taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762527	https://doi.org/10.3897/zookeys.1291.201892.figure1	Figure 1. Freshly preserved colouration of Ilyoplax kagga sp. nov., Aghanashini River Estuary, west coast of India. A, B. Dorsal and frontal views of holotype male (4.2 × 2.5 mm) (CASAU CR 1048), respectively; C, D. Dorsal and ventral views of paratype ovigerous female (4.5 × 2.9 mm) (CASAU CR 1047), respectively.	Figure 1. Freshly preserved colouration of Ilyoplax kagga sp. nov., Aghanashini River Estuary, west coast of India. A, B. Dorsal and frontal views of holotype male (4.2 × 2.5 mm) (CASAU CR 1048), respectively; C, D. Dorsal and ventral views of paratype ovigerous female (4.5 × 2.9 mm) (CASAU CR 1047), respectively.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762528	https://doi.org/10.3897/zookeys.1291.201892.figure2	Figure 2. Ilyoplax kagga sp. nov. (A, C, E, G, I) holotype male (4.2 × 2.5 mm) (CASAU CR 1048), Aghanashini River Estuary, west coast of India. Ilyoplax gangetica (Kemp, 1919) (B, D, F, H, J) male (5.6 × 4.5 mm) (ZRC 2001.2339), Ranong, Thailand. A, B. Dorsal view of carapace and carpus of chelipeds; C, D. Frontal view of cephalothorax; E. Stridulatory ridge with fungiform corneous tubercles, left side in ventral view; F. Stridulatory ridge with pectinated tubercles, right side in ventro-frontal view; G, H. Anterolateral teeth of carapace, dorsal view; I, J. Maxilliped 3, outer view.	Figure 2. Ilyoplax kagga sp. nov. (A, C, E, G, I) holotype male (4.2 × 2.5 mm) (CASAU CR 1048), Aghanashini River Estuary, west coast of India. Ilyoplax gangetica (Kemp, 1919) (B, D, F, H, J) male (5.6 × 4.5 mm) (ZRC 2001.2339), Ranong, Thailand. A, B. Dorsal view of carapace and carpus of chelipeds; C, D. Frontal view of cephalothorax; E. Stridulatory ridge with fungiform corneous tubercles, left side in ventral view; F. Stridulatory ridge with pectinated tubercles, right side in ventro-frontal view; G, H. Anterolateral teeth of carapace, dorsal view; I, J. Maxilliped 3, outer view.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762529	https://doi.org/10.3897/zookeys.1291.201892.figure3	Figure 3. SEM micrographs of Ilyoplax kagga sp. nov. (A – F) female (5.0 × 3.5 mm), (G – I) male (5.5 × 3.1 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast of India. A. Pterygostomial region showing evenly spaced setae, suborbital margin and stridulatory ridge; B, C, G, H. Stridulatory ridge starting outside lateral edge of external orbit with long papposerrate; D. Bifurcated stridulatory ridge and suborbital region, and outer (lower) half of main stridulatory ridge showing 6 fungiform corneous tubercles with 9 conical tubercles; E, F. Short stellate setae in different magnifications; I. Magnified view of papposerrate setae on external orbital tooth, near to anterolateral teeth.	Figure 3. SEM micrographs of Ilyoplax kagga sp. nov. (A – F) female (5.0 × 3.5 mm), (G – I) male (5.5 × 3.1 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast of India. A. Pterygostomial region showing evenly spaced setae, suborbital margin and stridulatory ridge; B, C, G, H. Stridulatory ridge starting outside lateral edge of external orbit with long papposerrate; D. Bifurcated stridulatory ridge and suborbital region, and outer (lower) half of main stridulatory ridge showing 6 fungiform corneous tubercles with 9 conical tubercles; E, F. Short stellate setae in different magnifications; I. Magnified view of papposerrate setae on external orbital tooth, near to anterolateral teeth.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762530	https://doi.org/10.3897/zookeys.1291.201892.figure4	Figure 4. Ilyoplax kagga sp. nov. (A, E, G) holotype male (4.2 × 2.5 mm) (CASAU CR 1048), (C) paratype male with subequal chelae (6.2 × 3.8 mm) (CASAU CR 1047 c), Aghanashini River Estuary, west coast of India. I. gangetica (Kemp, 1919) (B, D, F, H) male (5.6 × 4.5 mm) (ZRC 2001.2339), Ranong, Thailand. A – D. Outer view of cheliped; E, F. P 2 – P 4, dorsal view; G, H. Pleon, telson and sternum.	Figure 4. Ilyoplax kagga sp. nov. (A, E, G) holotype male (4.2 × 2.5 mm) (CASAU CR 1048), (C) paratype male with subequal chelae (6.2 × 3.8 mm) (CASAU CR 1047 c), Aghanashini River Estuary, west coast of India. I. gangetica (Kemp, 1919) (B, D, F, H) male (5.6 × 4.5 mm) (ZRC 2001.2339), Ranong, Thailand. A – D. Outer view of cheliped; E, F. P 2 – P 4, dorsal view; G, H. Pleon, telson and sternum.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762531	https://doi.org/10.3897/zookeys.1291.201892.figure5	Figure 5. SEM micrographs of Ilyoplax kagga sp. nov. (A, B, E – H) male (5.5 × 3.1 mm), (C, D) female (5.0 × 3.5 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast of India. A, B. Male cheliped, row of rounded, smaller and larger granules on surface of palm; C, D. Female cheliped outer view, carpus rounded, granulated margin, and with single row of small rounded granules on palm towards proximal half of fixed finger; E – H. Male P 3 with tuft of plumose setae, merus upper margin denticulated, tympani present minutely, dactylus with longitudinal grooves.	Figure 5. SEM micrographs of Ilyoplax kagga sp. nov. (A, B, E – H) male (5.5 × 3.1 mm), (C, D) female (5.0 × 3.5 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast of India. A, B. Male cheliped, row of rounded, smaller and larger granules on surface of palm; C, D. Female cheliped outer view, carpus rounded, granulated margin, and with single row of small rounded granules on palm towards proximal half of fixed finger; E – H. Male P 3 with tuft of plumose setae, merus upper margin denticulated, tympani present minutely, dactylus with longitudinal grooves.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762532	https://doi.org/10.3897/zookeys.1291.201892.figure6	Figure 6. SEM of gastric teeth and microstructures of the associated setae of Ilyoplax kagga sp. nov., male (5.5 × 3.1 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast of India. A. Overall view of gastric mill; B. Smooth median tooth with 3 spines on each side; C, D. Lateral tooth anterior margin with row of short bristle-like setae, and lateral edge with tuft of serrulate setae, right side, dorsal view; E, F. 7 branched palm-like structure with scaled macrosetae present on posterior margin of lateral tooth; G – I. Lateral tooth anterior margin with row of short bristle-like setae, and lateral edge with tuft of serrulate setae, and 7 branched palm-like structure in posterior margin and cusp-like structure on anterior margin; H, I. Magnified view of cusp-like structure and long, curved, tuft of serrate setae.	Figure 6. SEM of gastric teeth and microstructures of the associated setae of Ilyoplax kagga sp. nov., male (5.5 × 3.1 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast of India. A. Overall view of gastric mill; B. Smooth median tooth with 3 spines on each side; C, D. Lateral tooth anterior margin with row of short bristle-like setae, and lateral edge with tuft of serrulate setae, right side, dorsal view; E, F. 7 branched palm-like structure with scaled macrosetae present on posterior margin of lateral tooth; G – I. Lateral tooth anterior margin with row of short bristle-like setae, and lateral edge with tuft of serrulate setae, and 7 branched palm-like structure in posterior margin and cusp-like structure on anterior margin; H, I. Magnified view of cusp-like structure and long, curved, tuft of serrate setae.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762533	https://doi.org/10.3897/zookeys.1291.201892.figure7	Figure 7. A, C, E, G, H, I. Ilyoplax kagga sp. nov., paratype, female (5.4 × 3.4 mm) (CASAU CR 1047 d), Aghanashini River Estuary, west coast of India; B, D, F. Ilyoplax gangetica Kemp, (1919), paratype, ovigerous female (5.5 × 3.6 mm) (ZSI 9808 / 10), Matla Riverbank, opposite Port Canning, West Bengal, India; A, B. Dorsal view; C, D. Frontal view of cephalothorax; E – G. Sterno-pleonal view; H, I. SEM micrograph of sternum and vulvae.	Figure 7. A, C, E, G, H, I. Ilyoplax kagga sp. nov., paratype, female (5.4 × 3.4 mm) (CASAU CR 1047 d), Aghanashini River Estuary, west coast of India; B, D, F. Ilyoplax gangetica Kemp, (1919), paratype, ovigerous female (5.5 × 3.6 mm) (ZSI 9808 / 10), Matla Riverbank, opposite Port Canning, West Bengal, India; A, B. Dorsal view; C, D. Frontal view of cephalothorax; E – G. Sterno-pleonal view; H, I. SEM micrograph of sternum and vulvae.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762534	https://doi.org/10.3897/zookeys.1291.201892.figure8	Figure 8. A – E. Ilyoplax kagga sp. nov., holotype male (4.2 × 2.5 mm) (CASAU CR 1048); F – J. Ilyoplax gangetica (Kemp, 1919), male (5.6 × 4.5 mm) (ZRC 2001.2339), Ranong, Thailan; A, F. Left G 1, dorsal view; B, G. Left G 1, ventral view; C, H. Left G 1, distal tip, dorsal view; D, I. Left G 1, distal tip, ventral view; E. Right G 2, dorsal view; J. Left G 2, dorsal view. Scale bars: 0.2 mm (A – D, F – I); 0.1 mm (E, J).	Figure 8. A – E. Ilyoplax kagga sp. nov., holotype male (4.2 × 2.5 mm) (CASAU CR 1048); F – J. Ilyoplax gangetica (Kemp, 1919), male (5.6 × 4.5 mm) (ZRC 2001.2339), Ranong, Thailan; A, F. Left G 1, dorsal view; B, G. Left G 1, ventral view; C, H. Left G 1, distal tip, dorsal view; D, I. Left G 1, distal tip, ventral view; E. Right G 2, dorsal view; J. Left G 2, dorsal view. Scale bars: 0.2 mm (A – D, F – I); 0.1 mm (E, J).	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762535	https://doi.org/10.3897/zookeys.1291.201892.figure9	Figure 9. SEM of G 1 of Ilyoplax kagga sp. nov., paratype, male (5.5 × 3.1 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast India. All images are from left G 1, same position at different levels of higher magnification. A. Dorsal view of G 1; B, C. Distal region of G 1 with stiff simple setae; D. Swollen subdistal region with 2 lobes formed by a median groove.	Figure 9. SEM of G 1 of Ilyoplax kagga sp. nov., paratype, male (5.5 × 3.1 mm) (CASAU CR 1047), Aghanashini River Estuary, west coast India. All images are from left G 1, same position at different levels of higher magnification. A. Dorsal view of G 1; B, C. Distal region of G 1 with stiff simple setae; D. Swollen subdistal region with 2 lobes formed by a median groove.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762536	https://doi.org/10.3897/zookeys.1291.201892.figure10	Figure 10. Live colour of Ilyoplax kagga sp. nov. A. Male (4.6 × 3.1 mm) (CASAU CR 1049), Karwar, near Kali Matha Temple, from Island mangroves with Rhizophora; B. Female with brownish carapace; C. Waving behaviours of male individual; D. Crab feeding near to its burrow with tuft of setae on first two ambulatory legs clearly visible; E, F. Burying in sandy-muddy habitat; G, H. Submerged in water and feeding; I. Pterygostomial region, inner side of ambulatory legs pigmented with mottled red pattern; B – I. Specimens were not collected.	Figure 10. Live colour of Ilyoplax kagga sp. nov. A. Male (4.6 × 3.1 mm) (CASAU CR 1049), Karwar, near Kali Matha Temple, from Island mangroves with Rhizophora; B. Female with brownish carapace; C. Waving behaviours of male individual; D. Crab feeding near to its burrow with tuft of setae on first two ambulatory legs clearly visible; E, F. Burying in sandy-muddy habitat; G, H. Submerged in water and feeding; I. Pterygostomial region, inner side of ambulatory legs pigmented with mottled red pattern; B – I. Specimens were not collected.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
409E777A864855539CCF56B0EFBA13D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1762537	https://doi.org/10.3897/zookeys.1291.201892.figure11	Figure 11. Habitat of Ilyoplax kagga sp. nov. at Aghanashini River Estuary, a RAMSAR site. A. Upper mudflat bordered by Rhizophora mangroves. B. Newly planted Rhizophora along upper mudflat area; C. Soft-sediment mudflat area at low tide; D. Crab (white dotted circle) feeding on surface among pneumatophores of Rhizophora; E. Mudflat area at low tide with young Rhizophora; F, G. River bordered by mangroves and paddy field of endemic Kagga rice variety adjacent to the mangroves.	Figure 11. Habitat of Ilyoplax kagga sp. nov. at Aghanashini River Estuary, a RAMSAR site. A. Upper mudflat bordered by Rhizophora mangroves. B. Newly planted Rhizophora along upper mudflat area; C. Soft-sediment mudflat area at low tide; D. Crab (white dotted circle) feeding on surface among pneumatophores of Rhizophora; E. Mudflat area at low tide with young Rhizophora; F, G. River bordered by mangroves and paddy field of endemic Kagga rice variety adjacent to the mangroves.	2026-09-01	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.		Zenodo	biologists	Prema, Mani;Kakati, Vithal S.;Ng, Peter K. L.			
