taxonID	type	description	language	source
C80DC93ED36868233992FD6AFADADFAF.taxon	materials_examined	Type material: Holotype (Fig. 1): Male, “ STS- 85454, Sejila Mountain Research Station, Bayi District, Linzhi City (Nyingchi), Xizang, China, 29.64849 ° N, 94.70981 ° E, 22 July-August 2021, altitude 3900 m (collected by Chen Enyong) ”. Paratype (Fig. 2): 1 male, “ STS- 110067, Sejila Mountain Research Station, Bayi District, Linzhi City (Nyingchi), Xizang, China, 29.64849 ° N, 94.70981 ° E, 22 July-August 2021, altitude 3900 m (collected by pest forecast lamp) ”.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36868233992FD6AFADADFAF.taxon	description	Description: Wingspan 27 mm (Figs. 1 A, 1 B). Head blackish brown, palpi projecting before the frons: male antennae thinly quadripectinate. Forewings with blackish brown markings enlarged, transverse lines thin and whitish, sometimes patchily enlarged between veins. The longest lobe of the medial area is relatively short, reaching about the middle of the distal third of the wing. The apical streak is weak and grey. Fringe blackish on the forewing, lighter grey on hindwing. Hindwings greyish, gradually becoming darker grey distally, medial line strongly curved and grey, delimiting the darker medial area, with the discal spot present. Forewing underside darker than hindwing underside, discal spot linear on the forewing, as long as the discocellular vein, and punctate on the hindwing. Male genitalia (Fig. 1 C): Uncus is as long as the tegumen from the anterior edge to the base of the uncus. Vinculum trapezoidal, massive, with a straight anterior edge; tegumen well sclerotised, approximately triangular. The saccular part of the valva triangular, its dorsal edge not heavily sclerotised, slightly concave, ending in an inward-pointing hook from a moderately wide step. Valvula rounded, broader than the sacculus, setose. The sclerite of the costa broadens distally, its ventral edge raised in the basal half of the valva. Aedeagus trumpet-shaped (Fig. 1 D), shorter than the valva, with three sets of black, pencil-shaped cornuti (longer than the aedeagus sheath), and the base of the vesica is wide. No cornuti on vesica. Labides roundish, setose.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36868233992FD6AFADADFAF.taxon	diagnosis	Diagnosis: Heterothera sejilaensis sp. n, with its dark markings enlarged on the forewing, is similar to Heterothera incerta from Taiwan and Heterothera obscurata from western Nepal. It differs from H. incerta in the structure of the male genitalia: the valva apex of H. sejilaensis is relatively obtuse and rounded, and it possesses cornuti only at the base of the vesica. Heterothera pailongensis has cornuti on the vesica membrane and at its base. H. obscurata has a smaller number of cornuti on the aedeagus vesica in the male genitalia.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36868233992FD6AFADADFAF.taxon	biology_ecology	Ecological characteristics: This species was collected from the Xizang Linzhi (Nyingchi) Alpine Forest Ecosystem National Field Science Observation and Research Station, within a fir forest at an altitude of 3900 meters. The area belongs to a typical subalpine temperate semi-humid climate zone, with the main vegetation type being mountain temperate dark coniferous forest. The most common species in this forest are Picea likiangensis var. linzhiensis (Linzhi spruce) and Abies georgei var. smithii (Smith’s fir). Other main vegetation includes Pinus densata (alpine pine) and Quercus semecarpifolia (Himalayan oak).	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36868233992FD6AFADADFAF.taxon	etymology	Etymology. This species is named after Sejila Mountain, the location where the holotype was collected.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36A68253992FF36FF7EDDB6.taxon	materials_examined	Type material: Holotype (Fig. 3): Male, “ STS- 24210, Eastern Slope of Sejila Mountain, Bayi District, Linzhi City (Nyingchi), Xizang, China, 26 May 2020, altitude 2550 m (collected by Xian Chunlan) ”; Paratypes (Fig. 4): 1 female, “ STS- 4856, 106 k, Linzhi City (Nyingchi), Xizang, China, 29.999877 ° N, 94.884378 ° E, 2 October 2018, altitude 2356 m (collected by Zhou Runfa) ”. 1 female, “ STS- 57395, Laohuzui, Linzhi (Nyingchi) City, Xizang, China, 18 July 2021, (collected by Pan Zhaohui, Jiang Xinrui, Chen Le) ”.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36A68253992FF36FF7EDDB6.taxon	description	Description: Wingspan 31 mm (Figs. 3 A, 3 B). Head grey-brown. Palpi projecting about half the length of the eye diameter; antennae of males quadripectinate with thin, long rami. Rami branch from the apical and basal parts of antennal segments. Forewing dark ochreous brown, transverse fasciae and the lining on veins thin, more or less contrasting dark grey, the medial area is costally darker, its last segment at the hindmargin of wing is black as in the previous species. Hindwing whitish, distally suffused with greyish with the cell-spot dark grey, the transverse fascia is fairly greyish, accentuated by grey streaks on veins, straight from the costa to vein M 3, from where it turns rectangularly to the hindmargin of the wing. Male genitalia (Figs. 3 C, 3 D): The male genital armature is well sclerotised and massive. The valvula is roundish, a bit broader than the sacculus. The saccular part of valva is distally straight-edged, with a small process. The dorsal edge of costa is straight, before the apex obliquely cut. The ventral edge of the costal sclerite is rib-like and raised. The vinculum is broader than the tegumen, rectangular, ventrally with straight or slightly concave edge. The aedeagus is shorter than the valva, distally wide, with three sets of black, pencil-shaped distally and a set of smaller thorns on the vesica. Female genitalia (Figs. 4 B, 4 D): The ovipositor lobes are relatively large, anterior apophyses filiform, half the length of the posterior apophyses. The antrum is large and semi-spherical. The pyriform bursa copulatrix is membranous, without a signum.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36A68253992FF36FF7EDDB6.taxon	diagnosis	Diagnosis: Heterothera pailongensis sp. nov. is most similar to H. taigana from Siberia and H. etes from Assam in its simple dark markings on the forewings, but differs in the presence of hindwing discal spots and in three white spots that divide the hindmargin into four equal parts. The black streak of the middle area on the hindmargin of the forewing is shared with Heterothera sejilaensis sp. nov. There are short cornuti on the vesica and long pencil-shaped cornuti at the base of the vesica.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36A68253992FF36FF7EDDB6.taxon	biology_ecology	Ecological characteristics: This species was collected near Lulang and Pailong Town on the eastern slope of Sejila Mountain at an altitude of 2550 meters, which is located in the evergreen broad-leaved forest and coniferousbroadleaved mixed forest zone. The area has a subtropical climate with relatively high temperature and humidity. The vegetation is dominated by Fagaceae and Theaceae, along with plants from Ericaceae. The flora is rich in species and contains various warm and humid components. The broad-leaved mixed forests include arbor and shrub species such as Quercus lanata (Tongmai oak), Quercus semecarpifolia (Himalayan oak), and Machilus nanmu (nanmu).	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36A68253992FF36FF7EDDB6.taxon	etymology	Etymology. This species is named after Pailong Town, where the holotype was collected near Linzhi (Nyingchi), Xizang.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36D68263992F98EFC1CD9C6.taxon	materials_examined	Type material (Fig. 5): “ 1 male, STS- 79496, 62 K, Medog County, Linzhi (Nyingchi) City, Xizang Autonomous Region, China, 29.709333 ° N, 95.582222 ° E, 20 July 2022, altitude 2788 m (collected by Pan Zhaohui). ”	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36D68263992F98EFC1CD9C6.taxon	description	Description: The species is well characterised in the original description. Males have bipectinate antennae. The male has a wingspan of 28 – 29 mm (Figs. 5 A, 5 B). The frons and vertex are brown, interspersed with grey scales; the labial palpi are brown, with more than 1 / 3 extending beyond the frons; the thoracic dorsum is brown, with slightly lighter coloration at the base of both sides; the abdominal dorsum is dark brown, while the abdominal venter is dry greyish-brown. The forewings are dark grey to greyish-white, with a greyish-white ground color; the wing base and median band are dark grey; the subbasal line and inner line are indistinct; the median line jagged; the discal spot is a short black streak; the outer line is strongly convex and serrated, indented inward at the M 3 vein; an irregular serrated median band is formed between the median line and inner line; the area from the apex to the outer margin is dark grey; the cilia are blackish-grey. The hindwings are dirty white to dirty yellow; the discal spot is dark grey; the outer line is a thin, weak dark grey wavy stripe; the hind margin and outer margin are dirty yellow, scattered with some dark grey scales. Male genitalia (Figs. 5 C, 5 D): The uncus is sharp tipped and spiniform; the tegumen is short, small, and arc-shaped (rounded-arch); the juxta is a sclerotized lamella; the juxta lateral processes are rounded pestle-shaped, with numerous long, slender hair tufts on their surface; the saccus is short and broad, with a flat base; the valvae are broad; the sacculus (ventro-basal part of valva) is sclerotized, and the terminal process of sacculus is short, small, and triangular. The valva apex is arc-shaped; the basal half of the valva dorsum (the costa) has a strongly convex, sharp process at middle, which is weakly sclerotized. The aedeagus is long and slender; the caudal part of the aedeagus has a ring of cornuti; the aedeagal vesica membrane is smooth.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
C80DC93ED36D68263992F98EFC1CD9C6.taxon	distribution	Distribution: H. hoenei is recorded for the first time in Xizang.	en	Yang, Shuang, Yu, Xia, Viidalepp, Jaan, Pan, Zhaohui (2026): Two new species of the genus Heterothera Inoue, 1943 (Lepidoptera, Geometridae, Larentiinae) from Xizang, China. Zootaxa 5763 (4): 492-500, DOI: 10.11646/zootaxa.5763.4.2, URL: https://doi.org/10.11646/zootaxa.5763.4.2
