identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CA3E87E5FE4E55117E4CF94FFC23FA14.text	CA3E87E5FE4E55117E4CF94FFC23FA14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carcinione platypleura Bourdon 1983	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> CARCINIONE PLATYPLEURA BOURDON, 1983</p>
            <p>(FIGS 1, 2, 10)</p>
            <p> ‘ Seventeen specimens of epicarids … collected from four genera of crabs (including  Pseudohapalocarcinus )’ Monod &amp; Serène, 1976: 24 (in part) [ex  Pseudohapalocarcinus ransoni Fize &amp; Serène, 1956a , Indonesia, examined in this study]. </p>
            <p> ? ‘bopyrid parasites’ Takeda &amp; Tamura, 1980: 54–55 59, pl. 5 [ex  P. ransoni Fize &amp; Serène, 1956a , Ryukyu Islands, Japan]. </p>
            <p> Carcinione platypleura Bourdon, 1983: 854–855 , fig. 6 [ex  P. ransoni Fize &amp; Serène, 1956a , Banda Neira, Indonesia, examined in the present study]. – van der Meij, 2014: 45 [ex  Opecarcinus cathyae van der Meij, 2014 , Halmahera, Indonesia, examined in this study]. – van der Meij, 2015: fig. 7B [ex  Fungicola syzygia van der Meij, 2015 , Bunaken Island, Indonesia, examined in the present study]. – van der Meij, Reijnen &amp; Reimer, 2017: fig. 1F [ex  Lithoscaptus sp. , Odo Beach, Okinawa, Japan, examined in the present study]. </p>
            <p> ‘bopyrid isopod’ Kropp, 1994: 534 [ex  P. ransoni Fize &amp; Serène, 1956a , Banda Neira, Moluccas, examined in the present study]. </p>
            <p> ‘unidentified bopyrid’ Williams &amp; Boyko, 2012: fig. 1G [ex  Lithoscaptus helleri (Fize &amp; Serène, 1957) , Indonesia, examined in the present study]. </p>
            <p>Type material</p>
            <p> Indonesia: Mature female holotype (2.0 mm) [MNHN-IU-2007-3670 (= MNHN-EP 336)] and cryptoniscus paratype (0.6 mm) [MNHN-IU-2017-8 (= MNHN-Ep 336)], ex female  P. ransoni Fize &amp; Serène, 1956a (3.3 mm long × 3.4 mm wide) (MNHN-IU-2016–10934), parasites under right branchiostegite, ex  Pavona cactus (Forskål, 1775) (  Agariciidae ), Banda Neira, Amboine (= Ambon), 29 January 1975, collected by Rumphius II (R. Serène &amp; T. Monod) [crab and coral originally identified by R. Serène, with the coral only identified to genus; crab identification confirmed and coral identified to species by S. E. T. van der Meij). </p>
            <p>Additional material</p>
            <p> Indonesia: Immature female (1.5 mm) (MNHN-IU-2017-9), ex right branchiostegite of male  P. ransoni Fize &amp; Serène, 1956a (2.4 mm long × 2.3 mm wide) (MNHN-IU-2009-5734 [= MNHN-B12674]), ex  Pavona cactus (Forskål, 1775) (  Agariciidae ), southwest corner of Naira Island, at southern entrance to channel between Gunung Api Island and Naira Island, coral, Amboine (= Ambon), 29 January 1975, collected by Rumphius II [this specimen is not part of the type series, although it was collected at the same time, in the same place, and from the same host species as the holotype, but it was not mentioned in the original description by Bourdon (1983)]; mature non-ovigerous female (1.8 mm) and mature male (0.5 mm) (RMNH. Crus.I.7719), ex left branchiostegite, mature ovigerous female (1.7 mm), mature male (0.7 mm) (RMNH. Crus.I.7720), ex right branchiostegite, ex juvenile female  Dacryomaia japonica (Takeda &amp; Tamura, 1981) (2.7 mm long × 2.0 mm wide) (RMNH.Crus.D.54193), ex  Leptastrea cf. transversa Klunzinger, 1879 (  Scleractinia incertae sedis), Teluk Makawide Reef, Lembeh Strait, 01°29′05″N, 125°14′26″E, 9 February 2012, coll. S. E. T. van der Meij; mature non-ovigerous female (3.3 mm) and mature male (0.9 mm) (RMNH. Crus.I.7721), ex right branchiostegite of mature non-ovigerous female  F. syzygia van der Meij, 2015 (5.0 mm long × 4.3 mm wide) (RMNH.Crus.D.53224), ex  Pleuractis granulosa (Klunzinger, 1879) (  Fungiidae ), Tanjung Pasir Putih Reef, Ternate, 00°51′50″N, 127°20′37″E, 2 November 2009, coll. S. E.T.van der Meij; mature ovigerous female, non-ovigerous (3.6 mm) and mature male (1.1 mm) (RMNH.Crus.I.7722), ex right branchiostegite, cryptoniscus larva (0.6 mm) (RMNH. Crus.I.7723), ex left branchiostegite, ex mature non-ovigerous female  F. syzygia van der Meij, 2015 (4.3 mm long × 3.7 mm wide) (RMNH.Crus.D.56481), ex  Pleuractis paumotensis (Stutchbury, 1833) (  Fungiidae ), Alung Banua Reef, Bunaken Island, 01°37′07″N, 124°45′30″E, 19 December 2008, coll. S. E. T. van der Meij; mature non-ovigerous female (2.3 mm) (RMNH. Crus.I.7724), ex right branchiostegite of male  F. syzygia van der Meij, 2015 (3.3 mm long × 3.2 mm wide) (RMNH.Crus.D.56482), ex  Pleuractis paumotensis (Stutchbury, 1833) (  Fungiidae ), Siladen I Reef, Siladen Island, 01°37′31″N, 124°47′54″E, 18 December 2008, coll. S. E. T. van der Meij; mature ovigerous female (3.6 mm), mature male (1.3 mm) (RMNH.Crus.I.7725), ex right branchiostegite of male  Lithoscaptus prionotus Kropp, 1994 (5.3 mm long × 3.5 mm wide) (RMNH. Crus.D.54171), ex  Oulophyllia crispa (Lamarck, 1816) (  Merulinidae ), Tanjung Kuning Reef, Lembeh Strait, 01°23′10″N, 125°10′23″E, 11 February 2012, coll. S. E. T. van der Meij; mature ovigerous female (4.3 mm), mature male (1.5 mm) (RMNH.Crus.I.7726), ex left branchiostegite of non-ovigerous female  L. helleri (Fize &amp; Serène, 1957) (5.7 mm long × 4.2 mm wide) (RMNH. Crus.D.54057), ex  Favites cf. complanata (Ehrenberg, 1834) (  Merulinidae ), Siladen I Reef, Siladen Island, 01°37′31″N, 124°47′54″E, 18 December 2008, coll. S. E. T. van der Meij; mature ovigerous female (2.5 mm), mature male (1.0 mm), cryptoniscus larva (0.7 mm) (RMNH.Crus.I.7727), ex right branchiostegite of male  L. helleri (Fize &amp; Serène, 1957) (3.5 mm long × 2.6 mm wide) (RMNH. Crus. D.56121), ex  Favites cf. complanata (Ehrenberg, 1834) (  Merulinidae ), Lekuan I Reef, Bunaken Island, 01°35′44″N, 124°46′12″E, 11 December 2008, coll. S. E. T. van der Meij; mature ovigerous female (2.2 mm), mature male (0.9 mm) (RMNH.Crus.I.7728), ex right branchiostegite of male  Lithoscaptus cf. bani (Fize &amp; Serène, 1957) (telson abnormally wide) (3.2 mm long × 2.3 mm wide) (RMNH. Crus.D.56121), ex  Goniastrea pectinata (Ehrenberg, 1834) (  Merulinidae ), Negiri Reef, Manado Tua Island, 01°36′58″N, 124°41′39″E, 5 December 2008, coll. S. E. T. van der Meij; mature ovigerous female (3.3 mm), mature male (0.9 mm) (RMNH.Crus.I.7729), ex right branchiostegite of non-ovigerous female  Lithoscaptus sp. 1 (A) (4.3 mm long × 2.9 mm wide) (RMNH.D.54067), ex  Astrea curta Dana, 1846 (  Merulinidae ), Ron’s Point, Bunaken Island, 01°36′21″N, 124°44′13″E, 9 December 2008, coll. S. E.T. van der Meij; mature ovigerous female (3.1 mm), mature male (1.3 mm) (RMNH.Crus.I.7730), ex right branchiostegite, mature ovigerous female (3.1 mm), mature male (1.1 mm) (RMNH.Crus.I.7731), ex left branchiostegite of non-ovigerous female  Lithoscaptus sp. 2 (B) (4.0 mm long × 3.3 mm wide) (RMNH.Crus.D.54106), ex  Plesiastrea versipora (Lamarck, 1816) (  Scleractinia incertae sedis), Lekuan III, Bunaken Island, 01°36′20″N, 124°46′08″E, 12 December 2008, coll. S. E. T. van der Meij; mature ovigerous female (2.7 mm), mature male (1.0 mm) (RMNH.Crus.I.7732), ex left branchiostegite of juvenile female  Lithoscaptus sp. 2 (B) (4.0 mm long × 2.5 mm wide) (RMNH.Crus.D.54172), ex  P. versipora (Lamarck, 1816) (  Scleractinia incertae sedis), Pulau Abadi, Lembeh Strait, 01°26′01″N, 125°12′22″E, 10 February 2012, coll. S. E. T. van der Meij; cryptoniscus larvae (0.6 mm) (RMNH. Crus.I.7733), ex dorsal carapace of feminized male  Opecarcinus sp. 1 (F) (3.0 mm long × 2.3 mm wide) (RMNH.Crus.D.53990; host also parasitized by one  Sacculina quadrialata sp. nov. , RMNH.Crus.C.10249), ex  Leptoseris mycetoseroides Wells, 1954 (  Agariciidae ), South Pilongga, Tidore, 00°42′44″N, 127°28′47″E, 12 November 2009, coll. S. E. T. van der Meij; mature ovigerous female (4.0 mm), mature male (1.5 mm) (RMNH.Crus.I.7734), ex right branchiostegite of non-ovigerous female  Opecarcinus pholeter Kropp, 1989 (5.3 mm long × 4.3 mm wide) (RMNH.Crus.D.54000), ex  Pavona explanulata (Lamarck, 1816) (  Agariciidae ), Sulamadaha II, Ternate, 00°52′02″N, 127°19′46 ″E, 6 November 2009, coll. S. E. T. van der Meij; immature female (1.8 mm), mature male (0.8 mm) (RMNH. Crus.I.7735), ex right branchiostegite, immature female (1.5 mm), mature male (0.7 mm) (RMNH.Crus.I.7736), ex left branchiostegite of male  O. pholeter Kropp, 1989 (3.3 mm long × 2.7 mm wide) (RMNH.Crus.D.54000), ex  P. explanulata (Lamarck, 1816) (  Agariciidae ), Sulamadaha II, Ternate, 00°52′02″N, 127°19′46″E, 6 November 2009, coll. S. E. T. van der Meij; mature ovigerous female (2.8 mm), mature male (1.2 mm) (RMNH.Crus.I.7737), ex right branchiostegite of male  Opecarcinus cathyae van der Meij, 2014 (4.0 mm long × 2.7 mm wide) (RMNH.Crus.D.53923), ex  Pavona clavus (Dana, 1846) (  Agariciidae ), South Lela, Gura Ici, 00°01′51″S, 127°15′03″E, 10 November 2009, coll. S. E. T. van der Meij; mature non-ovigerous female (1.5 mm), ex right brachiostegite, mature non-ovigerous female (1.5 mm), ex left brachiostegite, mature male (0.8 mm), ex unknown brachiostegite (RMNH. Crus.I.7738), ex juvenile female  Xynomaia sheni (Fize &amp; Serène, 1956b) (2.7 mm long × 1.8 mm wide) (RMNH.Crus.D.54115), ex  Mycedium elephantotus (Pallas, 1766) (  Merulinidae ), Lobangbatu Besar, Lembeh Strait, 01°25′49″N, 125°11′26″E, 7 February 2012, coll. S. E. T. van der Meij; mature ovigerous female (2.4 mm), mature male (1.2 mm) (RMNH. Crus.I.7739), ex right branchiostegite of male  X. sheni (Fize &amp; Serène, 1956b) (4.0 mm long × 2.7 mm wide) (RMNH.Crus.D.54085), ex  Pectinia sp. (  Merulinidae ), Mandolin, Bunaken Island, 01°36′44″N, 124°43′57″E, 9 December 2008, coll. S. E. T. van der Meij. </p>
            <p> Malaysia: Mature ovigerous female (5.0 mm), mature male (1.5 mm) (RMNH.Crus.I.7740), ex left branchiostegite of non-ovigerous female  L. prionotus Kropp, 1994 (7.1 mm long × 5.2 mm wide) (RMNH. Crus.D.53719), ex  Oulophyllia crispa (Lamarck, 1816) (  Merulinidae ), Erzherzog Reef, Semporna, Malaysia, 04°14′26″N, 118°23′35″E, 1 December 2010, coll. S. E. T. van der Meij. </p>
            <p> Maldives: Immature female (1.7 mm) ex right branchiostegite, immature female (1.6 mm) ex left branchiostegite (RMNH.Crus.I.7700), ex female  F. syzygia van der Meij, 2015 (3.3 mm long × 3.1 mm wide) (RMNH.Crus.D.57063), ex  P. granulosa (Klunzinger, 1879) (  Fungiidae ), Sunny Reef, Faafu Atoll, 03°08′40″N, 73°00′45″E, 19 February 2015, coll. S. E. T. van der Meij; mature non-ovigerous female (1.5 mm), mature male (0.7 mm) (RMNH.Crus.I.7701), ex left branchiostegite of female  Opecarcinus sp. (2.0 mm long × 1.7 mm wide) (RMNH.Crus.D.57064), ex  L. mycetoseroides Wells, 1954 (  Agariciidae ), Wallino, Faafu Atoll, 03°05′12″N, 72°57′23″E, 21 February 2015, coll. S. E. T. van der Meij; mature female with brood of epicaridean larvae (3.3 mm), mature male (1.1 mm) ex right branchiostegite, mature non-ovigerous female (3.1 mm), mature male (1.2 mm) (RMNH.Crus.I.7704) ex left branchiostegite of female  Lithoscaptus sp. Z (5.4 mm long × 3.7 mm wide) (RMNH.Crus.D.57066); immature female (1.1 mm), mature male (0.9 mm) (RMNH.Crus.I.7704) ex right branchiostegite of male  Lithoscaptus sp. Z (RMNH. Crus.D.57066), ex  Dipsastraea cf. vietnamensis (Veron, 2000) (  Merulinidae ), Wallino, Faafu Atoll, 03°05′13″N, 72°57′24″E, 26 February 2015, coll. S. E. T. van der Meij; mature ovigerous female (1.5 mm), mature male (0.8 mm) (RMNH.Crus.I.7703), ex left branchiostegite of male  Lithoscaptus sp. Z (3.6 mm long × 2.3 mm wide) (RMNH.Crus.D.57066), ex  D. cf. vietnamensis (Veron, 2000) (  Merulinidae ), Wallino, Faafu Atoll, 03°05′13″N, 72°57′24″E, 26 February 2015, coll. S. E. T. van der Meij; mature ovigerous female (3.3 mm), mature male (1.4 mm) (RMNH.Crus.I.7702), ex left branchiostegite of female  Lithoscaptus sp. (5.2 mm long × 3.0 mm wide) (RMNH.Crus.D.57065), ex  Favites aff. flexuosa (Dana, 1846) (  Merulinidae ), Beyrufushi, Faafu Atoll, 03°06′29″N, 73°01′07″E, 26 February 2015, coll. S. E. T. van der Meij. </p>
            <p> Japan: Immature female (1.1 mm) (RMNH. Crus.I.7741), ex right branchial chamber of male  Dacryomaia sp. (2.3 mm long × 1.5 mm wide) (RMNH. Crus.D.57231), ex  Leptastrea sp. (  Scleractinia incertae sedis), Red Beach, Okinawa, 26°26′47″N, 127°54′43″E, 14 April 2016, coll. S. E. T. van der Meij; mature female (3.5 mm) (RMNH.Crus.I.7742) with immature female  Cabirops (2.1 mm) and cryptoniscus larva (1.3 mm) (RMNH.Crus.I.7743) in marsupium, immature male (1.7 mm), ex right branchial chamber of female  Lithoscaptus sp. (5.2 mm long × 3.8 mm wide) (RMNH.Crus.D.57232), ex  Coelastrea aspera (Verrill, 1866) (  Merulinidae ), Odo Beach, Okinawa, 26°05′22″N, 127°42′33″E, 22 April 2016, coll. S. E. T. van der Meij; ovigerous female (2.6 mm) (RMNH.Crus.I.7744) with three associated  Cabirops cryptoniscus larvae (1.2, 1.2, 1.3 mm) (RMNH.Crus.I.7745–7746), mature male (1.1 mm), (RMNH.Crus.I.7747), ex right branchial chamber, mature female (2.9 mm) with ovigerous  Cabirops female (2.9 mm long × 1.5 mm wide) and cryptoniscus larva (1.0 mm) (RMNH.Crus.I.7748) in brood chamber, mature male (1.0 mm), ex left branchial chamber of male  Xynomaia sp. (4.3 mm long × 3.8 mm wide) (RMNH.Crus.D.57233), ex  Pectinia sp. (  Merulinidae ), Blue Tombs, Henza Island, Okinawa, 26°21′46″N, 127°59′50″E, 29 April 2016, coll. S. E. T. van der Meij. </p>
            <p>Redescription</p>
            <p>Female (Fig. 1): Body length 4.3 mm, maximal width 2.3 mm, head length 0.9 mm, head width 0.9 mm. Pereon nearly straight, head not deflected (Fig. 1A, B). All body regions and pereomeres distinctly segmented.</p>
            <p>Head subrectangular, approximately as broad as long, usually overlapping entire first pereomere medially (Fig. 1A), but sometimes only overlapping in part Fig. 1C), with moderately broad frontal lamina (Fig. 1A, C). Eyes minute, distolaterally placed. Antennule of four articles; antenna of two articles (Fig. 1D). First oostegite (Fig. 1E) anterior lobe globular, posterior lobe slightly smaller than anterior lobe, ovate with rounded margins, distal projection lacking but with medially directed smooth lobe, internal ridge thick, smooth. Oostegites completely enclosing the marsupium. Maxilliped (Fig. 1F) with short, rounded fleshy spur; palp lacking, anterior segment triangular, rounded, posterior segment irregularly subquadrate. Barbula smooth (Fig. 1G).</p>
            <p>Pereon composed of seven pereomeres (Fig. 1A, C), broadest across pereomeres III–V, only slightly tapering anteriorly and posteriorly; pereomere I posterior margin usually under head, II anteromedially concave, III weakly anteromedially concave to weakly convex, IV anteromedially straight to weakly convex, V–VII weakly convex. Coxal plates very reduced, largest on pereomeres I, II, large dorsolateral bosses on pereomeres I–IV or I–V (those on V sometimes much smaller than other pairs). Anterior pereopods (Fig. 1H) much smaller than posterior pairs (Fig. 1I), posterior pairs about two times as large.</p>
            <p>Five pleomeres plus pleotelson (Fig. 1A, C), lateral plates well developed, edges crenulated, those on pereopods I–III about twice as broad as other pairs (Figs 1A–C, 2A). Pleomere I with biramous pleopods, II–V with uniramous pleopods all subequal in size, uniramous uropods resembling lateral plates, edges crenulated (Fig. 2A).</p>
            <p>Male (Fig. 2B): Length 1.5 mm, maximal width 0.5 mm, head length 0.2 mm, head width 0.3 mm, pleon length 0.5 mm.</p>
            <p>Head ovate, widest posteriorly, distinct from first segment of pereon (Fig. 2B); eyes present posterolaterally. Antennule of four articles (Fig. 2D), distally setose, not extending beyond margin of head; antenna of three articles, terminal article less than half as large as second segment, distally setose (Fig. 2D).</p>
            <p>Pereomere III–V broadest, others gradually tapering anteriorly and posteriorly. All pereomeres slightly concave anteriorly, convex posteriorly, distal margins rounded. All pereopods (Fig. 2C) subequal, all articles distinctly separated.</p>
            <p>Pleon with five segments plus pleotelson. All pleomeres directed laterally (Fig. 2B, C), with distolateral margins rounded. Midventral tubercles, pleopods and uropods lacking; small anal cone mediodistally on pleotelson (Fig. 2C).</p>
            <p>Remarks</p>
            <p> Bourdon (1983) described this species on the basis of the single female mentioned obliquely by Monod &amp; Serène (1976); although he had a cryptoniscus larva, he did not describe it. The male has not been described previously. The characters of the female, particularly the wide anterior lateral plates, made Bourdon (1983) unsure of the subfamily in which to place this species. He indicated that he did not think it belonged to the ‘Céponiens’ (now  Keponinae ), but that the female somewhat resembled those of  Pseudioninae . Given the lack of female characters that clearly placed this genus in any particular subfamily, Boyko et al. (2013) somewhat arbitrarily placed the genus with the other brachy-uran parasites in  Keponinae . However, the discovery of the male provides characters that are very similar to those of most males in  Pseudioninae and not at all like males belonging to  Keponinae . Therefore,  Carcinione is here placed with confidence in  Pseudioninae . </p>
            <p> This species appears to be the most common parasite of gall crabs, with a wide distribution in the Indo-West Pacific. The fact that it was described from only a single mature female and a cryptoniscus larva and has subsequently been reported only once before the present study (Sanabe &amp; Tsuchiya, 2005) is explained by limited sampling of hosts rather than genuine scarcity of the parasites. This conclusion is supported by the data of Sanabe &amp; Tsuchiya (2005), where 10.64% of  P. ransoni (N = 357) collected in the Ryukyus, Japan, were infested with a bopyrid identified as  C. platypleura . Although we have not seen any of Sanabe &amp; Tsuchiya’s specimens, we are confident in their identification of the bopyrid species. </p>
            <p> The parasitized  Xynomaia sp. from Okinawa might well hold the record for the most parasites and hyperparasites associated with a single host. This male crab has an ovigerous female  C. platypleura with a mature male in the right branchial chamber and a mature, non-ovigerous  C. platypleura in the left branchial chamber. The bopyrid in the right chamber, despite being ovigerous, co-occurs with three  Cabirops cryptoniscus larvae, one of which is in the marsupium amongst the eggs. The bopyrid in the left branchian chamber contains an ovigerous  Cabirops in the marsupium and a cryptoniscus larva. As if this were not enough energy burden on the host, there is an immature female bopyrid (RMNH.Crus.I.7750) under the abdomen, which belongs to a new bopyrid genus and species described in the next subsection. This single host, therefore, has an astounding five bopyrid parasites, belonging to two species, and five hyperparasitic cryptoniscoids associated with it. The  Cabirops species , being hyperparasitic and found associated with  C. platypleura from two hosts in Okinawa, will be described in a future paper. </p>
            <p>Known hosts</p>
            <p> Dacryomaia japonica (Takeda &amp; Tamura, 1981) ,  Dacryomaia sp. ,  F. syzygia van der Meij, 2015 , L i t h o s c a p t u s c f. b a n i (F i z e &amp; S e r è n e, 1 9 5 7),  L. helleri (Fize &amp; Serène, 1957) ,  L. prionotus Kropp, 1994 ,  Lithoscaptus sp. ,  Lithoscaptus sp. 1 (A),  Lithoscaptus sp. 2 (B),  Lithoscaptus sp. Z,  Opecarcinus cathyae van der Meij, 2014 ,  Opecarcinus pholeter Kropp, 1989 ,  Opecarcinus sp. 1 (F),  P. ransoni Fize &amp; Serène, 1956a ,  X. sheni (Fize &amp; Serène, 1956b) ,  Xynomaia sp.</p>
            <p>Distribution</p>
            <p>Known from localities in Indonesia, Malaysia, Maldives and Japan.</p>
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	https://treatment.plazi.org/id/CA3E87E5FE4E55117E4CF94FFC23FA14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE4955107EC2FA40FE10FCDE.text	CA3E87E5FE4955107EC2FA40FE10FCDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spathione Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> SPATHIONE GEN. NOV.</p>
            <p>urn:lsid:zoobank.org:act: B47EFD34-1336-48F2- B0AD-E3C0B6CD0584</p>
            <p>Diagnosis</p>
            <p>Female: Body compact, little deflected except for head with slight sinistral deflection. Head about as long as wide, overlapping almost entire first pereomere medially, inflated dorsoventrally, small lateral eyes present, frontal lamina present but very thin. Antennule of four segments; antenna of two segments. All pereomeres distinct, body highly vaulted ventrally, coxal plates very small, pronounced tergal projections on pereomeres one to seven, those on one to four irregular in shape, those on five to seven smooth and rounded at ends (similar to appearance of lateral plates); five pairs of well-developed oostegites on pereomeres I–V, completely closing marsupium. Maxilliped palp lacking. Five pleomeres plus pleotelson, lateral plates well developed, tubular, rounded distally and recurved; biramous tubular pleopods on pleomeres I–IV as ventrally directed extensions of lateral plates; uniramous pleopods on pleomere V, bulbous uniramous uropods.</p>
            <p>Male: Longer than broad, all segments clearly separated; abdomen not abruptly narrower than pereon, tapering anteriorly and posteriorly. Head ovate, eyes present. Antennae of four segments; antennules of two segments. Anterior pereomeres straight; posterior pereomeres and pleomeres curved posterolaterally (in dorsal view) and curved ventrally (in lateral view). Dactyli and propodi of all pereopods subequal. Four pleomeres plus pleotelson, distal margins rounded, no midventral tubercles on pleon; pleopods and uropods lacking.</p>
            <p>Type and only species</p>
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	https://treatment.plazi.org/id/CA3E87E5FE4955107EC2FA40FE10FCDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE4855107DB8FCDCFBACF8D2.text	CA3E87E5FE4855107DB8FCDCFBACF8D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spathione asprosdovrima Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Spathione asprosdovrima sp. nov.</p>
            <p>Remarks</p>
            <p> The finding of a bopyrid parasite in the marsupium of a gall crab was unexpected, as only species of  Rhopalione Pérez, 1920 were known to be adapted to this environment, and gall crabs and pinnotherids, although both thoracotreme brachyurans, are not closely related (Wetzer, Martin &amp; Boyce, 2009; van der Meij &amp; Schubart, 2014; Tsang et al., 2014). However,  Spathione gen. nov. is indeed very closely related to  Rhopalione , which contains four species (and one nomen nudum), all of which are parasitic in the marsupia of pinnotherid crabs. This is not, therefore, a case of convergent evolution of the bopyrids towards marsupial pouch parasitism, but rather a diversification of a prior preference towards marsupium parasitism from the original host to other crabs with similar structures. The only such crabs where females bear enlarged marsupial abdomens are species in the Pinnotheroidea (  Aphanodactylidae and  Pinnotheridae ) and Cryptochiroidea (  Cryptochiridae ). Which taxon was the original host for the common ancestor of  Rhopalione and  Spathione gen. nov. cannot be determined or even suggested, because there is no lengthy fossil record for gall crabs (but see below), and the oldest pinnotherid (a species of  Viapinnixa Schweitzer &amp; Feldmann, 2001 ) is only Danian (Paleocene, 61.6–66 Mya; Brösing, 2008) and appears to belong to Pinnothereliinae, species whose females do not have the requisite marsupial pouch needed for  Rhopalione to be able to occupy it. Schweitzer et al. (2010) did later move  Viapinnixa to  Pinnotherinae but without giving any rationale, and it is not clear that this is the proper placement (S. T. Ahyong, personal communication). Recently, De Angeli &amp; Ceccon (2015) described a putative fossil gall crab,  Montemagrechirus tethysianus , from Italian Eocene deposits, but this is not a cryptochirid (see Klompmaker &amp; Boxshall, 2015; Klompmaker, Portell &amp; van der Meij, 2016). Currently, gall crabs in the fossil record date back only to the Pleistocene and Pliocene and are known only from their crescentic pits as an ichnotaxon (Klompmaker, Portell &amp; van der Meij, 2016). </p>
            <p> Superficially,  Rhopalione and  Spathione gen. nov. species appear very similar, but the two genera can be distinguished easily by numerous characters, as follows: female head subquadrate in  Spathione gen. nov. vs. head rounded in  Rhopalione ; female and male antennae of four articles each, antennules of two articles in  Spathione gen. nov. vs. five articles and three articles, respectively, in  Rhopalione ; first oostegite with rounded posterior margin in  Spathione gen. nov. vs. with sharp point in  Rhopalione ; coxal plates small in  Spathione gen. nov. vs. well developed in  Rhopalione ; female first perepods approximately half as large as seventh pereopods in  Spathione gen. nov. vs. pereopods isomorphic in  Rhopalione ; tergal projections of pereomeres IV–VII similar in shape to lateral plates of pleomeres vs. tergal projections of pereomeres IV– VII not extended and not resembling lateral plates of pleomeres in  Rhopalione ; female pleopods biramous as two separate tubular stalk-like rami extending from ventral surface of lateral plate in  Spathione gen. nov. vs. pleopods as biramous as pair of flattened oval structures arising from common junction at base of ventral lateral plate in  Rhopalione ; males with four pleomeres, lacking pleopods, plus pleotelson in  Spathione gen. nov. vs. males with five pleomeres, rounded pleopods on each segment, plus pleotelson in  Rhopalione ; and female uropods as short, rounded lateral lobes of pleotelson in  Spathione gen. nov. vs. posteriorly extended and foliaceous in  Rhopalione . </p>
            <p> Rhopalione has been placed in Ioninae sensu lato (=  Keponinae Boyko, Moss, Williams &amp; Shields, 2013 ; e.g. Markham, 1992; Boyko et al., 2013; An et al., 2014), but several characters, particularly the biramous pleopods of the female and lack of midventral tubercles and presence of a non-bifurcated pleotelson of the male, support its transfer herein to  Pseudioninae .  Spathione gen. nov. is likewise placed in  Pseudioninae and shows the additional character of the male having fewer than five pleomeres; this character is variable in  Pseudioninae , whereas males of species in  Keponinae have five pleomeres. </p>
            <p>Etymology</p>
            <p> The generic name is a derived from a combination of the Greek σπάθη (spáthē), meaning saber, because of the extended, non-lamellar, lateral plates on the pleon, and  Ione , a genus of bopyrid itself and also a common suffix for other bopyrid genera. </p>
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	https://treatment.plazi.org/id/CA3E87E5FE4855107DB8FCDCFBACF8D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE47551F7DDEFF66FAACF9CA.text	CA3E87E5FE47551F7DDEFF66FAACF9CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spathione asprosdovrima Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> SPATHIONE ASPROSDOVRIMA SP. NOV.</p>
            <p>(FIGS 3– 5, 10E)</p>
            <p>urn:lsid:zoobank.org:act: 94BB49B0-A076-485B- BAA4-9F1E60CB5B77</p>
            <p>Type material</p>
            <p> Indonesia: Mature non-ovigerous holotype female (3.48 mm), mature allotype male (1.8 mm) (RMNH. Crus.I.7749), ex abdominal marsupium of female  Lithoscaptus tri (Fize &amp; Serène, 1956b) (4.0 mm long × 2.5 mm wide) (RMNH.Crus.D.54029), ex  Echinopora lamellosa (Esper, 1795) (  Merulinidae ), Sulamadaha I Reef, Ternate, 00°52′03″N, 127°19′33″E, 6 November 2009, coll. S. E. T. van der Meij. </p>
            <p> Japan: Immature paratype female (2.0 mm) (RMNH. Crus.I.7750), ex abdomen of male  Xynomaia sp. (4.3 mm long × 3.8 mm wide) (RMNH.Crus.D.57234), ex  Pectinia sp. (  Merulinidae ), Blue Tombs, Henza Island, Okinawa, 26°21′46″N, 127°59′50″E, 29 April 2016, coll. S. E. T. van der Meij. </p>
            <p>Description</p>
            <p>Female (Fig. 3): Body length 3.4 mm, maximal width 2.6 mm, head length 0.71 mm, head width 0.78 mm. Pereon nearly straight, head slightly deflected sinistrally (Fig. 3A, B). All body regions and pereomeres distinctly segmented.</p>
            <p>Head subrectangular, approximately as broad as long, overlapping almost entire first pereomere medially, inflated dorsoventrally with very thin frontal lamina (Fig. 3A, C). Eyes minute, lateral. Antennule of four articles; antenna of two articles (Fig. 3D). First oostegite (Fig. 3E) anterior lobe globular, posterior lobe slightly smaller than anterior lobe, ovate with rounded margins, distal projection lacking but with medially directed smooth lobe, internal ridge thick, smooth. Oostegites completely enclosing marsupium. Maxilliped (Fig. 3F) with short subacute fleshy spur; palp lacking, anterior segment rounded, posterior segment triangular. Barbula of three short, rounded fleshy lobes (Fig. 3G).</p>
            <p>Pereon composed of seven pereomeres, broadest across pereomere V, gradually tapering anteriorly and posteriorly; pereomere I posterior margin under head, II weakly concave, III–VII medially concave. Coxal plates very small on anterior pereomers, pronounced, separated, tergal projections on pereomeres one to seven, those on one to four irregular in shape, those on five to seven smooth and rounded at ends (similar to appearance of lateral plates). Anteriormost pair of pereopods (Fig. 3H) about half as large as other pairs (Fig. 3I).</p>
            <p>Five pleomeres plus pleotelson, lateral plates well developed, tubular, rounded distally and recurved (Fig. 3J). Pleomeres I–IV with biramous, widely spaced, tubular pleopods as ventrally directed extensions of lateral plates; uniramous pleopods on pleomere V, bulbous uniramous uropods.</p>
            <p>Male (Fig. 5): Length 1.8 mm, maximal width 0.82 mm, head length 0.23 mm, head width 0.47 mm, pleon length 0.46 mm.</p>
            <p>Head ovate, widest medially, distinct from first segment of pereon (Fig. 5A, B); eyes present. Antennule of four articles (Fig. 5C), distally setose, scarcely extending beyond margin of head; antenna of two articles, terminal article stout, distally setose (Fig. 5C).</p>
            <p>Pereomere V broadest, others gradually tapering anteriorly and posteriorly. Pereomeres I–V straight, VI and VII posteriorly concave, distal and ventral margins of all pereomeres rounded. All pereopods (Fig. 5B, D, E) subequal; carpus and merus nearly fused, all other articles distinctly separated.</p>
            <p>Pleon with four segments plus pleotelson. All pleomeres directed posterolaterally (in dorsal view; Fig. 5A), with distolateral margins rounded; extended ventrally (in lateral view). Midventral tubercles, pleopods and uropods lacking (Fig. 5B).</p>
            <p>Etymology</p>
            <p> From a combination of the Greek απροσδόκητος (aprosdókiti), meaning unexpected, and εύρημα(evrima),meaning finding, because of the unexpected finding of a sister taxon to  Rhopalione on a species of gall crab (  Cryptochiridae ). </p>
            <p>Remarks</p>
            <p>See under Remarks for the genus. The juvenile specimen (RMNH.Crus.I.7750; Fig. 4) already shows the overall body shape and pleopod formation seen in the adult female, and little additional modification to the body seems to occur between the immature and mature female forms. The occurrence of the juvenile female on a male host is perplexing, because the holotype female occupies the entirety of the female’s marsupium, hence the paratype female appears to have made a poor host choice because the male abdomen will not develop into an appropriate structure to hold the adult female. There is no evidence in the literature that bopyrids can markedly feminize their hosts à la rhizocephalans.</p>
            <p>Known hosts</p>
            <p> Lithoscaptus tri (Fize &amp; Serène, 1956b) ,  Xynomaia sp.</p>
            <p>Distribution</p>
            <p>Known from Indonesia and Japan.</p>
            <p>CRYPTONISCOIDEA KOSSMANN, 1880</p>
            <p> CRYPTONISCIDAE KOSSMANN, 1880</p>
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	https://treatment.plazi.org/id/CA3E87E5FE47551F7DDEFF66FAACF9CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE47551B7EFFF912FD27FCA7.text	CA3E87E5FE47551B7EFFF912FD27FCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danalia cervix Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> DANALIA CERVIX SP. NOV.</p>
            <p>(FIGS 6A, 11F)</p>
            <p>urn:lsid:zoobank.org:act: C5720E4D-F79B-4AAE- 9B2C-20AEF77AC607</p>
            <p>‘cryptoniscine isopod parasites’ Kropp &amp; Manning, 1987: 10, 12 [Belize, examined herein].</p>
            <p>Type material</p>
            <p> Curaçao: Immature holotype female (9.0 mm), two paratype cryptoniscus larvae (1.05 mm each) (RMNH. Crus. I. 7751), ex abdomen of female  Opecarcinus hypostegus (Shaw &amp; Hopkins, 1977) (5.6 mm long × 4.5 mm wide) (RMNH.D.56099), ex  Agaricia lamarcki H. Milne Edwards &amp; Haime, 1851 (  Agariciidae ), Playa Lagun, 12°19′ 69″N, 69°09′00″W, 35 m depth, 29 October 2013, coll. S. E. T. van der Meij. </p>
            <p> Belize: Two immature paratype females (0.6, 0.7 mm) (USNM 1437664), ex abdomen of female  Opecarcinus hypostegus (Shaw &amp; Hopkins, 1977) (2.0 mm long × 1.5 mm wide) (USNM 231677), ex  Agaricia sp. (  Agariciidae ), Carrie Bow Cay, Dangrega District, Belize, 16°48′12″N, 88°04′30″W, 15.2 m depth, 20 April 1981, coll. G. Hendler. </p>
            <p>Description</p>
            <p>Female (Fig. 6A): 9 mm long; mature female stout, body strongly recurved, proximal end truncate, distal end rounded, surface smooth but with indentations showing indications of four or five segments dorsally, internal segmentation not visible; anteroventral shield small. Trunk thick and strongly extended, inserted into body at an angle of ~45°, extending out from body slightly anterior to midventral indentation, attachment lobes missing (presumed thin and fragile). Directly parasitizing cryptochirid host.</p>
            <p>Etymology</p>
            <p> The species name is from the Latin  cervix , meaning nozzle, and refers to the large attachment process trunk that is partly fused with the body. It is used as a noun in apposition. </p>
            <p>Remarks</p>
            <p> The thick attachment trunk, partly fused with the body and not terminally positioned, is unique to this species. It is therefore difficult to know to which other species of  Danalia this new species is most closely related. This is the only species of  Danalia known from the western Atlantic;  Danalia frassei Nierstrasz &amp; Brender à Brandis, 1925, also from Curaçao, was shown to belong to  Cabirops by Boyko (2013). </p>
            <p>Known host</p>
            <p> Opecarcinus hypostegus (Shaw &amp; Hopkins, 1977) . </p>
            <p>Distribution</p>
            <p>  Known from Curaçao (type locality)  and Belize. </p>
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	https://treatment.plazi.org/id/CA3E87E5FE47551B7EFFF912FD27FCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE43551B7C09FCFAFA3EFB60.text	CA3E87E5FE43551B7C09FCFAFA3EFB60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danalia falsicrura Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> DANALIA FALSICRURA SP. NOV.</p>
            <p>(FIGS 6B–E, 11C)</p>
            <p>urn:lsid:zoobank.org:act: CEA5CFEA-C2FE-41FA- B5AB-5A2C14EB765F</p>
            <p> ‘three small sac-like parasites’ Utinomi, 1944: 696–697, fig. 4 {ex  Pseudocryptochirus viridis Hiro, 1938 , ex  Turbinaria contorta Bernard, 1896 [=  T. frondens (Dana, 1846) ] (  Dendrophyllidae ), Tanabe Bay, Japan}. </p>
            <p> ‘  Danalia sp. Utinomi, 1944 ’ Boyko, 2015: 76. </p>
            <p>Type material</p>
            <p> Guam: Immature holotype female (6.6 mm) (USNM 1461192), ex abdomen of female  Opecarcinus crescentus (Edmondson, 1925) (2.1 mm long × 1.8 mm wide) (USNM 234257), ex  Pavona duerdeni Vaughan, 1907 (  Agariciidae ), main patch reef, double reef, 24 February 1984, coll. R. K. Kropp. </p>
            <p> Indonesia: Immature paratype female (4.8 mm) (RMNH.Crus.I.7752), ex abdomen of female  Fungicola utinomi (Fize &amp; Serène, 1956a) (4.3 mm long × 2.5 mm wide) (RMNH.Crus.D.54222), ex  Lithophyllon repanda (Dana, 1846) (  Fungiidae ), North Pulau Dua, Lembeh Strait, 01°23′28″N, 125°12′58″E, 13 February 2012, coll. S. E. T. van der Meij. </p>
            <p>Description</p>
            <p>Female (Fig. 6B–E): 6.6 mm long; mature female cylindrical, body strongly recurved, proximal and distal ends rounded, surface smooth without indications of segments, internal segmentation not visible; anteroventral shield large. Trunk thick but very short, variably inserted into body either subterminally (Fig. 6B) or terminally (Fig. 6D, E), attachment lobes variable (probably owing to development), either from two short, rounded lobes terminally on stalk (Fig. 6E) or four thin, flat lobes extending from central stalk (Fig. 6C). Directly parasitizing cryptochirid host.</p>
            <p>Etymology</p>
            <p>The species name is a combination of falsum and crura, meaning ‘false legs’, which refers to the stumplike processes emanating from the centre of the ventral surface that resemble the prolegs of caterpillars (Lepidoptera). It is used as a noun in apposition.</p>
            <p>Remarks</p>
            <p> Utinomi (1944) found two female  Pseudocryptochirus viridis infested with one or two parasites on each of their ventral abdominal surfaces, and his description and drawings leave little doubt that his specimens from Japan are conspecific with  D. falsicrura sp. nov. The shape of the body is nearly identical, and the presence of four small foot-like ventral projections is unique to this species. Utinomi’s (1944) females all had the attachment process centrally located on the proximal end of each parasite, which is also seen in the specimen from Indonesia. The Guam female, however, has the attachment process set back from the proximal end and positioned more ventrally than in the other specimens. Utinomi’s (1944) fig. 4B appears to show indistinct surface lobes indicating the presence of five segments to the body, but this is not seen in his fig. 1A or in the present specimens. </p>
            <p>Known hosts</p>
            <p> Fungicola utinomi (Fize &amp; Serène, 1956a) ,  Opecarcinus crescentus (Edmondson, 1925) ,  Pseudocryptochirus viridis Hiro, 1938 . </p>
            <p>Distribution</p>
            <p>  Known from Guam (type locality)  , Indonesia and Japan. </p>
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	https://treatment.plazi.org/id/CA3E87E5FE43551B7C09FCFAFA3EFB60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE43551A7EF2FAABFD87FA8F.text	CA3E87E5FE43551A7EF2FAABFD87FA8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danalia galea Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> DANALIA GALEA SP. NOV.</p>
            <p>(FIGS 6F, G, 11A, B)</p>
            <p>u r n: l s i d: z o o b a n k. o r g: a c t: 2 2 A 4 6 5 B F - D 0 C 5 - 4E2A-A8A4-F88C175439D4</p>
            <p>Type material</p>
            <p> Indonesia: Mature non-ovigerous holotype female (6.3 mm) (RMNH.Crus.I.7753), ex ventral abdomen of female  Lithoscaptus paradoxus A. Milne-Edwards, 1862 (6.0 mm long × 3.3 mm wide) (RMNH.Crus.D.53188), ex  Platygyra lamellina (Ehrenberg, 1834) (  Merulinidae ), Timur I, Bunaken Island, 01°36′38″N, 124°46′59″E, 11 December 2008, coll. S. E. T. van der Meij. </p>
            <p> Papua New Guinea: Mature non-ovigerous paratype female (1.7 mm long × 1.0 mm wide) (RMNH. Crus.I.7705), between fourth and fifth pereopod, left side, inserted into fifth pereopod base of female  Dacryomaia sp. (3.5 mm long × 1.9 mm wide) (RMNH.Crus.D.57067), ex  Psammocora cf. digitata H. Milne Edwards &amp; Haime, 1851 (  Psammocoridae ), Steffen Channel, Kavieng, 02°43′22″S, 150°34′28″E, 18 August 2014, coll. F. Benzoni. </p>
            <p>Description</p>
            <p>Female (Fig. 6F, G): 6.3 mm long; mature females very broad in dorsal view, body robust, width nearly two-thirds of length, intersection of proximal and distal portions of body giving a laterally indented appearance in dorsal view, body with slight ventral indentation, proximal and distal ends rounded, surface without lobes but with irregular indentations dorsally, some of surface with areas of cuticular lines, internal segmentation not visible; anteroventral shield very small. Trunk short and barely extending from body, inserted into body subterminally at proximal end, attachment lobes missing (presumed thin and fragile). Directly parasitizing cryptochirid host, either on the ventral surface of the abdomen or on posterior pereopods.</p>
            <p>Etymology</p>
            <p> The species name is from the Latin  galea , meaning helmet, for the overall appearance of the female body. It is used as a noun in apposition. </p>
            <p>Remarks</p>
            <p> The distinctive helmet-like shape is unique to this species and the attachment stalk perhaps the shortest known in species of the genus. The attachment position of the paratypes on posterior pereopods of the host is unusual, but other species in the genus (e.g.  Danalia hapalocarcini , see next subsection) parasitize locations other than the ventral surface of the abdomen. However, given that this species was found in two locations on hosts, it is unclear whether one is the preferred location or if both are equally suitable to the parasite. </p>
            <p>Known hosts</p>
            <p> Dacryomaia sp. ,  L. paradoxus A. Milne-Edwards, 1862 . </p>
            <p>Distribution</p>
            <p>Indonesia and Papua New Guinea.</p>
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	https://treatment.plazi.org/id/CA3E87E5FE43551A7EF2FAABFD87FA8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE42551A7DDCFACFFC39F98A.text	CA3E87E5FE42551A7DDCFACFFC39F98A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danalia hapalocarcini Fize 1955	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> DANALIA HAPALOCARCINI FIZE, 1955</p>
            <p>(FIGS 6H–J, 11D)</p>
            <p> Danalia hapalocarcini Fize, 1955: 2444–2447 , figs. 1–4. – Fize, 1956: 22–28, figs. 15–18. – Altes, 1982: 28–29. – Grygier, 1993: 189–190. – Boyko, 2015: 76, fig. 2H. </p>
            <p>Material examined</p>
            <p> Malaysia: Mature non-ovigerous female (3.0 mm), mature female (2.9 mm) (RMNH.Crus.I.7754), on carapace of female  Hapalocarcinus marsupialis Stimpson, 1859 (2.9 mm long × 2.8 mm wide) (RMNH. Crus.D.53703), ex  Seriatopora cf. caliendrum Ehrenberg, 1834 (  Pocilloporidae ), Horn Reef, Semporna, 04°14′32″N, 118°26′25″E, 1 December 2010, coll. S. E. T. van der Meij. </p>
            <p> Indonesia: Cryptoniscus larva (1.33 mm) (RMNH. Crus.I.7755), on carapace of female  H. marsupialis Stimpson, 1859 (2.6 mm long × 2.6 mm wide) (RMNH.Crus.D.53189), ex  Seriatopora cf. caliendrum Ehrenberg, 1834 (  Pocilloporidae ), Tanjung Sidangolo, Halmahera, 00°53′40″N, 127°29′28″E, 5 November 2009, coll. S. E. T. van der Meij. </p>
            <p> New Caledonia: Immature female (0.9 mm) (MNHN-IU-2017-10), on lower left carapace of immature female  H. marsupialis Stimpson, 1859 (2.5 mm long × 3.0 mm wide) (MNHN-IU-2009-5733), ex  Stylophora sp. (  Pocilloporidae ), New Caledonia, Ilot Maître, Lagoon, 5 m depth, 22°19′48″S, 166°25′06″E, 24 April 1995, coll. P. Castro. </p>
            <p>Redescription</p>
            <p>Female (Fig. 6H–J): 3.0 mm long; mature female very broad in dorsal view, body weakly cuticularized, sac like; body moderately recurved, proximal and distal ends rounded; surface without lobes but with irregular folds dorsally, internal segmentation not visible; anteroventral shield small. Trunk short, inserted into body terminally at proximal end; attachment lobes extremely long and slender, covering large portion of ventral surface of host carapace. Directly parasitizing cryptochirid host on ventral surface of carapace.</p>
            <p>Remarks</p>
            <p> The new records of this species are the first since the species was described based on specimens from Vietnam. The attachment processes are the most elaborately extended of any species in the genus and reach nearly across the entire ventral surface of the host’s carapace, presumably to maximize nutrient absorption. This was the only species of  Danalia known to parasitize through the dorsal surface of the carapace of its hosts before the discovery of  Danalia vesica sp. nov. (see next subsection). It can be distinguished from  D. vesica sp. nov. by the shape of body (sac like but recurved in  D. hapalocarcini vs. sac like but disc shaped in  D. vesica sp. nov. ) and in the length of the attachment processes (long in  D. hapalocarcini vs. short in  D. vesica sp. nov. ). </p>
            <p>Known host</p>
            <p> Hapalocarcinus marsupialis Stimpson, 1859 . </p>
            <p>Distribution</p>
            <p> Vietnam (type locality) , Malaysia, Indonesia and New Caledonia. </p>
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	https://treatment.plazi.org/id/CA3E87E5FE42551A7DDCFACFFC39F98A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE4255187EECF9DFFE5DFB3A.text	CA3E87E5FE4255187EECF9DFFE5DFB3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danalia vesica Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> DANALIA VESICA SP. NOV.</p>
            <p>(FIG. 7A, B)</p>
            <p>urn:lsid:zoobank.org:act: 728A215D-468D-433C- A36E-9380E436C9A8</p>
            <p>Type material</p>
            <p> Mature non-ovigerous holotype female (3.3 mm long × 2.5 mm wide) (RMNH.Crus. I.1742), on carapace of female  H.marsupialis Stimpson, 1859 (3.0 mm long × 3.0 mm wide), (RMNH.Crus.D.57235), ex unknown host coral (probably  Pocilloporidae , because it is the only known host family for this gall crab species), Mamba Bawi, east coast of Zanzibar, 19 September 1970, coll. A. J. Bruce. </p>
            <p>Description</p>
            <p>Female (Fig. 7A, B): 3.3 mm long; mature female nearly circular in dorsal view, body weakly cuticularized, sac like; body not recurved; convex dorsally and concave ventrally, conforming to the dorsal surface of the host carapace; surface without lobes but with irregular folds dorsally; internal segmentation not visible; anteroventral shield small. Trunk short, inserted into body near centre of ventral surface; attachment lobes, apparently only two, short and slender, covering small portion of ventral surface of host carapace. Directly parasitizing cryptochirid host on ventral surface of carapace.</p>
            <p>Etymology</p>
            <p>The species name is derived from the Latin for blad- der or balloon, in reference to the female resembling a deflated balloon.</p>
            <p>Remarks</p>
            <p> This species, known presently only from the female holotype, can be distinguished from its closest relative,  D. hapalocarcini , by the characters given in the Remarks for that species. The presence of only two attachment processes instead of the usual four is unusual but, because there is only the single specimen known, needs confirmation as a species-specific character through collection and examination of additional specimens. </p>
            <p>Known host</p>
            <p> Hapalocarcinus marsupialis Stimpson, 1859 . Distribution </p>
            <p>Known only from Zanzibar.</p>
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	https://treatment.plazi.org/id/CA3E87E5FE4255187EECF9DFFE5DFB3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE5F55057ECEF9E5FEF3F8CA.text	CA3E87E5FE5F55057ECEF9E5FEF3F8CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cabirnalia Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> CABIRNALIA GEN. NOV.</p>
            <p>urn:lsid:zoobank.org:act: 61D0157E-D349-4BC4- B24C-9F4C30AD41CF</p>
            <p>Diagnosis</p>
            <p>Mature female: strongly recurved, circular in outline. Strongly convex dorsal margin without lobes but with cuticular indentations indicating segmentation; ventral sides meeting medially, with several small, plate-like areas demarcated at midpoint. Attachment trunk short, with single attachment lobe, medially divided and expanded into two broad sections. Parasitic on abdomen or pereopods of crab hosts.</p>
            <p>Type species</p>
            <p> Cabirnalia nausicaa sp. nov.</p>
            <p>Etymology</p>
            <p> The generic name is a combination of  Cabirops and  Danalia , because the genus has characters found in both genera. The gender is feminine. </p>
            <p>Remarks</p>
            <p> This genus is most similar to  Danalia in the overall shape of the mature females and the presence of an attachment trunk. However, the medioventral area of the holotype of the type species shows several small plate-like structures, very similar to those seen on species of  Cabirops and not found in any species of  Danalia except  D. caulleryi Nierstrasz &amp; Brender à Brandis, 1923 (see Boyko, 2015). As  Cabirops species do not possess any attachment structures and  Danalia species do not possess medioventral plates, this new genus is necessary to accommodate the new species described below.  Danalia caulleryi is also placed in  Cabirnalia gen. nov. , as  Cabirnalia caulleryi comb. nov. , because the unique holotype has mediovental plates and shows evidence of a now-lost attachment trunk (see Boyko, 2015). Even though the structure of the attachment trunk of  C. caulleryi comb. nov. is not known, it is not likely to be conspecific with  C. nausicaa gen. nov., sp. nov. because (1) the cuticular ridges of  C. caulleryi comb. nov. are much more robust than those found on the type species and (2) the host of  C. caulleryi comb. nov. is a galatheid, not a gall crab.  Cabirnalia gen. nov. is therefore unique among cryptoiscoids in containing species found parasitizing both brachyuran and anomuran hosts. </p>
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	https://treatment.plazi.org/id/CA3E87E5FE5F55057ECEF9E5FEF3F8CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE5D55037E9AFADBFD0CFA5E.text	CA3E87E5FE5D55037E9AFADBFD0CFA5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cabirnalia nausicaa Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> CABIRNALIA NAUSICAA SP. NOV.</p>
            <p>(FIGS 8, 12C–F)</p>
            <p>urn:lsid:zoobank.org:act: 27FFC15A-5D05-41F4- A6B3-B26FCC86DBF0</p>
            <p>Type material</p>
            <p> New Caledonia: Mature holotype female (14.1 mm) (MNHN-IU-2017-11), two immature paratype females (1.6 mm each) (MNHN-IU-2017-12), ex abdomen of female  H. marsupialis Stimpson, 1859 (4.0 mm long × 4.0 mm wide) (MNHN-IU-2009–5735), ex  Pocillopora sp. (  Pocilloporidae ), New Caledonia, lagoon, Passe de Boulari, 10–20 m depth, 22°29′54″S, 166°26′33″E, 28 April 1995, coll. P. Castro. </p>
            <p> Malaysia: Three mature paratype females (5.0, 5.5 and 6.3 mm), paratype cryptoniscus larva (1.3 mm) (RMNH.Crus.I.7757), ex posterior pereopods (females) and abdomen (larva) of female  L. helleri (Fize &amp; Serène, 1957) (4.2 mm long × 2.7 mm wide) (RMNH. Crus.D.53737), ex  Favites cf. halicora (Ehrenberg, 1834) (  Merulinidae ), Batura Reef, Semporna, 04°30′49″N, 118°48′31″E, 7 December 2010, coll. S. E. T. van der Meij. </p>
            <p> Japan: Mature female (4.8 mm long × 3.5 mm wide) (RMNH.Crus.I.7758), inserted ventromedially between second and third sternites of female  Lithoscaptus sp. 2 (B) (RMNH.Crus.D.57236), ex  P. versipora (  Scleractinia incertae sedis), Mizugama, Okinawa, 26°21′35″N, 127°44′18″E, 15 April 2016, coll. S. E. T. van der Meij. </p>
            <p>Other material</p>
            <p> Thailand: Immature female (2.2 mm), ex third pereopod of male  L. paradoxus A. Milne-Edwards, 1862 , 1862 (2.5 mm long × 1.5 mm wide) (ZRC 2016.0435), ex unknown coral, coral reef, Phuket, December 1998, coll. P. K. L. Ng; immature female (1.6 mm), ex abdomen of female  L. paradoxus A. Milne-Edwards, 1862 (4.8 mm long × 3.3 mm wide) (ZRC 2016.0443), ex unknown coral, Ao Thang Khen, Phuket, May 2000, coll. P. K. L. Ng &amp; K. L. Yeo. </p>
            <p>Description</p>
            <p>Mature female (Fig. 8): 4.8–14.1 mm long; strongly recurved, circular in outline (viewed laterally); body robust, strongly convex dorsal margin without lobes but with cuticular indentations indicating segmentation; body with slight ventral indentation; ventral sides meeting medially, with several small, plate-like areas demarcated at midpoint. Attachment trunk short, with single attachment lobe, medially divided and expanded into two broad sections. Directly parasitizing cryptochirid hosts on ventral surface of abdomen, on posterior pereopods, or on sternites.</p>
            <p>Etymology</p>
            <p> The species is named after the main character in the film 風の谷のナウシカ (  Nausicaä of the Valley of the Wind) by Hayao Miyazaki (Toei 1984), because the attachment processes greatly resembles the gas mask worn by Nausicaä when entering the Toxic Jungle. </p>
            <p>Remarks</p>
            <p> See under Remarks for the genus. Note that this parasite can grow to be larger than its host (e.g. RMNH.Crus.I.7758, from Okinawa), and this would seem to limit the mobility of the host; however, gall crabs, particularly females, probably do not move outside their dwellings.The non-type material is provisionally assigned to this species. The smaller specimen from Thailand is a non-descript sac, with an attachment stalk somewhat resembling that of  D. cervix sp. nov. from the Atlantic, but the larger specimen, which is still immature, was partly moulted and shows a smooth outer cuticle covering a segmented inner cuticle. This pattern of metamorphosis from smooth to segmented sac is also seen in species of  Cabirops (see Reverberi &amp; Catalano, 1963), but  Cabirops species lack attachment stalks. </p>
            <p>Known hosts</p>
            <p> Hapalocarcinus marsupialis Stimpson, 1859 (type host),  L. helleri (Fize &amp; Serène, 1957) ,  Lithoscaptus sp. 2 (B); probably also on  L. paradoxus A. Milne-Edwards, 1862 . </p>
            <p>Distribution</p>
            <p>  Known from New Caledonia (type locality)  , Malaysia and Japan; probably also from Thailand. </p>
            <p>CIRRIPEDIA BURMEISTER, 1834</p>
            <p>RHIZOCEPHALA MÜLLER, 1862</p>
            <p> SACCULINIDAE LILLJEBORG, 1861</p>
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	https://treatment.plazi.org/id/CA3E87E5FE5D55037E9AFADBFD0CFA5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
CA3E87E5FE5B55027C20F9F4FE49FC6A.text	CA3E87E5FE5B55027C20F9F4FE49FC6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sacculina quadrialata Boyko & Van Der Meij 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> SACCULINA QUADRIALATA SP. NOV.</p>
            <p>(FIGS 9, 12A, B)</p>
            <p>urn:lsid:zoobank.org:act: A646BB1F-4BA5-46EE-96C1-AF7A4BDE1FCB</p>
            <p>Type material</p>
            <p> Indonesia: mature holotype externa (1.6 mm long × 1.2 mm wide) (RMNH.Crus.C.10250), ex abdomen of feminized male  Opecarcinus sp. 1 (F) (3.0 mm long × 2.3 mm wide) (RMNH.Crus.D.53990), ex  L. mycetoseroides Wells, 1954 (  Agariciidae ), South Pilongga, Tidore, 00°42′44″N, 127°28′47″E, 12 November 2009, coll. S. E. T. van der Meij; two mature paratype externae (2.0 mm long × 3.4 mm wide, 1.4 mm long × 2.0 mm wide) (RMNH.Crus.C.10251), ex abdomen of female  Fizesereneia heimi (Fize &amp; Serène, 1956a) (5.3mmlong×4.5mmwide)(RMNH.Crus.D.54018), ex  Lobophyllia cf. radians (H. Milne Edwards &amp; Haime, 1849) (Lobophyliidae), Tanjung Sidangolo, Halmahera, 00°53′40″N, 127°29′28″E, 5 November 2009, coll. S. E.T.van der Meij; mature paratype externa (1.4 mm long × 1.9 mm wide) (RMNH.Crus.C.10252), ex abdomen of feminized male  Lithoscaptus cf. bani (Fize &amp; Serène, 1957) (3.1 mm long × 2.0 mm wide) (RMNH.Crus.D.53993), ex  Goniastrea edwardsi Chevalier, 1971 (  Merulinidae ), West Mansuar, Raja Ampat, 00°30′41″S, 130°33′35″E, 9 December 2007, coll. S. E. T. van der Meij; immature paratype externa (0.7 mm long × 1.0 mm wide) (RMNH.Crus.C.10253), ex abdomen of female  Opecarcinus cf. sierra Kropp, 1989 (1.9 mm long × 1.4 mm wide) (RMNH.Crus.D.54208), ex  Pavona cf. varians Verrill, 1864 (  Agariciidae ), North Tanjung Pandean, Lembeh Strait, 01°24′21″N, 125°10′04″E, 14 February 2012, coll. S. E. T. van der Meij; mature paratype externa (0.8 mm long × 1.1 mm wide) (RMNH.Crus.C.10254), ex abdomen of female  X. sheni (Fize &amp; Serène, 1956b) (1.9 mm long × 1.3 mm wide) (RMNH.Crus.D.54166), ex  M. elephantotus (Pallas, 1766) (  Merulinidae ), Tanjung Labuhankompeni, Lembeh Strait, 01°25′55″N, 125°11′10″E, 4 February 2012, coll. S. E. T. van der Meij. </p>
            <p>Description</p>
            <p>Externa of immature specimens with rounded subequal lateral lobes; mature (ovigerous) externae with lateral lobes produced anteriorly and posteriorly into wing-like extensions, sometimes subequal but often of differing degrees of extension. Mantle opening mediodistally located opposite stalk. Mantle opening ranging from smooth to crenulated, raised off the surface of the externa. Stalk short but externa raised off the surface of the host abdomen. Cuticle with varying degrees of wrinkling, particularly prominent in larger specimens.</p>
            <p>Etymology</p>
            <p>The species name alludes to the four wing-like lobes (quadri + alata) of the mature externa.</p>
            <p>Remarks Vervoort(inMartin&amp;Davis,2001)statedthat‘Sacculinidae must be ascribed to Lilljeborg (1860)’. However, nowhere in Martin &amp; Davis (2001) was the actual bibliographic reference to Lilljeborg, ‘1860’ given. After careful analysis of all the publications of Lilljeborg dealing with rhizocephalans, we must conclude that the date was given incorrectly by Vervoort, and is, in fact, Lilljeborg (1861).</p>
            <p> Although it would be desirable to perform histology on at least some of these specimens, the hosts were not preserved with rhizocephalan histology in mind, and it is doubtful that any internal characters can be gleaned from such treatment. Additionally, the utility of many of Boschma’s internal characters in terms of showing evolutionary relationships has yet to be demonstrated; some of these characters may be homoplasies, but this remains to be tested. Rybakov &amp; Høeg (2002) showed the utility of studying the retinacula found on the mantle of externae, but there are no such structures in this new species nor are they known in many of the species of  Sacculina . However, this new species can be described and named because the presence of the four laterally extended wing-like lobes on the mature externae is unlike the externae shapes seen in other species of  Sacculina . The most similar species to this new species is  S. rugosa Van Kampen &amp; Boschma, 1925 , but the shapes of the externae are only vaguely similar. Additionally, there are no previous records of any rhizocephalans from gall crab hosts. </p>
            <p>Known hosts</p>
            <p> Opecarcinus sp. 1 (F) (type host),  F. heimi (Fize &amp; Serène, 1956a) ,  Lithoscaptus cf. bani (Fize &amp; Serène, 1957) ,  Opecarcinus cf. sierra Kropp, 1989 ,  X. sheni (Fize &amp; Serène, 1956b) . </p>
            <p>Distribution</p>
            <p>Known only from Indonesia.</p>
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	https://treatment.plazi.org/id/CA3E87E5FE5B55027C20F9F4FE49FC6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boyko, Christopher B.;Van Der Meij, Sancia E. T.	Boyko, Christopher B., Van Der Meij, Sancia E. T. (2018): A trifecta of Swiftian symbioses: stony corals, gall crabs and their parasites (Scleractinia; Brachyura: Cryptochiridae; Isopoda: Epicaridea and Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 184: 304-329
