taxonID	type	description	language	source
CA729441C20CFF9BFEAFFBEAFDB1FB60.taxon	type_taxon	TYPE SPECIES. — Cerithium adansonii Bruguière, 1792, by subsequent designation (ICZN opinion 1109, 1978).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	description	(Figs 9 D-F; 11) urn: lsid: zoobank. org: act: 8 A 430 C 74 - 7 DDA- 4 FEE- 922 E- 7 C 9 F 2 FD 0 D 3 D 8	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98401 (Letourneux coll.); H 20.1 mm (Fig. 11 A-C). Paratypes. France • 1 spm; same data locality as the holotype; Plio-Pleistocene; MNHN. F. A 98402 (Letourneux coll.); H 19.1 mm (Fig. 11 D-F) • 1 spm; Vaihuoa, near archeological platform (sample N 106); Late Holocene; MNHN. F. A 98404 (Salvat coll.); H 15 mm (Fig. 11 G, H) • idem; MNHN. F. A 98405 (Salvat coll.); H 15.5 mm (Fig. 11 I, J), • 1 spm; same locality and age (sample N 108); MNHN. F. A 98403 (Salvat coll.); H 20 mm (Fig. 11 K, L) • 1 spm; idem (sample TP 1 A); MNHN. F. A 98406 (Salvat coll.): H 20.8 mm (Fig. 11 M, N) • 1 spm; idem (samples TP 1 A); MNHN. F. A 98407 (Salvat coll.); H 16 mm (Fig. 11 O, P) • 1 spm; idem (samples TP 1 B); MNHN. F. A 98408 (Salvat coll.); H 23 mm (Fig. 11 Q, R) • 1 spm; idem (samples TP 1 B); MNHN. F. A 98409 (Salvat coll.); H 19.9 mm (Fig. 11 S, T) • 1 spm; idem (samples TP 1 B); MNHN. F. A 98410 (Salvat coll.); H 14.6 mm (Fig. 11 U, V). ADDITIONAL MATERIAL. — France • 148 spm; same as the holotype; Plio-Pleistocene; MNHN. F. A 98414 (Letourneux coll.) • 1 spm; Aturona (sample N 58); Mid Holocenee; MNHN. F. A 98417 (Salvat coll.) • 2 spm; idem (sample N 57); MNHN. F. A 98418 (Salvat coll.) • 4 spm; idem (sample N 63); MNHN. F. A 98420 (Salvat coll.) • 1 spm; Vaihuoa, near archeological platform (sample N 90); Late Holocene; MNHN. F. A 98419 (Salvat coll.) • 4 spm; idem (sample N 91); MNHN. F. A 98425 (Salvat coll.) • 16 spm; idem (sample N 106); MNHN. F. A 98424 (Salvat coll.) • 8 spm; idem (sample TP 1 A); MNHN. F. A 98427 (Salvat coll.) • 11 spm; idem (sample TP 1 B); MNHN. F. A 98428 (Salvat coll.) • 1 spm; idem (sample N 105); MNHN. F. A 98411 (Salvat coll.); Fig. 9 D • 1 spm; idem (sample 105); MNHN. F. A 98412 (Salvat coll.); Fig. 9 E • 1 spm; idem (sample 105); MNHN. F. A 98413 (Salvat coll.); Fig. 9 F • 11 spm; idem (sample N 105); MNHN. F. A 98423 (Salvat coll.) • 8 spm; idem (sample N 93); MNHN. F. A 98415 (Salvat coll.) • 1 spm; idem (sample N 108); MNHN. F. A 98421 (Salvat coll.) • 1 spm, idem (sample N 104); MNHN. F. A 98422 (Salvat coll.) • 3 spm; idem (sample N 94); MNHN. F. A 98416 (Salvat coll.) • 1 spm; idem (sample N 103); MNHN. F. A 98426 (Salvat coll.).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	etymology	ETYMOLOGY. — From the type locality, Niau Atoll.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage), seeFigs 7; 8. TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	distribution	DISTRIBUTION. — Only know from the type locality, from Early Pliocene to Late Holocene.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	description	DESCRIPTION Shell medium in size, subangulate, slightly cyrtoconoid. Protoconch unknown. Apical angle 34 °. Height of last whorl: 43.5 % of shell lenght. Sculpture on spire consisting of weak to moderately strong, slightly curved axial ribs that are swollen near the posterior suture. On last whorl, one varical node in opposite side of the outer lip. Early whorls with three primary beaded cords (P 1 - P 3), P 2 being more prominent. Secondary cords s 1 and s 3 occasionnal. Last whorl with three slightly beaded primary cords (P 1 - P 3) more prominent than the anterior cords (P 4 - P 7). Secondary cords s 1, s 2, s 3 and s 4 occasionnal. Under UV light, beaded cords fairer than the background. Height of aperture: 32.5 % of shell lenght. Outer lip slightly thickened. Inner lip callused, weakly detached. Anal notch moderately deep. Parietal callus weak.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	discussion	COMPARISONS The sculpture of Cerithium niauense Merle, n. sp. is variable. In Plio-Pleistocene material, axial ribs can be more or less prominent and rows of cords on P 1 to P 3 can be more or less beaded. This variation allows to distinguish two extreme morphotypes, one characterized by well formed axial ribs (e. g., Fig. 11 A-C) and one characterized by a beaded sculpture with low ribs (e. g., Fig. 11 D-F). Transitional forms link these two morphotypes, showing that they belong to a same species. Holocene material displays a similar variation with two same extreme morphototypes (e. g., Fig. 11 I, J; 11 K, L) and no diagnostic characters allowing to distinguish a different species. All shells of C. niauense Merle, n. sp. are regularly slightly cyrtoconoid, their axial ribs are slightly curved and P 2, the most prominent cord, is well distinct from P 3. Plio-Pleistocene shells are creamy and sediment sometimes included in these shells is whitish, whereas Holocene shells are withish pale grey and display a dark grey sediment within their shell when present. Cerithium tenellum G. B. Sowerby II, 1855, originally described from Philippines, is reminiscent to C. niauense Merle, n. sp., but differs by having a conical shape, straigh axial ribs and P 3 almost fused with P 2, and no varical nodes. Cerithium filosum G. B. Sowerby II, 1865, also originally described from Philippines, displays a prominent beaded secondary cord s 1 and has varical nodes lacking on the spire of C. niauense Merle, n. sp.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FC70F80EFA5AF86A.taxon	type_taxon	TYPE SPECIES. — Drupa cornus Röding, 1798 by original designation.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FEA0FC09FA92FED0.taxon	description	(Fig. 12 A, B)	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FEA0FC09FA92FED0.taxon	materials_examined	TYPE MATERIAL. — Syntype. Philippines • Maxwell et al. (2021: 100) wrote that Röding (1798: 62, n ° 786 - 788) designated a holotype refering to the illustration of Martini (1777: pl. 27, fig. 794), but Röding (1798: 62) did not designate a holotype. TYPE LOCALITY. — Cebu Island, Philippines (Abbott 1960: 144, fig. 8). FIGURED MATERIAL. — France • 1 spm; French Polynesia, Niau Atoll, La Piscine ”: 6 ° 08 ’ 22.32 S, 146 ° 23 ’ 14.09 W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98429 (Letourneux coll.); H 47.1 mm (Fig. 12 A, B). OTHER MATERIAL. — France • 21 spm; French Polynesia, Niau Atoll, La Piscine ”: 6 ° 08 ’ 22.32 S, 146 ° 23 ’ 14.09 W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98430 (Letourneux coll.) • 3 spm; same; MNHN. F. A 98431 (Letourneux coll.).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FEA0FC09FA92FED0.taxon	distribution	DISTRIBUTION. — Plio-Pleistocene, French Polynesia (this work) to today, Philippines, eastern Indonesian Archipelago, northeastern Papua New Guinea and associated Island chains (Maxwell et al. 2021: 104, fig. 11).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FEA0FC09FA92FED0.taxon	description	DESCRIPTION See Maxwell et al. (2021: 99).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FEA0FC09FA92FED0.taxon	discussion	COMMENTS The figured material displays a uniform pinkish-yellow background interrupted by around seven small grey spiral bands. This residual pattern suggests that the coloured background was interrupted by unpigmented spiral bands. The colour pattern of Gibberulus gibbosus is very variable (see Maxwell et al. (2021: figs 8 - 10), but it is also interrupted by white spiral bands. These white bands correspond to the small grey bands observed under UV light and indicate that the residual colour pattern on the Plio-Pleistocene fossils is strongly reminiscent of the colour pattern of the extant Gibberulus gibbosus.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FEA0FD6AFCECFCC6.taxon	type_taxon	TYPE SPECIES. — Strombus gibberulus Linnaeus, 1758 by monotypy.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • French Polynesia, Niau Atoll, La Piscine ”: 6 ° 08 ’ 22.32 S, 146 ° 23 ’ 14.09 W; Plio-Pleistocene (molluscs assemblage); MNHN-IM- 2000 - 23289; H 34.5 mm. PARATYPE. — France • 1 spm; same as the holotype; EPHE coll.; H 36 mm • 1 spm; idem; Letourneux coll.; H 35 mm • 1 spm; idem; MNHN. F. A 98433 (Tröndlé coll.); H 40 mm • 1 spm; idem; G. Kronenberg coll.; H 36 mm. FIGURED MATERIAL. — 1 spm; idem; MNHN. F. A 98432; H 31.3 mm (Fig. 12 C, D). TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7 - 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	distribution	DISTRIBUTION. — Only know from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	description	DESCRIPTION See Tröndlé & Salvat (2010: 616) and Dekkers & Maxwell (2018: 224).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	discussion	COMMENTS This species is close to Striatostrombus micklei (Ladd, 1972) from the Late Miocene of Marshall Archipelago (Dekkers & Maxwell 2018). When S. blanci was described, traces of a colour pattern were not researched. Under UV light, the studied material displays a uniform grey background (Fig. 12 D) that suggests that the species was probably poorly pigmented and devoid of colour pattern.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C210FF87FCEBFE08FBFFFD83.taxon	type_taxon	TYPE SPECIES. — Striatostrombus blanci (Tröndlé & Salvat, 2010) by original designation.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C217FF81FF48F9CBFE55FDF1.taxon	description	(Fig. 12 E-G)	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C217FF81FF48F9CBFE55FDF1.taxon	materials_examined	TYPE MATERIAL. — Lectotype. Australia • Designated by Emerson & d’Attilio (1981: 81); Lord Howe Island; BMNH 1981146. FIGURED MATERIAL. — France • 1 spm; French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98434 (Letourneux coll.); H 22.9 mm (Fig. 12 E, F) • 1 spm; idem; MNHN. F. A 98435 (Letourneux coll.); H 25.1 mm (Fig. 12 G). OTHER MATERIAL. — France • 4 spm; same data as the Figured material; MNHN. F. A 98465 (Letourneux coll.). TYPE LOCALITY. — Australia, Tasman Sea, Lord Howe Island.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C217FF81FF48F9CBFE55FDF1.taxon	distribution	DISTRIBUTION. — Plio-Pleistocene (molluscs assemblage), French Polynesia, Niau Atoll to today, Indo-West-Pacific.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C217FF81FF48F9CBFE55FDF1.taxon	description	DESCRIPTION See Tröndlé & Houart (1992: 85).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C217FF81FF48F9CBFE55FDF1.taxon	discussion	COMMENTS Two specimens display a residual colour pattern, but with a different degree of diagenesis. One is almost pristine (Fig. 12 E, F) and its colour pattern is similar to that found in extant specimens. The other specimen exhibits yellow spots where the nodules are, that indicates strongly pigmented nodules (Fig. 12 G). Unpigmented areas correspond to spiral grooves and intervarical spaces. This pattern is reminiscent to living specimens having brown or black nodules interrupted by fairer or white spiral grooves.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	description	(Fig. 13 A-D) urn: lsid: zoobank. org: act: 5 DDF 566 A- 05 C 5 - 43 D 7 - B 2 D 5 - BB 49 C 5189 FC 7	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98436 (Letourneux coll.); H 70.5 mm.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	etymology	ETYMOLOGY. — Dedicated to Jean Letourneux.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7; 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	distribution	DISTRIBUTION. — Only known from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	description	DESCRIPTION Shell moderately large, with very low conical spire. Apical angle 118 °. Teleoconch of 9 whorls, whorl tops smooth delimited by a shoulder very slightly lower than the suture. Suture moderately impressed. Subsutural sinus deep, strongly curved, strongly asymmetrical. Last whorl with concave subsutural platform, slighlty angled at shoulder placed below suture, regularly conical below, position of maximum diameter close below shoulder, not constricted at base. Siphonal fasciole weakly developed. Weak, close set spaced cords from the base to the center of last whorl and fasciole. Aperture relatively narrow with parallel sides, hardly widening abapically; siphonal canal short. Colour pattern poorly preserved consisting large dark triangular spots surrounded by light yellow fluorescent bands under UV light.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	discussion	COMPARISONS The residual colour pattern of Conus (Conus) letourneuxi n. sp. (Fig. 13 C, D) is reminicent to that of Conus (Conus) marmoreus (Linneaus, 1758), Conus (Conus) bandanus Hwass in Bruguière, 1792 and Conus (Eugeniconus) marchionatus Hinds, 1843. The shells of C. (C.) marmoreus and C. (C.) bandanus have a crenulated spire, whereas that of C. (C.) letourneuxi n. sp. is smooth. C. (E.) marchionatus displays a smooth spire, but it is flatter than that of C. (C.) letourneuxi n. sp. and the shape of its triangular spots is more rounded.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FC01FD29FA5BFC86.taxon	type_taxon	TYPE SPECIES. — Conus tessulatus Born, 1778 by original designation.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF81FE86FC8AFE48FC01.taxon	type_taxon	TYPE SPECIES. — Conus marmoreus Linnaeus, 1758, by subsequent designation (Children 1823).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	description	(Figs 13 E-Q; 14 A-L; 15 A-D) urn: lsid: zoobank. org: act: 00915 CC 5 - 39 F 5 - 478 D- 84 BA- 87551894 B 565	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98437 (Letourneux coll.); H 43.4 mm (Figs 13 E-H; 15 A, B). Paratypes. France • 1 spm; same data as the holotype; MNHN. F. A 98438; H 39.4 mm (Fig. 13 I, J) • 1 spm; idem; MNHN. F. A 98439; H 89.6 mm (part of the last whorl, Fig. 13 K) • 1 spm; idem; MNHN. F. A 98440; H 35.9 mm (Fig. 13 L-O) • 1 spm; idem; MNHN. F. A 98441; H 34 mm (Fig. 13 P, Q) • 1 spm; idem; MNHN. F. A 98442; H 29.6 mm (Fig. 14 A, B) • 1 spm; idem; MNHN. F. A 98443; H 28.1 mm (Fig. 14 C) • 1 spm; idem; MNHN. F. A 98444; H 22 mm (Fig. 14 D, E) • 1 spm; idem; MNHN. F. A 98445; H 36.7 mm (Fig. 14 F, G) • 1 spm; idem; MNHN. F. A 98446; H 35.6 mm (Fig. 14 H, I; 15 C, D) • 1 spm; idem; MNHN. F. A 98447; H 23 mm • 1 spm; idem; MNHN. F. A 98448; H 23.6 mm (Fig. 14 J-L). All paratypes from Letourneux coll. ADDITIONAL MATERIAL. — France • 3 spm; same data as the holotype; MNHN. F. A 98449 (Letourneux coll.).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	etymology	ETYMOLOGY. — Dedicated to Georges Richard.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7; 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	distribution	DISTRIBUTION. — Only know from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	description	DESCRIPTION Shell large (MNHN. F. A 98439 (Fig. 13 K) part of the last whorl = 89.6 mm), with low coeloconoid to conical spire. Apical angle 135 °. Teleoconch of 7 - 8 whorls, whorl tops smooth except for close-set, comma-shaped growth lines, concave, delimited by shoulder raised very slightly below suture. Suture weakly impressed. Subsutural sinus shallow, slightly curved, moderately asymmetrical. Last whorl with horizontal, flat subsutural platform, smoothy angled at shoulder placed almost at suture, regularly conical below, position of maximum diameter close below shoulder, not constricted at base. Siphonal fasciole weakly developed; a few weak, widely spaced cords over base and fasciole, strengthening slightly abapically. Aperture relatively narrow with parallel sides, widening abapically; siphonal canal short. Under UV light, residual colour pattern consisting of two components: (1) pale, yellow-white fluorescent spots, spirally aligned and (2) dark, non-fluorescent background. Small rectangular spots of unequal size, distributed on the last whorl in four zones from shoulder to base. Zone a (first posterior quarter of whorl): 3 - 4 rows of sparse dots; zone b (second posterior quarter of whorl): 3 - 6 rows of close-set dots; zone c (first anterior quarter of whorl: 2 - 3 spaced rows of sparse dots; zone c (second anterior quater of whorl): 4 - 8 rows of more or less spaced dots (Fig. 15 A-D).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	discussion	COMPARISONS Among French Polynesian Conidae bearing spiral rows of dots, C. (Tesselliconus) georgesrichardi n. sp. may be compared with Conus (Elisaconus) litteratus (Linnaeus, 1758), C. (Tesselliconus) tessulatus Born, 1778 and C. (T.) eburneus (Hwass in Bruguière, 1792). C. (E.) litteratus differs in having a planispiral spire, a narrow, concave subsutural platform and a shoulder placed above the suture. Conus (E.) litteratus displays around 20 - 25 row of dots, instead 15 - 18 in C. (T.) georgesrichardi n. sp. In addition, the rows are regularly aligned from the shoulder to the base on last whorl, whereas those of C. (T.) georgesrichardi n. sp. are organised in four zones (a to d). The shell shape of Conus (T.) tessulatus is reminiscent to that C. (T.) georgesrichardi n. sp. and as in the Plio-Pleistocene species, the rows of dots are organised in four zones (Fig. 15 E, F). However, Conus (T.) tessulatus species differs clearly by having in elongated larger dark dots. The colour pattern of C. (T.) erburneus consists in four similar zones of spiral rows of dots as in C. (T.) georgesrichardi n. sp. (Fig. 15 G, H) However, the dots of C. (T.) georgesrichardi n. sp. are smaller and more close set in the zone b. Moreover, the spire of C. (T.) georgesrichardi n. sp. can be more conical, the shell shape is wider and some gerontic individuals can reach a larger size (H more than 89 mm).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FC75F9ECFBC9F95C.taxon	type_taxon	TYPE SPECIES. — Bulla ampulla Linnaeus, 1758 by subsequent designation (Gray 1847).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FE91F8ECFF11F86A.taxon	type_taxon	TYPE SPECIES. — Buccinum maculatum Linnaeus, 1767 by original designation.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FCFBFF48FC44FA0C.taxon	description	(Fig. 14 M-P)	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FCFBFF48FC44FA0C.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN-IM- 2000 - 23684; H 96.5 mm. Paratypes. France • 1 spm; same data as the holotype; EPHE coll.; H 89.55 mm • 9 spm; idem; MNHN-IM- 2000 - 23686; H 78 mm, 81.6 mm, 74 mm, 62.7 mm, 62 mm, 60.4 mm, 55 mm, 50.7 mm, 48.9 mm • 1 spm; idem; Jean Tröndlé coll.; H: 61.2 mm • 1 spm; idem; Letourneux coll.; H 86.1 mm (Fig. 19 N-P) • 1 spm; idem; G. Paulay coll.; H 84 mm • 1 spm; idem; A. Waren coll.; H 79.1 mm • 1 spm; idem; Mairie de Niau coll.; H 87.5 mm • 1 spm; idem; M. Boutet coll.; H 87.8 mm • 1 spm; idem; G. Kronenberg coll.; H 60.6 mm. TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7 - 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W. FIGURED MATERIAL. — One paratype (Letourneux coll.), Fig. 19 M-P.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FCFBFF48FC44FA0C.taxon	distribution	DISTRIBUTION. — Only known from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FCFBFF48FC44FA0C.taxon	description	DESCRIPTION See Tröndlé & Letourneux (2011: 88).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF83FCFBFF48FC44FA0C.taxon	discussion	COMMENTS Tröndlé & Letourneux (2011) noted that UV light revealed only a few spots under the suture of the holotype. On the examined paratype, a dark band is also present under the suture of the penultimate and the last whorl of the specimen examined. These authors found strong shell similarities with Oxymeris gouldi (Deshayes, 1857), today Hawaii. O. niauensis looks also allied to Oxymeris caledonica (G. B. Sowerby III, 1909), today New Caledonia. Therefore, O. niauensis seems better placed in the genus Oxymeris Dall, 1903.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	description	(Fig. 16) urn: lsid: zoobank. org: act: 8 C 774 DB 3 - F 7 BE- 4 DD 8 - 838 C- 01 E 30 AA 984 D 9	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98450 (Letourneux coll.); H 25.4 mm (Fig. 16 A-E). Paratypes. France • 1 spm; same data as the holotype; MNHN. F. A 98452 (Letourneux coll.); H 25.7 mm • 1 spm; idem; MNHN. F. A 98451 (Letourneux coll.), H 24.5 mm (Fig. 16 F, G). ADDITIONAL MATERIAL. — France • 3 spm; same data as the holotype; MNHN. F. A 98453 (Letourneux coll.).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	etymology	ETYMOLOGY. — From the type locality Niau (French Polynesia).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7; 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”, 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	distribution	DISTRIBUTION. — Only know from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	description	DESCRIPTION Shell thick, shape subquadrangular, wider anteriorly. Parietal callus thin; collumella thick. Spire aperture wide. Outer lip straight posteriorly, rounded anteriorly. Under UV light, residual pattern composed of two levels of residual pigmentation: a lighter background and dark stripes from wavy to zigzagging.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C214FF8EFC75F88CFD71FAAF.taxon	discussion	COMPARISONS Bulla ampulla (Linnaeus, 1758) resembles Bulla niauensis Merle, n. sp., but differs by having a subquadrangular shape. The colour pattern displays mainly a fair background and darker dots and tending to collapse in small dots of irregular shape. Bulla vernicosa A. Gould, 1859 is more quadrangular than Bulla niauensis n. sp. and its outer lip is straighter anteriorly. The colour pattern of Bulla vernicosa is fair brown, with darker small dots. In reverse UV light, the pattern of Bulla niauensis n. sp. is also fair brown, but it bears dark stripes from wavy to zigzagging, lacking in Bulla vernicosa (see Malaquias & Reid 2008: fig. 27 L-Q).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8EFEA1F9ACFCE8F904.taxon	type_taxon	TYPE SPECIES. — Cardium retusum Linnaeus, 1767 by monotypy.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	description	(Fig. 17 A-H) urn: lsid: zoobank. org: act: 24555983 - F 607 - 4 A 6 A-A 048 - 4767 EA 62 CCEE	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 spm (right valve); French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 99068 (Letourneux coll.); DVL 18 mm (Fig. 17 A, B, G). Paratypes. France • 1 spm (right valve); same data as the holotype; MNHN. F. A 99821 (Letourneux coll.); DVL 21.8 mm (Fig. 17 E, F) • 1 spm (left valve); MNHN. F. A 99822 (Letourneux coll.); DVL 29.9 mm (Fig. 17 C, H) • 1 spm (left valve); MNHN. F. A 99823 Letourneux coll.); DVL 28.7 mm (Fig. 17 D) • 6 spm (2 right valves, 4 left valves); MNHN. F. A 99055 (Letourneux coll.); DVL 21.8 mm.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	etymology	ETYMOLOGY. — From the Latin adjective distortus (meaning distorted), because of the strongly curved the posterior area of the shell.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7; 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”, 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	distribution	DISTRIBUTION. — Only known from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	description	DESCRIPTION Shell moderately thin, strongly inflated in older specimens; strongly oblique, with a strong posterior sulcus. Shell up to 29.9 in DVL and 23.5 mm in APL. Anterior margin obliquely rounded; ventral margin oblique, descending toward carina; posterior margin strongly sinuate concave in its center. Carina sharply expressed, rounded and smooth. Shell equivalve and inequilateral with angulate, weakly prosogyrous umbones placed in anterior half and extending above dorsal line. Shell surface densely radially sculptured, with 13 - 18 anterior ribs and 10 - 12 porterior ribs. Anterior ribs low, rather flat, slightly split by a central groove and separated by narrow interspaces less than 3 / 4 rib in breadth. Posterior ribs low becoming larger near the ventral margin. Pallial line entire, positioned moderately close to the ventral margin and connecting two medium-sized subovate scars. Shell margin finely crenulate ventrally, but with coarser crenulations on posterior margin. Posterior part well delineated from anterior by carinal groove. Hinge line strong and rather thick, with two cardinal teeth and two prominent lateral teeth. Posterior lateral tooth in right valve and anterior lateral in left valve strong.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C219FF8FFEA4F94CFB72F86D.taxon	discussion	COMPARISONS The Plio-Pleistocene assemblage from Niau contains two species of Fraginae, Fragum fragum (Linnaeus, 1758) and? Lunulicardia distorta Merle, n. sp. Fragum fragum is less carinate, its posterior margin is slightly convex and the surface of its ribs is squamous (seeFig. 9 M-O). In the living fauna from French Polynesia, Boutet et al. (2020) report three other species of Fragum Röding, 1798: F. mundum (Reeve, 1845), F. sueziense (Issel, 1869) and F. whitleyi Iredale, 1919. They differ easily from? Lunulicardia distorta Merle, n. sp. by their weak carina and their convex posterior margin. Members of Lunulicardia J. E. Gray, 1853 (type species: Cardium retusum Linnaeus, 1767 by monotypy, present day: Indo-West Pacific from Japan to Australia to South Africa) are reminiscent of? Lunicardia distorta Merle, n. sp. in their angulate carina, but have a concave posterior margin. Americardia Stewart, 1930 (type species: Cardium medium Linnaeus, 1758 by original designation, present day, Western Atlantic) has a weakly angulate carina, more ribs (22 - 25 anterior ribs), but is slightly reminiscent of? Lunulicardia distorta Merle, n. sp. in its weakly concave posterior margin. Americardia lightbourni Lee & Huber, 2012 from Bermuda Island displays a very concave posterior margin as it is in? L. distorta n. sp., but differs by many other characters (weakly angulate carina, numerous ribs with a squamous surface). Finally,? L. distorta n. sp. is a very peculiar species for the Tuamotu archipelago. Its attribution to Lunulicardia is also uncertain as its posterior margin is strongly concave, whereas it is convex in species usually attributed to this genus.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF8AFEA0F928FD59F86A.taxon	description	(Fig. 18 F-L) urn: lsid: zoobank. org: act: AF 800 AD 8 - 3 ACD- 488 C- 8306 - 9 EF 6 C 41342 BE Soletellina sp. – Salvat et al. 2018: table 1.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF8AFEA0F928FD59F86A.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 complete spm (right and left valves); French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 99067 (Letourneux coll.); APL 93.2 mm (Fig. 18 F-I). Paratypes. France • 1 complete spm; same data as the holotype; MNHN. F. A 99069 (Letourneux coll.); APL 99.8 mm (Fig. 18 J, K) • 1 complete spm; MNHN. F. A 99824 (Letourneux coll.); APL 104 mm (Fig. 18 L) • 1 complete spm with a broken right vave; MNHN. F. A 99978 (Letourneux coll.) • 5 spm; MNHN. F. A 99151 (Letourneux coll.).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF8AFEA0F928FD59F86A.taxon	etymology	ETYMOLOGY. — From the type locality, Fakaau: Paumotu name for Niau.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF8AFEA0F928FD59F86A.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7 - 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, “ La Piscine ”: 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF8AFEA0F928FD59F86A.taxon	distribution	DISTRIBUTION. — Only know from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF8AFEA0F928FD59F86A.taxon	description	DESCRIPTION Larger shell extending to 104 mm in APL and to 55 mm in DVL, elongate, moderately inequilateral, equivalved, subsolid, rather compressed and with weak flexure posteriorly. Umbo slightly behind middle, rather inconspicuous and blunt. Anterior margin broadly rounded; ventral margin elongate, slightly convex, rising arcuately posteriorly; anterior dorsal margin straight to gently convex, fairly elongate, and gently descending; posterior dorsal margin elongate, slightly descending and weakly convex; posterior margin weakly biangulate, concave ventrally. Surface of shell smooth with only concentric growing striae. Ligament area thin and shallow on anterior dorsal margin and deeper and slightly wider on posterior dorsal margin; weakly developed escutcheon; lunule poorly developed. Commissure line of the ventral margin slightly twisted in its posterior part. Hinge line well developed. In left valve, cardinal complex consisting of an anterior, strong bifid tooth with subequal lobes, and of a posterior, elongate and thin laminate tooth; anterior lateral tooth obsolete; posterior lateral tooth obsolete, distal to cardinal complex; cardinal hinge plate extensive, particularly anteriorly. In right valve, cardinal complex consisting of a posterior, slightly skewed, well-developed, bifid cardinal tooth with subequal lobes, and of anterior slightly thickened, laminate, cardinal tooth; anterior lateral tooth small; posterior lateral tooth obsolete and distal to cardinal complex; anterior lateral tooth closer to cardinal complex. Muscle scars moderately well impressed; anterior and posterior adductor muscles scar irregularly semilunate. Pallial sinus subequal in opposite valves; pallial line poorly distinct. COMPARISONS This species was first identified as a Psammobiidae J. Fleming, 1828 of the genus Soletellina Blainville, 1824, a junior synonym of Hyatula Modeer, 1793 (type species Hiatula dyphos (Linnaeus, 1771) by subsequent designation. Hyatula diphos (= Soletellina radiata Blainville, 1824) is a present-day Indo-West Pacific species and is reminiscent of Indotellina in its smooth surface and in its elongate and inequilateral shape with a posterior margin longer than the anterior margin. Hiatula diphos differs from Indotellina by lacking lateral tooth and by having a moderately long nymph. Moreover, its posterior margin is less angular and more convex than that of Indotellina. Until now, Indotellina was only known by its type species, Indotellina chariessa A. E. Salisbury, 1934 [junior synonym of Tellina elegans W. Wood, 1828, not Deshayes (1824)]. It is a present-day Indo-West Pacific species but there is an unpublished record in the Pliocene from Niué atoll in Western Polynesia (see https: // www. gbif. org / occurrence / 1064827107 and Paulay & Spencer 1992 for the geological context). According to Boutet et al. (2020), this species is missing in the present-day Polynesian fauna. Indotellina chariessa is very similar to Indotellina fakaauensis Merle, n. sp. and both species share an angulate posterior margin and a hinge with lateral tooth. However, Indotellina fakaauensis can be distinguished by its more rectilinear ventral margin, its less angulate posterior margin and its more convex dorso-posterior margin. Indotellina fakaauensis is located just outside the geographical range of I. chariassae and it is possible that its presence in the Tuamotu Archipelago results from a Pliocene speciation phase separating both species and followed by its extinction.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF88FF02F989FF16F900.taxon	type_taxon	TYPE SPECIES. — Tellina chariessa A. E. Salisbury, 1934 by original designation.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF88FE85FEC8FE3FFE42.taxon	type_taxon	TYPE SPECIES. — Tellina virgata Linnaeus, 1758, by subsequent designation (Schmidt F. C. 1818).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF88FF58FE67FCE2FA51.taxon	description	(Fig. 18 A-C)	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF88FF58FE67FCE2FA51.taxon	materials_examined	FIGURED MATERIAL. — France • 1 left valve; French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98454 (Letourneux coll.); APL 69.7 mm (Fig. 18 A-C). OTHER MATERIAL EXAMINED. — France • 2 spm; same data as Figured material; MNHN. F. A 98455 (Letourneux coll.)	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF88FF58FE67FCE2FA51.taxon	description	DESCRIPTION Shell rather large for family. Shape ovate-slightly elongate and subequilateral. Sculpture of commarginal striae becoming slightly oblique near posterior end; posterior slope with well distinct commarginal striae; anterior lateral tooth elongate, rather thick, moderately close to cardinals; posterior lateral tooth rather thick, distant from cardinals; pallial sinus poorly visible. Residual colour pattern uniform showing poorly defined concentric bands crossed by several very weak transverse lines more visible on the anterior part of valve.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21FFF88FF58FE67FCE2FA51.taxon	discussion	COMMENTS This shell (Fig. 18 A-C) resembles living Tellinella virgata (see Fig. 18 D-E) but its shape is more ovate with a higher dorso-ventral lenght. In addition, its residual colour pattern don’t display well large transverse dark bands, as observed in the syntype (P-Z 0010893) of the Linnean collection [https: // www. linnean. org / research-collections / linnaean-collections] and as in other specimens (Fig. 18 D, E). Thus, the attribution of this specimen to Tellinella virgata is provisional and is proposed with some doubts.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8AFC60F9EAFBF7F960.taxon	type_taxon	TYPE SPECIES. — Venus pectinata Linnaeus, 1758 by subsequent designation (Dall 1902).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	description	(Fig. 19) urn: lsid: zoobank. org: act: 2 AB 9871 C- 6 D 0 C- 4858 - A 136 - 0 CD 4 E 1 AF 64 E 4	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	materials_examined	TYPE MATERIAL. — Holotype. France • 1 left valve; French Polynesia, Niau Atoll, “ La Piscine ”; 6 ° 08 ’ 22.32 ” S, 146 ° 23 ’ 14.09 ” W; Plio-Pleistocene (molluscs assemblage); MNHN. F. A 98456 (Letourneux coll.); APL 53 mm (Fig. 19 A-D). Paratypes. France • 10 left valves, 6 right valves, one complete specimen (two valves); same data as the holotype; MNHN. F. A 98458 (Letourneux coll.) • 1 spm, MNHN. F. A 98457 (Letourneux coll.); APL 52 mm (Fig. 19 E, F).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	etymology	ETYMOLOGY. — Dedicated to Jean Tröndlé.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	materials_examined	TYPE HORIZON. — Plio-Pleistocene (molluscs assemblage) (see Figs 7; 8). TYPE LOCALITY. — French Polynesia, Niau Atoll, La Piscine ”: 6 ° 08 ’ 22.32 S, 146 ° 23 ’ 14.09 W.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	distribution	DISTRIBUTION. — Only known from the type locality.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	description	DESCRIPTION Shell weakly suborbicular, solid, thick; umbones flat, prosogyrate; lunule elongate heart-shaped, poorly raised, weakly delineated by incised line; escutcheon poorly defined, narrow; anterodorsal margin short, rounded terminally; ventral margin convex slightly angulate near posterior margin; posterodorsal margin moderately convex, angulate at its posterior termination. Sculpture radially, divaricately ribbed at early stages of growth and of dense concentric ribs at later stage of growth. Hinge of left valve with anterior lateral tooth solid, thick, peg-like; anterior cardinal very thin, separated from median cardinal by narrow V-shaped socket; median cardinal thick, oblique, raised; posterior cardinal free, thin, elongate and oblique; posterior lateral thin, parallel to nymph. Hinge of right valve with broad anterior pit, anterior lateral short, anterior cardinal short, slightly oblique, median cardinal, thick, peg-like, slightly oblique; posterior cardinal oblique, peg-like; posterior lateral, parallel to nymph. Anterior muscle adductor scars trapezoidally shaped, posterior adductor scars ovate. Pallial line wide; pallial sinus diminutive. Residual colour pattern formed of radial, narrow and more or less wavy yellow fluorescent lines (Fig. 18 C).	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
CA729441C21DFF8BFC12F889FC20FD11.taxon	discussion	COMPARISONS This enigmatic species was previoulsy identified as Circe plicatina (Lamarck, 1818) in collection and as Gafrarium sp. (Salvat et al. 2018). Valves of Circe plicatina are more trigone in shape, their sculpture is less divaricate near the umbo, their cardinal teeth are less solid, less thick and their anterior pits are less broad. Early stages of growth of shells of Circe scripta (Linnaeus, 1758), type species of Circe Schumacher, 1817 resemble those of Circe plicatina and differ from those of? Gafrarium troendlei n. sp. in having only concentric ribs. In addition, both species of Circe differs by having more compressed shells. The early stages of growth of? Gafrarium troendlei Merle, n. sp. are reminscent to G. harteae and other species of Gafrarium Röding, 1798 in having a dominant radial sculpture of ribs tending sometimes to be divaricated. However,? G. troendlei Merle, n. sp. can be easily distinguished from G. harteae Huber, 2010 and the type species of the Gafrarium Röding, 1798, Gafrarium pectinatum (Linnaeus, 1758), by its a dominant concentric sculpture and by its smooth, not crenulated ventral margin (see Fig. 9 J-L). Gafrarium tumidum Röding, 1798, currently present on part of the south-west coast of Tahiti from Port Phaëton to Vairao, differs by its stronger radial sculpture and by the shape of its cardinal plate and muscle impressions. Among other species of Gafrarium, a dominant concentric sculpture is found in G. eaquivocum (Holten, 1802). Therefore, we prefer attribute? Gafrarium troendlei Merle, n. sp. to Gafrarium than Circe, but this attribution is provisional, because the characters of this species are not fully typical of Gafrarium.	en	Salvat, Bernard, Merle, Didier, Letourneux, Jean, Tröndlé, Jean, Montaggioni, Lucien François, Pons-Branchu, Edwige, Dumoulin, Jean-Pascal, Bacchet, Philippe (2026): Evolutionary history of reef-dwelling and associated lagoonal mollusc assemblages of the uplifted Niau Atoll (Tuamotu, French Polynesia) since the Pliocene. Geodiversitas 48 (6): 87-125, DOI: 10.5252/geodiversitas2026v48a6
