taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384229/files/figure.png	https://doi.org/10.5281/zenodo.19384229	FIG. 9. — Species assemblage of Late Holocene sediments deposited above the caliche (sample N105, Vaihuoa,near Archeological platform),Niau Atoll (Tuamotu, French Polynesia):A-C, Cerithium columna G. B. Sowerby I, 1834;A, MNHN.F.A98459; B, MNHN.F.A98460; C, MNHN.F.A98461; D-F, Cerithium niauense Merle, n. sp.: D, MNHN.F.A98411; E, MNHN.F.A98412; F, MNHN.F.A98413; G-I, Cerithium rostratum G. B. Sowerby II, 1855:G, MNHN.F.A98462; H, MNHN.F.A98463; I, MNHN.F.A98464; J-L, Gafrarium pectinatum (Linnaeus, 1758): J, MNHN.F.C02086; K, MNHN.F.C02087; L, MNHN.F.C02088; M-O, Fragum fragum (Linnaeus, 1758): M, MNHN.F.C02089; N, MNHN.F.C02090; O, MNHN.F.C02091; P-R, Ctena bella (Conrad, 1837): P, MNHN.F.C02092; Q, MNHN.F.C02093; R, MNHN.F.C02094. Scale bars: 10 mm. Credit: J.-P. Pointier.	FIG. 9. — Species assemblage of Late Holocene sediments deposited above the caliche (sample N105, Vaihuoa,near Archeological platform),Niau Atoll (Tuamotu, French Polynesia):A-C, Cerithium columna G. B. Sowerby I, 1834;A, MNHN.F.A98459; B, MNHN.F.A98460; C, MNHN.F.A98461; D-F, Cerithium niauense Merle, n. sp.: D, MNHN.F.A98411; E, MNHN.F.A98412; F, MNHN.F.A98413; G-I, Cerithium rostratum G. B. Sowerby II, 1855:G, MNHN.F.A98462; H, MNHN.F.A98463; I, MNHN.F.A98464; J-L, Gafrarium pectinatum (Linnaeus, 1758): J, MNHN.F.C02086; K, MNHN.F.C02087; L, MNHN.F.C02088; M-O, Fragum fragum (Linnaeus, 1758): M, MNHN.F.C02089; N, MNHN.F.C02090; O, MNHN.F.C02091; P-R, Ctena bella (Conrad, 1837): P, MNHN.F.C02092; Q, MNHN.F.C02093; R, MNHN.F.C02094. Scale bars: 10 mm. Credit: J.-P. Pointier.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C20CFF87FEB2FA89FD9CFD90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19392543/files/figure.png	https://doi.org/10.5281/zenodo.19392543	FIG. 11. — Cerithium niauense Merle,n. sp. from the Plio-Pleistocene (A-F) and the Late Holocene at Vaihuoa near archeological platform (G-V) of Niau Atoll (Tuamotu, French Polynesia): A, holotype, MNHN.F.A98401, ventral face; B, idem in ventral face under UV light; C, idem, dorsal face; D, paratype MNHN.F.A98402, ventral face;E, idem,dorsal face,F, idem,dorsal face under UV light; G-H, paratype (sample N106),MNHN.F.A98404, ventral (G) and dorsal (H) faces;I, J, paratype (sample N106), MNHN.F.A98405, ventral (I) and dorsal (J) faces; K, L, paratype (sample N108), MNHN.F.A98403, ventral (K) and dorsal (L) faces; M, N, paratype (sample TP1A), MNHN.F.A98406, ventral (M) dorsal (N) faces; O, P, paratype (sample TP1A), MNHN.F.A98407, ventral (O) and dorsal (P) faces; Q, R, paratype (sample TP1B), MNHN.F.A98408, ventral (Q) and dorsal (R) faces; S, T, paratype (samples TP1B), MNHN.F.A98409, ventral (S) and dorsal (T) faces; U, V, paratype (sample TP1B), MNHN.F.A98410, ventral (U) and dorsal (V) faces; W, X, Cerithium tenellum G. Sowerby II, 1855 (Limpelaer coll.), Pelotes Point, Philippines, present day, ventral (X) and dorsal (Y) faces for comparison with C. niauense Merle, n. sp. Scale bars: 5 mm. Credits: A-F, P. Loubry; G-Y, L. Cazes. B and F, photos taken under UV, other photos taken in natural light. A-F, M-V, material from Letourneux coll.; G-L, material from Salvat coll.	FIG. 11. — Cerithium niauense Merle,n. sp. from the Plio-Pleistocene (A-F) and the Late Holocene at Vaihuoa near archeological platform (G-V) of Niau Atoll (Tuamotu, French Polynesia): A, holotype, MNHN.F.A98401, ventral face; B, idem in ventral face under UV light; C, idem, dorsal face; D, paratype MNHN.F.A98402, ventral face;E, idem,dorsal face,F, idem,dorsal face under UV light; G-H, paratype (sample N106),MNHN.F.A98404, ventral (G) and dorsal (H) faces;I, J, paratype (sample N106), MNHN.F.A98405, ventral (I) and dorsal (J) faces; K, L, paratype (sample N108), MNHN.F.A98403, ventral (K) and dorsal (L) faces; M, N, paratype (sample TP1A), MNHN.F.A98406, ventral (M) dorsal (N) faces; O, P, paratype (sample TP1A), MNHN.F.A98407, ventral (O) and dorsal (P) faces; Q, R, paratype (sample TP1B), MNHN.F.A98408, ventral (Q) and dorsal (R) faces; S, T, paratype (samples TP1B), MNHN.F.A98409, ventral (S) and dorsal (T) faces; U, V, paratype (sample TP1B), MNHN.F.A98410, ventral (U) and dorsal (V) faces; W, X, Cerithium tenellum G. Sowerby II, 1855 (Limpelaer coll.), Pelotes Point, Philippines, present day, ventral (X) and dorsal (Y) faces for comparison with C. niauense Merle, n. sp. Scale bars: 5 mm. Credits: A-F, P. Loubry; G-Y, L. Cazes. B and F, photos taken under UV, other photos taken in natural light. A-F, M-V, material from Letourneux coll.; G-L, material from Salvat coll.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C210FF87FEA0FC09FA92FED0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384237/files/figure.png	https://doi.org/10.5281/zenodo.19384237	FIG. 12. — Strombidae (A-D) and Muricidae (E-G) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, Gibberulus gibbosus (RÖding, 1798), MNHN.F.A98429: A, ventral face under UV light; B, dorsal face under UV light; C, D, Striatostrombus blanci (TrÖndlé & Salvat, 2010), MNHN.F.A98432: C, ventral face in natural light;D, ventral face under UV light;E-G, Drupella margariticola (Broderip,1833):E, F, MNHN.F.A98434, ventral and dorsal faces under UV light; G, MNHN.F.A98435, ventral face under UV light. Scale bars: 10 mm. Credits P. Loubry (MNHN/CNRS).All material from Letourneux coll. housed at MNHN.	FIG. 12. — Strombidae (A-D) and Muricidae (E-G) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, Gibberulus gibbosus (RÖding, 1798), MNHN.F.A98429: A, ventral face under UV light; B, dorsal face under UV light; C, D, Striatostrombus blanci (TrÖndlé & Salvat, 2010), MNHN.F.A98432: C, ventral face in natural light;D, ventral face under UV light;E-G, Drupella margariticola (Broderip,1833):E, F, MNHN.F.A98434, ventral and dorsal faces under UV light; G, MNHN.F.A98435, ventral face under UV light. Scale bars: 10 mm. Credits P. Loubry (MNHN/CNRS).All material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384237/files/figure.png	https://doi.org/10.5281/zenodo.19384237	FIG. 12. — Strombidae (A-D) and Muricidae (E-G) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, Gibberulus gibbosus (RÖding, 1798), MNHN.F.A98429: A, ventral face under UV light; B, dorsal face under UV light; C, D, Striatostrombus blanci (TrÖndlé & Salvat, 2010), MNHN.F.A98432: C, ventral face in natural light;D, ventral face under UV light;E-G, Drupella margariticola (Broderip,1833):E, F, MNHN.F.A98434, ventral and dorsal faces under UV light; G, MNHN.F.A98435, ventral face under UV light. Scale bars: 10 mm. Credits P. Loubry (MNHN/CNRS).All material from Letourneux coll. housed at MNHN.	FIG. 12. — Strombidae (A-D) and Muricidae (E-G) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, Gibberulus gibbosus (RÖding, 1798), MNHN.F.A98429: A, ventral face under UV light; B, dorsal face under UV light; C, D, Striatostrombus blanci (TrÖndlé & Salvat, 2010), MNHN.F.A98432: C, ventral face in natural light;D, ventral face under UV light;E-G, Drupella margariticola (Broderip,1833):E, F, MNHN.F.A98434, ventral and dorsal faces under UV light; G, MNHN.F.A98435, ventral face under UV light. Scale bars: 10 mm. Credits P. Loubry (MNHN/CNRS).All material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C210FF87FCC6FDA8FBD3F90C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C217FF81FF48F9CBFE55FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384237/files/figure.png	https://doi.org/10.5281/zenodo.19384237	FIG. 12. — Strombidae (A-D) and Muricidae (E-G) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, Gibberulus gibbosus (RÖding, 1798), MNHN.F.A98429: A, ventral face under UV light; B, dorsal face under UV light; C, D, Striatostrombus blanci (TrÖndlé & Salvat, 2010), MNHN.F.A98432: C, ventral face in natural light;D, ventral face under UV light;E-G, Drupella margariticola (Broderip,1833):E, F, MNHN.F.A98434, ventral and dorsal faces under UV light; G, MNHN.F.A98435, ventral face under UV light. Scale bars: 10 mm. Credits P. Loubry (MNHN/CNRS).All material from Letourneux coll. housed at MNHN.	FIG. 12. — Strombidae (A-D) and Muricidae (E-G) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, Gibberulus gibbosus (RÖding, 1798), MNHN.F.A98429: A, ventral face under UV light; B, dorsal face under UV light; C, D, Striatostrombus blanci (TrÖndlé & Salvat, 2010), MNHN.F.A98432: C, ventral face in natural light;D, ventral face under UV light;E-G, Drupella margariticola (Broderip,1833):E, F, MNHN.F.A98434, ventral and dorsal faces under UV light; G, MNHN.F.A98435, ventral face under UV light. Scale bars: 10 mm. Credits P. Loubry (MNHN/CNRS).All material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384244/files/figure.png	https://doi.org/10.5281/zenodo.19384244	FIG. 13. — Conus (Conus) letourneuxi Merle, n. sp. (A-D) and Conus (Tesselliconus) georgesrichardi Merle, n. sp. (E-Q) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98436 (Letourneux coll.), ventral (A) and dorsal (B) faces in natural light, profil view under UV light (C) and in inverse UV light (D); E-H, holotype, MNHN.F.A98437, ventral face in natural light (E) and under UV light (F), dorsal face in natural light (G) and under UV light (H); I, J, paratype, MNHN.F.A98438, ventral (I) and dorsal (J) faces under UV light; K, paratype, MNHN.F.A98439, broken specimen in ventral face under UV light; L-O, paratype, MNHN.F.A98440, ventral face in natural light (L) and under UV light (M), dorsal face in natural light (N) and under UV light (O); P-Q, paratype, MNHN.F.A98441, ventral (P) and dorsal (Q) faces under UV light. Scale bars: A-D, K, 20 mm; E-J, L-Q, 10 mm. Credits: A, B, E, F, L-N, L. Cazes (MNHN/CNRS); C, D, F, H, I, J, K-M, O, P. Loubry (MNHN/CNRS). All material from Letourneux coll. housed at MNHN.	FIG. 13. — Conus (Conus) letourneuxi Merle, n. sp. (A-D) and Conus (Tesselliconus) georgesrichardi Merle, n. sp. (E-Q) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98436 (Letourneux coll.), ventral (A) and dorsal (B) faces in natural light, profil view under UV light (C) and in inverse UV light (D); E-H, holotype, MNHN.F.A98437, ventral face in natural light (E) and under UV light (F), dorsal face in natural light (G) and under UV light (H); I, J, paratype, MNHN.F.A98438, ventral (I) and dorsal (J) faces under UV light; K, paratype, MNHN.F.A98439, broken specimen in ventral face under UV light; L-O, paratype, MNHN.F.A98440, ventral face in natural light (L) and under UV light (M), dorsal face in natural light (N) and under UV light (O); P-Q, paratype, MNHN.F.A98441, ventral (P) and dorsal (Q) faces under UV light. Scale bars: A-D, K, 20 mm; E-J, L-Q, 10 mm. Credits: A, B, E, F, L-N, L. Cazes (MNHN/CNRS); C, D, F, H, I, J, K-M, O, P. Loubry (MNHN/CNRS). All material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF81FF4CFC6BFA47FDF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384244/files/figure.png	https://doi.org/10.5281/zenodo.19384244	FIG. 13. — Conus (Conus) letourneuxi Merle, n. sp. (A-D) and Conus (Tesselliconus) georgesrichardi Merle, n. sp. (E-Q) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98436 (Letourneux coll.), ventral (A) and dorsal (B) faces in natural light, profil view under UV light (C) and in inverse UV light (D); E-H, holotype, MNHN.F.A98437, ventral face in natural light (E) and under UV light (F), dorsal face in natural light (G) and under UV light (H); I, J, paratype, MNHN.F.A98438, ventral (I) and dorsal (J) faces under UV light; K, paratype, MNHN.F.A98439, broken specimen in ventral face under UV light; L-O, paratype, MNHN.F.A98440, ventral face in natural light (L) and under UV light (M), dorsal face in natural light (N) and under UV light (O); P-Q, paratype, MNHN.F.A98441, ventral (P) and dorsal (Q) faces under UV light. Scale bars: A-D, K, 20 mm; E-J, L-Q, 10 mm. Credits: A, B, E, F, L-N, L. Cazes (MNHN/CNRS); C, D, F, H, I, J, K-M, O, P. Loubry (MNHN/CNRS). All material from Letourneux coll. housed at MNHN.	FIG. 13. — Conus (Conus) letourneuxi Merle, n. sp. (A-D) and Conus (Tesselliconus) georgesrichardi Merle, n. sp. (E-Q) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98436 (Letourneux coll.), ventral (A) and dorsal (B) faces in natural light, profil view under UV light (C) and in inverse UV light (D); E-H, holotype, MNHN.F.A98437, ventral face in natural light (E) and under UV light (F), dorsal face in natural light (G) and under UV light (H); I, J, paratype, MNHN.F.A98438, ventral (I) and dorsal (J) faces under UV light; K, paratype, MNHN.F.A98439, broken specimen in ventral face under UV light; L-O, paratype, MNHN.F.A98440, ventral face in natural light (L) and under UV light (M), dorsal face in natural light (N) and under UV light (O); P-Q, paratype, MNHN.F.A98441, ventral (P) and dorsal (Q) faces under UV light. Scale bars: A-D, K, 20 mm; E-J, L-Q, 10 mm. Credits: A, B, E, F, L-N, L. Cazes (MNHN/CNRS); C, D, F, H, I, J, K-M, O, P. Loubry (MNHN/CNRS). All material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384251/files/figure.png	https://doi.org/10.5281/zenodo.19384251	FIG. 14. — Conus (Tesselliconus) georgesrichardi Merle,n.sp.(A-L) and Oxymeris niauensis (TrÖndlé & Letourneux,2011) (M-P) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, paratype,MNHN.F.A98442, ventral(A) and dorsal(B) faces under UV light;C, paratype,MNHN.F.A98443, ventral face underUV light; D, E, paratype,MNHN.F.A98444, ventral (D) and dorsal (E) faces under UV light;F, G, paratype,MNHN.F.A98445, ventral face under UV light (F) and in inverse UV light (G); H, I, paratype,MNHN.F.A98446, ventral (H) and dorsal (I) faces under UV light;J-L, paratype,MNHN.F.A98448 dorsal face under UV light (J) and ventral face in natural light (K) and under UV light (L); M-P, paratype,Letourneux coll., ventral face in natural light (M) and under UV light (N) and dorsal face in natural light (O) and under UV light (P). Scale bars:A-M,10 mm;N-P,20 mm.Credits:P.Loubry (MNHN/CNRS).A-L,material from Letourneux coll.housed at MNHN;M-O,material from Letourneux coll.,at Tahiti.	FIG. 14. — Conus (Tesselliconus) georgesrichardi Merle,n.sp.(A-L) and Oxymeris niauensis (TrÖndlé & Letourneux,2011) (M-P) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, paratype,MNHN.F.A98442, ventral(A) and dorsal(B) faces under UV light;C, paratype,MNHN.F.A98443, ventral face underUV light; D, E, paratype,MNHN.F.A98444, ventral (D) and dorsal (E) faces under UV light;F, G, paratype,MNHN.F.A98445, ventral face under UV light (F) and in inverse UV light (G); H, I, paratype,MNHN.F.A98446, ventral (H) and dorsal (I) faces under UV light;J-L, paratype,MNHN.F.A98448 dorsal face under UV light (J) and ventral face in natural light (K) and under UV light (L); M-P, paratype,Letourneux coll., ventral face in natural light (M) and under UV light (N) and dorsal face in natural light (O) and under UV light (P). Scale bars:A-M,10 mm;N-P,20 mm.Credits:P.Loubry (MNHN/CNRS).A-L,material from Letourneux coll.housed at MNHN;M-O,material from Letourneux coll.,at Tahiti.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384255/files/figure.png	https://doi.org/10.5281/zenodo.19384255	FIG. 15. — The fourth areas (a, b, c, d) of pigmentation in Conus (Tesselliconus) georgesrichardi Merle, n. sp. (A-D), Conus (Tesselliconus) tessulatus Born, 1778 (E, F) and Conus (Tesselliconus) eburneus Hwas, 1792 (G, H): A, B, holotype, MNHN.F.A98437 (Letourneux coll.), ventral face (A) and dorsal (B) faces under UV light, Plio-Pleistocene, Niau Atoll (Tuamotu, French Polynesia); C, D, paratype, MNHN.F.A98446 (Letourneux coll.), ventral (C) and dorsal (C) faces under UV light, Plio-Pleistocene, same locality; E, F, MNHN.IM (TrÖndlé coll.), ventral (E) and dorsal faces (F), present day, Papeete (Faaone), Tahiti island (Tuamotu, French Polynesia);G, H, MNHN.IM (Pierrot coll.), ventral (G) and dorsal faces (H), present day,Tahiti island.Scale bars: 20 mm. Credits:P. Loubry (MNHN/CNRS).	FIG. 15. — The fourth areas (a, b, c, d) of pigmentation in Conus (Tesselliconus) georgesrichardi Merle, n. sp. (A-D), Conus (Tesselliconus) tessulatus Born, 1778 (E, F) and Conus (Tesselliconus) eburneus Hwas, 1792 (G, H): A, B, holotype, MNHN.F.A98437 (Letourneux coll.), ventral face (A) and dorsal (B) faces under UV light, Plio-Pleistocene, Niau Atoll (Tuamotu, French Polynesia); C, D, paratype, MNHN.F.A98446 (Letourneux coll.), ventral (C) and dorsal (C) faces under UV light, Plio-Pleistocene, same locality; E, F, MNHN.IM (TrÖndlé coll.), ventral (E) and dorsal faces (F), present day, Papeete (Faaone), Tahiti island (Tuamotu, French Polynesia);G, H, MNHN.IM (Pierrot coll.), ventral (G) and dorsal faces (H), present day,Tahiti island.Scale bars: 20 mm. Credits:P. Loubry (MNHN/CNRS).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C216FF83FCC0FCC9FDAAF94C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF83FCFBFF48FC44FA0C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384251/files/figure.png	https://doi.org/10.5281/zenodo.19384251	FIG. 14. — Conus (Tesselliconus) georgesrichardi Merle,n.sp.(A-L) and Oxymeris niauensis (TrÖndlé & Letourneux,2011) (M-P) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, paratype,MNHN.F.A98442, ventral(A) and dorsal(B) faces under UV light;C, paratype,MNHN.F.A98443, ventral face underUV light; D, E, paratype,MNHN.F.A98444, ventral (D) and dorsal (E) faces under UV light;F, G, paratype,MNHN.F.A98445, ventral face under UV light (F) and in inverse UV light (G); H, I, paratype,MNHN.F.A98446, ventral (H) and dorsal (I) faces under UV light;J-L, paratype,MNHN.F.A98448 dorsal face under UV light (J) and ventral face in natural light (K) and under UV light (L); M-P, paratype,Letourneux coll., ventral face in natural light (M) and under UV light (N) and dorsal face in natural light (O) and under UV light (P). Scale bars:A-M,10 mm;N-P,20 mm.Credits:P.Loubry (MNHN/CNRS).A-L,material from Letourneux coll.housed at MNHN;M-O,material from Letourneux coll.,at Tahiti.	FIG. 14. — Conus (Tesselliconus) georgesrichardi Merle,n.sp.(A-L) and Oxymeris niauensis (TrÖndlé & Letourneux,2011) (M-P) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, paratype,MNHN.F.A98442, ventral(A) and dorsal(B) faces under UV light;C, paratype,MNHN.F.A98443, ventral face underUV light; D, E, paratype,MNHN.F.A98444, ventral (D) and dorsal (E) faces under UV light;F, G, paratype,MNHN.F.A98445, ventral face under UV light (F) and in inverse UV light (G); H, I, paratype,MNHN.F.A98446, ventral (H) and dorsal (I) faces under UV light;J-L, paratype,MNHN.F.A98448 dorsal face under UV light (J) and ventral face in natural light (K) and under UV light (L); M-P, paratype,Letourneux coll., ventral face in natural light (M) and under UV light (N) and dorsal face in natural light (O) and under UV light (P). Scale bars:A-M,10 mm;N-P,20 mm.Credits:P.Loubry (MNHN/CNRS).A-L,material from Letourneux coll.housed at MNHN;M-O,material from Letourneux coll.,at Tahiti.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF83FCFBFF48FC44FA0C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384274/files/figure.png	https://doi.org/10.5281/zenodo.19384274	FIG. 19. —?Gafrarium troendlei Merle, n. sp. from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98456, left valve, internal view in natural light (A), external view in natural light (B), under UV light (C) and in inverse UV light (D); E, F, paratype, MNHN.F.A98457, right valve, internal view (E) and external view (F) in natural light. Scale bars: 10 mm. Credits: A-C, P. Loubry (MNHN/CNRS); E, F, L. Cazes (MNHN/CNRS). Material from Letourneux coll. housed at MNHN.	FIG. 19. —?Gafrarium troendlei Merle, n. sp. from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98456, left valve, internal view in natural light (A), external view in natural light (B), under UV light (C) and in inverse UV light (D); E, F, paratype, MNHN.F.A98457, right valve, internal view (E) and external view (F) in natural light. Scale bars: 10 mm. Credits: A-C, P. Loubry (MNHN/CNRS); E, F, L. Cazes (MNHN/CNRS). Material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF83FCFBFF48FC44FA0C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF83FCFBFF48FC44FA0C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF8EFC75F88CFD71FAAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384259/files/figure.png	https://doi.org/10.5281/zenodo.19384259	FIG. 16. — Bulla niauensis Merle,n.sp.from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-E, holotype,MNHN.F.A98450 (Letourneux coll.), ventral face in natural light (A) and under UV light (B), dorsal face in natural light (C), under UV light (D) and in inverse UV light (E); paratype,MNHN.F.A98451 (Letourneux coll.), profil view under UV light (F) and in inverse UV light (G). Scale bars: 10 mm. Credits: P. Loubry (MNHN/CNRS).	FIG. 16. — Bulla niauensis Merle,n.sp.from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-E, holotype,MNHN.F.A98450 (Letourneux coll.), ventral face in natural light (A) and under UV light (B), dorsal face in natural light (C), under UV light (D) and in inverse UV light (E); paratype,MNHN.F.A98451 (Letourneux coll.), profil view under UV light (F) and in inverse UV light (G). Scale bars: 10 mm. Credits: P. Loubry (MNHN/CNRS).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF8EFC75F88CFD71FAAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C214FF8EFC75F88CFD71FAAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C219FF8FFEA4F94CFB72F86D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384262/files/figure.png	https://doi.org/10.5281/zenodo.19384262	FIG. 17. —?Lunulicardia deformata Merle,n. sp.from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, holotype,MNHN.F.A99068, right valve, internal (A) and external viewes (B); C, paratype, MNHN.F.A99822, left valve, internal view; D, paratype, MNHN.FA99823, left valve, external view; E, F, paratype, MNHN.A.99821, external (E) and internal (F) views; G, enlarged view of the hinge ot the holotype; H, enlarged view of the hinge of the paratype MNHN.F.A99822. Scale bars: A-F, 10 mm, G, H, 5 mm. Credits: P. Loubry. Material from Letourneux coll. housed at MNHN.	FIG. 17. —?Lunulicardia deformata Merle,n. sp.from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A, B, holotype,MNHN.F.A99068, right valve, internal (A) and external viewes (B); C, paratype, MNHN.F.A99822, left valve, internal view; D, paratype, MNHN.FA99823, left valve, external view; E, F, paratype, MNHN.A.99821, external (E) and internal (F) views; G, enlarged view of the hinge ot the holotype; H, enlarged view of the hinge of the paratype MNHN.F.A99822. Scale bars: A-F, 10 mm, G, H, 5 mm. Credits: P. Loubry. Material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C219FF8FFEA4F94CFB72F86D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C219FF8FFEA4F94CFB72F86D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21FFF8AFEA0F928FD59F86A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384267/files/figure.png	https://doi.org/10.5281/zenodo.19384267	FIG. 18. — Tellinidae (A-L) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-C, Tellinella cf.virgata(Linnaeus,1758), MNHN.F.A98454, left valve, internal view in natural light (A), external view in natural light (B) and under UV light (C); D, E, Tellinella virgata (Linnaeus, 1758) for comparison, MNHN.F.C02108 (Salvat coll.), present day, Rurutu island (Tuamotu,French Polynesia),left valve (D) and right valve (E); F-L, Indotellina fakaauensis Merle,n. sp.; F-I, holotype MNHN.F.A99067, external view of the right valve (F), external view of the left valve (G), view of the posterior margin (H), view of the ventral margin (I); J, K, paratype MNHN.F.A99069, internal view of the right valve (J), internal view of the left valve (K); L, paratype MNHN.F.A99824, external view of the right valve. Scale bars: 10 mm. Credits: A-C, F-L, P. Loubry (MNHN/CNRS); D-E, L. Cazes (MNHN/CNRS). A-C, F-L, material from Letourneux coll. housed at MNHN.	FIG. 18. — Tellinidae (A-L) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-C, Tellinella cf.virgata(Linnaeus,1758), MNHN.F.A98454, left valve, internal view in natural light (A), external view in natural light (B) and under UV light (C); D, E, Tellinella virgata (Linnaeus, 1758) for comparison, MNHN.F.C02108 (Salvat coll.), present day, Rurutu island (Tuamotu,French Polynesia),left valve (D) and right valve (E); F-L, Indotellina fakaauensis Merle,n. sp.; F-I, holotype MNHN.F.A99067, external view of the right valve (F), external view of the left valve (G), view of the posterior margin (H), view of the ventral margin (I); J, K, paratype MNHN.F.A99069, internal view of the right valve (J), internal view of the left valve (K); L, paratype MNHN.F.A99824, external view of the right valve. Scale bars: 10 mm. Credits: A-C, F-L, P. Loubry (MNHN/CNRS); D-E, L. Cazes (MNHN/CNRS). A-C, F-L, material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21FFF8AFEA0F928FD59F86A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21FFF8AFEA0F928FD59F86A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21FFF88FF58FE67FCE2FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384267/files/figure.png	https://doi.org/10.5281/zenodo.19384267	FIG. 18. — Tellinidae (A-L) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-C, Tellinella cf.virgata(Linnaeus,1758), MNHN.F.A98454, left valve, internal view in natural light (A), external view in natural light (B) and under UV light (C); D, E, Tellinella virgata (Linnaeus, 1758) for comparison, MNHN.F.C02108 (Salvat coll.), present day, Rurutu island (Tuamotu,French Polynesia),left valve (D) and right valve (E); F-L, Indotellina fakaauensis Merle,n. sp.; F-I, holotype MNHN.F.A99067, external view of the right valve (F), external view of the left valve (G), view of the posterior margin (H), view of the ventral margin (I); J, K, paratype MNHN.F.A99069, internal view of the right valve (J), internal view of the left valve (K); L, paratype MNHN.F.A99824, external view of the right valve. Scale bars: 10 mm. Credits: A-C, F-L, P. Loubry (MNHN/CNRS); D-E, L. Cazes (MNHN/CNRS). A-C, F-L, material from Letourneux coll. housed at MNHN.	FIG. 18. — Tellinidae (A-L) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-C, Tellinella cf.virgata(Linnaeus,1758), MNHN.F.A98454, left valve, internal view in natural light (A), external view in natural light (B) and under UV light (C); D, E, Tellinella virgata (Linnaeus, 1758) for comparison, MNHN.F.C02108 (Salvat coll.), present day, Rurutu island (Tuamotu,French Polynesia),left valve (D) and right valve (E); F-L, Indotellina fakaauensis Merle,n. sp.; F-I, holotype MNHN.F.A99067, external view of the right valve (F), external view of the left valve (G), view of the posterior margin (H), view of the ventral margin (I); J, K, paratype MNHN.F.A99069, internal view of the right valve (J), internal view of the left valve (K); L, paratype MNHN.F.A99824, external view of the right valve. Scale bars: 10 mm. Credits: A-C, F-L, P. Loubry (MNHN/CNRS); D-E, L. Cazes (MNHN/CNRS). A-C, F-L, material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21DFF8BFC12F889FC20FD11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384274/files/figure.png	https://doi.org/10.5281/zenodo.19384274	FIG. 19. —?Gafrarium troendlei Merle, n. sp. from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98456, left valve, internal view in natural light (A), external view in natural light (B), under UV light (C) and in inverse UV light (D); E, F, paratype, MNHN.F.A98457, right valve, internal view (E) and external view (F) in natural light. Scale bars: 10 mm. Credits: A-C, P. Loubry (MNHN/CNRS); E, F, L. Cazes (MNHN/CNRS). Material from Letourneux coll. housed at MNHN.	FIG. 19. —?Gafrarium troendlei Merle, n. sp. from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu, French Polynesia): A-D, holotype, MNHN.F.A98456, left valve, internal view in natural light (A), external view in natural light (B), under UV light (C) and in inverse UV light (D); E, F, paratype, MNHN.F.A98457, right valve, internal view (E) and external view (F) in natural light. Scale bars: 10 mm. Credits: A-C, P. Loubry (MNHN/CNRS); E, F, L. Cazes (MNHN/CNRS). Material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21DFF8BFC12F889FC20FD11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384220/files/figure.png	https://doi.org/10.5281/zenodo.19384220	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	FIG. 7. — Topographical profile and morphological units of the Vaiohua sector from the ocean to the lagoon, Niau Atoll (Tuamotu, French Polynesia).	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21DFF8BFC12F889FC20FD11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384225/files/figure.png	https://doi.org/10.5281/zenodo.19384225	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	FIG. 8. — Topographic profile of the Ofare-Vaiohua sector, northwest sector of Niau Atoll (Tuamotu, French Polynesia) with mollusk assemblages: 1, Plio-Pleistocene; 2, Holocene beds overlying the caliche; 3, Holocene deposits accessible from taro culture pits; 4, Modern lagoon. Caliche in red. The assumption is made that the level of the lagoon is the same as that 0 m of the ocean.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21DFF8BFC12F889FC20FD11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384267/files/figure.png	https://doi.org/10.5281/zenodo.19384267	FIG. 18. — Tellinidae (A-L) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-C, Tellinella cf.virgata(Linnaeus,1758), MNHN.F.A98454, left valve, internal view in natural light (A), external view in natural light (B) and under UV light (C); D, E, Tellinella virgata (Linnaeus, 1758) for comparison, MNHN.F.C02108 (Salvat coll.), present day, Rurutu island (Tuamotu,French Polynesia),left valve (D) and right valve (E); F-L, Indotellina fakaauensis Merle,n. sp.; F-I, holotype MNHN.F.A99067, external view of the right valve (F), external view of the left valve (G), view of the posterior margin (H), view of the ventral margin (I); J, K, paratype MNHN.F.A99069, internal view of the right valve (J), internal view of the left valve (K); L, paratype MNHN.F.A99824, external view of the right valve. Scale bars: 10 mm. Credits: A-C, F-L, P. Loubry (MNHN/CNRS); D-E, L. Cazes (MNHN/CNRS). A-C, F-L, material from Letourneux coll. housed at MNHN.	FIG. 18. — Tellinidae (A-L) from the Plio-Pleistocene assemblage of Niau Atoll (Tuamotu,French Polynesia):A-C, Tellinella cf.virgata(Linnaeus,1758), MNHN.F.A98454, left valve, internal view in natural light (A), external view in natural light (B) and under UV light (C); D, E, Tellinella virgata (Linnaeus, 1758) for comparison, MNHN.F.C02108 (Salvat coll.), present day, Rurutu island (Tuamotu,French Polynesia),left valve (D) and right valve (E); F-L, Indotellina fakaauensis Merle,n. sp.; F-I, holotype MNHN.F.A99067, external view of the right valve (F), external view of the left valve (G), view of the posterior margin (H), view of the ventral margin (I); J, K, paratype MNHN.F.A99069, internal view of the right valve (J), internal view of the left valve (K); L, paratype MNHN.F.A99824, external view of the right valve. Scale bars: 10 mm. Credits: A-C, F-L, P. Loubry (MNHN/CNRS); D-E, L. Cazes (MNHN/CNRS). A-C, F-L, material from Letourneux coll. housed at MNHN.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
CA729441C21DFF8BFC12F889FC20FD11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19384229/files/figure.png	https://doi.org/10.5281/zenodo.19384229	FIG. 9. — Species assemblage of Late Holocene sediments deposited above the caliche (sample N105, Vaihuoa,near Archeological platform),Niau Atoll (Tuamotu, French Polynesia):A-C, Cerithium columna G. B. Sowerby I, 1834;A, MNHN.F.A98459; B, MNHN.F.A98460; C, MNHN.F.A98461; D-F, Cerithium niauense Merle, n. sp.: D, MNHN.F.A98411; E, MNHN.F.A98412; F, MNHN.F.A98413; G-I, Cerithium rostratum G. B. Sowerby II, 1855:G, MNHN.F.A98462; H, MNHN.F.A98463; I, MNHN.F.A98464; J-L, Gafrarium pectinatum (Linnaeus, 1758): J, MNHN.F.C02086; K, MNHN.F.C02087; L, MNHN.F.C02088; M-O, Fragum fragum (Linnaeus, 1758): M, MNHN.F.C02089; N, MNHN.F.C02090; O, MNHN.F.C02091; P-R, Ctena bella (Conrad, 1837): P, MNHN.F.C02092; Q, MNHN.F.C02093; R, MNHN.F.C02094. Scale bars: 10 mm. Credit: J.-P. Pointier.	FIG. 9. — Species assemblage of Late Holocene sediments deposited above the caliche (sample N105, Vaihuoa,near Archeological platform),Niau Atoll (Tuamotu, French Polynesia):A-C, Cerithium columna G. B. Sowerby I, 1834;A, MNHN.F.A98459; B, MNHN.F.A98460; C, MNHN.F.A98461; D-F, Cerithium niauense Merle, n. sp.: D, MNHN.F.A98411; E, MNHN.F.A98412; F, MNHN.F.A98413; G-I, Cerithium rostratum G. B. Sowerby II, 1855:G, MNHN.F.A98462; H, MNHN.F.A98463; I, MNHN.F.A98464; J-L, Gafrarium pectinatum (Linnaeus, 1758): J, MNHN.F.C02086; K, MNHN.F.C02087; L, MNHN.F.C02088; M-O, Fragum fragum (Linnaeus, 1758): M, MNHN.F.C02089; N, MNHN.F.C02090; O, MNHN.F.C02091; P-R, Ctena bella (Conrad, 1837): P, MNHN.F.C02092; Q, MNHN.F.C02093; R, MNHN.F.C02094. Scale bars: 10 mm. Credit: J.-P. Pointier.	2026-04-02	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe		Zenodo	biologists	Salvat, Bernard;Merle, Didier;Letourneux, Jean;Tröndlé, Jean;Montaggioni, Lucien François;Pons-Branchu, Edwige;Dumoulin, Jean-Pascal;Bacchet, Philippe			
