taxonID	type	description	language	source
C94B87AFFFF79256FF5DF8BEB237A484.taxon	materials_examined	UZBEKISTAN • 2 ♀♀, 2 ♂♂; Pamir-Alai Mountains, Surkhandarya area, Kuhitang-Tau mountain ridge, Bagly-Dara valley; 25 – 27 May 1985; Tanasevitch leg. and det.; SMF 34846.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFEFDB335A286.taxon	materials_examined	ISRAEL • 1 ♀; Wadi Grar; 20 Feb. 2007; E. Gavish-Regev leg. and det.; priv. coll. E. Gavish-Regev 4266 - 4267 • 1 ♂; Tal Or.; 19 Feb. 2007; E. Gavish-Regev leg. and det.; priv. coll. E. Gavish-Regev 3934 - 3936 • 1 ♀; Sede Teiman; 11 – 18 Feb. 2007; T. Plüss leg. and det.; priv. coll. E. Gavish-Regev (3232) • 2 ♂♂; Eshkol; 12 – 19 Feb. 2007; T. Plüss leg. and det.; priv. coll. E. Gavish-Regev (3695 to 3702).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFE21B37FA139.taxon	materials_examined	ALGERIA • 1 ♀, 1 ♂; Wil. El Tarf, Lac Oubeira; 1 May 1990; R. Bosmans leg., det. and coll.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFDCDB70FA192.taxon	materials_examined	AUSTRIA – 1 ♀; Styria, Seckauer Alpen, Ringkogel; 11 – 21 Jun. 1993; H. Brunner leg.; C. Kropf det.; NMBE Ar 3652 • 1 ♂; Salzburg, Gastein, Weissenbachtal; 29 May – 19 Jun. 1993, V. Relys leg. and det.; NMBE Ar 3203.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFD14B2A1A068.taxon	materials_examined	AZERBAIDJAN • 1 ♂ (paratype); Shemaka district, Pirkuli State Reserve; 19 Nov. 1984; D. Logunov leg.; Tanasevitch det.; SMF 33781. Females were coded according to Tanasevitch (1987).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFC92B793A0EF.taxon	materials_examined	SWITZERLAND • 2 ♀♀, 2 ♂♂; Bern, Bremgarten, Riederenwaldmoos; 2 – 6 Jun. 1980; A. Hänggi leg. and det.; NMBE Ar 1428.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFC18B50BA765.taxon	materials_examined	AZERBAIDJAN • 2 ♀♀, 1 ♂ (paratypes); Caucasus, Shemakha District, Pirkuli State Reserve; 3 Jun. 1984; D. Logunov leg.; Tanasevitch det.; SMF 33822.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFB81B70FA7DE.taxon	materials_examined	AUSTRIA – 1 ♀; Styria, Bad Gleichenberg, Rudorfkogel; 8 Jul. 1991; C. Kropf leg. and det.; NMBE Ar 3836 • 2 ♀♀, 3 ♂♂; Salzburg, Gastein, Kötschachtal; 19 Apr. – 10 May 1994; V. Relys leg. and det.; NMBE Ar 3186.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFAE8B301A672.taxon	materials_examined	FRANCE • 1 ♀, 1 ♂; Col du Galibier; 23 Sep. 1983; H. G. Müller leg.; J. Wunderlich det.; SMF 33257.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DFA75B7A7A6F7.taxon	materials_examined	TANZANIA • 1 ♀, 1 ♂; Uzungwe mountains, Kigogo forest, Mufindi; 10 Oct. 1984; N. Scharff leg. and det.; ZMUC 00006708.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DF9F3B301A56B.taxon	materials_examined	IRAN • 1 ♀, 1 ♂; Lorestan, Dizgaran; 25 Jun. 1974; A. Senglet leg.; A. Tanasevitch det.; MHNG 7444.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DF99CB50DA5E1.taxon	materials_examined	RUSSIA • 1 ♂ (paratype); Krasnodar province, Caucasian State Reserve, mount Chugush; 22 – 28 Jun. 1975; V. Ovcharenko leg.; A. Tanasevitch det.; SMF 33788.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DF905B285A478.taxon	materials_examined	UZBEKISTAN • 1 ♀, 1 ♂; W-Tien-Shang mountains, Tashkent area, Ugamsky mountain ridge close to Sidjak, Kainar-Sai valley; 22 – 25 Apr. 1983; Zonstein leg.; A. Tanasevitch det.; SMF 34850.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF69257FF5DF883B5E0A4FF.taxon	materials_examined	KYRGYZSTAN • 1 ♀, 1 ♂; Kirghizia, N-Tien-Shang mountains, Issyk-Kul area, close to Dolinka; 31 Aug. 1979; Zonstein leg.; A. Tanasevitch det.; SMF 34851.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFEFDB4A8A242.taxon	materials_examined	AUSTRIA • 1 ♂; Obergurgl; 3 Sep. 1965; K. Thaler leg. and det.; NMBE Ar 51359 • 1 ♀; same data as for preceding; 17 Sep. 1968; NMBE Ar 51358.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFE65B7FDA2DA.taxon	materials_examined	RUSSIA • 2 ♀♀, 2 ♂♂; Evenkia, Taimura river mouth of Chambe river; 1982; Eskov leg.; Y. Marusik det.; SMF 39587.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFDEDB47FA06A.taxon	materials_examined	AUSTRIA • 2 ♂♂; Styria, Graz; 10 Jul. 1995; W. Paill and O. Winder leg.; C. Kropf det.; NMBE Ar 6602. DENMARK • 1 ♀; Jaegersborg Dyrehave; 9 Apr. 2003; J. Pedersen leg.; O. Gudik-Sørensen det.; ZMUC 9996. GERMANY • 2 ♀♀; Thuringia, Südharz, Ilfeld, Brandesbachtal; 12 – 19 May 1996; Taeger leg.; B. von Broen det.; NMBE Ar 1189 • 1 ♂; Berlin, Berlin-Friedrichsfelde, Tierpark; 28 Apr. 1985; B. von Broen leg. and det.; NMBE Ar 637.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFC9CB77AA759.taxon	materials_examined	AUSTRIA • 1 ♀; Styria, Graz; 10 Jul. 1995; W. Paill and O. Winder leg.; C. Kropf det.; NMBE Ar 6610. DENMARK • 1 ♂; Rise Skov; 22 Apr. 2003; H. Liljehult and J. Pedersen leg.; J. - B. Schmidt det.; ZMUC 11430 • 1 ♀; Hestehaven, Rønde; 1 Oct. 1994; P. de P. Bjørn et al. leg.; P. de P. Bjørn det.; ZMUC 11213. GERMANY • 1 ♂; Lower Saxony, Fischbeker Heide; 21 Apr. 1993; A. Lisken-Kleinmans leg.; NMB 1507 e.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFB62B301A7FD.taxon	materials_examined	AUSTRIA • 1 ♀, 1 ♂; Salzburg, Gastein; 28 Jul. – 17 Aug. 1993; V. Relys leg. and det.; NMBE Ar 3234.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFB09B687A675.taxon	materials_examined	ITALIA • 1 ♂, 1 ♀; Abruzzo, L’Aquila, La Camosciara; 24 Sep. 2000; G. Osella leg.; M. Isaia det. and coll.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DFA71B468A6CD.taxon	materials_examined	GEORGIA • 2 ♀♀, 2 ♂♂; Caucasus, Abkhazia, Myussera state reserve; 8 – 10 Jun. 1983; Golovatch leg.; Tanasevitch det.; SMF 33790.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DF9FEB34DA561.taxon	materials_examined	LITHUANIA • 1 ♀, 2 ♂♂; Vilnius; 18 Apr. – 13 May 1998; V. Relys leg. and det.; NMBE Ar 2281.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DF985B23FA41E.taxon	materials_examined	LITHUANIA • 1 ♀; Vilnius, Botanical garden; 18 Apr. – 13 May 1998; V. Relys leg. and det.; NMBE Ar 2281. Subspecies included in the analysis to account for morphological differences to D. connatus. We agree that it is a synonym of D. connatus as proposed by Sherwood (2024). SWITZERLAND • 1 ♂; location unknown; Tr. 180; De Lessert leg.; MHNG.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF19250FF5DF8A8B5E2A495.taxon	materials_examined	AUSTRIA • 1 ♀; Styria, Laufnitzdorf; May 1995; W. Paill and O. Winder leg.; C. Kropf det.; NMBE Ar 67172 • 2 ♂♂; same data as for preceding; NMBE Ar 6668.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFEFDB334A2A0.taxon	materials_examined	GERMANY • 2 ♀♀, 1 ♂; Berlin, Köpenick; 1 Oct. 1983; B. von Broen leg. and det.; NMBE Ar 1187. SWITZERLAND • 1 ♂; Bern, Köniz, Wabern; 5 May 1996; C. Kropf leg. and det.; NMBE Ar 138.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFE5AB77AA138.taxon	materials_examined	SWITZERLAND • 1 ♀, 2 ♂♂; Vaud, Domaine de Changins; 15 – 22 May 1996; G. Blandenier leg.; NMB 2598 a.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFDC0B322A1BD.taxon	materials_examined	AUSTRIA • 1 ♀; Styria, Oppenberg; 12 Jul. 1995; C. Kropf leg. and det.; NMBE Ar 1264 • 1 ♀, 2 ♂♂; Salzburg, Gastein, Hinteres Weissenbachtal, 3 – 16 Oct. 1993; V. Relys leg. and det.; NMBE Ar 3235.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFD4FB46EA057.taxon	materials_examined	AUSTRIA – 1 ♂; Styria, Laufnitzdorf; Jun. 1995; W. Paill and O. Winder leg.; C. Kropf det.; NMBE Ar 6699 • 2 ♂♂; Styria, Unterzeiring; 13 Oct. 1994; C. Kropf leg. and det.; NMBE Ar 3662 • 1 ♀; Styria, Oppenberg; 10 May 1995; C. Kropf leg. and det.; NMBE Ar 48 • 1 ♂; Styria, Oppenberg. Gulling bank, SW Rottenmann; 12 Jul. 1995; C. Kropf leg. and det.; NMBE Ar 1262 • 1 ♀; same data as for preceding; 17 Jul. 1995; NMBE Ar 1261.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFC50B2E3A0CA.taxon	materials_examined	GERMANY • 2 ♀♀, 1 ♂; Westfalen, Bismarck-cave; Wunderlich det.; SMF 37577.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFBF2B55CA74E.taxon	materials_examined	RUSSIA • 2 ♀♀, 1 ♂ (paratypes); Siberia, Magadan area, Sibit-Tyellakh, Bolshoy Annachag mountain; Aug. 1985; Y. Marusik leg.; Eskov det.; SMF 36878.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFB79B7ADA7C3.taxon	materials_examined	RUSSIA • 1 ♀, 1 ♂ (paratypes); Siberia, Sibit-Tyellakh, Kolyma river; 10 Sep. 1987; Y. Marusik leg.; Eskov det.; SMF 36881.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFAE4B555A647.taxon	materials_examined	RUSSIA • 1 ♀, 1 ♂ (paratypes); Siberia, Magadan area, Sibit-Tyellakh, Bolshoy Annachag mountain; 6 Sep. 1985; Y. Marusik leg.; Eskov det.; SMF 36879.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DFA60B463A51D.taxon	materials_examined	ITALY • 1 ♂; Piemont, Cuneo, Gindo; 10 Oct. 2004; A. Maisi leg.; M. Isaia det. and coll. SWITZERLAND • 1 ♀; Ticino, Genestrerio close to Mendrisio; 6 – 11 Oct. 1994; N. Patocchi leg.; A. Hänggi det.; NMB 2577 b.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DF9AFB2FFA5BF.taxon	materials_examined	GERMANY • 2 ♀♀, 1 ♂; close to Wuppertal; 1984; Wunderlich det.; SMF 33430.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DF948B33BA456.taxon	materials_examined	AUSTRIA • 1 ♂; Styria, Rothleiten; 3 May – 4 Jun. 1995; C. Kropf leg. and det.; NMBE Ar 95 • 2 ♀♀; Styria Laufnitzdorf; Jun. 1995; W. Paill and O. Winder leg.; C. Kropf det.; NMBE Ar 6694 • 1 ♂; Styria Graz; 3 Jun. 1995; W. Paill and O. Winder leg.; C. Kropf det.; NMBE Ar 6645. SWITZERLAND • 1 ♂; Bern, Wohlen; 4 – 7 Jul. 1925; Bartels leg.; A. Hänggi det.; NMBE Ar 2629.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF09251FF5DF851B300A4C9.taxon	materials_examined	ITALY • 1 ♂; Piemont, Cuneo, Alpi Maritimi, Valdieri; M. Isaia and Berelutta leg.; M. Isaia det. and coll.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFEFDB7E7A2A2.taxon	materials_examined	AUSTRIA • 1 ♂ (holotype); Tyrol, Ötztaler Alps; Aug. 1930; Schenkel leg. and det.; NMB 1509 a • 1 ♀ (paratype); Tyrol, south side of the Amthorspitze at the Brenner; Aug. 1949; Schmötzer leg.; Schenkel det.; NMB 1509 a.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFE5AB348A2C4.taxon	materials_examined	RUSSIA • 2 ♀♀, 2 ♂♂; river Norilsk Kharealath; 29 Aug. 1983; Eskov leg. and det.; SMF 36882.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFDE7B5AAA159.taxon	materials_examined	TURKEY • 1 ♀, 1 ♂ (paratypes); Burdur, Insuyu magarasi, grotta; 13 Aug. 1967; P. Brignoli and V. Sbordoni leg.; P. Brignoli det.; MHNG, MHNG-ARTO- 0027059.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFD6DB7AAA1D1.taxon	materials_examined	RUSSIA • 1 ♀, 1 ♂ (paratypes); Siberia, Magadan area, Sibit-Tyellakh; 23 Aug. 1984; Y. Marusik leg.; Eskov det.; SMF 36887.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFCD5B70EA0CF.taxon	materials_examined	DENMARK • 1 ♂, 1 ♀; Zealand, Enemaerket, v. Naesbyholm; 19 Feb. 1997; J. Pedersen leg.; N. Scharff det.; ZMUC, NHMD 1785383. GERMANY • 1 ♀, 2 ♂♂; Berlin, Berlin-Friedrichshain; 20 Mar. 1977; B. von Broen leg. and det.; NMBE Ar 1164.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFBFBB7BCA7D0.taxon	materials_examined	GERMANY – 2 ♂♂; Brandenburg, Uckermark, Lychen; 18 May 1998; B. von Broen leg. and det.; NMBE Ar 2881 • 1 ♀; Dresden, Moritzburger Teiche; May 1979; S. Heimer leg. and det.; MHNG, MHNG-ARTO- 0030876. SWITZERLAND • 1 ♀; Aargau, Siggenthal; 1974; R. Maurer leg. and det.; NMB 792 f (misidentified as Glyphesis servulus).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFAEAB46EA68D.taxon	materials_examined	SWITZERLAND • 3 ♀♀, 2 ♂♂; Grisons, Alp Flix, Salategnas. 28 Oct. 2003 – 24 May 2004; H. Frick leg. and det.; NMBE Ar 5604 • 1 ♀; Grisons, Alp Flix, Salategnas; 24 Jul. 2005 – 21 Aug. 2005; P. Muff leg. and det.; NMBE Ar 7656.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DFA3EB470A505.taxon	materials_examined	GERMANY • 1 ♀, 1 ♂; Wesel; 18 Mar. 1956; Casemir leg.; H. Wiehle det.; SMF 18756. Male prosoma according to Wiehle (1960).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DF9A1B581A59E.taxon	materials_examined	UKRAINE • 2 ♀♀, 1 ♂ (paratypes); Cherkassy area, Kanev state reserve; 25 – 29 May 1988; Golovatch and Penev leg.; Esjunin and Efimik det.; Carpinus forest; SMF 38882.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DF928B54DA416.taxon	materials_examined	SWITZERLAND • 2 ♀♀, 2 ♂♂; Basel; E. Schenkel leg. and det.; NMB 792 a • 2 ♂♂, 1 ♀; Aargau, Siggenthal; 1974: R. Maurer leg. and det.; NMB 792 f.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF39252FF5DF890B2E7A4EF.taxon	materials_examined	AUSTRIA • 1 ♂ (paratype); Gamig / Ötscher; Oct.; Ressel leg.; Wunderlich det.; SMF 23978. Females scored according to Wunderlich (1969), Loksa (1981) and Blick & Szinetar (1996).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DF88AB46EA4C8.taxon	materials_examined	SWITZERLAND • 1 ♀, 1 ♂; Grisons, Sur, Alp Flix, Salategnas; 28 Oct. 2003 – 24 May 2004; H. Frick leg. and det.; NMBE Ar 4630.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFEFDB2E2A26E.taxon	materials_examined	AUSTRIA • 1 ♀, 1 ♂; Tyrol; 1937; Reimoser det.; ex coll. L. Koch; SMF RII 7033.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFE98B5CDA2E6.taxon	materials_examined	RUSSIA • 2 ♀♀, 2 ♂♂ (paratypes); Siberia, Taimyr autonomous Okrug (Area), Putorana Plateau, Lake Ayan; 16 Aug. 1983; K. Eskov leg. and det.; coll. A. Tanasevitch.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFE03B301A153.taxon	materials_examined	AUSTRIA • 1 ♀; Gastein, Kötschachtal; 13 Jun. – 5 Jul. 1994; V. Relys leg. and det.; NMBE Ar 3290 • 1 ♂; Steiermark, Grimming; 22. Sept. 1985; C. Kropf leg.; A. Rohner det.; NMBE Ar 2188 • 1 ♂; Gastein, Kötschachtal, Salzburg, Prossau; 25 May – 17 Jun. 1993; V. Relys leg. and det.; NMBE Ar 3289.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFD57B35DA02B.taxon	materials_examined	NEPAL • 1 ♂; southern Annapurna massive; 10 – 14 Dec. 1969; Martens leg.; Wunderlich det.; SMF 31715 • 1 ♀; Thakkola, Chadziou Khola; Oct. 1969; Martens leg.; Wunderlich det.; SMF 31714.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFCDFB778A0A4.taxon	materials_examined	NEPAL • 1 ♀, 1 ♂ (paratypes); Khumbu, Lughla; 22 – 23 Oct. 1970; Martens leg.; Wunderlich det.; SMF 31712.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFC46B79CA73C.taxon	materials_examined	POLAND • 2 ♀♀, 1 ♂; Gorce region, valley of Jaszcze; 2 Sep. 1986; Jedryczkowski and Starega leg.; Starega det.; SMF 36804.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFBC9B5A9A752.taxon	materials_examined	USA • 5 ♀♀, 5 ♂♂; Alaska, Alaska, Kodjak Island; NMBE Ar 6744.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFB54B2C3A7F7.taxon	materials_examined	GERMANY • 1 ♀, 2 ♂♂; Mecklenburg, Göldenitzer moor; Rabeler leg.; NMB 1611 c.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFAF2B55AA640.taxon	materials_examined	SPAIN • 1 ♂ (holotype); Málaga, 7 Oct. 1972; Harms leg.; J. Wunderlich det.; SMF 29187 • 2 ♀♀ (paratypes); same data as for preceding; SMF 29188.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DFA7AB778A6D9.taxon	materials_examined	RUSSIA • 1 ♀, 1 ♂; Komi Aut. Republic, Vorgashor; Jul. 1984; Tanasevitch leg.; Eskov det.; SMF 36885.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DF9EDB358A551.taxon	materials_examined	RUSSIA • 1 ♂ (paratype); Sakhalin, Kunashir Island, Otrandnoye; 19 Sep. 1987; A. M. Basarukin leg.; coll. Y. M. Marusik • 1 ♀; Southeast port, Mereya river; 28 Jul. 2001; Y. M. Marusik leg. and coll.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFF29253FF5DF955B778A429.taxon	materials_examined	RUSSIA • 2 ♀♀, 2 ♂♂; NE Sibiria, 25 km E of Magadan; 18 Sep. 1990; Marusik leg. and det.; SMF 39671.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFDC8B477A1B6.taxon	materials_examined	GERMANY • 3 ♀♀, 2 ♂♂; Thuringia, Südharz, Brandesbachtal; 12 – 27 May 1996; Taeger leg.; B. von Broen det.; NMBE Ar 1192.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFD30B2D3A02B.taxon	materials_examined	SWITZERLAND • 4 ♀♀, 4 ♂♂; Bern, Bremgartenwald; H. Frick. leg., det. and coll.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFCDFB211A0A4.taxon	materials_examined	AUSTRIA • 1 ♂; Styria, Gleichenberg; 1 Jun. 1991; C. Kropf leg. and det.; NMBE Ar 3771 • 2 ♀♀; Lower Austria, Eckhardsau; 7 Jun. 1985; C. Kropf leg. and det.; NMBE Ar 3990.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFC47B419A73A.taxon	materials_examined	AUSTRIA • 3 ♂♂; Styria, Rothleiten; 4 Jun. 1995; C. Kropf leg. and det.; NMBE Ar 172 • 3 ♀♀; same data as for preceding; NMBE Ar 173.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFBCCB2C4A79F.taxon	materials_examined	AUSTRIA • 1 ♀; Styria, Oppenberg; 12 Jul. 1995; NMBE Ar 1265. GERMANY • 1 ♂; Mecklenburg, Poppendorf / Rostock; 16 Jun. 1984; NMBE Ar 1097.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFB2BB43CA616.taxon	materials_examined	GERMANY • 2 ♀♀, 3 ♂♂; Brandenburg, Criewen, Uckermark; 5 – 25 May 1995; M. Sommer leg.; B. von Broen det.; NMBE Ar 553.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFEFDB79BA242.taxon	materials_examined	AUSTRIA • 1 ♀; Styria, Northeast Frohnleiten; 4 Aug. 1995; NMBE Ar 271 • 1 ♂; same data as for preceding; NMBE Ar 272.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFA96B5BCA6EB.taxon	materials_examined	SWITZERLAND • 1 ♂; Grisons, Sur, Alp Flix, Salategnas; 16 May – 14 Jun. 2003; H. Frick leg. and det.; NMBE Ar 4869 • 1 ♀; same data as for preceding; NMBE Ar 4860.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFE64B453A2DA.taxon	materials_examined	AUSTRIA • 1 ♀; Styria, Graz; 10 Jul. 1995; Paill and Winder leg.; NMBE Ar 6654 • 1 ♂; same data as for preceding; NMBE Ar 6661.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFFD925CFF5DFA1CB2D1A561.taxon	materials_examined	GERMANY • 1 ♂; Berlin, Friedrichsfelde; 8 Dec. 1993; B. von Broen leg. and det.; NMBE Ar 620 • 1 ♀; Brandenburg, Eberswalde; 13 May 1996; B. von Broen leg. and det.; NMBE Ar 1130.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	description	Figs 1 B, 2 A, 4 B, 5 B, 7 A, 9 B, 10 C – D, 12 C – D, 14 B, 15 E, 16 A – F, 18 C – E, 19 E – F, 20 B, D, F, 21 A, C; Table 2	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	type_taxon	Type species Araeoncus humilis (Blackwall, 1841).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	diagnosis	Diagnosis In males, the presence of a spoon-like retrolateral distal lobe on the palpal tibia (character 38, Fig. 1 B) in several species within node 63 (A. anguineus, A. caucasicus, A. crassiceps, A. humilis) and an autapomorphic cone-like lobe lobe in A. galeriformis (character 42; Fig. 20 D) as well as the long, robust and widely curved embolus (character 109) are two reliable characters to identify species of Araeoncus. Additionally, the massive kinked ventral radical process (character 135; Fig. 16 D) with a feather-like shape and the cephalic region raised forwardly in many cases covered with setae in the front allow to separate Araeoncus from similar genera. Females can be distinguished from similar taxa such as Diplocephalus by the more elongated spermathecae with longer copulatory ducts. In Diplocephalus spermathecae have a more rounded shape and shorter copulatory ducts. Synapomorphies Four former species of Diplocephalus, i. e., Araeoncus dentatus (Tullgren, 1995) comb. nov., Araeoncus helleri (L. Koch, 1869) comb. nov., Araeoncus marijae (Bosmans, 2010) comb. nov. and Araeoncus protuberans (O. Pickard-Cambridge, 1875) comb. nov. show simple conformation of the male palp that characterise Araeoncus (with the long and curved embolus). They belong to Araeoncus (node 59; Bremer support of 1 (MP analysis; Fig. 26) and bootstrap of 73 (ML analysis; Fig. 27 )) which is hereby defined based on the following putative synapomorphies (Fig. 31): the presence of a retrolateral distal lobe on the palpal tibia (character 38, state 1; Fig. 1 B), which is broader than long in A. dentatus, A. helleri and A. protuberans (node 60; character 40, state 1; Fig. 19 F) and longer than broad in the remaining species (Araeoncus, node 63; character 40, state 0; Fig. 1 B). The presence of a long, robust, and widely curved embolus (character 109, state 1; Fig. 16 C, A. dentatus) and the restricted embolus velum (character 122, state 1; Fig. 2 A), although this is an autapomorphy of A. humilis. Additionally, some homoplastic characters support Araeoncus (Fig. 31): among those are the lack of a protegulum (character 72) which is replaced by a tegular sac (character 75; Fig. 5 B, A. humilis in clade 63; Fig. 31 C) and the ventral radical process that is much larger than the radix (character 133; Fig. 16 D, A. crassiceps), also present in Hilaira excisa (see discussion of character 133 for more details).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	discussion	Remarks Simon (1884) based his description of Araeoncus mainly on characters concerning the eye pattern and the legs. However, he also pointed out that all considered species have a specific forward-facing cephalic lobe of variable length, lacking lateral incisions. Additionally, the tibia is split into two apophyses of which the median one (the retrolateral distal lobe, character 38) is enlarged. Most authors after Simon have added taxa to this genus based on the specific forward-facing cephalic lobe, covered with setae frontally and the two tibial apophyses (the prolateral tibial apophysis and the retrolateral distal lobe). This is the only genus out of the three most species rich genera within the Savignia - group that was recovered as monophyletic in all analyses independent of taxon sampling or characters used (e. g., Figs 25, 27, Supp. file 2, Supp. file 3). Three species formerly belonging to Diplocephalus are recovered as a clade sister to Araeoncus (Figs 25, 27). Since these species share many characters with Araeoncus, we hereby transfer them to Araeoncus and redefine the genus (see Table 2).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	description	Araeoncus with uncertain species group assignments Based on the available figures it is not possible to place A. vorkutensis with confidence, but it seems close to A. anguineus or A. victorianyanzae. The form of the tibial appendices (Tanasevitch 1984: fig. 5) and the female genitalia argue for the victorianyanzae- group (Tanasevitch 1984: figs 7 – 8). Tullgren (1955) described four species, from Sweden and / or Estonia according to females only (A. convexus, A. curvatus Tullgren, 1955, A. dispar Tullgren, 1955 and A. tuberculatus Tullgren, 1955). Kronestedt (2001) mentions in the list of Swedish spiders that Tullgren (1955) described a couple of species that have later been considered junior synonyms and does not list these four species of Araeoncus as present in Sweden. Checking this material is out of the scope of this work. Out of these four species A. curvatus and A. dispar most probably belong to the anguineus - group (node 67) based on the globular posterior part and the curved anterior part of the spermathecae. Araeoncus convexus is closely related to A. humilis and A. tuberculatus is uncertain concerning its assignment (see next section). Tanasevitch (2011) described two species from the Mediterranean, A. cypriacus Tanasevitch, 2011 and A. rhodes Tanasevitch, 2011, characterised by the shape of two anterior radical processes and the embolic division. Araeoncus rhodes males present a peculiar shorter and thicker embolus and in the females, oblong spermathecae are well visible. Based on the cephalic shape they seem close to A. crassiceps. Misplaced species of Araeoncus Araeoncus tuberculatus could not be assigned to any of the newly defined groups and may not be an Araeoncus at all due to the very simple globular spermathecae that are not found in other species of Araeoncus apart from A. vorkutensis (Tanasevitch 1984: figs 7 – 8). Araeoncus sicanus Brignoli, 1979 is close to Araeoncus humilis and to Erigonella ignobilis concerning the vulva (Brignoli 1979). Brignoli (1979) assigned it to Araeoncus based on the determination keys in Locket & Millidge (1953) and Wiehle (1960). Since this simple type of globular spermathecae are found also outside Araeoncus, it might belong to another genus within the Savignia- group as well. Araeoncus macrophthalmus Miller, 1970 does not belong to Araeoncus due to the lack of a retrolateral distal tibial lobe and the lack of a massive, kinked ventral radical process (Miller 1970: figs 1 – 6). Araeoncus gertschi Caporiacco, 1949 is known from females only and is misplaced in this genus (Holm 1962; Scharff 1990) since its epigyne has no median fissure (Holm 1962).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF959236FF0AF96EB40BA50F.taxon	description	A well-supported group of mainly African taxa with a massively enlarged, kinked ventral radical process (character 135, state 3) that exceeds the tip of the cymbium (e. g., Holm 1962: fig. 25 a – b; Tanasevitch 1987: figs 63 – 64). Their prolateral tibial apophysis and retrolateral distal lobe (characters 7, 38) are indistinct and equally sized and arise from a longitudinally elongated male tibia (e. g., Tanasevitch 1987: figs 61, 63). They all have relatively simple cephalic lobes bearing all eyes on a forward to upwards facing lobe (e. g., Tanasevitch 1987: fig. 59) except for A. subniger Holm, 1962 with an AME and a PME lobe (Jocqué 1985: fig. 1). The ventral plates of the females have two nearly parallel inner margins that form a right angle at the posterior edge. Their spermathecae are round to oval and are very close to each other right below the median margin of the ventral plates. The copulatory ducts enter the spermathecae at the lateral (ectal) side and are often also directed towards the lateral side of the vulva (e. g., Holm 1962: fig. 27 a). This group includes two species from Asia (A. caucasicus and A. banias Tanasevitch, 2013) and eight from Africa (A. etinde Bosmans & Jocqué, 1983, A. obtusus Bosmans & Jocqué, 1983, A. subniger and A. victorianyanzae) of which four are only known from females (A. impolitus Holm, 1962, A. malawiensis Jocqué, 1981, A. picturatus Holm, 1962 and A. viphyensis Jocqué, 1981). Araeoncus femineus (Roewer, 1942) from Equatorial Guinea is also known by females only and resembles females of A. victorianyanzae with respect to their body and eye size according to Bosmans & Jocqué (1983). The close relation of the African species was already noted by Holm (1962), who presumed a recent common ancestor for A. victorianyanzae, A. picturatus, A. impolitus and A. subniger due to their similarity. Holm (1962) also mentioned the close relation of A. victorianyanzae (as A. praeceps Holm, 1962) to A. crassiceps and A. anguineus based on the palpal conformation and the cephalic lobe, respectively. In our analysis the representatives of this group (A. caucasicus and A. victorianyanzae) are sister taxa of node 67, which also includes A. crassiceps and A. anguineus. Potentially A. vorkutensis Tanasevitch, 1984 belongs to this clade but see discussion of species of Areaoncus with uncertain assignment.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF979237FF0AF9BBB7CAA1CB.taxon	description	So far this clade includes four species formerly assigned to Diplocephalus, i. e., A. dentatus comb. nov., A. helleri comb. nov., A. marijae comb. nov. and A. protuberans comb. nov. Together they form node 60, including D. picinus which also has a simple palp and appeared in different positions on the different analyses (see Wild card taxa). This group includes taxa that presumably have ancestral character states to species of Araeoncus within node 63. They have relatively simple male genital characters, i. e., their retrolateral distal lobe on the male palpal tibia is broader than long (character 40, synapomorphy of node 60, the dentatus - group). It seems to be not much more than a broadened margin but in ventral view the spoon-like form is well visible (Fig. 19 E). The radical appendices are also relatively simple: they either lack a ventral radical process (character 130; Fig. 16 C, A. dentatus comb. nov.) or have a ventral radical process shaped like a ridged sail (character 135, state 2; Fig. 16 B, A. protuberans comb. nov.; synapomorphy of node 60). This sail is presumably an ancestral version of the massive, kinked ventral radical process (character 135, state 3) found in the Araeoncus clade corresponding to node 63, the sister-clade of the dentatus - group. Additionally, they have a PME lobe (character 204) which is absent in the Araeoncus clade of node 63. Araeoncus marijae comb. nov. is similar to A. dentatus comb. nov. based on the elongated curved embolus and the darker areas at the corners of the ventral plates of the epigyne. In addition, based on the shape of the palpal tibia with a retrolateral lobe and the elongated spermatechae it seems to belong to Araeoncus, and therein to the dentatus - group. Potentially, also A. hyalinus Song & Li, 2010 and A. longispineus Song & Li, 2010 described from China belong into this group. Besides the long, robust embolus diagnostic for Araeoncus, they show a similarity in the shape of the ventral radical process with A. protuberans comb. nov. Also, they have a PME lobe typical for this group.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF969237FF0AFCFFB498A780.taxon	description	This group includes two species, one exclusively known from the type locality in Iran, A. mitriformis (Tanasevitch 2008) and one from the Caucasian mountains, A. galeriformis. They are very distinct from members of the other groups: they lack a retrolateral distal lobe (character 38) but instead have a dorsal cone-like lobe (character 42; Fig. 20 D); they lack the specific massive, kinked ventral radical process (character 135) but instead have a laterally flattened ventral radical process with a dorsal facing sclerotised tip (character 135, state 5; Fig. 16 F) and additionally a laterally flattened sail-like dorsal radical process (Fig. 16 F); their cephalic lobe is also extraordinary with respect to the other species of Araeoncus of node 63: it is more pointed and either bears no eyes on top of it in A. mitriformis (Tanasevitch 2008: figs 1 – 2), or only the AME, in A. galeriformis (Tanasevitch 1987: fig. 123). Tanasevitch (1987) mentioned similarities in the form of the epigyne of A. galeriformis with A. altissimus Simon, 1884, A. crassiceps and A. vorkurtensis.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF969237FF0AFB3BB549A4EF.taxon	description	Currently, Araeoncus humilis, A. convexus Tullgren, 1955, A. longiusculus (O. Pickard-Cambridge, 1875) and two species from northern Africa (A. hanno and A. martinae) are assigned to this group. Bosmans (1996) already mentioned the close resemblance of three of these species (A. humilis, A. hanno and A. martinae). This group is defined based on the form of the embolus, which is narrowly curved at its tip (character 125; Fig. 9 B, but present in many other taxa) but additionally bears a velum restricted to the narrowly curved section of the embolus (character 122; Figs 2 A, 9 B, Araeoncus humilis). The retrolateral side of the tibia, right below the retrolateral distal lobe (character 38) bears several edges in A. humilis (Bosmans 1996: fig. 5) and even a distinct sclerotised appendix in A. hanno and A. martinae (e. g., Bosmans 1996: figs 21 – 22). Araeoncus humilis and A. martinae present special hooked setae found dorsally on tibia I and ventrally on tibia II (character 228). Araeoncus longiusculus is presumably similar to A. humilis (see Simon 1884: fig. 458) based on the setose cephalic lobe, the shape of the spiral embolus and the setose lobe-like tibial apophysis (see Simon 1884: fig. 459; Pantini & Sassu 2009: figs 1 – 2) that might be a retrolateral distal lobe (character 38). The females of this group have largely elongated spermathecae forming an outward-facing curve starting at the posterior side of the vulva and a kind of spiral at the anterior end (Wiehle 1960: fig. 424). These elongated spermathecae are also found in A. convexus (Tullgren 1955: fig. 25 b), a species that is most probably conspecific with A. humilis. The latter is known to have spermathecae variable in size (e. g., Deltshev 1987: figs 12 – 13) and a variable epigyne shape (e. g., Roberts 1987). However, according to Tullgren (1955) A. convexus differs in many aspects from A. humilis. A comparison of type material is wished for to synonymise A. convexus with A. humilis.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF919230FF0AFEFDB722A09D.taxon	description	This group includes nine species with an elongated tegular sac (character 76; Fig. 20 F). This character is a synapomorphy of node 67, including all three species that were scored from the anguineus - group. Additionally, they have a specific narrowly curved embolic tip (Fig. 16 A). The spermathecae are rounded at the posterior end and curved outwards at the anterior ends (one exception is A. altissimus; Bosmans 1996: fig. 81). Two subgroups can be recognised: the first one (a) includes A. altissimus, A. anguineus, A. clavatus Tanasevitch, 1987 and A. toubkal Bosmans, 1996. The males have a distinctly forward facing cephalic lobe (Wiehle 1960: fig. 440) and the posterior margin of the epigyne exceeds the epigastric furrow (Wiehle 1960: fig. 442). The second group (b) includes A. clivifrons Deltshev, 1987, A. crassiceps, A. discedens (Simon, 1881) and A. vaporariorum. These species have less distinct male cephalic lobes, usually arising diagonally and the posterior margin of the epigyne does not protrude over the epigastric furrow (Wiehle 1960: figs 435 – 436, respectively). Deltshev (1987) mentioned that the cephalothorax of A. clivifrons is close to A. discedens and the palpal tibia and embolus to A. altissimus and A. anguineus. Araeoncus tauricus Gnelitsa, 2004 is somewhat intermediate, having a distinct forward facing cephalic lobe and an epigyne with a posterior margin not protruding over the epigastric furrow (Gnelitsa 2004: figs 5 – 8, respectively). The current phylogenetic analysis supports the anguineus - group while the relationships within this group are not clear.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF909232FF5DF93AB7F2A7E4.taxon	description	The genus Dicymbium is morphologically very homogeneous and the definition of the two species groups is based on the morphology of the male palpal tibia. However, Di. libidinosum is quite exceptional in its male genital characters. Therefore, the characters that support node 30 (i. e., the nigrum - group) are present also in many members of the facetum - group. Node 30 is unambiguously supported by the ventral internal tooth on the prolateral tibial apophysis (character 21) which is also found in, e. g., Di. yaginumai Eskov & Marusik, 1994 (Eskov & Marusik 1994: fig. 1), the mesal second pointy tip on the inner suprategular apophysis (character 86) and the whip-like distal suprategular apophysis (character 90). It is also ambiguously supported by the pointed outer suprategular apophysis that is also present in D. connatus and D. connatus jacksoni (character 83), the ridged cheliceral stridulatory striae (character 218) and the presence of a trichobothrium on metatarsus four (character 263) that are lacking in all other members of the Savignia - group (including Di. libidinosum).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF909231FDB9FE41B301A580.taxon	description	Figs 1 C – D, 2 B – C, 4 C, 5 C – D, 7 B, 9 C – D, 10 E – G, 11 A, 12 E – F, 14 C – D, 17 F, 18 F, 19 M; Table 3	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF909231FDB9FE41B301A580.taxon	type_taxon	Type species Dicymbium nigrum (Blackwall, 1834).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF909231FDB9FE41B301A580.taxon	diagnosis	Diagnosis Diagnostic characters for males are: the spiral embolus in a sagittal plane, very typical in all species of Dicymbium (character 108; Fig. 5 D); the highly elongated palpal patella (character 3); the hook-like distal part of the prolateral tibial apophysis (Fig. 1 C) and the distinct lobe-like retrolateral process on the paracymbium (character 62; Fig. 4 C). The females are recognisable by the lip-like bisection margins (character 183; Fig. 10 E – G); the fertilization ducts with a spiral ending; the long oblong spermathecae separated by a slit around the middle and the typical separated conformation of the ventral and dorsal plate of the epigyne seen in aboral view (character 189; Fig. 10 F). Synapomorphies Dicymbium is well supported (node 29), with a Bremer support of 10, a jackknife support of 99 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27). It is supported by the following synapomorphies (Fig. 29): the massive, thin to leaf-like inner suprategular apophysis with a longitudinal backing (character 85, state 7) and the lip-like bisection margins of the epigyne (character 183). Due to fast optimisation (ACCTRAN), the latter state is shown already one step before, at node 27, including Saloca, for which this character is inapplicable. Additionally, this genus is supported by several homoplastic characters among which are: the male palpal patella that is more than 3.5 times as long as broad (character 3, state 2); the macrosetae on the cymbial retrobasal thin apophysis (character 55); the distinct retrolateral process on the paracymbium (character 62) that is found a second time in node 23; the distinct lobe-like ventral process on the paracymbium (character 64, state 1) that is also found in node 37; the spiraled radical tailpiece (character 161, state 1); the separation between the dorsal and the ventral plate of the epigyne (character 189, state 1) and the scaly or ridged cheliceral stridulatory striae (character 218, states 0 and 1). Characters 64 and 189 also appear one step before, at node 27, including Saloca for which these characters are inapplicable.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF909231FDB9FE41B301A580.taxon	discussion	Remarks Menge (1868) described this genus based on the elongated palpal tibia, the spiral embolus, the ventral radical tailpiece, the posteriorly separated ventral plate of the epigyne and a few other less conspicuous characters. A more detailed description of Dicymbium is given in Song et al. (2006). We included three out of the ten species in our analysis (see Table 3). Dicymbium emerged as monophyletic in all analyses.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF909231FDB9FE41B301A580.taxon	description	Species of Dicymbium with uncertain group assignments Dicymbium libidinosum cannot be placed into the above mentioned groups but expresses most of the diagnostic characters of Dicymbium. This species is rather exceptional in the presence of two distinct male palpal apophyses and the presence of a blunt instead of a pointed ventral radical process like all other species of Dicymbium. Dicymbium libidinosum is also the only species in this genus that has a dorsal trichobothrium on metatarsus four. The shape of the genitalia “ seems extremely closely related ” to Di. yaginumai according to Eskov & Marusik (1994). Together with the geographic distribution the presence of a second tibial apophysis points to a closer relation to Di. facetum. Dicymbium elongatum (Emerton, 1882) seems to have some derived characters compared to the other species of Dicymbium, concerning the elongated tip of the prolateral tibial apophysis and the expanded spiral (rather than spiral in one plane) embolus. Additionally, the cephalic region is extended forward (only slight to no cephalic elevations in the other species of the genus) and the ventral radical process is much thinner than in all other species (see Paquin & Dupérré 2003: figs 894 – 896).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF939232FF0AF9F5B71AA5C6.taxon	description	Most members of this group are distributed in Asia and have at least one extra apophysis besides the prolateral apophysis on the male palpal tibia. Usually, the prolateral apophysis is accompanied by a pointy apophysis (e. g., Song & Li 2008: fig. 7) or a more pronounced blunt apophysis in, e. g., Di. yaginumai (Eskov & Marusik 1994: fig. 2). This group contains Di. facetum (L. Koch, 1879), Di. pingqianense Irfan, Wang & Zhang, 2023, Di. sinofacetum Tanasevitch, 2006, Di. salaputinum Saito, 1986 and Di. yaginumai.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF939232FF0AFB07B7CDA6F1.taxon	description	This group includes the European species, namely Di. nigrum, Di. brevisetosum Locket, 1962 and Di. tibiale, corresponding to node 30. Since Di. nigrum and Di. brevisetosum were identical in the scoring, Di. brevisetosum was omitted from the trees. This group is established based on the simple male palpal tibia that only bears a prolateral apophysis and a ventral inside tooth.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	description	Figs 1 F, 2 E, 4 E, 5 F, 7 D, 9 F, 11 C, 12 H, 14 F, 16 G, 18 A, H, 19 A – B, I, L, 20 A, G, 21 B, F, 22 F, 23 A – C, 24 A; Table 4.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	type_taxon	Type species Diplocephalus cristatus (Blackwall, 1833).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	diagnosis	Diagnosis Diplocephalus is not easy to recognise based on the below mentioned synapomorphies apart from one very typical character in the males: the large dorsal radical process (character 142) which is considered most reliable to assign species to Diplocephalus. This is a large, elongated dorsal radical process with a curved tip (Figs 18 A, 21 B, F). Additionally, males usually have a cephalic PME lobe (e. g., Fig. 12 H); a retrolaterally directed prolateral tibial apophysis that runs parallel to the distal margin of the tibial apophysis and a tibial apophysis with a squarish (e. g., Fig. 19 B; Roberts 1987: fig. 36 d) rather than triangular shape (e. g., Fig. 19 C) as usually seen in Savignia but also in Diplocephalus permixtus that has distal spines on a retrolateral thickening of the palpal tibial (character 27; Fig. 19 L). Another character that distinguishes Diplocephalus from closely related genera is the short embolus with a blunt tip (characters 105 and 124; Figs 21 F, 24 A). No unambiguous diagnostic characters could be found for females. However, the lateral lobes of the ventral plate of their epigyne is often T-shaped and create a triangle posteriorly. Also, their spermathecae are usually rounded to curved. Synapomorphies Diplocephalus is restricted to the taxa in node 53 which is supported by all analyses independent of the weighting scheme and by a Bremer support of 4, jackknife support of 69 (MP analysis; Fig. 26) and a bootstrap of 95 (ML analysis; Fig. 27). The only unambiguous synapomorphy of this clade is the retrolateral distal thickening on the palpal tibia (character 37; Fig. 19 I) that is present in most taxa of node 53 (Fig. 31). The only exceptions are Diplocephalus cristatus (see, e. g., Roberts 1987: fig. 36 d) and D. alpinus, which have a setose outgrowth, that might be a vestigial version of this distal thickening which was considered as inapplicable in these species since a lobe or sac was coded as absent (character 36). However, Diplocephalus is also supported by a few ambiguous synapomorphies among which are the twisted prolateral tibial apophysis (character 20), that is plane in D. caucasicus, the short embolus (character 105) and the narrowly curved embolic tip (character 125).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	discussion	Remarks We included 23 out of the 52 described species (World Spider Catalog 2025) in the present analysis. Our analysis revealed that eleven of these taxa belong to other genera in the Savignia - group (see Table 9). This does not come as a surprise as Diplocephalus was used as a container for many species that could not be placed easily in any of the Savignia - group genera. We hereby try to define the genus based on the phylogenetic analysis and discuss the placements of species formerly assigned to Diplocephalus with reference to the new definition and its potential placement (see Table 4).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	description	Species and groups of Diplocephalus with uncertain assignment Many species currently placed in Diplocephalus do not belong to Diplocephalus in the strict sense following the synapomorphies and diagnostic characters described above. As revising the whole genus Diplocephalus on species level is out of the scope of this paper, we discuss potential groups, single wild card taxa of the current analysis and additional taxa with respect to their potential placement within the Savignia genus group. Out of the 52 species that were assigned to this genus (World Spider Catalog 2025) 21 are not possible to place based on the current analysis and / or literature data. Most of these taxa are discussed in this section but are provisionally kept in Diplocephalus.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	description	Species of Diplocephalus with uncertain assignment Diplocephalus permixtus is also a “ Diplocephalus ” in the strict sense following our diagnostic characters, i. e., having the specific enlarged dorsal radical process (character 142). However, it emerged between the cristatus - and the pavesii - group, sharing the narrow cephalic PME lobe with the cristatus- group. Diplocephalus turcicus, which emerged at node 51, might also be associated with the graecus - group by having a less distinct ventral radical process and curved spermathecae, located at the anterior side of the epigyne. Additionally, D. turcicus has a relatively special prolateral tibial apophysis with a tooth emerging from it at its retrolateral side (Brignoli 1972: fig. 15) and superficially similar to Savignia kawachiensis (Oi 1960: fig. 157). Bosmans et al. (2009: figs 7 – 12) illustration of D. turcicus seems to correspond to A. rhodes based on the male palp and the epigyne conformation, and the location record also from an island in Greece. Diplocephalus picinus (node 61) appears within the Araeoncus clade (node 59) (Figs 25 – 26). However, this species is problematic as it changes position through different analyses (Supp. file 2, Supp. file 3), most likely because of its singular palp morphology. As its position is unstable, we leave it provisionally in Diplocephalus. Within node 39 are two well-supported nodes corresponding to Savignia (node 45) and Hemistajus (node 43). The three remaining taxa in this clade are problematic: D. uliginosus, D. lusiscus and D. subrostratus. Diplocephalus lusiscus appears within node 39 (Fig. 25) which might be correct at least based on the embolic division that shares some similarities with other members in node 39, especially the flattened embolus. However, D. lusiscus is a cave dweller (Wiehle 1963) with morphological adaptations to that habitat that have led to some derived characters that are difficult to homologise. The same is true for the cave dwelling Savignia rostellatra Song & Li, 2009 with very derived characters of the embolic division (see Savignia section) but no obvious morphological adaptations to cave dwelling (Song & Li 2009). Diplocephalus uliginosus seems to be a simpler type of Hemistajus concerning the form of the cephalic lobe that resembles many members of Erigonella (e. g., Fig. 15 C, F) while the other species in Hemistajus have diagonally flattened cephalic fronts (Fig. 15 B). Also, the simpler prolateral tibial apophysis and the unmodified triangular dorsal radical process underline the basal position of D. uliginosus with respect to the other species of Hemistajus. Diplocephalus subrostratus probably belongs to Hemistajus, and was thought to be very similar to D. picinus (e. g., Eskov 1988) but differs in its embolic division. In our phylogeny, both species emerge well separated from each other (Fig. 25). Diplocephalus subrostratus is also very similar to D. uliguinosus but even simpler; it lacks any radical processes on the embolic division, has smaller suprategular apophyses and lacks a protruding inner margin of the epigyne (Eskov 1988: figs 42, 44). However, the curved, slightly elongated spermathecae, the diagonally flattened cephalic front and the inwards turned prolateral tibial apophysis (Eskov 1988: figs 40, 43) resemble very much to what is found in, e. g., D. montanus (Eskov 1988: figs 25, 28). According to Denis (1952), D. caecus Denis, 1952, a cave dweller from Romania, is very close to D. lusiscus. The form of the cephalic lobes, the arrangement of the eyes in the males of both species, and the male prolateral tibial apophysis are very similar. There are also two eyeless species from caves from Morocco, D. inanis Tanasevitch, 2014 and D. bosmansi Lecigne, 2025. Diplocephalus bosmansi resembles D. picinus but with a broader epigynal fissure. Diplocephalus inanis males can be distinguished by the broad prolateral tibia and a curved embolus and females by the thick seminal ducts and wide epigynal fissure in the proximal part of the ventral plate, similar to D. picinus. Diplocephalus protuberiscus Wunderlich, 2022 is known from the male and is named based on the similarities to A. protuberans comb. nov. and D. lusiscus. It has a cephalic part raised with a setose clypeus like in Hemistajus and Savignia, a palpal tibia with a retrolateral branch, and a thick and curved embolus. The associations of D. montaneus Tanasevitch, 1992, D. bifurcatus Tanasevitch, 1989, D. hispidulus Saito & Ono, 2001 and D. sphagnicola Eskov, 1988, that are likely not closely related to each other, remain obscure. Tanasevitch (1989) mentions the close resemblance of D. montaneus with D. cristatus. However, the lack of an enlarged dorsal radical process argues against a close relationship. Diplocephalus sphagnicola has a very distinct ventral radical process and a particularly long embolus (Eskov 1988: fig. 33) with a corresponding long copulatory duct in the female (Eskov 1988: fig. 37). Both characters are not found in any other group of taxa apart from maybe Araeoncus to which D. sphagnicola is presumably not closely related due to the lack of synapomorphic characters. Diplocephalus circularis Irfan, Zhang & Peng, 2025 and D. tongrenensis Irfan, Zhang & Peng, 2025 are two species recently described from China. Based on the median fissure, the wide copulatory opening of the epigyne and the long and spiral embolus in D. circularis, they probably belong to another genus of the Savignia genus group. Diplocephalus parentalis Song & Li, 2010 is closely related to D. hispidulus based on the palpal tibial apophysis shape, which is also covered by papillae (similar to several species of Erigonella) and the complicated dorsal plate of the epigyne (Song & Li 2010: figs 9 – 11). It was placed in Diplocephalus based on the long copulatory ducts and it has a particular embolic division with a large radix with a sclerotised margin with no apophyses. The lack of a radical process on the radix and the different shape of the ventral plate with a “ column-shaped complex ” anteriorly, indicates that it does not belong to Diplocephalus and probably corresponds to another genus of the Savignia genus group. Diplocephalus machadoi Bosmans & Cardoso, 2010 males occur in two forms. In one, the cephalic PME lobe is very similar to D. picinus and the other has a pointed AME lobe that resembles to the one of A. galeriformis. It shares characters with species from different genera of the Savignia genus group and cannot be assigned to any based on the available figures. Species of Diplocephalus misplaced in the Savignia genus group Diplocephalus gravidus Strand, 1906 does not belong to the Savignia genus group due to the lack of a bisected epigyne (Ono et al. 2009: fig. 238) that is synapomorphic for this node (21). Therefore, it does not belong to Diplocephalus and presumably not to any other genus in the Savignia- group. Only the female is known from this species and compared to the other taxa included in the current analysis, it has a slightly elevated cephalothorax (Ono et al. 2009: fig. 236). Diplocephalus bicurvatus Bösenberg & Strand, 1906 also lacks the bisected epigyne (Bösenberg & Strand 1906: pl. 12 fig. 272 e) and the prolateral tibial apophysis is more pronounced than in other Savignia- group members and is initially facing dorsally. The embolic division is also very different from what is found in any Savignia- group species (Ono et al. 2009: figs 230 – 234). The epigyne of D. culminicola Simon, 1884 seems to lack a median fissure (Denis 1953: figs 6 – 7) and therefore is most probably not a Savignia- group member. The current information on D. tiberinus (Caporiacco, 1936) is restricted to one figure showing the male palp in Caporiacco (1936: fig. 3) and a description in Latin, which are not sufficient for any discussion of generic assignment.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9D923CFF0AFEC2B79AA62A.taxon	description	This group corresponds to node 57. With a Bremer support of 10, jackknife support of 99 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27) this node is one of the best-supported nodes in the current analysis. Synapomorphies are the delicate sigmoid type of distal suprategular apophysis (character 90, state 2), the typical embolus that is a secondarily shortened “ elongated ” lamella with a tip fused to the radix (character 114) and the flat, simple dorsal radical process with a simple pointed tip (character 144; Fig. 31). Moreover, several homoplastic characters support this group. Among those are the ventral tibial process (character 47), the narrow cymbial retrobasal thin apophysis (character 56; state 2) and the lateral dark spot or stripe (character 214, lacking in D. alpinus). Additionally, all these species have both a cephalic PME lobe and an AME lobe. Two subgroups can be recognised. The first subgroup (a; node 58) is defined by the presence of the retrolateral blunt sclerotised structure on the male palpal tibia (character 45) and the lack of a distinct tooth emerging from the retrolateral side of the dorsal radical process (character 146). The following species belong to this subgroup: D. alpinus and D. cristatus. The second subgroup (b), represented by only one species in the current analysis, D. crassilobus, lacks the retrolateral blunt sclerotised structure on the male palpal tibia (character 45, state 0) but instead has a distinct tooth emerging from the retrolateral side of the dorsal radical process (character 146). This tooth is an autapomorphy of D. crassilobus. However, it is also present in the following species that also belong to this subgroup and that were not included in the current analyses: D. hungaricus Kulczyński, 1915 and D. pseudocrassilobus. Gnelitsa (2006) already pointed out the close relationship of these three species and mentions that Millidge (1979) even thought that D. crassilobus and D. hungaricus are synonyms. Comparison of type material is therefore needed to solve this taxonomic problem which is out of the scope of the current analysis. Diplocephalus komposchi Milasowszky, Bauder & Hepner, 2017 might belong to the cristatus- group based on the shape of the palpal tibia with a narrow ventral tibial process, the short lamella-like embolus and the distinct radical process. The epigyne resembles to the one of D. pseudocrassilobus. However, D. komposchi has a wider dorsal radical process and it lacks the lateral teeth characteristic for the species of the cristatus- group (b).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9D923EFF0AFADFB77FA774.taxon	description	This group includes a few species with very derived characters and corresponds to node 54. It is supported by a Bremer support of 5, a jackknife support of 79 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27) and the following synapomorphies (Fig. 31): the distinctly forward curved marginal suprategular apophysis (character 80), the large flat and ventrally folded dorsal radical process (character 143), the basal tooth on the radical tailpiece (character 167) and the paired inner longitudinal narrow lobes of the epigyne (character 187). Two subgroups can be differentiated; first, the Caucasian one (a) that is represented only by D. caucasicus in the phylogenetic analysis but also includes D. transcaucasicus Tanasevitch, 1990. They have a bifurcate ventral radical process with a thin sclerotised distal tip (character 135, state 6) that is distinct in both species and connected to the prolateral margin of the dorsal radical apophysis (Fig. 16 G). The tip of the dorsal radical process faces dorsally in these species. The second subgroup (b) includes node 55. It is supported by a Bremer support of 10, a jackknife support of 99 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27) and the following unambiguous characters: the double folded prolateral tibial apophysis (character 18), the cymbial retrobasal striated glabrous bump (character 58) and the massively broadened radical tailpiece tip with an inner thickening (character 165). Additionally, their dorsal radical process is highly enlarged and points ventrally and bears retrolateral striations. The males of all these species have a PME lobe (except for D. guidoi Frick & Isaia, 2012 which has a small post PME lobe instead). Based on the dorsal radical process and the double folded prolateral tibial apophysis we assigned the following species to this species group: D. arnoi, D. guidoi, D. longicarpus (Simon, 1884), D. pavesii and D. procer (Simon, 1884).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9F923EFF0AFA32B265A4AF.taxon	description	This group includes six Mediterranean species that were not scored for the current phylogenetic analysis and lack the above described large dorsal radical process: D. graecus (O. Pickard-Cambridge, 1872), D. lancearius (Simon, 1884), D. mystacinus (Simon, 1884), D. algericus Bosmans, 1996, D. altimontanus Deltshev, 1984 and D. toscanaensis Wunderlich, 2011. Four species form a group based on the simple and broad prolateral tibial apophysis (e. g., Bosmans 1996: figs 46, 54): D. graecus, D. lancearius, D. mystacinus and D. toscanaensis. The cephalic lobe of D. graecus is very similar to D. picinus and D. algericus but different from what is found in D. lancearius and D. mystacinus (Bosmans 1996: fig. 52; 2002: fig. 67). Three of the taxa with the broad prolateral tibial apophysis (D. graecus, D. lancearius and D. mystacinus) have distinct ventral radical processes, like D. altimontanus and D. algericus and presumably also D. turcicus (as far as it can be judged from illustrations; see Table 4). Additionally, all these species have a flattened and curved to spiral embolus. The females have a relatively simple vulva with spermathecae situated at the posterior to central part of the epigyne.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF98923AFD8BFEFCB417A628.taxon	description	Figs 1 E, I – J, 2 D, 3 B – C, 4 D, H, 5 E, 6 C, G, 7 C, 9 E, H – I, 10 H – I, L – M, 11 B, D – F, 12 G, 13 C – D, 14 E, H – J, 15 C, F – G, 17 E, 19 H, O, 21 H – I, 23 D – K, 24 B – L; Table 5	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF98923AFD8BFEFCB417A628.taxon	type_taxon	Type species Erigonella hiemalis (Blackwall, 1841).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF98923AFD8BFEFCB417A628.taxon	diagnosis	Diagnosis The diagnosis does not include E. ignobilis, which is discussed in section “ uncertain assignments ”. Diagnostic characters for males are the presence of a prolateral and a retrolateral tibial apophysis where the prolateral apophysis is sometimes papillate (character 14, state 2); the short coiled embolus with a blunt tip, the presence of ventral, central or dorsal radical processes, the pointed (in E. hiemalis and E. subelevata) or blunt (in E. latifrons comb. rev. and E. ignobilis) inner suprategular apophysis and the simple straight to slightly curved distal suprategular apophysis (character 90; state 4) with a blunt to slightly to sharply pointed tip (character 93). Additionally, they present a punctuated sternum (character 196), which is only found in E. hiemalis, E. subelevata (node 34) and E. ignobilis within the members of the Savignia - group. Females usually present a triangularly split on the posterior side of the epigyne (character 181) (also present in Diplocephalus) and spermathecae with an L-shape (e. g., E. hiemalis and E. subelevata) in contrast to the rounded spermathecae of similar Diplocephalus females (e. g., D. cristatus). Synapomorphies This genus corresponds to node 33 that is supported by a Bremer support of 1 (MP analysis; Fig. 26) and a bootstrap of 97 (ML analysis; Fig. 27) and includes two species-groups: the hiemalis - group (node 34) and the connatus - group (node 35 excluding E. ignobilis). Species of Erigonella shared only one unambiguous synapomorphy (Fig. 30): the cramp-like dorsal radical process (character 150) that was inapplicable in the hiemalis - group and E. ignobilis. Additional supportive homoplastic characters include the retrolateral tibial apophysis (character 28) that is lacking in E. ignobilis and only also present in Glyphesis and Entelecara; the outer suprategular apophysis (character 82); the transverse spiraled embolus (character 108; state 1) and the rough irregular cephalothorax surface (character 217). They also have in common the simple straight to slightly curved distal suprategular apophysis (character 90, state 4), but this character appears as a synapomorphy one step before, at node 32, despite it is only present in all Erigonella taxa (node 33). The retrolateral tibial apophysis is glabrous and initially distal facing only in Erigonella (characters 31, 32).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF98923AFD8BFEFCB417A628.taxon	discussion	Remarks The genus Erigonella was described by Dahl (1901) based on the punctuated sternum that differs from Lophomma punctatum (Blackwall, 1841) by additional rugose areas in between (Fig. 11 D – F) and the lack of punctuations on the cephalothorax. The generic definition and assignment of taxa did not change much since. Dahl (1901) included E. hiemalis, E. subelevata, E. ignobilis and one species that was later transferred to Diplocephalus but belongs to Erigonella according to our phylogenetic analysis and is therefore a revived combination: E. latifrons comb. rev. Since then, E. groenlandica Strand, 1905, E. stubbei Heimer, 1987 and E. subelevata pyrenaea Denis, 1965 were added. The current analysis suggests that at least three more taxa belong to this genus (E. connatus comb. nov., E. latifrons comb. rev. and E. producta comb. nov.) and the genus definition should include more characters than only the punctuated sternum. Wunderlich (1970) suggested a synonymy with Diplocephalus, which was not followed by later authors. The similarity of species from Erigonella with representatives of Diplocephalus is very high, especially concerning the shape of the cephalic lobes. However, instead of lumping all these genera into one (Savignia by priority) we prefer to keep most of the genera defined within the Savigniagroup and instead revise the genus Diplocephalus by transferring some of its species to other genera. In our analysis, we included three of the six species previously described as Erigonella. Erigonella turned out to be a relatively well-supported clade when two species from Diplocephalus and one from Savignia are included (see Table 5).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF98923AFD8BFEFCB417A628.taxon	description	Species of Erigonella with uncertain assignment Strand (1905) described E. groenlandica from one subadult female based to the punctuated sternum that is not found in any other member of the Savignia - group. However, this character is also found in other linyphiid taxa, e. g., Lophomma punctatum (Hormiga 2000: pl. 47 a – b). Meanwhile, this species has been considered a nomen dubium. The assignment of E. stubbei that is known from females only is also uncertain and considered as preliminary by Heimer (1987). He mentions the similarity of the vulva with Tapinocyba Simon, 1884 but assigned this species to Erigonella based on the habitus, the leg spination and the position of the metatarsal trichobothria. The globular spermathecae are similar to E. ignobilis and according to the figures, the epigyne is bisected. However, a punctuated sternum was not mentioned, so, until males are available clear generic assignment for this species is not possible. Erigonella ignobilis is very difficult to place within Erigonella and even may not belong to this genus. It emerged within Erigonella in all analyses but it emerged between Glyphesis and Dicymbium in Frick et al. (2010) rather than between E. latifrons comb. rev. and E. producta comb. nov. Problematic is the lack of derived characters in this species. Erigonella ignobilis lacks a cephalic lobe and has a central radical process that is otherwise only present in Glyphesis and a very simple conformation of the male palpal tibia. However, it shares the retrolateral glabrous edge on the male palpal tibia (character 35) and the punctuated sternum (character 196). Erigonella ignobilis might change its position within the Savignia- group (to which it certainly belongs) in future phylogenetic studies.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9B923AFF0AFAA0B566A56B.taxon	description	This group corresponds to node 34 plus E. subelevata pyrenaea. It is ambiguously supported by several characters among which are: the papillate prolateral tibial apophysis (character 14, state 2), a thin, long inner suprategular apophysis with a blunt tip (character 85, state 4) and the punctuated sternum (character 196). Erigonella subelevata and E. subelevata pyrenaea are very similar but differ in the size of the AME lobe and the PME lobe (Denis 1965: fig. 2; Thaler 1971: fig. 6) and the shape of the retrolateral tibial apophysis (Denis 1965: fig. 3; Thaler 1971: fig. 3).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9B923BFF0AF99FB7A7A18A.taxon	description	This group is defined based on the cramp-like dorsal radical process (character 150) that is only present in E. latifrons comb. rev., E. producta comb. nov. and E. connatus comb. nov., which together form this group. The triangularly split posterior side of the epigyne (character 181) is also typical in this group and the presence of a more or less distinct AME lobe (character 209). Erigonella latifrons comb. rev. is basal to the remaining species. Erigonella producta comb. nov., E. connatus comb. nov., and E. connatus jacksoni comb. nov. form node 37 which is supported by a Bremer support of 6 a jackknife support of 99 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27) and the following synapomorphies (Fig. 30): the sharply pointed simple to slightly curved type of distal suprategular apophysis (character 93, state 1) and the proximally directed and pointed dorsal to retrolateral radical tailpiece (character 169, state 3) and as ambiguous synapomorphy, the pointed inner suprategular apophysis that is flush with the distal suprategular apophysis (character 85, state 6). Additionally, the flat cramp-like dorsal radical process (character 151) that is only found in these three species but is inapplicable in the other members of Erigonella, appears as a synapomorphy of node 36. They also have a distinct lobe-like ventral process on the paracymbium (character 64) that is otherwise only present in Dicymbium. Holm (1977) assigned E. producta comb. nov. to Savignia due to its great similarity of the cephalic lobe even though he pointed out that secondary sexual characters such as the cephalic lobes are not reliable for genus delimitations. In the current analysis this species is the sister taxon of E. connatus comb. nov. plus E. connatus jacksoni comb. nov. Meanwhile, the latter subspecies has been synonymised with E. connatus comb. nov.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	description	Figs 3 D, 4 I, 6 D, 8 A, 9 J, 10 N, O, 13 E, 14 K, 15 A, I, 16 H, 17 C, 19 K, N, P, 22 G; Table 6	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	type_taxon	Type species Glyphesis servulus (Simon, 1882).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	diagnosis	Diagnosis The males of Glyphesis can best be recognised by the cymbial prolateral basal glabrous apophysis (character 60; Fig. 6 D) and the central radical process (character 152; Fig. 6 D) in combination with the clypeal lobe (lacking in the former Paraglyphesis) (character 208; Fig. 13 E), the usually ridged or scaled pro- (character 14; Fig. 19 N) and retrolateral tibial apophysis (character 31; Fig. 3 D) (the retrolateral tibial apophysis directs towards the inside in the cottonae - group) and the presence of a prolateral sickle on the prolateral tibial apophysis (character 13; Fig. 19 P). Some species of Glyphesis (like G. servulus and G. taoplesius) can also be recognised by the long strong inter pro- to retrolateral tibial apophysis macrosetae (character 25; Fig. 19 P). The females can be recognised by the oval-shaped epigyne (character 184; Fig. 10 N) and the anteriorly turned posterior tip on the ventral plate of the epigyne (character 190) (lacking in the basal cottonae - group). Synapomorphies We follow the suggestion of Eskov (1991 b) in his description of Paraglyphesis, where he pointed out the close similarity to Glyphesis. We agree with Eskov (1991 b) in this respect and hereby synonymise Paraglyphesis with Glyphesis. Simon (1926) described Glyphesis based on the cephalic clypeal lobe (character 208), the tibial apophyses with thickened retrolaterally curved setae and the semicircular epigyne. These characters are still valuable to describe Glyphesis. This genus corresponds to node 22 and is monophyletic in the current analysis, supported by a Bremer support of 6, a jackknife support of 98 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27). Species of Glyphesis share the following synapomorphies (Fig. 29): the ridged or scaled texture of the prolateral tibial apophysis (character 14, states 3 and 4; are also found in Entelecara), the inter pTA-rTA dorsal protuberance (character 43), the cymbial prolateral basal glabrous apophysis (character 60), the blunt, small, thin and short ventral radical process (character 135, state 8) and the retro- to prolateral bulb-like central radical process (character 153). Additionally, a few more ambiguous characters support this genus. Among these are the lack of a prolateral trichobothrium on the male palpal tibia (character 4); the general conformation of the palpal tibia showing two short fingers on a broad base (character 6; state 1); the presence of a retrolateral tibial apophysis (character 28); the small distal suprategular apophysis with a blunt or pointed tip (character 90, state 11); the cephalic clypeal lobe (character 208, reduced in former Paraglyphesis) and the single proximal dorsal trichobothrium on the female palpal tibia (character 246). The scored taxa also had a ventral radical process that was either blunt, small, thin and short (character 135, state 8) or a pointed small tooth (character 135, state 9). Additional characters but present also in other taxa are the scaled texture of the retrolateral tibial apophysis (character 31). The females of Glyphesis excluding the cottonae - group have an oval-shaped epigyne (character 184).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	discussion	Remarks We included four out of the seven known species of Glyphesis (see Table 6). Our results revealed that Paraglyphesis should be synonymized with Glyphesis, adding another three taxa (see Table 9).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	description	Species of Glyphesis with uncertain species group assignments Glyphesis nemoralis is intermediate between the servulus - and the polaris - group. It has a less developed retrolateral sickle than seen in the servulus - group but it is not lacking as in the polaris - group. It lacks an inter pro- to retrolateral tibial apophysis macrosetae found in the servulus - group but shares the elongated spermathecae with it, while the polaris - group has nearly globular spermathecae. This intermediate position is also reflected by our analyses where it emerges between these two groups in the implied weighting analyses.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849225FF0AFE39B2F7A1D3.taxon	description	This group contains the early branching species of Glyphesis. It is represented by G. cottonae in our analysis. Glyphesis cottonae and G. asiaticus Eskov, 1989, that also belongs to this species group, share a few characters that are not found in any other species group within Glyphesis. The males have a highly sclerotised dorsal facing retrolateral tibial apophysis (character 32) and the comparatively broad and highly sclerotised retrolateral facing prolateral tibial apophysis (e. g., Eskov 1989: fig. 2). The females have a bisected epigyne with flush lateral lobes of the ventral plate until the posterior end of the epigyne (e. g., Eskov 1989: fig. 3) like most other Savignia genus group members. This is another argument for the common origin of the bisected epigyne in the Savignia genus group (node 21).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849225FF0AFCD7B252A7BF.taxon	description	This group is equivalent with the former Glyphesis. It can be recognised by the long strong inter pro- to retrolateral tibial apophysis macrosetae (character 25). There are two distinct subgroups within this group: first (a), G. servulus and G. taoplesius with a pronounced retrolateral sickle (character 13; G. nemoralis with a smaller one), a well-developed cephalic clypeal lobe (character 208; e. g., Wunderlich 1969: fig. 23) and a well visible retrolaterally directed retrolateral tibial apophysis. The second subgroup (b) includes the North American taxa of this genus G. idahoanus (Chamberlin, 1949) and G. scopulifer (Emerton, 1882). They reduced both the pro- and retrolateral tibial apophysis in size compared to the European taxa of the servulus - group, lack a distinct cephalic clypeal lobe and have a more pronounced (towards the posterior end of the epigyne) posterior tip of the epigyne. The central radical process is much more pronounced distally than in the other species of this subgroup.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849225FF0AFB48B264A6C5.taxon	description	This group corresponds to the former genus Paraglyhesis and includes the very homogenous species G. lasiargoides (Eskov, 1991) comb. nov., G. monticola (Eskov, 1991) comb. nov. and G. polaris (Eskov, 1991) comb. nov. Eskov (1991 b) diagnosed Paraglyphesis by the “ unmodified male carapace with neither postocular pits nor any clypeal projections ” a character that was shown to be highly variable in other well-defined genera (e. g., see discussion of E. connatus comb. nov. and E. connatus jacksoni comb. nov.). He also noticed the well-separated posterior tips of the lateral lobes of the ventral plate that are touching each other in the servulus - group. Members of this group lack the long strong inter pro- to retrolateral tibial apophysis macrosetae and have globular spermathecae.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	description	Figs 15 B, 16 I, 17 D, 18 B, 19 G, 22 D – E; Table 7.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	type_taxon	Type species Hemistajus rostratus (Schenkel, 1934) comb. rev.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	diagnosis	Diagnosis Hemistajus males can be distinguished from similar genera by the broad prolateral tibial apophysis ending in a broad cut off and papillate tip (character 15; Fig. 19 G). The males of this group have a diagonally flattened cephalic front (character 199; Fig. 15 B) that is densely covered with setae and even setose in females. The females can be recongised by the rebordered, sclerotised median margins of the epigyne (character 185; Fig. 22 E), but also present in D. mirabilis and S. frontata. Within Hemistajus, H. montanus comb. nov., H. barbiger comb. nov. and H. marusiki comb. nov. present the most derived characters concerning the retrolateral protuberance (character 149) arising from the triangular dorsal radical process (character 147) and the elongated dagger- to dirk-like distal suprategular apophyses (character 90, state 1). Hemistajus rostratus comb. rev. is closely related to H. barbiger comb. nov. based on genital morphology (Schenkel 1934; Thaler 1970). Thaler (1970) underlines the isolated position of these species within the “ Diplocephalus - group ” (which corresponds to the Savignia - group later defined by Millidge 1977) and that the subgenus Hemistajus should be kept in mind in a future revision. We agree with Thaler (1970) in this respect and the description of a few more taxa with similar characters by Eskov (1988) seems to justify the new generic rank of Hemistajus. Synapomorphies This definition corresponds to node 43 in the preferred tree (Fig. 25). The ventral ridge on the triangular dorsal radical process (character 148) is the only unambiguous character that supports this clade. However, it is also supported by a Bremer support of 4, jackknife support of 85 (MP analysis; Fig. 26) and a bootstrap of 99 (ML analysis; Fig. 27) and several homoplastic characters among which are the following (Fig. 30): the ventral facing distal part of the prolateral tibial apophysis (character 19); the very large and flat inner suprategular apophysis formed like an equilateral triangle (character 85, state 11) and the supplementary clypeal setae in females (character 234). Additionally, the prolateral tibial apophysis is broadened and broadly cut off at its tip (e. g., Eskov 1988: figs 9, 15, 28).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	discussion	Remarks Hemistajus was described by Schenkel (1934) as a subgenus of Diplocephalus with the type species H. rostratus (Schenkel, 1934) comb. rev. Schenkel (1934) established this genus to account for the quite different morphology of Diplocephalus rostratus and D. barbiger (Roewer, 1955). Hemistajus was later elevated to genus in Janetschek (1956) and then synonymised to Diplocephalus by Thaler (1970) by the transfer of its type species. We included three out of four species of Hemistajus in the phylogenetic analysis (see Table 7). All four species were transferred here from Diplocephalus: H. barbiger (Roewer, 1955) comb. nov., H. marusiki (Eskov, 1988) comb. nov., H. montanus (Eskov, 1988) comb. nov. and revive the combination H. rostratus (Schenkel, 1934) comb. rev. (see Table 9).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	description	Figs 15 H, J, 17 G, 19 C, 20 H, 21 D, G, 22 B – C; Table 8	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	type_taxon	Type species Savignia frontata Blackwall, 1833.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	diagnosis	Diagnosis Males of Savignia are well recognisable by the specific embolic division, where the embolus forms a so called “ transparent outgrowth ” (character 116; Fig. 21 G) and also by the radical processes including an unique tooth-like basal radical process (character 159; Fig. 21 G) and the dorsal radical process having a fold (character 141; Fig. 21 D). Homoplastic but also present in all species of node 45 is the elongated tip of the embolus bearing a slim projection (character 123; Lasut et al. 2009: figs 4, 8, 12) and the massive inner suprategular apophysis (character 85; Fig. 17 G). Additionally, males often have a triangular palpal tibia with a hook-like prolateral tibial apophysis ending in a pointed tip (Fig. 19 C). Females are best recognised by the roundly broadened anterior end of the ventral plate (character 182), the straight parallel interior margins of the ventral plate, the triangular shape of the ventral plate and oval-shaped spermathecae, oblique to parallel to each other. However, this is modified in certain species like S. rostellatra (Song & Li 2009: figs 33, 35) with very derived characters. Synapomorphies Savignia (node 45; Bremer support of 2, jackknife support of 92 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27 )) is supported by the following synapomorphies (Fig. 30): the column apophysis (character 98; Fig. 21 D); the transparent outgrowth at the embolus (character 116; Fig. 21 D); the dorsal radical fold (character 141; Fig. 21 D) and the tooth-like basal radical process (character 159; Fig. 21 D). Additionally, some homoplastic characters support Savignia. Among these are the massive, highly sclerotised inner suprategular apophysis (character 85) and the twisted embolus (character 118). Node 46 corresponds mainly to what Eskov (1988) considered as Savignia excluding S. mirabilis comb. nov. It is worth mentioning this clade since it is supported by many characters that account for most distal species of Savignia and are therefore diagnostic for most species of Savignia (i. e., in the current analysis for all but S. mirabilis comb. nov. with simpler palpal conformations). Node 46 is unambiguously supported by the roundly broadened anterior end of the ventral plate (character 182). Several homoplastic characters also support this clade. Among them are the highly sclerotised, massive, blunt distal suprategular apophysis (character 90, state 10; Fig. 17 G); the distal tooth on the radical tailpiece (character 170; Fig. 20 H) and the AME lobe (character 209; Fig. 15 J). Another character that is found in all Savignia but was not scored in the current analysis is the retrolaterally curved, usually twisted prolateral tibial apophysis of the male palp (e. g., Eskov 1988: fig. 85).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	discussion	Remarks This genus was described based on male specimens of S. frontata from England according to the six eyes (overlooking the AME on the AME lobe) and other eye characters, which are not diagnostic anymore. The spelling of Savignia was a cause for debate (e. g., Bonnet 1958; Bosselaers & Hendrickx 2002; summary of spellings used since the original description in World Spider Catalog 2025). Even though Blackwall (1833) dedicated this genus name to Jules-César Savigny he spelled it as Savignia throughout the original description (two times). Bonnet (1958: 3937, note 39) referring to the rules of nomenclature (art. 8, par. H, note b) neglected the spelling of Savignia (Blackwall 1833) and Savignyia (Simon 1926) and proposed Savignya as the correct spelling. However, according to the International Code of Zoological Nomenclature (1999), Savignia is an original spelling (art. 32.1.); therefore, the correct spelling of this genus is Savignia as spelled in the original work of Blackwall (1833). Eskov (1988) redescribed Savignia according to the slightly curved and backwards directed embolus; the curved distal suprategular apophysis and the leaf-like broadened inner suprategular apophysis; the broadly rounded, terminally unciform prolateral tibial apophysis; the distinctly emarginated anterior part of the epigyne and the pronounced dorsal plate covering half of the oval spermathecae. The current analysis agrees with most of his diagnostic characters but found some more that are discussed hereafter. The genus Savignia is not monophyletic in the current analysis due to two taxa, S. producta and S. harmsi that are both proposed to be transferred to other genera. However, many more taxa that we did not include in our analysis likely do not belong to Savignia as defined by Eskov (1988). We agree with Eskov (1988) that Savignia has a few very derived characters that are not found in any other genus within the Savigniagroup. According to our analysis this is true for node 46 that includes all Far East Savignia species plus S. frontata and is well supported by a Bremer support of 6, a jackknife support of 99 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27). We hereby include one more species, S. mirabilis comb. nov., that is a simpler type of Savignia (node 45) but shows most of the derived characters also. This is why Eskov (1988) described the species as a Diplocephalus rather than a Savignia. In the current analysis we included six out of 24 known species of the genus Savignia (see Table 8). Out of all Savignia taxa, eleven occur in Russia (one also in Alaska), four in the Far East (Japan, China or Korea), four are known from Europe only and two are occurring all over the Palaearctic. Another one from Australia (S. erythrocephalus (Simon, 1908 )) can be excluded from the Savignia- group. One from the Comoro Islands (S. kartalensis Jocqué, 1985) and one from Ethiopia (S. ericola Tanasevitch, 2023) both have uncertain assignments. We included the two Palaearctic species, one that is only found in Europe and three from the most Eastern regions of Russia (around Magadan). The Russian species are underrepresented in our matrix and the three species from Japan, China and Korea are not represented. Since the Russian species are very closely related to each other it seems that we cover most of the known variety within the Russian species. Since the three species scored from Russia occur in the most Eastern regions (around Magadan), the species from Japan, China and Korea would not have added generally new morphological features (e. g., figures in Eskov 1988, 1991 a).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	description	Savignia with uncertain species group assignments The cave dwelling S. rostellatra cannot be assigned to any species group based on morphology alone due to its very derived male copulatory organs. It clearly belongs to Savignia having the unique tooth-like basal radical process (character 159; Fig. 21 G) but its inner suprategular apophysis is fused to the suprategulum (distal part), covered with papillae and bearing a tooth (Song & Li 2009: fig. 28). Its cephalic lobe, however, is a simple PME lobe that would argue for the mirabilis - group. Savignia ericola as mentioned by the author (Tanasevitch 2023 a) is conditionally placed in Savignia until a female is found, based on the different shape of the distal suprategular apophysis, lacking the broad inner suprategular apophysis and the distinct shape of the male palpal tibia bearing three long strong setae. We agree with the genus assignment, as S. ericola has the radical conformations, including the unique tooth-like basal radical process (character 159; Fig. 21 G). However, the species group assignments are more difficult. Interpretating the large sclerotised part of the suprategulum as inner suprategular apophysis rather than a distal one, would argue for the mirabilis - group. This would also be supported by the presence of an AME lobe like in S. frontata and lacking any other cephalic lobes. Misplaced species of Savignia Paik (1978) assigned S. pseudofrontata Paik, 1978 to Savignia based on its close resemblance to S. frontata. We do not agree with that: the vulva is spiral and much more complex than the globular spermathecae found in S. frontata and the radix lacks the typical processes. The radical processes and the form of the embolus (Paik 1978: figs 48, 49, 51) are somewhat similar to S. harmsi (Wunderlich 1980 a: figs 48 – 49) and D. lusiscus (Wiehle 1963: fig. 30). The cephalic lobe looks like one found in Araeoncus (e. g., Tanasevitch 1987: fig. 59) and the prolateral tibial apophysis resembles what is found in some species of Diplocephalus (e. g., Isaia 2005: fig. 4). Based only on these observations and without phylogenetic analyses, it is not possible to assign S. pseudofrontata to any other genus, but it clearly belongs to the Savignia - group. The position of S. harmsi is ambiguous in the phylogenetic analysis but always outside Diplocephalus. It does not have the diagnostic characters of males or females of Savignia. We agree with Wunderlich (1980 a), who mentioned the close resemblance of the male cephalic lobe and the male secondary genital organs to S. fronticornis. Both seem to be intermediate between Araeoncus and Diplocephalus while S. harmsi seems to be closer to Diplocephalus and S. fronticornis closer to Araeoncus. Based on figures (e. g., in Bosmans 1996: fig. 38), S. fronticornis seems to have a lobe emerging from the retrolateral side of the male palpal tibia similar to Araeoncus. Since it is not possible to solve the appropriate generic assignment without a phylogeny that includes both taxa plus some more potential relatives, we prefer not to draw systematic conclusions for these two species here. One of these potential close relatives is S. naniplopi as discussed in Bosselaers & Henderickx (2002). This species is probably very close to S. harmsi and S. fronticornis based on the radix and its appendices. The cephalic lobe however bears the PME rather than the AME. Also, the epigyne and vulva of S. naniplopi (Bosselaers & Henderickx 2002: fig. 8) are similar to S. fronticornis (Bosmans 1996: fig. 40). Simon (1908) described S. erythrocephalus from Western Australia as similar to the Mediterranean S. fronticornis and Coreorgonal monoceros (Keyserling, 1884) from North America (Millidge 1981: fig. 147), based on the eyes and the palp but differing from them by the simpler clypeus that is neither elongated nor a small bump. No native erigonines are known from Australia and this species does not belong to the Savignia- group nor to Savignia. Thaler (1984) underlined the problematic position of S. superstes Thaler, 1984 within the erigonines but placed it in the Savignia- group based on the form of the suprategulum and the chaetotaxy despite the lack of a bisected epigyne. The cephalic lobe resembles some species of Diplocephalus (e. g., E. latifrons comb. rev.; Fig. 15 F) but differs in the position of the AME (Thaler 1984) and the epigyne is close to Saloca. Due to the problematic relationships within the Savignia- group he preferred to place in its “ typical ” genus rather than in another monotypic genus. The current phylogenetic analysis showed that the bisected epigyne (character 178) is synapomorphic for the Savignia genus group (node 21). Savignia is deeply nested in this group so that S. superstes does not belong to Savignia. However, a pointed radical tailpiece (character 163; Fig. 8 B) is found in S. superstes (Thaler 1984: fig. 12) that was only observed in three taxa in the outgroup (Walckenaeria and “ Saloca ” in node 10) pointing to its exclusion from the Savignia- group also. Oi (1960) placed S. kawachiensis Oi, 1960 in Savignia due to the close resemblance of the palp with S. frontata but also mentions the similarity of the cephalic lobe with certain species of Araeoncus. Unfortunately, the figures and text do not allow discussing this assignment. The genus assignment of S. kartalensis is also difficult. It does not belong to Savignia. Based on the literature data only, it is not possible to discuss its assignment, however, we agree with Jocqué (1985) that it belongs to the Savignia-group.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF809221FF0AFB2BB312A572.taxon	description	This group includes taxa with a massive inner suprategular apophysis that has a highly sclerotised distal facing tip (character 85, state 8; Fig. 17 G) but lacking the massive highly sclerotised proximal facing tip (character 87). They have relatively simple cephalic lobes compared with the zero - group, i. e., a PME lobe (e. g., Eskov 1988: figs 18, 76, S. mirabilis comb. nov. and S. ussurica Eskov, 1988) but sometimes with an additional small extension of some sort (e. g., Eskov 1991 a: pl. 2 fig. 1, pl. 3 fig. 1, S. badzhalensis Eskov, 1991 and S. centrasiatica Eskov, 1991). It includes two species that were considered in the phylogenetic analysis, S. mirabilis comb. nov. and S. frontata, the type species of Savignia. However, S. frontata is somewhat intermediate of the mirabilis- and the zero - group, as it has an inner suprategular apophysis with only a distal branch (mirabilis - group) but also a very long AME lobe (zero - group). Three more taxa can be assigned to this group based on literature data: S. badzhalensis, S. centrasiatica and S. ussurica.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF809222FF0AF974B4F8A2A5.taxon	description	This group includes species restricted to the Far East. These taxa have both a massive highly sclerotised distal and proximal facing tip on the inner suprategular apophysis (characters 85, 87). Additionally, they have very derived cephalic lobes, i. e., have an AME lobe and a PME lobe (e. g., Eskov 1988: figs 67, 83, S. saitoi and S. zero) or at least a very long AME lobe (Eskov 1988: fig. 53, S. basarukini Eskov, 1988). It includes: S. amurensis Eskov, 1991, S. basarukini, S. birostra, S. borea Eskov, 1988, S. bureensis Tanasevitch & Trilikauskas, 2006, S. saitoi and S. zero. Savignia eskovi Marusik, Koponen & Danilov, 2001 should also belong to the zero - group according to the form of the cephalic lobe, which Marusik et al. (2001: fig. 51) discussed as being closest to S. borea. The same reasoning concerns S. yasudai (Saito, 1986) that was assigned to Savignia from Diplocephalus by Eskov (1988) due to the cephalic lobe that is most similar to that of S. borea. The male palpal tibial apophysis, the embolus, the tooth-like basal radical process, the dorsal radical fold and the cephalic lobes of both species are typical for Savignia.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF829223FF5DFC24B5BCA67A.taxon	type_taxon	Type species: Diastanillus pecuarius (Simon, 1884).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF829223FF5DFC24B5BCA67A.taxon	description	This genus was established mainly based on the form of the male cephalic lobe, leg and eye characters and its association with ants (Simon 1926) and includes only the type species Dia. pecuarius. It emerged as sister of node 50 that includes Diplocephalus, Araeoncus and related taxa. Its genital and somatic morphology shows characters found in one or the other of these genera, but it is clearly different to all of them. Despite the problems of monotypic genera (e. g., Platnick 1976; Prószyński 1986) this species clearly does not belong to other genera considered in this study and should keep its status. The phylogenetic position of this species is difficult to reconstruct with morphology alone due to its very derived characters that developed as adaptation to life in ant-nests.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF829223FF5DFA8CB59FA435.taxon	type_taxon	Type species: Janetschekia monodon (O. Pickard-Cambridge, 1873).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF829223FF5DFA8CB59FA435.taxon	description	This genus includes two species, J. monodon and J. necessaria Tanasevitch, 1985. They have a very special male cephalic lobe that is an autapomorphy of J. monodon in our phylogenetic analysis and only found in this genus in the current study: the inter AME – PME lobe (character 207; Fig. 15 L). This genus is the earliest branching representative of the Savignia - group in our preferred tree (Fig. 25). It has a very simple form of circular suprategular apophysis lacking any secondary appendices like the inner or outer suprategular apophysis (but still has a marginal suprategular apophysis) and has an unmodified distal suprategular apophysis with a rounded tip. Species of this genus have a sclerotised embolic membrane (like Saloca) and have a massively broadened (at least the type species, unknown to the authors if this is also true for J. necessaria) radical tailpiece (Millidge 1977: fig. 143).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8D922CFF5DFE47B363A079.taxon	type_taxon	Type species: Caucasopisthes procurvus (Tanasevitch, 1987).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8D922CFF5DFE47B363A079.taxon	description	This is a monotypic genus including the species Caucasopisthes procurvus. Tanasevitch (1987) described this species tentatively as Dactylopisthes and assigned it to the Savignia - group based on the well-developed suprategular apophysis. Its position was unclear since it showed associations to Alioranus pauper concerning the embolic division, to Diplocephalus and Erigonella ignobilis concerning the palpal tibia and had a unique female copulatory organ (Tanasevitch 1987). The embolic division and the suprategular apophyses are similar to Dactylopisthes at least based on literature data (compare Tanasevitch 1987: figs 91 – 92 with, e. g., Tanasevitch 1985 b: figs 3 – 4). Based on our phylogenetic analysis and comparing the simpler cephalic lobe, the simpler tibial apophysis, the lack of an inner suprategular apophysis and the simpler epigyne Caucasopisthes might be a close relative of Dactylopisthes.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8D922CFF5DFC82B7E5A491.taxon	type_taxon	Type species: Dactylopisthes digiticeps (Simon, 1882).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8D922CFF5DFC82B7E5A491.taxon	description	This genus includes 11 species of which Da. locketi and Da. mirabilis (Tanasevitch, 1985) are particularly similar based on several ambiguous characters from the phylogenetic analysis and the unambiguous synapomorphy (Fig. 29): the anteriorly folded ventral plate (character 191; Fig. 22 A). Among the supporting homoplastic characters are the short, robust, blunt and twisted triangular inner suprategular apophysis (character 85, state 1); the distal suprategular apophysis that protrudes and covers the tegulum and exceeds the cymbium (character 90, state 0) and the prolateral radical apophysis (character 157). Another nine species, of which five have recently been described, are currently assigned to Dactylopisthes. Based on the figures available, Da. ramit Tanasevitch, 2023 seems to fit well to the above mentioned species (Tanasevitch 2023 b: figs 1 – 12, 14 – 15). According to Zhao & Li (2014: figs 30 – 31) this is also true for Da. separatus Zhao & Li, 2014 only known from females. Dactylopisthes video (Chamberlin & Ivie, 1947) (Tanasevitch 1984: pl. 5, figs 3 – 10) and Da. digiticeps (Bosmans et al. 2009: figs 1 – 6) can also be assigned to this group of species based on figures of the prolateral radical process, the form of the distal suprategular apophysis, the thin but long tibial apophysis and the anteriorly curved posterior ends of the dorsal plate in females. Dactylopisthes khatipara Tanasevitch, 2017 also shares most characters with Da. video according to Tanasevitch (2017: figs 1 – 8). The assignments of Da. mirificus (Georgescu, 1976) and Da. diphyus (Heimer, 1987) are more difficult for the females since the anteriorly folded posterior tips of the ventral plate are not detectable on the figures. However, the males of Da. mirificus (Georgescu 1976: figs 1 – 13) and Da. diphyus (Heimer 1987: figs 4 – 8; Song et al. 1999: fig. 91 a – d) seem to have a prolateral radical process and the same type of tibial apophysis with a retrolateral appendix upon it. The cephalic PME lobes, even though smaller in Da. mirificus, are also similar to Da. video and Da. digiticeps. Dactylopisthes dongnai Tanasevitch, 2018 and Da. marginalis Tanasevitch, 2018 form a separate species group as already noted by Tanasevitch (2018 a, 2018 b). They are morphologically quite outstanding from the other species of Dactylopisthes, lacking most of the typical characters (Tanasevitch 2018 a: figs 1 – 7, 2018 b: figs 1 – 10).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8C922EFF5DFA87B5AEA26F.taxon	type_taxon	Type species: Archaraeoncus prospiciens (Thorell, 1875).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8C922EFF5DFA87B5AEA26F.taxon	description	This genus currently includes four species of which Ar. alticola Tanasevitch, 2008 and Ar. proscipiens are presumably closely related based on the very similar tibial apophysis and inner suprategular apophysis (character 85, state 5). The preferred tree proposes Dactylopisthes as the closest relative of Archaraeoncus (node 19). This is also supported by the cephalic PME lobe of Ar. alticola that is very similar to Da. video and Da. mirificus. The radical process in Ar. proscipiens was coded as a ventral radical process; however, it might be a prolateral one, which would argue again for a closer relation to Dactylopisthes. Archaraeoncus sibiricus Eskov, 1988 shows many conformations intermediate to Archaraeoncus, Dactylopisthes and Dactylopisthoides. The male cephalic lobe is closest to Ar. proscipiens and the radical process to Dactylopisthoides, at least according to literature data (compare Eskov 1988: fig. 3 with Eskov 1990: fig. 2). A closer comparison of the epigyne and the vulva of these genera might enlighten their relationships. According to Eskov (1988), Archaraeoncus is closer to the Leptorhoptrum / Lophomma group of Millidge (1977) and does not belong to the Savignia - group. This cannot be answered without considering any member of that group, but Millidge (1977) already mentions the close relations of this group with Hilaria, Tapinocyba and Savignia. In the current phylogenetic analysis Archaraeoncus emerged closer to Hilaira than to Savignia.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	type_taxon	Type species: Saloca diceros (O. Pickard-Cambridge, 1871).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	description	The genus Saloca currently includes six species. However, only two very similar species, Sa. diceros and Sa. kulczynskii, belong to this genus in the current analysis. They emerge as sister species (node 28) in the phylogenetic analyses and are well supported by a Bremer support of 16, a jackknife support of 100 (MP analysis; Fig. 26) and a bootstrap of 100 (ML analysis; Fig. 27) with the following synapomorphies (Fig. 29): the simple broad distal suprategular apophysis with an incised tip (character 90, state 9); the twisted ventral radical process (character 131) and the longitudinally and densely packed inter AMEPME setae (character 198). Additionally, some ambiguous characters support this clade. Among these characters are the cymbial retrobasal process (character 53) that is also present in Bolyphantes L. Koch, 1837, the sclerotised embolic membrane (character 100); the ventral radical hook (character 135, state 0); the massively broadened tip of the radical tailpiece (character 164) that is only basally constricted in this clade; the anteriorly extending dorsal plate of the epigyne (character 188, supporting node 8 also) and the single proximal dorsal trichobothrium on the female palpal tibia (see character 246). Saloca is also the only taxon within the Savignia - group that lacks a marginal suprategular apophysis (character 78). Currently four more species are assigned to Saloca, which are not closely related to Saloca or any other Savignia - group member. Of these, Sa. ryvkini was considered as “ extremely close to Sa. nigra ” “ by the shape of the male carapace and genitalia ” (Eskov & Marusik 1994). Saloca nigra was synonymised with Walckenaeria stylifrons (O. Pickard-Cambridge, 1875) by Wunderlich (1972). This close resemblance can be seen in the drawings of, e. g., Eskov & Marusik (1994: figs 54, 57) and Roberts (1987: figs 3 d, 8 e). We agree that Sa. ryvkini is morphologically more similar to W. stylifrons than to any other species currently assigned to Saloca. The form of the embolus and the two bunches of merged setae on the male cephalic lobe and the form of the tibial apophysis for example. Consequently, Sa. ryvkini is hereby transferred to Walckenaeria (see Table 9). According to Wunderlich (2011), Saloca elevata Wunderlich, 2011, described from Turkey, is probably closely related to the European species of Saloca as it also bears tiny denticles on the retrolateral side of the prolateral tibial apophysis. Saloca gorapaniensis and Sa. khumbuensis (node 12) were also questioned to belong to Saloca (Eskov & Marusik 1994). In the current analyses, they show affiliations with Walckenaeria (node 10) supported by the general conformation of the embolic division (the radical tailpiece, radix and embolus form a spiral; unambiguous synapomorphy of node 10; character 172) and with members of the Pelecopsis - group (node 13) rather than with the Savignia - group (Fig. 28). This confirms the isolated morphological position of these two species. Based on the characters scored for the current analysis, these two species belong to the Pelecopsis - group. Homoplastic characters like the membranous distal suprategular apophysis (character 90, state 13; convergently also present in Diplocephalus turcicus) and the spine formula 1111 (2211 in the Savignia - group) account for this. These two species should be considered in a similar project as the current one on the Pelecopsis - group to synonymise them with an already existing genus or to establish a new genus for them. According to Wunderlich (1983), they are close to Horcotes strandi (Sytshevskaja, 1935) and Alioranus chiardolae while Eskov & Marusik (1994) argue that they neither belong to Saloca nor Horcotes Crosby & Bishop, 1933.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	type_taxon	Type species: Alioranus pauper (Simon, 1882).	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	description	We included three out of four species from this genus in the Savignia genus group phylogeny. Alioranus chiardolae always emerged outside the Savignia - group close to Hilaria, while Al. pauper the type species and Al. pastoralis either emerged in a clade with Al. chiardolae (equal weights; ML; implied weights k value 23 or higher) or else as sister clade to Dactylopisthes (implied weights with a k value lower than 11). Alioranus should be restricted to Al. pauper and Al. pastoralis, i. e., node 6, which is supported by a Bremer support of 3, a jackknife support of 78 (MP analysis; Fig. 26) and a bootstrap of 99 (ML analysis; Fig. 27) and by several homoplastic characters, among which are the following (Fig. 28): the prolateral radical process (character 157) and the nearly smooth booklung covers (character 225). The prolateral radical process is absent in most taxa; however, node 6, uniting Al. pauper and Al. pastoralis, has a unique thread-like prolateral radical process that does not occur in any other species in the current analysis and might be a diagnostic of Alioranus (character 158, state 0). Alioranus chiardolae has a very simple palpal conformation compared to the former species and is probably not very closely related with Alioranus. Wunderlich (1995) mentions that Al. chiardolae is most probably a new monotypic genus due to its long palpal tibial apophysis and the form of the epigyne. Alioranus diclivitalis Tanasevitch, 1990 lacks a thread-like prolateral radical process and therefore probably does not belong to Alioranus. However, its epigyne is quite similar to Al. pastoralis (Wunderlich 1980 b: fig. 14). It is also clearly different from Al. chiardolae (e. g., the tibial apophysis; Tanasevitch 1990: fig. 3) but cannot be assigned to any other taxon considered in this analysis.	en	Frick, Holger, Sanz-Laparra, Annui M., Kropf, Christian (2026): Systematic revision of the Savignia genus group (Araneae: Linyphiidae: Erigoninae) with notes on their cephalic lobe evolution. European Journal of Taxonomy 1071: 1-138, DOI: 10.5852/ejt.2026.1071.3296, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3296/14522
