very low second tooth by shallow concavity in I. gangetica; Figs 2 H, 7 B); (3) epistome slightly more elongated longitudinally, with the median lobe relatively narrower and more triangular (Figs 2 C, 4 A, 4 C) (vs. epistome longitudinally shorter and the median lobe proportionately broader and truncate in I. gangetica; Figs 2 D, 7 D); (4) stridulatory ridge starting outside lateral edge of orbit, bifurcated around median part of suborbital region, outer (lower) half of main stridulatory ridge with 22–24 subconical, corneous tubercles, inner half of ridge with 5–7 fungiform tubercles, last tubercle being largest, with cap of tubercle entirely corneous (Figs 2 C, 2 E, 3 D, 3 G, 3 H), upper part of bifurcated stridulatory ridge lined with subconical tubercles without corneous tip, additional short ridge of stridulatory granules below and subparallel to outer part of main stridulatory ridge, with 25 rounded tubercles in male (Fig. 3 G, H), being similar for the female except that the additional ridge has 30 tubercles which is longer and curves downwards along the inner part (Fig. 3 A, B, D) (vs. stridulatory ridge starting outside lateral edge of orbit, not bifurcated, with a single row of stridulatory ridge of 22–25 pectinated comb-like teeth, densely arranged, gradually becoming larger inwards and is more spaced out, without any trace of an additional short ridge in both males and females of I. gangetica; Fig. 2 F); (5) adult male chelipeds usually heterochelous (Fig. 4 C), the carpus being relatively broader and shorter with the inner distal spine more rounded, the palm being short, broad, with the lower margin lined with 2 rows of rounded granules, the lower margin of the fixed finger and the upper margin of the movable finger gently serrated (Figs 1 A, 1 B, 2 A, 4 A, 4 C, 5 A – D) (vs. adult male chelipeds subequal, carpus distinctly elongate and more slender, inner distal spine sharp, the palm being relatively longer, narrower, the lower margin with 2 rows of rounded, minute granules all the way to fixed finger, upper and lower margins of dactylus and propodal fingers minutely serrated in I. gangetica; Fig. 4 B, D; see also Kemp 1919: 341, fig. 20; Chhapgar and Borgaonkar 1985: 227, fig. 1); (6) ambulatory legs proportionately longer, merus slender with tufts of setae present on the carpus and propodus of P 2 and P 3 in male (Figs 1 A, 4 E, 5 E, 5 G), the setae being absent on P 2 in females and present but less dense on P 3 (Figs 1 C, 7 A) (vs. ambulatory legs proportionally shorter, merus broad with the tuft of setae absent or minutely present in I. gangetica; Figs 4 F, 7 B; see also Chhapgar and Borgaonkar 1985: 227, fig. 1); and (7) G 1 long, slender, sinuous, with the proximal third appearing gently bent at an angle, distal part prominently elongate, triangular, gently curved, bent obliquely outwards at angle of ca. 90 ° from longitudinal axis of G 1 distal portion, subdistal part just before bend prominently swollen, rounded (Fig. 8 A – D) (vs. G 1 very long, more slender, more strongly sinuous, distal part relatively short, triangular with straight margins, gently directed obliquely outwards at angle of ca. 70 ° from longitudinal axis of G 1 distal portion, subdistal part just before bend relatively low, obtusely triangular, rounded but not swollen in I. gangetica; Fig. 8 F – I; see also Chhapgar et al. 2004: 185, fig. 1).</p><p>Ilyoplax kagga sp. nov. shares some characteristics with other congeners. For instance, the cheliped, epistome and stridulatory ridge pattern of I. kagga sp. nov. resemble those of I. delsmani De Man, 1926, and I. stapletoni (De Man, 1908) . Ilyoplax delsmani, however, has a very long spine on the inner angle of the carpus of the cheliped (see De Man 1926: fig. 10), but this is completely absent in I. stapletoni (present study; see De Man 1908: pl. 18, fig. 1); in I. kagga sp. nov., the spine is short and rounded (Figs 1 A, 2 A). The corneous tubercles on the stridulatory ridges, with their sizes gradually becoming larger medially, are similar, but the shape of the tubercles is significantly different. The tubercles are conical with a pointed tip in I. delsmani (see De Man 1926: fig. 3), conical and rounded in I. stapletoni (present study), and fungiform in I. kagga sp. nov. (Figs 2 E, 3 A, 3 B, 3 D, 3 G, 3 H).</p></div>	https://treatment.plazi.org/id/409E777A864855539CCF56B0EFBA13D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.	Prema, Mani, Kakati, Vithal S., Ng, Peter K. L. (2026): A new species of intertidal crab of the genus Ilyoplax Stimpson, 1858 (Decapoda, Brachyura, Dotillidae) from the Aghanashini River Estuary, a RAMSAR site on the west coast of India. ZooKeys 1291: 19-38, DOI: 10.3897/zookeys.1291.201892
6D50370626EC5D55B34DA29A7DC9CBC9.text	6D50370626EC5D55B34DA29A7DC9CBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ilyoplax Stimpson 1858	<div><p>Genus Ilyoplax Stimpson, 1858</p><p>Type species.</p><p>Ilyoplax tenella Stimpson, 1858 (by monotypy). Gender of genus feminine.</p><p>Remarks.</p><p>Davie and Naruse (2010) have discussed the challenge with Ilyoplax, noting that the genus as presently defined is clearly polyphyletic, with several distinct lineages. They indicated that a detailed taxonomic revision is required (see also Ng et al. 2008: 235).</p></div>	https://treatment.plazi.org/id/6D50370626EC5D55B34DA29A7DC9CBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.	Prema, Mani, Kakati, Vithal S., Ng, Peter K. L. (2026): A new species of intertidal crab of the genus Ilyoplax Stimpson, 1858 (Decapoda, Brachyura, Dotillidae) from the Aghanashini River Estuary, a RAMSAR site on the west coast of India. ZooKeys 1291: 19-38, DOI: 10.3897/zookeys.1291.201892
