taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397278/files/figure.png	https://doi.org/10.5281/zenodo.21397278	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397280/files/figure.png	https://doi.org/10.5281/zenodo.21397280	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397289/files/figure.png	https://doi.org/10.5281/zenodo.21397289	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397291/files/figure.png	https://doi.org/10.5281/zenodo.21397291	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397303/files/figure.png	https://doi.org/10.5281/zenodo.21397303	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397313/files/figure.png	https://doi.org/10.5281/zenodo.21397313	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397320/files/figure.png	https://doi.org/10.5281/zenodo.21397320	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397345/files/figure.png	https://doi.org/10.5281/zenodo.21397345	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397355/files/figure.png	https://doi.org/10.5281/zenodo.21397355	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397387/files/figure.png	https://doi.org/10.5281/zenodo.21397387	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFFAA9234FD81F99FB5D9A56D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397369/files/figure.png	https://doi.org/10.5281/zenodo.21397369	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF979237FF0AF9BBB7CAA1CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF979237FF0AF9BBB7CAA1CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF969237FF0AFCFFB498A780.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF969237FF0AFCFFB498A780.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF969237FF0AFB3BB549A4EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF969237FF0AFB3BB549A4EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397280/files/figure.png	https://doi.org/10.5281/zenodo.21397280	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF919230FF0AFEFDB722A09D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF919230FF0AFEFDB722A09D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397278/files/figure.png	https://doi.org/10.5281/zenodo.21397278	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397280/files/figure.png	https://doi.org/10.5281/zenodo.21397280	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397289/files/figure.png	https://doi.org/10.5281/zenodo.21397289	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397291/files/figure.png	https://doi.org/10.5281/zenodo.21397291	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397303/files/figure.png	https://doi.org/10.5281/zenodo.21397303	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397313/files/figure.png	https://doi.org/10.5281/zenodo.21397313	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397316/files/figure.png	https://doi.org/10.5281/zenodo.21397316	Fig. 11. Scanning Electron Microscope pictures of the male sternum in ventral view. A. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). B. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). C. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). D. Erigonella hiemalis (Blackwall, 1841) (ZMUC). E. Erigonella hiemalis, detail (ZMUC). F. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars: A–D, F = 100 μm; E = 50 μm.	Fig. 11. Scanning Electron Microscope pictures of the male sternum in ventral view. A. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). B. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). C. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). D. Erigonella hiemalis (Blackwall, 1841) (ZMUC). E. Erigonella hiemalis, detail (ZMUC). F. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars: A–D, F = 100 μm; E = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397320/files/figure.png	https://doi.org/10.5281/zenodo.21397320	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397345/files/figure.png	https://doi.org/10.5281/zenodo.21397345	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF909231FDB9FE41B301A580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397382/files/figure.png	https://doi.org/10.5281/zenodo.21397382	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397278/files/figure.png	https://doi.org/10.5281/zenodo.21397278	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397280/files/figure.png	https://doi.org/10.5281/zenodo.21397280	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397289/files/figure.png	https://doi.org/10.5281/zenodo.21397289	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397291/files/figure.png	https://doi.org/10.5281/zenodo.21397291	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397303/files/figure.png	https://doi.org/10.5281/zenodo.21397303	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397316/files/figure.png	https://doi.org/10.5281/zenodo.21397316	Fig. 11. Scanning Electron Microscope pictures of the male sternum in ventral view. A. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). B. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). C. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). D. Erigonella hiemalis (Blackwall, 1841) (ZMUC). E. Erigonella hiemalis, detail (ZMUC). F. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars: A–D, F = 100 μm; E = 50 μm.	Fig. 11. Scanning Electron Microscope pictures of the male sternum in ventral view. A. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). B. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). C. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). D. Erigonella hiemalis (Blackwall, 1841) (ZMUC). E. Erigonella hiemalis, detail (ZMUC). F. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars: A–D, F = 100 μm; E = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397320/files/figure.png	https://doi.org/10.5281/zenodo.21397320	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397345/files/figure.png	https://doi.org/10.5281/zenodo.21397345	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397355/files/figure.png	https://doi.org/10.5281/zenodo.21397355	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397361/files/figure.png	https://doi.org/10.5281/zenodo.21397361	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397363/files/figure.png	https://doi.org/10.5281/zenodo.21397363	Fig. 23. Different types of suprategular apophyses of male palps in retrolateral (B, E, G, I, K) and distal (A, C–D, F, H, J) views. A–B. Diplocephalus cristatus (Blackwall, 1833), (Ar138; NMBE). C. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). D–E. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). F–G. Erigonella subelevata (L. Koch, 1869) (NMBE). H–I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). J–K. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar6699; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 23. Different types of suprategular apophyses of male palps in retrolateral (B, E, G, I, K) and distal (A, C–D, F, H, J) views. A–B. Diplocephalus cristatus (Blackwall, 1833), (Ar138; NMBE). C. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). D–E. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). F–G. Erigonella subelevata (L. Koch, 1869) (NMBE). H–I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). J–K. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar6699; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397365/files/figure.png	https://doi.org/10.5281/zenodo.21397365	Fig. 24. Different types of embolic divisions of male palps in distal (A–B, E, H, K), ventral (C, F, I, L) and proximal (D, G, J) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar138; NMBE). B–D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). E–G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar6699; NMBE). H–J. Erigonella subelevata (L. Koch, 1869) (NMBE). K–L. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 24. Different types of embolic divisions of male palps in distal (A–B, E, H, K), ventral (C, F, I, L) and proximal (D, G, J) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar138; NMBE). B–D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). E–G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar6699; NMBE). H–J. Erigonella subelevata (L. Koch, 1869) (NMBE). K–L. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397387/files/figure.png	https://doi.org/10.5281/zenodo.21397387	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397369/files/figure.png	https://doi.org/10.5281/zenodo.21397369	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF929233FDA3FDAEB27DA4F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923CFF0AFEC2B79AA62A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923CFF0AFEC2B79AA62A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923CFF0AFEC2B79AA62A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397387/files/figure.png	https://doi.org/10.5281/zenodo.21397387	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923EFF0AFADFB77FA774.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923EFF0AFADFB77FA774.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923EFF0AFADFB77FA774.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397387/files/figure.png	https://doi.org/10.5281/zenodo.21397387	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	Fig. 31. Continuation of Figure 30 showing nodes 49 to 68. Showing the position of Diastanillus Simon, 1926 (31A, node 49) and the redefined genus Diplocephalus Bertkau, 1883 (31B, node 53) and Araeoncus Simon, 1884 (31C, node 59, omitting D. picinus).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9D923EFF0AFADFB77FA774.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397278/files/figure.png	https://doi.org/10.5281/zenodo.21397278	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397280/files/figure.png	https://doi.org/10.5281/zenodo.21397280	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 2. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397284/files/figure.png	https://doi.org/10.5281/zenodo.21397284	Fig. 3. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). B. Erigonella hiemalis (Blackwall, 1841) (ZMUC). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 3. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). B. Erigonella hiemalis (Blackwall, 1841) (ZMUC). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397289/files/figure.png	https://doi.org/10.5281/zenodo.21397289	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397291/files/figure.png	https://doi.org/10.5281/zenodo.21397291	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397297/files/figure.png	https://doi.org/10.5281/zenodo.21397297	Fig. 6. Scanning Electron Microscope pictures of the male palp in ventral view. A. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Alioranus pastoralis (O. Pickard- Cambridge, 1872), embolic division (3695-3702; coll. E.G.). G. Erigonella ignobilis (O. Pickard- Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 6. Scanning Electron Microscope pictures of the male palp in ventral view. A. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Alioranus pastoralis (O. Pickard- Cambridge, 1872), embolic division (3695-3702; coll. E.G.). G. Erigonella ignobilis (O. Pickard- Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397303/files/figure.png	https://doi.org/10.5281/zenodo.21397303	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 7. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). B. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). C. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). D. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). E. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). F. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397313/files/figure.png	https://doi.org/10.5281/zenodo.21397313	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397316/files/figure.png	https://doi.org/10.5281/zenodo.21397316	Fig. 11. Scanning Electron Microscope pictures of the male sternum in ventral view. A. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). B. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). C. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). D. Erigonella hiemalis (Blackwall, 1841) (ZMUC). E. Erigonella hiemalis, detail (ZMUC). F. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars: A–D, F = 100 μm; E = 50 μm.	Fig. 11. Scanning Electron Microscope pictures of the male sternum in ventral view. A. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). B. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). C. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). D. Erigonella hiemalis (Blackwall, 1841) (ZMUC). E. Erigonella hiemalis, detail (ZMUC). F. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars: A–D, F = 100 μm; E = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397320/files/figure.png	https://doi.org/10.5281/zenodo.21397320	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397326/files/figure.png	https://doi.org/10.5281/zenodo.21397326	Fig. 13. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A–E, G–H) and frontal (F, I) views. A–B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). E. Glyphesis servulus (Simon, 1882) (792f; NMB). F. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). G. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). H–I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 13. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A–E, G–H) and frontal (F, I) views. A–B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). E. Glyphesis servulus (Simon, 1882) (792f; NMB). F. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). G. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). H–I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397355/files/figure.png	https://doi.org/10.5281/zenodo.21397355	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397363/files/figure.png	https://doi.org/10.5281/zenodo.21397363	Fig. 23. Different types of suprategular apophyses of male palps in retrolateral (B, E, G, I, K) and distal (A, C–D, F, H, J) views. A–B. Diplocephalus cristatus (Blackwall, 1833), (Ar138; NMBE). C. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). D–E. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). F–G. Erigonella subelevata (L. Koch, 1869) (NMBE). H–I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). J–K. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar6699; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 23. Different types of suprategular apophyses of male palps in retrolateral (B, E, G, I, K) and distal (A, C–D, F, H, J) views. A–B. Diplocephalus cristatus (Blackwall, 1833), (Ar138; NMBE). C. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). D–E. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). F–G. Erigonella subelevata (L. Koch, 1869) (NMBE). H–I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). J–K. Diplocephalus latifrons (O. Pickard- Cambridge, 1863) (Ar6699; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397365/files/figure.png	https://doi.org/10.5281/zenodo.21397365	Fig. 24. Different types of embolic divisions of male palps in distal (A–B, E, H, K), ventral (C, F, I, L) and proximal (D, G, J) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar138; NMBE). B–D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). E–G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar6699; NMBE). H–J. Erigonella subelevata (L. Koch, 1869) (NMBE). K–L. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 24. Different types of embolic divisions of male palps in distal (A–B, E, H, K), ventral (C, F, I, L) and proximal (D, G, J) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar138; NMBE). B–D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). E–G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar6699; NMBE). H–J. Erigonella subelevata (L. Koch, 1869) (NMBE). K–L. Erigonella hiemalis (Blackwall, 1841) (Ar1164; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF98923AFD8BFEFCB417A628.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397385/files/figure.png	https://doi.org/10.5281/zenodo.21397385	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9B923BFF0AF99FB7A7A18A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9B923BFF0AF99FB7A7A18A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9B923BFF0AF99FB7A7A18A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397385/files/figure.png	https://doi.org/10.5281/zenodo.21397385	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397284/files/figure.png	https://doi.org/10.5281/zenodo.21397284	Fig. 3. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). B. Erigonella hiemalis (Blackwall, 1841) (ZMUC). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 3. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). B. Erigonella hiemalis (Blackwall, 1841) (ZMUC). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397289/files/figure.png	https://doi.org/10.5281/zenodo.21397289	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397297/files/figure.png	https://doi.org/10.5281/zenodo.21397297	Fig. 6. Scanning Electron Microscope pictures of the male palp in ventral view. A. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Alioranus pastoralis (O. Pickard- Cambridge, 1872), embolic division (3695-3702; coll. E.G.). G. Erigonella ignobilis (O. Pickard- Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 6. Scanning Electron Microscope pictures of the male palp in ventral view. A. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Alioranus pastoralis (O. Pickard- Cambridge, 1872), embolic division (3695-3702; coll. E.G.). G. Erigonella ignobilis (O. Pickard- Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397307/files/figure.png	https://doi.org/10.5281/zenodo.21397307	Fig. 8. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Glyphesis servulus (Simon, 1882) (792f; NMB). B. Panamomops tauricornis (Simon, 1882) (NMBE). C. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 8. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Glyphesis servulus (Simon, 1882) (792f; NMB). B. Panamomops tauricornis (Simon, 1882) (NMBE). C. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397313/files/figure.png	https://doi.org/10.5281/zenodo.21397313	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397326/files/figure.png	https://doi.org/10.5281/zenodo.21397326	Fig. 13. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A–E, G–H) and frontal (F, I) views. A–B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). E. Glyphesis servulus (Simon, 1882) (792f; NMB). F. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). G. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). H–I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 13. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A–E, G–H) and frontal (F, I) views. A–B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). E. Glyphesis servulus (Simon, 1882) (792f; NMB). F. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). G. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). H–I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397361/files/figure.png	https://doi.org/10.5281/zenodo.21397361	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF9A9225FD87FA7FB362A25F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397382/files/figure.png	https://doi.org/10.5281/zenodo.21397382	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397335/files/figure.png	https://doi.org/10.5281/zenodo.21397335	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 16. Ventral radical process (vRP). Male palp in retrolateral (B, D–E, G–H), ventral (C) and prolateral (A, F, I) views. A. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). B. Diplocephalus protuberans (O. Pickard-Cambridge, 1875) (coll. M.I.). C. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). D. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). G. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). H. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). I. Diplocephalus montanus Eskov, 1988 (36879; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397345/files/figure.png	https://doi.org/10.5281/zenodo.21397345	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397361/files/figure.png	https://doi.org/10.5281/zenodo.21397361	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397369/files/figure.png	https://doi.org/10.5281/zenodo.21397369	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF849227FE0BF8C5B56AA2A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397385/files/figure.png	https://doi.org/10.5281/zenodo.21397385	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397355/files/figure.png	https://doi.org/10.5281/zenodo.21397355	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397361/files/figure.png	https://doi.org/10.5281/zenodo.21397361	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397385/files/figure.png	https://doi.org/10.5281/zenodo.21397385	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	Fig. 30. Continuation of Figure 29 showing nodes 32 to 48. Showing the redefined genus Erigonella Dahl, 1901 (30A, node 33), Hemistajus Schenkel, 1934 resurrected rank (30B, node 43) and Savignia Blackwall, 1833 (30B, node 45).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF869221FDB8FE47B5C1A7A1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397307/files/figure.png	https://doi.org/10.5281/zenodo.21397307	Fig. 8. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Glyphesis servulus (Simon, 1882) (792f; NMB). B. Panamomops tauricornis (Simon, 1882) (NMBE). C. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 8. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Glyphesis servulus (Simon, 1882) (792f; NMB). B. Panamomops tauricornis (Simon, 1882) (NMBE). C. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF809221FF0AFB2BB312A572.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF829223FF5DFC24B5BCA67A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF829223FF5DFA8CB59FA435.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397333/files/figure.png	https://doi.org/10.5281/zenodo.21397333	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	Fig. 15. Male cephalic lobes in lateral view. A. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2881; NMBE). D. Pelecopsis elongata (Wider, 1834) (Ar4869; NMBE). E. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). F. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar3662; NMBE). G. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). H. Savignia zero Eskov, 1988 (39671; SMF). I. Glyphesis servulus (Simon, 1882) (792a; NMB). J. Savignia frontata Blackwall, 1833 (1611c; NMB). K. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). L. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). Scale bars = 500 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF829223FF5DFA8CB59FA435.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397345/files/figure.png	https://doi.org/10.5281/zenodo.21397345	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF829223FF5DFA8CB59FA435.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397355/files/figure.png	https://doi.org/10.5281/zenodo.21397355	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 21. Radical process types. Male palp in retrolateral (A, C–D, G–H) and prolateral (B, E–F, I) views. A. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Araeoncus caucasicus Tanasevitch, 1987 (33781; SMF). D. Savignia birostra (Chamberlin & Ivie, 1947) (Ar6744; NMBE). E. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). F. Diplocephalus crassilobus (Simon, 1884) (Ar6668; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). H. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). I. Savignia producta Holm, 1977 (36885; SMF). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF829223FF5DFA8CB59FA435.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397369/files/figure.png	https://doi.org/10.5281/zenodo.21397369	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 25. Preferred tree, based on the implied weights analysis using a k-value of 23. It shows node numbers. Nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8D922CFF5DFE47B363A079.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397349/files/figure.png	https://doi.org/10.5281/zenodo.21397349	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 19. Male palpal tibia in retrolateral (A, D–I, K, N), prolateral (B) and dorsal (C, J, L–M, O–P) views. A–B. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). C. Savignia frontata Blackwall, 1833) (1611c; NMB). D. Savignia harmsi Wunderlich, 1980 (29187; SMF). E. Diplocephalus dentatus Tullgren, 1955 (2598a; NMB). F. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). G. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). H. Erigonella subelevata (L. Koch, 1869) (Ar5604; NMBE). I. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). J. Caucasopisthes procurvus (Tanasevitch, 1987) (33788; SMF). K. Glyphesis taoplesius Wunderlich, 1969 (23978; SMF). L. Diplocephalus permixtus (O. Pickard-Cambridge, 1871) (33430; SMF). M. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). N. Glyphesis cottonae (La Touche, 1946) (18756; SMF). O. Diplocephalus connatus Bertkau, 1889 (Ar2281; NMBE). P. Glyphesis servulus (Simon, 1882) (792a; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8D922CFF5DFC82B7E5A491.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397339/files/figure.png	https://doi.org/10.5281/zenodo.21397339	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 17. Different types of inner suprategular apophyses (iSTA). Male palp in retrolateral (B, E–F), ventral (C), prolateral (A, P) and distal (D) views. A. Linyphia triangularis (Clerck, 1757) (Ar272; NMBE). B. Dactylopisthes locketi (Tanasevitch, 1983) (34850; SMF). C. Glyphesis servulus (Simon, 1882) (792a; NMB). D. Diplocephalus rostratus Schenkel, 1934 (1509a; NMB). E. Savignia producta Holm, 1977 (36885; SMF). F. Dicymbium nigrum (Blackwall, 1834) (Ar6602; NMBE). G. Savignia frontata Blackwall, 1833 (1611c; NMB). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8D922CFF5DFC82B7E5A491.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397361/files/figure.png	https://doi.org/10.5281/zenodo.21397361	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 22. Female epigynes in ventral (A, C, F–G), lateral (B, D) and posterior (E) views. A. Dactylopisthes locketi (Tanasevitch, 1983) (36881; SMF). B–C. Diplocephalus mirabilis (Tanasevitch, 1985) (36881; SMF). D–E. Diplocephalus montanus Eskov, 1988 (36879; SMF). F. Diplocephalus pavesii Pesarini, 1996 (2577b; NMB). G. Paraglyphesis polaris Eskov, 1991 (coll. A.T.). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8D922CFF5DFC82B7E5A491.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397382/files/figure.png	https://doi.org/10.5281/zenodo.21397382	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397278/files/figure.png	https://doi.org/10.5281/zenodo.21397278	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397284/files/figure.png	https://doi.org/10.5281/zenodo.21397284	Fig. 3. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). B. Erigonella hiemalis (Blackwall, 1841) (ZMUC). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 3. Scanning Electron Microscope pictures of the male palp in retrolateral view. A. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). B. Erigonella hiemalis (Blackwall, 1841) (ZMUC). C. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397307/files/figure.png	https://doi.org/10.5281/zenodo.21397307	Fig. 8. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Glyphesis servulus (Simon, 1882) (792f; NMB). B. Panamomops tauricornis (Simon, 1882) (NMBE). C. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 8. Scanning Electron Microscope pictures of the male palp in prolateral view. A. Glyphesis servulus (Simon, 1882) (792f; NMB). B. Panamomops tauricornis (Simon, 1882) (NMBE). C. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397326/files/figure.png	https://doi.org/10.5281/zenodo.21397326	Fig. 13. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A–E, G–H) and frontal (F, I) views. A–B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). E. Glyphesis servulus (Simon, 1882) (792f; NMB). F. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). G. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). H–I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 13. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A–E, G–H) and frontal (F, I) views. A–B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). E. Glyphesis servulus (Simon, 1882) (792f; NMB). F. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). G. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). H–I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397345/files/figure.png	https://doi.org/10.5281/zenodo.21397345	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 18. Different types of distal suprategular apophyses (dSTA). Male palp in retrolateral (G), prolateral (B–F, H) and distal (A, I) views. A. Diplocephalus pavesii Pesarini, 1996 (coll. M.I.). B. Diplocephalus marusiki Eskov, 1988 (36878; SMF). C. Araeoncus crassiceps (Westring, 1861) (Ar1428; NMBE). D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Araeoncus victorianyanzae Berland, 1936 (00006708; ZMUC). F. Dicymbium libidinosum (Kulczyński, 1926) (39587; SMF). G. Janetschekia monodon (O. Pickard-Cambridge, 1873) (7033; SMF). H. Diplocephalus caucasicus Tanasevitch, 1987 (33790; SMF). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397353/files/figure.png	https://doi.org/10.5281/zenodo.21397353	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	Fig. 20. Different structures. Male palp in retrolateral (A, C, E, I), prolateral (D, F–H) and distal (B) views. A. Diplocephalus cristatus (Blackwall, 1833) (Ar1187; NMBE). B. Diplocephalus helleri (L. Koch, 1869) (Ar3235; NMBE). C. Savignia harmsi Wunderlich, 1980 (29187; SMF). D. Araeoncus galeriformis (Tanasevitch, 1987) (33822; SMF). E. Diastanillus pecuarius (Simon, 1884). F. Araeoncus anguineus (L. Koch, 1869) (Ar3203; NMBE). G. Diplocephalus arnoi Isaia, 2005 (coll. M.I.). H. Savignia frontata Blackwall, 1833 (1611c; NMB). I. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Abbreviations: see Material and methods. Scale bars = 100 µm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397382/files/figure.png	https://doi.org/10.5281/zenodo.21397382	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	Fig. 29. Continuation of Figure 28 showing nodes 16 to 31. Showing Dicymbium Menge, 1868 (node 29) and the redefined genus Glyphesis Simon, 1926 (node 22).	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922EFF5DFE98B53EA40F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397380/files/figure.png	https://doi.org/10.5281/zenodo.21397380	Fig. 28. Excerpt of the preferred tree, showing AccTran optimisation for nodes 1–7 (A) and 8–15 (B). Black squares account for unambiguous character changes (synapomorphies) and white squares for homoplastic character changes. Above the squares are the character numbers and below the state numbers. Node numbers are given in circles. The nomenclature corresponds to the genus assignments used until now.	Fig. 28. Excerpt of the preferred tree, showing AccTran optimisation for nodes 1–7 (A) and 8–15 (B). Black squares account for unambiguous character changes (synapomorphies) and white squares for homoplastic character changes. Above the squares are the character numbers and below the state numbers. Node numbers are given in circles. The nomenclature corresponds to the genus assignments used until now.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397278/files/figure.png	https://doi.org/10.5281/zenodo.21397278	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 1. Scanning Electron Microscope pictures of different male palpal tibial apophyses in dorsal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (2695-3702; coll E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836), retrolateral view (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863), (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). I. Erigonella hiemalis (Blackwall, 1841) (ZMUC). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397289/files/figure.png	https://doi.org/10.5281/zenodo.21397289	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	Fig. 4. Scanning Electron Microscope pictures of the male palp paracymbium in retrolateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). D. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). E. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). F. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841), (ZMUC). I. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 50 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397291/files/figure.png	https://doi.org/10.5281/zenodo.21397291	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 5. Scanning Electron Microscope pictures of the male palp in ventral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397297/files/figure.png	https://doi.org/10.5281/zenodo.21397297	Fig. 6. Scanning Electron Microscope pictures of the male palp in ventral view. A. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Alioranus pastoralis (O. Pickard- Cambridge, 1872), embolic division (3695-3702; coll. E.G.). G. Erigonella ignobilis (O. Pickard- Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 6. Scanning Electron Microscope pictures of the male palp in ventral view. A. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). B. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). C. Erigonella hiemalis (Blackwall, 1841) (ZMUC). D. Glyphesis servulus (Simon, 1882) (792f; NMB). E. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). F. Alioranus pastoralis (O. Pickard- Cambridge, 1872), embolic division (3695-3702; coll. E.G.). G. Erigonella ignobilis (O. Pickard- Cambridge, 1871) (Ar2880; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397311/files/figure.png	https://doi.org/10.5281/zenodo.21397311	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 9. Scanning Electron Microscope pictures of the male palp distal view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871) (Ar2880; NMBE). J. Glyphesis servulus (Simon, 1882) (792f; NMB). K. Hypomma bituberculatum (Wider, 1834) (Ar553; NMBE). L. Saloca diceros (O. Pickard- Cambridge, 1871) (Ar3289; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397313/files/figure.png	https://doi.org/10.5281/zenodo.21397313	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 10. Scanning Electron Microscope pictures of female epigyne types in ventral (A, C, E, G–H, J, L, N) and aboral (B, D, F, I, K, M, O) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3232; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E–F. Dicymbium nigrum (Blackwall, 1834) (9996; ZMUC). G. Dicymbium tibiale (Blackwall, 1836) (11213; ZMUC). H–I. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar161; NMBE). J–K. Entelecara erythropus (Westring, 1851) (Ar173; NMBE). L–M. Erigonella hiemalis (Blackwall, 1841) (ZMUC). N–O. Glyphesis servulus (Simon, 1882) (792f; NMB). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397320/files/figure.png	https://doi.org/10.5281/zenodo.21397320	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	Fig. 12. Scanning Electron Microscope pictures of the male cephalic lobes in lateral (A, C, E–I) and frontal (B, D) views. A–B. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). C–D. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). E. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). F. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). G. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). H. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). I. Dismodicus bifrons (Blackwall, 1841) (Ar1192; NMBE). Abbreviations: see Material and methods. Scale bars = 100 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397329/files/figure.png	https://doi.org/10.5281/zenodo.21397329	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	Fig. 14. Scanning Electron Microscope pictures of different male chelicera in lateral view. A. Alioranus pastoralis (O. Pickard-Cambridge, 1872) (3695-3702; coll. E.G.). B. Araeoncus humilis (Blackwall, 1841) (Ar3186; NMBE). C. Dicymbium nigrum (Blackwall, 1834) (Ar637; NMBE). D. Dicymbium tibiale (Blackwall, 1836) (11430; ZMUC). E. Diplocephalus latifrons (O. Pickard-Cambridge, 1863) (Ar1262; NMBE). F. Diplocephalus picinus (Blackwall, 1841) (Ar6645; NMBE). G. Entelecara erythropus (Westring, 1851) (Ar172; NMBE). H. Erigonella hiemalis (Blackwall, 1841) (ZMUC). I. Erigonella ignobilis (O. Pickard-Cambridge, 1871), ♂ (Ar2880; NMBE). J. Erigonella ignobilis (O. Pickard-Cambridge, 1871), detail (Ar2880; NMBE). K. Glyphesis servulus (Simon, 1882) (792f; NMB). L. Saloca diceros (O. Pickard-Cambridge, 1871) (Ar2188; NMBE). Scale bars: A–I, K–L = 100 μm; J = 10 μm.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397374/files/figure.png	https://doi.org/10.5281/zenodo.21397374	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	Fig. 26. Preferred tree as seen in Figure 25, showing nomenclatural changes resulting from the systematic revision (Table 9). Genera that are considered to belong to the Savignia genus group according to the current analysis are marked in grey and with bars. The marked groups define the new genus delimitations following the nomenclature established in the current analysis. It shows node numbers, Bremer and Jackknife support values respectively on the nodes.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397378/files/figure.png	https://doi.org/10.5281/zenodo.21397378	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	Fig. 27. Maximum likelihood tree with SH-aLRT and ultrafast bootstrap values respectively on the nodes. Genera assigned to the Savignia-group by Millidge (1977) are marked with colours and bars. The black bars indicate taxa from genera assigned to the Savignia genus group and those with grey bars to the other subgroups of the Savignia-group. The nomenclature corresponds to the genus assignments used until now. The different colours mark the new genus delimitations.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
C94B87AFFF8F922FFF5DF871B365A0F1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/21397380/files/figure.png	https://doi.org/10.5281/zenodo.21397380	Fig. 28. Excerpt of the preferred tree, showing AccTran optimisation for nodes 1–7 (A) and 8–15 (B). Black squares account for unambiguous character changes (synapomorphies) and white squares for homoplastic character changes. Above the squares are the character numbers and below the state numbers. Node numbers are given in circles. The nomenclature corresponds to the genus assignments used until now.	Fig. 28. Excerpt of the preferred tree, showing AccTran optimisation for nodes 1–7 (A) and 8–15 (B). Black squares account for unambiguous character changes (synapomorphies) and white squares for homoplastic character changes. Above the squares are the character numbers and below the state numbers. Node numbers are given in circles. The nomenclature corresponds to the genus assignments used until now.	2026-06-25	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian		Zenodo	biologists	Frick, Holger;Sanz-Laparra, Annui M.;Kropf, Christian			
