taxonID	type	description	language	source
CB4C87EFF138FFF0E222FE55D245FBEE.taxon	type_taxon	Type species Trochus matonii Payraudeau, 1826 (by subsequent designation of von Martens 1885: 52, in the Zoological Record), accepted as Jujubinus vulgaris (Risso, 1826).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFF0E222FE55D245FBEE.taxon	diagnosis	Diagnosis Shell small for the family, slightly turriculate, imperforate, frequently brightly coloured, with obsolete or blunt columellar notch; sculpture of incised spiral lines with or without prosocline lamellae between cords. Soft parts (Fig. 4) with simplified epipodium, simple neck lobes, three long epipodial tentacles with well-developed, basal macropapillae; complex mottled colouration of foot and epipodium. Radula rhipidoglossate (∞ + 5 + 1 + 5 + ∞: Fig. 5 A – B), typically trochid: rachidian with wide base, sharp, triangular primary cusp and 3 – 5 small, sharp secondary cusps; laterals gradually increasing in size outward, with triangular, sharp apex, and serrated internal and external edges; inner marginals with long, smooth apex, and median secondary cusps; outer marginals progressively pluricuspidate. Grammatical gender Masculine.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFEFE236FBC1D767FA75.taxon	description	SSH I – Fig. 6 A – F	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFEFE236FBC1D767FA75.taxon	diagnosis	Diagnosis Shell of large size for the genus, elevated, thick, 3 – 4 spaced spiral cords, elevated and granulated, broad interspaces with strong prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFEFE236FBC1D767FA75.taxon	materials_examined	Type material Lectotype of Trochus vulgaris Risso, 1826 FRANCE • dd; Nice; unknown locality (Nice, designated by Arnaud 1978); MNHN-IM- 2000 - 31374. Type (s) of Trochus matonii Payraudeau, 1827 CORSICA • “ L’Ile de Corse ”; lost (Virginie Héros, Muséum National d’Histoire Naturelle, France, pers. com., 27 Oct. 2019). Type (s) of Trochus unidentatus Philippi, 1844 SICILY • “ Siciliae ”, probably in MNdHN (Santiago de Chile). Syntypes of Jujubinus striatus var. monterosatoi Bucquoy, Dautzenberg & Dollfus, 1884 FRANCE • 2 dd; Roussillon; MNHN-IM- 2000 - 38199, MNHN-IM- 2000 - 38200. Holotype of Jujubinus africanus Nordsieck, 1973 TUNISIA • dd; Djerba; SMF 341822. Other material examined ALGERIA • 7 dd; Alger; MCZR-M- 11692 • 1 dd; Alger; (labelled “ Jujubinus quadrigranulatus ”); MCZR-M- 11708 • 6 dd; Alger, Bone; MCZR-M- 11720 • 1 dd; Oran; MCZR-M- 11756 • 10 dd; Oran; MCZR-M- 11775 • 2 dd; Oran; “ D’Oran ”; MCZR-M- 11757 • 4 dd; Algeria; MCZR-M- 11758. CORSICA • 3 lv; Petit Pain de Sucre; 42 ° 44.7 ′ N; 9 ° 28.00 ′ E; 19 m depth; used for DNA sequencing; MNHN-IM- 2019 - 4115 to 4117 • 3 lv; off Punta vecchia; 42 ° 59.8 ′ N, 9 ° 27.2 ′ E; 11 m depth; used for DNA sequencing; MNHN-IM- 2019 - 4187 to 4689 • 3 lv; Minelli; 42 ° 42.7 ′ N, 9 ° 27.5 ′ E; 17 m depth; used for DNA sequencing; MNHN-IM- 2019 - 4712 to 4714 • 1 lv; Grand Canyon; 42 ° 44.2 ′ N, 9 ° 28.0 ′ E; 29 m depth; used for DNA sequencing; MNHN-IM- 2019 - 4747 • 1 lv; North Bastia; 42 ° 44.4 ′ N, 9 ° 28.1 ′ E; 20 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5052 • 3 lv; La Louise; 42 ° 41.4 ′ N, 9 ° 27.2 ′ E; 19 – 18 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5366 to 5368 • 1 lv; Le Sphinx; 42 ° 43.3 ′ N, 9 ° 14.9 ′ E; 17 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5398 • 2 lv; same data as for preceding; MNHN-IM- 2019 - 5399, 5408 • 3 lv; Saint Florent; 42 ° 42.3 ′ N, 9 ° 19.2 ′ E; 3 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5497 to 5499 • 1 lv; North Bastia; 42 ° 44.4 ′ N, 9 ° 28.1 ′ E; 20 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5633 • 4 lv; Saint Florent; 42 ° 41.8 ′ N, 9 ° 18.3 ′ E; 10 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5641 to 5644 • 1 lv; Sec de Negru; 42 ° 46.1 ′ N, 9 ° 20.0 ′ E; 29 – 16 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5645 • 1 lv; Chapeau du Gendarme; 42 ° 43.0 ′ N, 9 ° 15.9 ′ E; 29 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5646 • 1 lv; Sec de Negru; 42 ° 46.1 ′ N, 9 ° 20.0 ′ E; 26 m depth; used for DNA sequencing; MNHN- IM- 2019 - 5649 • 2 lv; Saint-Florent; 42 ° 43.0 ′ N, 9 ° 18.6 ′ E; 30 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5669, 5670 • 1 lv; Sec de Giraglia; 43 ° 01.7 ′ N, 9 ° 24.4 ′ E; 21 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5687 • 1 lv; Centuri depths; 42 ° 59.3 ′ N, 9 ° 17.8 ′ E; 21 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5886 • 1 lv; Centuri; 42 ° 57.7 ′ N, 9 ° 20.1 ′ E; 30 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5894 • 9 lv; Sec de Giraglia; 43 ° 01.7 ′ N, 9 ° 24.4 ′ E; 21 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5900, 5901, 5914 to 5920 • 6 lv; Miomo; 42 ° 44.7 ′ N, 9 ° 28.0 ′ E; 16 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6313, 6314, 6343 to 6346 • 3 lv; Punta Di U Capicciolu; 41 ° 25.8 ′ N, 9 ° 15.8 ′ E; 5 – 10 m depth; used for DNA sequencing; MNHN- IM- 2019 - 13092, 13093, 13258 • 2 lv; South of Lavezzi Is.; 41 ° 18.6 ′ N, 9 ° 15.4 ′ E; 15 – 32 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13602, 13603 • 2 lv; off South coasts; 41 ° 21.8 ′ N, 9 ° 11.6 ′ E; 15 – 18 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13605, 13606 • 3 lv; Ficaghjola; 42 ° 15.5 ′ N, 8 ° 39.3 ′ E; 15 m depth; used for DNA sequencing; MNHN-IM- 2019 - 16476, 16477, 16488 • 1 lv; Cargèse Port; 42 ° 07.9 ′ N, 8 ° 35.8 ′ E; 7 m depth; used for DNA sequencing; MNHN-IM- 2019 - 16825 • 2 lv; Punta Scandola; 42 ° 43.2 ′ N, 9 ° 27.9 ′ E; 60 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17050, 17051 • 1 lv; Gulf de Girolata; 42 ° 19.2 ′ N, 8 ° 36 ′ E; 30 m depth; used for DNA sequencing; MNHN- IM- 2019 - 17651 • 1 lv; Punta Palazzu, Phoque-moine; 42 ° 22.6 ′ N, 8 ° 33.1 ′ E; 10 – 15 m depth; used for DNA sequencing; MNHN-IM- 2019 - 18107 • 3 dd; Provence and Corsica; “ Coste di Provenza e Corsica ” “ Jujubinus festivus ”; MCZR-M- 11716. CROATIA • 2 lv; Rab Island; 15 m depth, Posidonia; CS-PM • 3 dd; Dalmazia; (labelled “ Jujubinus mixtus ”); MCZR-M- 11702 • 17 dd; 1906; “ Adriatico ”; (labelled “ Jujubinus mixtus ”); MCZR-M- 11702 • 16 dd; Velirat; MCZR-M- 11707 • 2 dd; Velirat; 1910; MCZR-M- 11707 • 21 dd; Velirat; MCZR-M- 11724 • 9 dd; Velirat; (labelled “ Jujubinus alternans ”); MCZR-M- 11728 • 10 dd; Velirat; (labelled “ Jujubinus matoni mixta ”); MCZR-M- 11750 • 1 dd; Velirat; MCZR-M- 11770 • 5 dd; Dalmazia; (labelled “ Jujubinus monterosati ”); MCZR-M- 11724 • 4 dd; Dalmazia; MCZR-M- 11725 • 12 dd; Dalmazia; (labelled “ Jujubinus fusca ”); MCZR-M- 11728 • 12 dd; Melada; MCZR-M- 11740 • 6 dd; Dalmazia; (labelled “ Jujubinus peculiaris ”); MCZR-M- 11746 • 3 dd; Dalmazia; MCZR-M- 11747 • 1 dd; Puntebianche; (Dalmazia); MCZR-M- 11756 • 1 dd; Velirat; MCZR-M- 11770. FRANCE • 2 dd; Nice; MCZR-M- 11720 • 4 dd; Toulon; MCZR-M- 11745 • 6 dd; Toulon; MCZR-M- 11761 • 6 dd; Bandol; ‘ Jujubinus elenchoides ’; MCZR-M- 11771 • 8 dd; Cannes; MCZR-M- 11758 • 7 dd; Cannes; MCZR-M- 11769 • 1 dd; Corsica; MCZR-M- 11735 • 1 lv; Saint-Raphaël, Varo; used for DNA sequencing; BAU 2661.1 • 1 lv; Calanque de Figuerolles, La Ciotat; 36 ° 34 ′ 18.7 ″ N, 26 ° 30 ′ 14.3 ″ E; used for DNA sequencing; MNHN-IM- 2013 - 32744. GREECE • 9 dd; Gulf of Preveza; MCZR-M- 11748 • 2 dd; Greece; (labelled “ Jujubinus tricolor ”); MCZR-M- 11750 • 11 dd; Patras; MCZR-M- 11751 • 2 dd; Patras; MCZR-M- 11761 • 10 dd; Conemenos; MCZR-M- 11707 • 1 dd; Kefalonia Island, Assos, port; 3 m depth; LT • 2 lv; Analipsi, Astypalea Is.; 36.575946 ° N, 26.393070 ° E; 0 – 1 m depth Posidonia; 28 – 30 Jun. 2017; used for DNA sequencing; BAU 1374, 1376 • 2 lv; Marmàri, Astypalea Is.; 36.564676 ° N, 26.353297 ° E; 0.2 – 0.5 m depth Posidonia; 23 Jun. 2017; used for DNA sequencing; BAU 3038.1, 3028.2 • 1 lv; same data as for preceding; used for DNA sequencing; BAU 3039.1 • 1 lv; Analipsi, Astypalea Is.; 36.575946 ° N, 26.393070 ° E; 0 – 1 m depth Posidonia; 28 – 30 Jun. 2017; used for DNA sequencing; BAU 3042.1 • 1 lv; Halkidiki, Marmaras; 40 ° 04 ′ 46.2 ″ N, 23 ° 45 ′ 48.7 ″ E; used for DNA sequencing; MNHN-IM - 2013 - 32749. ITALY • 1 lv; Giannutri Is., Punta Secca, stn. GMM 2; 42 ° 15 ′ 41 ″ N, 11 ° 06 ′ 24 ″ E; 12 – 13 m depth; used for DNA sequencing; BAU 2668.1 • 2 lv; Giannutri Is.; 42 ° 15 ′ 7.05 ″ N, 11 ° 05 ′ 31.51 ″ E; used for DNA sequencing; RM 3 - Jex 2, RM 3 - Jex 3 • 4 lv; same data as for preceding; used for DNA sequencing; BAU 1368.1 to 1368.4 • 1 lv; same data as for preceding; used for DNA sequencing; RM 3 - Jex 1 • 2 lv; Giglio Is.; 42 ° 22 ′ 7.82 ″ N, 10 ° 55 ′ 5.11 ″ E; used for DNA sequencing; RM 3 - Jex 4, RM 3 - Jex 5 • 3 lv; Giglio Is., Cala Cupa; 42 ° 22 ′ 06 ″ N, 10 ° 55 ′ 12 ″ E; used for DNA sequencing; BAU 2851.1 to 2851.3 • 2 lv; same data as for preceding; used for DNA sequencing; BAU 2910.1, 3 • 1 lv; Argentario, La Caletta; 42 ° 26 ′ 40.73 ″ N, 11 ° 6 ′ 51.84 ″ E; used for DNA sequencing; RM 3 - Jex 10 • 1 lv; same data as for preceding; used for DNA sequencing; RM 3 - Jex 9 • 2 lv; Tarquinia; 42 ° 26 ′ 40.73 ″ N, 11 ° 06 ′ 51.84 ″ E; used for DNA sequencing; RM 3 - Jex 11, RM 3 - Jex 12 • 2 lv; Elba Is., Fetovaia; 42 ° 43 ′ 58.1 ″ N, 10 ° 09 ′ 13.1 ″ E; used for DNA sequencing; MNHN-IM- 2013 - 32666, 32667 • 13 dd; Elba Island, Lacona; 5 m depth; bioclastic sediment; CS-PM • 29 dd; Elba I., Porto Azzurro, Naregno; 3 m depth; bioclastic sediment; LT • 3 dd; Talamone, beached; CS-PM • 2 dd; Isola del Giglio; MCZR-M- 11761 • 13 lv; Giglio Island; 15 m depth; Posidonia meadow; CS-PM • 13 lv; Giannutri Island; 15 m depth; Posidonia meadow; CS-PM • 34 dd; Torre Valdaliga; 25 m depth; bioclastic sediment; CS-PM • 27 dd; Santa Marinella; 20 m depth; bioclastic sediment; CS-PM • 250 dd; Santa Marinella; 0 m depth; beached, in bioclastic sediments; CS-PM • 71 dd; Santa Marinella; fishing boat; CS-PM • 5 dd; Santa Marinella; fishing nets, Posidonia residues; LT • 600 dd; Torre Valdaliga, Civitavecchia; 25 m depth; CS-PM • 9 dd; San Felice Circeo; 30 m depth; fishing nets; LT • 4 dd; San Felice Circeo; fishing nets, Posidonia residues; LT • 1 dd; San Felice Circeo; 40 m depth; fishing nets; LT • 1 dd; Anzio; MCZR-M- 11733 • 50 dd; Ventotene Island; 18 – 20 m depth; CS-PM • 2 dd; Ventotene Island; fishing boat; CS-PM • 63 dd; Ventotene Island; 20 m depth; bioclastic sediment; CS-PM • 3 dd; Napoli; MCZR-M- 11684 • 5 dd; Napoli; MCZR-M- 11720 • 4 dd; Napoli; MCZR-M- 11723 • 1 dd; “ Napoli? ”; MCZR-M- 11724 • 1 dd; Napoli; (labelled “ Jujubinus histris ”); MCZR-M- 11724 • 12 dd; Napoli; (labelled “ Jujubinus histrionella ”); MCZR-M- 11733 • 3 dd; Napoli; (labelled “ Jujubinus matoni mitis ”); MCZR-M- 11756 • 1 dd; Napoli; MCZR-M- 11733 • 11 dd; Napoli; MCZR-M- 11733 • 1 dd; Napoli; “ Jujubinus matoni ”; MCZR-M- 11733 • 27 dd; Napoli; MCZR-M- 11755 • 3 dd; Napoli; amidst sponges (“ spugne ”); MCZR-M- 11758 • 1 dd; Napoli; 1918; “ Jujubinus micropictus ”; MCZR-M- 11760 • 1 dd; Napoli; MCZR-M- 11733 • 5 dd; Napoli; MCZR-M- 11760 • 5 dd; Napoli?; MCZR-M- 11776 • 1 dd; Napoli and Capri; MCZR-M- 11733 • 8 dd; Capri; MCZR-M- 11758 • 7 dd; Capri; MCZR-M- 11759 • 1 dd; Capri; “ Jujubinus matoni ”; MCZR-M- 11733 • 1 dd; Capri; MCZR-M- 11735 • 1 dd; Napoli; MCZR-M- 11733 • 4 dd; off Monte Conero; 0 m depth; beached; CS-PM • 67 dd + 13 dd; Marina di Camerota; 8 m depth; bioclastic sediment; LT • 1 dd; Brindisi; 3 m depth; algae; LT • 1 dd; Otranto; beached; CS-PM • 8 dd; Le Castella; 25 m depth; bioclastic sediment; CS-PM • 8 dd; Adriatic; “ Jujubinus similis ”; MCZR-M- 11740 • 3 dd; Adriatic; “ Adriatico ” “ Jujubinus mixtus ”; MCZR-M- 11769 • 4 dd; Adriatic; “ Jujubinus tristis ”; MCZR-M- 11756 • 6 dd; Adriatic; MCZR-M- 11758 • 1 dd; Adriatic?; “ Adriatico? ”; MCZR-M- 11762 • 3 dd; Trieste; MCZR-M- 11735 • 3 dd; Trieste; MCZR-M- 11747 • 1 dd; Porto Maurizio; “ Jujubinus insignis ”; MCZR-M- 11746 • 3 dd; Porto Maurizio; MCZR-M- 11746 • 1 dd; Porto Maurizio; 1917; MCZR-M- 11761 • 2 dd; Porto Maurizio; MCZR-M- 11766 • 4 dd; Chioggia; MCZR-M- 11740 • 34 dd; unidentified locality; “ Romagnolo ”; MCZR-M- 11691 • 2 dd; Molfetta; beached; CS-PM • 43 dd; Tricase Marina; 3 m depth; bioclastic sediment; CS-PM • 8 lv; Polignano; 20 m depth; amidst Posidonia; CS-PM • 80 dd; Santa Maria di Leuca; 15 m depth; CS-PM • 1 lv; S. Isidoro; 3 m depth; used for DNA sequencing; BAU 1678. MALTA • 14 lv; Gozo Island, Marsalforn, Xwejni Bay; 36 ° 04 ′ 47.83 ″ N, 14 ° 14 ′ 51.25 ″ E; CS-PM • 13 lv; Gozo Island, Xlendi Bay; 5 m depth; Cystoseira; CS-PM • 6 dd; Malta; MCZR-M- 11721 • 5 lv; Malta; used for DNA sequencing; BAU 2674.1 – 5. MOROCCO • 6 dd; Casablanca, Mazata, Mazaclon; MCZR-M- 11767. PORTUGAL • 5 dd; Algarve; fishing boat; CS-PM. SARDINIA • 4 lv; Capo Carbonara; 5 m depth; MO • 10 lv; Cala d’Ambra; 1 – 5 m depth; MO • 22 dd; Golfo d’Arzachena; beached; CS-PM • 35 dd; La Maddalena Island; 25 m depth; bioclastic sediment; CS-PM • 1 dd; Bay of Cagliari; (labelled “ Jujubinus limulosus ”); MCZR-M- 11712 • 1 dd; Alghero; MCZR-M- 11718 • 2 dd; Alghero; MCZR-M- 11756 • 1 dd; Cagliari; MCZR-M- 11720 • 2 dd; Sardinia; MCZR-M- 11733 • 1 dd; Sardinia; labelled “ undulata ”; MCZR-M- 11738 • 3 dd; Alghero; MCZR-M- 11740 • 5 dd; Sardinia; MCZR-M- 11742 • 1 dd; Sardinia; MCZR-M- 11746 • 14 dd; Sardinia; MCZR-M- 11749 • 8 dd; Sardinia; MCZR-M- 11757 • 8 dd; Sardinia; MCZR-M- 11761 • 4 dd; Sardinia; MCZR-M- 11761 • 3 dd; Sardinia; “ Jujubinus tricolor ”; MCZR-M- 11761 • 8 lv; San Pietro Is.; 3 m depth; on Posidonia; CS-PM • 4 dd; Cagliari; MCZR-M- 11733. SICILY • 22 dd; Palermo; “ Jujubinus tricolor ”; MCZR-M- 11709 • 44 dd; Palermo; (labelled “ Jujubinus matoni ”); MCZR-M- 11717 • 1 dd; Palermo; MCZR-M- 11718 • 94 dd; Port of Palermo; (labelled “ Jujubinus matoni ”); MCZR-M- 11719 • 6 dd; Palermo; (labelled “ Jujubinus giacobi ”); MCZR-M- 11720 • 3 dd; Palermo; “ fondo coralligeno ”; MCZR-M- 11723 • 1 dd; Palermo; MCZR-M- 11724 • 6 dd; Palermo; MCZR-M- 11735 • 15 dd; Palermo; MCZR-M- 11745 • 9 dd; Palermo; MCZR-M- 11749 • 1 dd; Palermo; MCZR-M- 11749 • 7 dd; Palermo; MCZR-M- 11758 • 2 dd; Palermo; (labelled “ Jujubinus tricolor ”); MCZR-M- 11763 • 2 dd; Sciacca; MCZR-M- 11773 • 3 dd; Favignana I.; beached; CS-PM • 7 dd; Favignana Is.; MCZR-M- 11749 • 1 dd; Linosa I.; 2 m depth; Posidonia meadow; CS-PM • 1 dd; Lampedusa; 1901; (“ spugne ”); MCZR-M- 11723 • 1 dd; Lampedusa; (labelled “ Jujubinus disperses ”); MCZR-M- 11731 • 5 dd; Lampedusa; beached; CS-PM • 3 dd; Pantelleria Is., Scauri; 13 m depth; bioclastic sediment; LT • 1 dd; Ognina; MCZR-M- 11721 • 1 dd; Ognina; MCZR-M- 11759 • 3 dd; Messina; (labelled “ Jujubinus matoni rupestris ”); MCZR-M- 11721 • 9 dd; Messina; (labelled “ Jujubinus rupestris ”); MCZR-M- 11721 • 2 dd; Messina; MCZR-M- 11748 • 1 dd; Messina; MCZR-M- 11749 • 2 dd; Messina; MCZR-M- 11769 • 3 dd; Messina; (labelled “ Jujubinus incertus ”); MCZR-M- 11776 • 16 dd; Lipari; MCZR, no number • 1 dd; Lipari; (labelled “ Jujubinus matoni grossesculpta ”); MCZR-M- 11738 • 20 dd; Lipari; (labelled “ Jujubinus inflammatus ”); MCZR-M- 11742 • 11 dd; Palermo; “ J. matoni Pay Palermo! = vulgaris Risso ”; MCZR-M- 11685 • 5 lv; San Vito Lo Capo; used for DNA sequencing; BAU 1677.2 – 6. SLOVENIJA • 2 lv; Izola; 45 ° 32 ′ 51.4 ″ N, 13 ° 42 ′ 12.9 ″ E; 4 m depth; on Posidonia meadow; used for DNA sequencing; RM 3 - Jex 7, RM 3 - Jex 8 • 1 lv; Izola; 45 ° 32 ′ 51.4 ″ N, 13 ° 42 ′ 12.9 ″ E; 4 m depth; on Posidonia meadow; used for DNA sequencing; BAU 2071.1. SPAIN • 1 lv; Cabo de Palos, Murcia; 30 m depth; CS-PM • 3 dd; Tarifa; intertidal; tide pools, crabbed; CS-PM • 5 dd; Ibiza; “ Jujubinus monterosatoi ”; MCZR-M- 11707 • 29 dd; Velirat and Valencia; vial with unique label, reading “ Jujubinus tricolor ”; MCZR-M- 11807 • 31 dd; Valencia; 1915; “ Jujubinus matoni ”; MCZR-M- 11734 • 13 dd; Valencia; 1915; “ Jujubinus matoni ”; MCZR-M- 11734 • 8 dd; Valencia; “ Jujubinus crenulatus ”; MCZR-M- 11749 • 2 dd; Mahón; MCZR-M- 11748 • 5 dd; Valencia; “ Jujubinus tricolor ”; MCZR-M- 11763 • 35 dd; Cádiz, Santa María; intertidal; tide beds; CS-PM • 1 lv; Benalmádena, Málaga; 36 ° 34.7 ′ N, 4 ° 30.9 ′ W; used for DNA sequencing; BAU 2665 • 1 lv; Cabo de Palos, Murcia; 37 ° 38 ′ 08.0 ″ N, 0 ° 41 ′ 30.4 ″ W; used for DNA sequencing; MNHN-IM- 2013 - 32680 • 1 lv; Almería; used for DNA sequencing; RM 3 - Jex 13 • 1 lv; Tarifa; 36 ° 0 ′ 14.16 ″ N, 5 ° 36 ′ 21.32 ″ W; used for DNA sequencing; RM 3 - Jex 6. TUNISIA • 3 dd; Sfax; MCZR-M- 11684 • 1 dd; “ Coste d’Africa ”; MCZR-M- 11746 • 8 dd; “ Coste d’Africa ”; MCZR-M- 11761 • 21 dd; Sfax; MCZR-M- 11761 •> 30 dd; North of Zarzis; 33 ° 36 ′ 28 ″ N, 11 ° 04 ′ 04 ″ E; 0 – 7 m depth; MO • 2 dd; Djerba; 1 m depth; MO • 49 dd; Kerkennah Islands, Sidi Yousef; beached; CS-PM • 89 dd; Djerba Is.; beached; CS-PM • 3 dd; Djerba Island; Beached; LT • 7 dd; Djerba Island; upper infralittoral; ex Katy Nicolay coll; LT • 6 dd; Kerkennah Islands, W Gharbi Island, Sidi Youssef; 0.5 m depth; LT • 3 dd; Djerba Island; ex Mauro Pizzini coll; LT • 2 lv; Gremdi Is.; 34 ° 45 ′ 50 ″ N, 11 ° 19 ′ 55 ″ E; used for DNA sequencing; BAU 2313.1, 2313.2 • 1 lv; Gharbi Is., Sidi Youssef; 34 ° 45 ′ 50 ″ N, 11 ° 19 ′ 55 ″ E; used for DNA sequencing; BAU 2315.1 • 1 lv; Djerba Is., Sidi Solimen; 34 ° 02 ′ N, 11 ° 05 ′ E; used for DNA sequencing; MNHN-IM- 2013 - 31944 • 1 lv; Djerba Is., Sidi Jmour; 33 ° 49.88 ′ N, 10 ° 44.83 ′ E; used for DNA sequencing; MNHN-IM- 2013 - 32107 • 1 lv; Djerba Is., Sidi Jmour; 33 ° 49.88 ′ N, 10 ° 44.83 ′ E; used for DNA sequencing; MNHN-IM- 2013 - 32121 • 2 lv; Kerkennah Is., Sharqi Is.; 34 ° 45 ′ 16.97 ″ N, 11 ° 20 ′ 31.00 ″ E; used for DNA sequencing; RM 3 - Jk 1 A, RM 3 - Jk 2 A. TURKEY • 2 lv; Bozcaada Island; 45 m depth; CS-PM • 4 lv; Datça, stn AKD 92 - 02; MO • 10 dd; Datça; 36 ° 42 ′ 16.6 ″ N, 27 ° 40 ′ 52.3 ″ E; 8 m depth; under stones and brown algae; LT • 12 dd; Datça; 36 ° 42 ′ 16.6 ″ N, 27 ° 40 ′ 52.3 ″ E; 3 – 7 m depth; on brown algae; LT. UNPRECISED GEOGRAPHIC DATA • 5 dd; Mediterranean Sea; (labelled “ Jujubinus matoni ”); MCZR-M- 11761 • 3 dd; no data; (labelled “ Jujubinus matoni ”); MCZR-M- 11750.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFEFE236FBC1D767FA75.taxon	description	Description (in parentheses data of the lectotype) SHELL. Shell of large size for the genus, height 5.9 – 12.6 (10.2) mm, width 4.7 – 7.3 (6.7) mm, height / width ratio 1.25 – 1.86, conical, robust. Protoconch seemingly smooth, of ca 1 whorl, 0.21 – 0.22 mm in width. Teleoconch of 6 – 7 (6) slightly convex whorls. Sculpture of 3 – 5 (3) minutely granulose little closely set abapical spiral cords with wide interspaces of about the same strength, and peripheral cord evident formed by 1 – 5 (4) merged cordlets. Interspaces wide with strong prosocline lamellae. Suture incised. Base convex, with 6 – 9 concentric reddish or brown spiral cords white dotted and regularly spaced, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls reddish-greyish; remaining whorls and base white or whitish background with reddish brown to dark-brown or black prosocline or opisthocline lines or blotches of varying size irregularly arranged. SOFT PARTS. Epipodial tentacles grayish with a whitish stripe, covered by scattered very tiny whitish speckles. Foot background brownish or reddish mottled with longitudinal brown or red stripes and covered by white speckles; sole whitish or creamy. Cephalic tentacles grayish, with two dorsal darker stripes and covered by small light whitish speckles. Eye stalks brownish dorsally, whitish ventrally, covered by brown speckles; eyes black encircled by lighter crown, sometimes white ventrally. Snout brownish to reddish with a creamy terminal band.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFEFE236FBC1D767FA75.taxon	distribution	Distribution In the Atlantic it is known with certainty only from Morocco, and Algarve (Portugal). In almost the entire Mediterranean Sea (e. g., Barash & Danin 1992: 34 – 35, fig. 31 a; Portalatina 2008). Almost all records under Jujubinus exasperatus, living specimens at dephts from 1 to 35 m, rarely down to 60 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF138FFEFE236FBC1D767FA75.taxon	discussion	Remarks Records in the Atlantic from Scotland and the southernmost British Isles (Poppe & Goto 1991) to Azores, Madeira, Canary Islands (Bucquoy et al. 1882; Nordsieck 1968) are probably based on misidentifications. This is the commonest species of the genus in the Mediterranean Sea, very variable morphologically and chromatically. The name Jujubinus exasperatus has been largely employed for it, but Trochus exasperatus Pennant, 1777 was based on a different species (SSH II, see below). Therefore, many literature records of ‘ Jujubinus exasperatus ’ cannot be unequivocally ascribed to either species. Trochus vulgaris Risso, 1826 is the first available name for this species. Trochus matonii Payraudeau, 1827 is also this same species, and is the type of the genus Jujubinus: the type material (now lost) from Corsica was described as “ maculis longitudinalibus angulosis albis et fuscis, aut albis et rubris picta ” (Payraudeau 1827: 126 – 127), clearly referring to two of the most common chromatic forms, with angular longitudinal blotches coloured in either dark and white, or red and white. For the date of publication of Payraudeau’s “ Catalogue descriptif ... ” we refer to Falkner et al. (2002). The two figured specimens of Trochus monterosatoi (Bucquoy, Dautzenberg & Dollfus, 1884: pl. xliii figs 18 – 19) are clearly J. vulgaris, and the very B. D. D. put in the synonymy of their taxon also Trochus exasperatus var. excavata Monterosato, 1880. Jujubinus unidentatus (Philippi, 1844) and Jujubinus africanus Nordsieck, 1973 were based on a particular morphotype from Tunisia with light shell, ivory-whitish background with dark-brown to brown-reddish prosocline elongated flames, 5 – 8 weakly to minutely granulose spiral cords of varying size, base with 7 – 10 spiral cords with brown-reddish lines, and a very strong notch on the columella. The animal of this form has the same general chromatic pattern as the typical one but is homogeneously lighter and much fainter. The morphotype ‘ unidentatus attains a height of 7.8 – 10.1 mm, width 4.7 – 6.2 (ratio 1.42 – 1.86). See under J. exasperatus and J. pseudogravinae, for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF127FFE8E253FA57D14FF860.taxon	diagnosis	Diagnosis Shell of large size for the genus, thick, elevated, 3 – 4 spaced spiral cords, elevated and granulated, broad interspaces with strong prosocline lamellae, peripheral cord evident, high height / width ratio, colour reddish with whitish sagittal flammule.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF127FFE8E253FA57D14FF860.taxon	materials_examined	Type material Trochus exasperatus Pennant, 1777 The original description was based on Martini Lister’s illustration (Martini Lister 1770: pl. 616 fig. a 2) and the relevant material (from the type locality ‘ England’: Martini Lister marked the figure with “ A ”, as for an English shell) is now lost (A. Salvador, Natural History Museum, pers. com., 28 Jul. 2015; M. Carnall, Oxford University Zoology Museum, pers. com. 18 May 2018). A neotype designation based on a genotyped English specimen is desirable to stabilize the use of the name. Syntypes of Jujubinus corallinus Monterosato, 1884 SICILY • 5 dd; Palermo; NMW. 1910.29.02042. Syntypes of Jujubinus igneus Sturany, 1896 CROATIA • 3 dd; Pelagruza, Pola Expedition stn 260; 128 m depth; NHMW 72399. Other material examined CANARY ISLANDS • 1 dd; Gran Canaria, Sardina; 20 m depth; MO. CORSICA • 1 lv; Miomo; 42 ° 45.2 ′ N, 9 ° 28.4 ′ E; 23 m depth; used for DNA sequencing; MNHN- IM- 2019 - 4178 • 2 lv; Minelli; 42 ° 42.7 ′ N, 9 ° 27.5 ′ E; 17 m depth; used for DNA sequencing; MNHN- IM- 2019 - 4711, 4713 • 1 lv; East Cap Corse; 42 ° 53.6 ′ N, 9 ° 30.3 ′ E; 40 m depth; used for DNA sequencing; MNHN-IM- 2019 - 4737 • 1 lv; La Louise, 42 ° 41.4 ′ N, 9 ° 27.2 ′ E, 19 – 18 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5369 • 2 lv; Gulf de Saint-Florent; 42 ° 44.1 ′ N, 9 ° 19.3 ′ E; 60 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5677, 5887 • 2 lv; Ficaghjola; 42 ° 41.4 ′ N, 9 ° 27.1 ′ E; 18 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6341, 6342 • 1 lv; North Cap Corse; 43 ° 02.6 ′ N, 9 ° 19.9 ′ E; 60 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6377 • 1 lv; Est Ile Forana, 41 ° 33.7 ′ N, 9 ° 23.7 ′ E; 45 – 46 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13063 • 1 lv; Tête de Cheval; 41 ° 20.3 ′ N, 9 ° 14.5 ′ E; 44 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13551 • 1 lv; Cabo Di Feno; 41 ° 23.4 ′ N, 9 ° 06.2 ′ E; 37 m depth; used for DNA sequencing; MNHN- IM- 2019 - 13691 • 2 lv; Punta Di U Capicciolu; 41 ° 25.7 ′ N, 9 ° 15.9 ′ E; 12 – 36 m depth; used for DNA sequencing; MNHN-IM- 2019 - 14094, 14095 • 1 lv; Madonetta lighthouse; 41 ° 23.2 ′ N, 9 ° 08.4 ′ E, 37 m depth; used for DNA sequencing; MNHN-IM- 2019 - 14626 • 2 lv; Gulf de Girolata, 42 ° 19.8 ′ N, 8 ° 33.7 ′ E, 46 – 46 m depth; used for DNA sequencing; MNHN-IM- 2019 - 16457, 16565 • 1 lv; South of Cargèse; 42 ° 07.2 ′ N, 8 ° 35.4 ′ E; 45 – 48 m depth; used for DNA sequencing; MNHN-IM- 2019 - 16897 • 1 lv; Punta Scandola; 42 ° 20 ′ N, 8 ° 33.6 ′ E; 35 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17047 • 1 lv; N of Gargalo Is.; 42 ° 22.3 ′ N, 8 ° 32.4 ′ E; 26 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17270 • 1 lv; Gulf de Girolata, Ruzaghiu; 42 ° 20.6 ′ N, 8 ° 34.9 ′ E; 60 m depth; used for DNA sequencing; MNHN- IM- 2019 - 17535 • 2 lv; Punta Palazzu; 42 ° 22.8 ′ N, 8 ° 33.2 ′ E; 50 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17613, 17614 • 2 lv; Gulf de Girolata, Ruzaghiu; 42 ° 20.6 ′ N, 8 ° 34.9 ′ E; 60 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17645, 17821. CROATIA • 27 dd; Velirat; “ Juj. Igneus v. coccinella Velirat 1910 tra laminarie e coralligeno ”; MCZR-M- 11733. CYPRUS • 5 dd; Agia Napa; Cape Greco; 34 ° 58 ′ 49.0 ″ N, 34 ° 01 ′ 02.0 ″ E; 3 m depth; MO. FRANCE • 1 lv; Brittany, Guérhéon, near Roscoff; used for DNA sequencing; BAU 2067.5, 2067.6 • 5 lv; Brittany, Pointe de l’Arcouest, Ploubazlanec; 48 ° 49 ′ 10 ″ N 3 ° 00 ′ 26 ″ W; used for DNA sequencing; BAU 2660.1 to 2660.5 • 3 dd; Brest; MCZR-M- 11718 • 2 dd; St Lunaire; MCZR-M- 11718 • 1 lv; Saint-Raphaël (Varo); BAU 2661.2 • 1 lv; La Ciotat, Calanque de Figuerolles; 36 ° 34 ′ 18.7 ″ N, 26 ° 30 ′ 14.3 ″ E; used for DNA sequencing; MNHN-IM - 2013 - 32743 • 1 lv; Saint-Raphaël (Varo); used for DNA sequencing; BAU 2661.2. GREAT BRITAIN • 4 dd; England; MCZR-M- 11718 • 8 dd; Guernsey I.; MCZR-M- 11718 • 4 dd; Herm I.; MCZR-M- 11749. GREECE • 1 dd.; Astypalea I.; 10 m depth; fishing nets; LT • 2 lv; Halkidiki, Marmaras; 40 ° 04 ′ 46.2 ″ N, 23 ° 45 ′ 48.7 ″ E; MNHN-IM- 2013 - 32746, 32751 • 4 lv; Astypalea I., Analipsi; 36.575946 ° N, 26.393070 ° E; 0 – 1 m depth; Posidonia oceanica meadow; 28 – 30 Jun. 2017; used for DNA sequencing; BAU 3040.1 to 3040.4 • 2 lv; same data as for preceding; used for DNA sequencing; BAU 3041.1, 3041.2 • 5 lv; same data as for preceding; used for DNA sequencing; BAU 3044.1 to 3044.4, 3044.6 • 1 lv; Astypalea I., Stenò; 36.574897 ° N, 26.367907 ° E; 2 – 8 m depth; Posidonia oceanica meadow; 29 Jun. 2017; used for DNA sequencing; BAU 3043. ITALY • 1 lv; Elba I., Fetovaia; 42 ° 43 ′ 58.1 ″ N, 10 ° 09 ′ 13.1 ″ E; used for DNA sequencing; MNHN- IM- 2013 - 32663 • 1 dd; Elba I., Lacona; 5 m depth; bioclastic sediment; CS-PM • 3 dd; Elba I., Porto Azzurro Naregno; 3 m depth; bioclastic sediment; LT • 2 lv; Giglio I.; 42 ° 22 ′ 7.82 ″ N, 10 ° 55 ′ 5.11 ″ E; used for DNA sequencing; RM 3 - Jec 3, RM 3 - Jec 4 • 1 lv; Giglio I., Cala Cupa; 42 ° 22 ′ 06 ″ N, 10 ° 55 ′ 12 ″ E; BAU 2910.2 • 2 lv; Argentario, Secca del Corallo; 42 ° 24 ′ 3.90 ″ N, 11 ° 5 ′ 31.57 ″ E; used for DNA sequencing; RM 3 - Jec 1, RM 3 - Jec 2 • 6 dd; Santa Marinella; fishing nets Posidonia residues; LT • 20 dd; Santa Marinella; 20 m depth; bioclastic sediment; CS-PM • 8 dd; Torre Valdaliga; 25 m depth; bioclastic sediment; CS-PM • 2 lv; Secche di Tor Paterno MPA; 41 ° 36 ′ 27.01 ″ N, 12 ° 20 ′ 49.97 ″ E; used for DNA sequencing; RM 3 - Jec 5, RM 3 - Jec 6 • 1 dd; San Felice Circeo; fishing nets; LT • 3 dd; San Felice Circeo; fishing nets Posidonia residues; LT • 18 dd; Capo D’Anzio, 3 miles off; 40 m depth; bioclastic sediment; CS-PM • 4 lv; Giglio I.; 15 m depth; Posidonia; CS-PM • 7 lv; Giannutri I.; 15 m depth; Posidonia; CS-PM • 2 dd; Ventotene I.; 20 m depth; bioclastic sediment; CS-PM • 2 dd; Marina di Camerota; fishing boat; CS-PM • 16 dd; Porto Cesareo; 20 m depth; fishing nets; LT • 1 dd; Tricase Marina; 3 m depth; bioclastic sediment; CS-PM • 2 dd; Argentario, Capo d’Uomo; 35 m depth; MO • 2 dd; Giannutri I., Punta Secca; 35 m depth; MO • 10 dd; Tor Paterno MPA; 15 – 70 m depth; MO • 7 sh; Ventotene I.; 40 m depth; fishing nets residuals; MO • 1 dd; Ventotene I., Sconciglie shoal; 31 m depth; MO • 3 lv; Punta Prosciutto; 40 ° 17 ′ 37 ″ N, 17 ° 45 ′ 47 ″ E; used for DNA sequencing; BAU 2658.2, 2658.3, 2658.5 • 2 lv; Torre Colimena; 40 ° 17 ′ 40 ″ N, 017 ° 45 ′ 20 ″ E; 2 – 7 m depth; Aug. 2007; under rocks; used for DNA sequencing; BAU 1726.1, 1726.3 • 1 lv; Scilla; used for DNA sequencing; BAU 2357.1 • Santa Maria al Bagno; 40 ° 07 ′ 40.9 ″ N, 17 ° 59 ′ 37.5 ″ E; used for DNA sequencing; BAU 3617 • 1 lv; S. Isidoro, Lecce; used for DNA sequencing; BAU 3620.1. MALTA • 3 lv; Gozo I., Xlendi Bay; 10 m depth; Posidonia oceanica meadow; CS-PM. MOROCCO • 1 dd; Al Hoceima, Cala Iris; 15 m depth; MO. SARDINIA • 7 lv; Alghero; 40 – 80 m depth; fishing nets residuals; MO • 1 lv; Bonifacio Strait; 80 m depth; MO • 8 dd; Capo Caccia, ‘ Ennio Falco’ cave; 5 – 15 m depth; MO • 7 dd; Bosa Marina; 40 m depth; bioclastic sediment; CS-PM • 25 dd; La Maddalena I.; 25 m depth; bioclastic sediment; CS-PM • 4 dd; Asinara I., Punta La Nave; 15 – 30 m depth; MO • 5 dd; La Maddalena I.; 80 m depth; fishing nets; LT. SICILY • 10 dd; Lampedusa I.; 50 m depth; MO • 3 lv; Salina I.; 100 – 150 m depth; fishing nets residuals; MO • 4 dd; Lampedusa I.; 130 m depth; MO • 1 dd; Favignana I.; beached; CS-PM • 4 dd; Siracusa; 70 m depth; fishing nets residuals; MO • 6 dd; Levanzo I.; 31 m depth; MO • 171 dd; Palermo; “ J. igneus M. apud Sturany Palermo ”, “ JuJ. Igneus Monts vide Sturany Zool. Ergebu. VII, 1896 p. 28, t. II, f. 45 Palermo ”; unnumbered sample (Monterosato coll.) • 26 dd; Palermo; “ Sbarra ”, (locality in the Palermo area, currently not identifiable); MCZR-M- 11723. SLOVENIJA • 1 lv; Izola; 45 ° 32 ′ 51.4 ″ N, 13 ° 42 ′ 12.9 ″ E; 4 m depth; on Posidonia meadow; used for DNA sequencing; RM 3 - Jec 8. SPAIN • 1 dd; Alborán I., “ Mediterraneo- 92 ” stn ALB A; 35 ° 56.94 ′ N, 003 ° 00.90 ′ W; 32 m depth; MO • 18 dd; Getares, 4 km S of Algeciras; 20 m depth; LT • 1 lv; Almería; used for DNA sequencing; RM 3 - Jec 7 • 5 lv; Torre de Calahonda, Cabopino; 36 ° 28.3 ′ N, 4 ° 42.4 ′ W; BAU 2666.1 to 2666.5. TURKEY • 3 dd; Dardanelles Channel; 65 m depth; CS-PM • 1 dd; Istanbul, Yesilkoy; 55 m depth; CS-PM • 2 dd; Dardanelles Channel; 65 m depth; CS-PM.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF127FFE8E253FA57D14FF860.taxon	description	Description SHELL. Shell of large size for the genus, height 6.0 – 12.2 mm, width 4.6 – 8.2 mm, height / width ratio 1.32 – 1.78, conical, robust. Protoconch seemingly smooth, of ca 1 whorl, 280 – 290 μm in width. Teleoconch of 5.5 – 6.5 slightly convex whorls. Sculpture of 3 – 4 strong granulose spiral cords of about the same strength, regularly spaced, and a broad peripheral cord formed by 2 – 3 merged cordlets, very granulose; suture well-marked. Interspaces with strong prosocline lamellae regularly set. Base convex, with 6 – 8 concentric reddish and white spiral cords regularly spaced, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish or pinkish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls glossy red or pinkish; remaining whorls and base with alternating whitish-reddish spiral blotches more or less of the same size. SOFT PARTS. Epipodial tentacles rosy to reddish, densely covered by red speckles. Foot background reddish mottled with creamy irregularly shaped blotches containing white speckles, dorsally delimited by a brown streak, sole rosy. Cephalic tentacles reddish densely covered by red speckles, with a dorsal rosy stripe. Eye stalks reddish; eyes black encircled by white. Snout reddish with a creamy terminal band.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF127FFE8E253FA57D14FF860.taxon	distribution	Distribution Atlantic Ocean, from the British Isles to Morocco, Azores, Madeira, Selvagens and Canary Islands (e. g., Rolán 1983; Segers et al. 2009; Gofas et al. 2011; Hernández et al. 2011). Entire Mediterranean (e. g., Conti & Rossini 1985; Brunet Navarro & Capdevila 2005; Portalatina 2008; Albano & Sabelli 2009; Vazzana 2010; Manousis 2012; Donnarumma et al. 2018; Wölfling et al. 2019), living specimens from intertidal down to 60 m, occasionally (in very clear waters) down to 80 – 100 m deep (deeper records from fishing nets residuals need confirmation).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF127FFE8E253FA57D14FF860.taxon	discussion	Remarks SSH-I and SSH-II were already regarded as distinct species by Monterosato (who used the specific epithets corallinus and igneus for this species, referring to its most frequent colouration). However, most specialists of Mediterranean molluscs have pooled them into a single very polymorphic Trochus exasperatus. Pennant (1777: 126) based the description explicitly on Martini Lister’s figure in the Historiae Conchyliorum (Martini Lister 1770: pl. 616 fig. a 2), which may apply to both species. However, the description (“ ruberrimus fasciis crebris exasperates ”) clearly indicates SSH-II, in its most typical morphotype, and Jujubinus vulgaris seems to be absent in Great Britain from where we have seen no record. Jujubinus exasperatus differs from J. vulgaris in its strongly tuberculate spiral cords and narrow interspaces vs weaker tubercles and broader interspaces in J. vulgaris; the colouration is also diagnostic, red-coralline or brownish background with whitish or yellowish vertical stripes in J. exasperatus, vs a cream or whitish background with all variation of vertical stripes and blotches (including reddish or brownish streaks) in J. vulgaris.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF123FFEAE276FE90D76CFE02.taxon	description	SSH III – Fig. 6 J – L	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF123FFEAE276FE90D76CFE02.taxon	diagnosis	Diagnosis Shell of medium size for the genus, thick, 2 – 3 spaced spiral cords, elevated and granulated on the first whorls, broad interspaces with strong prosocline lamellae and obvious growth lines, less evident on the last whorls, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF123FFEAE276FE90D76CFE02.taxon	materials_examined	Type material Syntype AZORES • 1 dd; S. Miguel, Ponta Delgada; SMF 341875. Other material examined AZORES • 18 dd; unprecised locality; MCZR-M- 11761 • 13 dd; S. Miguel, Cais da Ribeirinha; 10 m depth; on stones; CS-PM • 3 dd; Santa Cruz de Flores; tidal pools; MO • 2 lv; São Miguel Is., Caloura; 37 ° 42 ′ 24 ″ N, 25 ° 30 ′ 31 ″ W; 7 m depth; used for DNA sequencing; BAU 771.1, 771.3. MADEIRA • 34 dd; unprecised locality; labelled “ J. Madrensis Monts, copia a Issel ”; MCZR-M- 30087 • 6 dd; unprecised locality; MCZR-M- 30087. MID ATLANTIC SEAMOUNTS •> 200 dd; Gorringe Bank, Gettysburg; 36 º 31 ′ N, 11 º 34 ′ W; 71 – 88 m depth; MO.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF123FFEAE276FE90D76CFE02.taxon	description	Description (in parentheses data of the syntype) SHELL. Shell of medium size for the genus, height 5.8 – 8.4 (5.0) mm, width 3.7 – 8.4 (3.9) mm, height / width ratio 1.31 – 1.62, conical, solid. Protoconch, of ca 1.2 whorls 0.28 – 0.29 mm in width, sculptured with sparse micropustules. Teleoconch of 5.5 – 6.5 (5.5) slightly and solid convex whorls. Sculpture of 2 – 3 (3) abapical spiral cords of different size and strength and irregularly arranged, the first granulose, and a major peripheral cord formed by 3 – 5 (3) merged cordlets of different sizes. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set. Base convex, with 7 – 10 (7) regularly spaced, basal spiral cords flat and well separated umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish; remaining whorls and base greyish or reddish, with alternating reddish and whitish spiral cords of different size, with the upper part of the brownish-reddish or whitish-reddish stripe dotted with white. SOFT PARTS. Epipodial tentacles purplish-brown. Foot cream to reddish-brownish, mottled with white and reddish blotches mostly on the posterior half; sole creamy whitish. Cephalic tentacles dark-brown dorsally, whitish ventrally. Eye stalks whitish or creamy ventrally, dorsally reddish, eyes black. Snout light or dark brown with a creamy terminal band.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF123FFEAE276FE90D76CFE02.taxon	distribution	Distribution Azores Islands (Nordsieck 1973; Curini-Galletti 1982 b; Gofas 2005; Ávila et al. 2006, 2011; Mariottini et al. 2013; Borges et al. 2016; Uribe et al. 2017; Scaillet & Delongueville 2019) and Gorringe Bank (new record); the records from Madeira based on historica materials need confirmation. Empty shells from a few metres to a depth of 80 m, but living specimens from shallower depth (7 m, our material).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF123FFEAE276FE90D76CFE02.taxon	discussion	Remarks Jujubinus pseudogravinae differs from J. exasperatus in its 2 – 3 spiral cords, of irregular shape and size, sculptured with coarse tubercles, occasionally visible on the first teleoconch whorl, v ersus spiral cords well separated by regular interspaces, sculptured with finer tubercles in J. exasperatus. Jujubinus vulgaris can be easily separated by its weaker and more regular sculpture, and by the colour, always with a cream or whitish background, very rarely present in J. pseudogravinae.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF122FFE4E27EFD9DD637FCC9.taxon	description	SSH IV – Fig. 6 M – O	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF122FFE4E27EFD9DD637FCC9.taxon	diagnosis	Diagnosis Shell of medium size for the genus, not very solid, 5 – 6 spaced spiral cords, elevated, broad interspaces with prosocline lamellae and fine growth lines, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF122FFE4E27EFD9DD637FCC9.taxon	materials_examined	Type material Holotype MADEIRA • dd; Porto Moniz; 33 ° N, 17 ° W; DSAT (Pisa, Italy). Paratypes MADEIRA • 3 dd; same data as for holotype; DSAT (Pisa, Italy) (paratypes B – D) • 2 dd; same data as for holotype; labelled “ C. striatum L. Madeira ”; MCZR-M-TYPE- 00018 / P (paratypes A, E). Other material examined MADEIRA • 34 lv; unprecised locality; MCZR-M- 11715 • 1 dd; Porto Moniz; 1912; Tomlin leg.; MCZR-M- 11684 • 14 dd; Madeira; MCG collection • 1 dd; unprecised locality; 1912; Tomlin leg.; MCZR-M- 11684.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF122FFE4E27EFD9DD637FCC9.taxon	description	Description (in parentheses data of the holotype) SHELL. Shell of medium size for the genus, height 6.5 – 8.0 (6.9) mm, width 5.0 – 6.0 (6.0) mm, height / width ratio 1.15 – 1.33, conical, light. Protoconch seemingly smooth, of ca 1 whorl ca 0.20 mm in width. Teleoconch of 6 – 10 (7) slightly convex whorls. Sculpture of 5 – 6 (7) flat, spaced abapical spiral cords of about the same strength, and an evident peripheral cord formed by 2 – 3 (3) merged cordlets; peripheral cord visibly rippled on first two / tree whorls of teleoconch, flat on the others remaining. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set. Base convex, with 5 – 6 (6) regularly spaced, basal spiral cords large and flat, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous.. COLOURATION. Colour of protoconch and initial teleoconch whorls greyish-whitish; remaining whorls and base cream with alternating dull pinkish-brownish-reddish longitudinal and opisthocline flammules of different size and irregularly arranged. SOFT PARTS. Not observed.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF122FFE4E27EFD9DD637FCC9.taxon	distribution	Distribution Madeira (Curini-Galletti 1990; Cretella 1992), empty shells in shallow waters or beached.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF122FFE4E27EFD9DD637FCC9.taxon	discussion	Remarks Monochrome specimens are known (Curini-Galletti 1990). Jujubinus vexationis can be distinguished from J. vulgaris, J. exasperatus and J. pseudogravinae by its spiral cords without tubercles and the interspaces furrowed by dense prosocline lines; additionally, J. vexationis has the sutural cord tuberculate in the first 2 – 3 teleoconch whorls, a feature never present in the three other species. See under J. gravinae, J. errinae Smriglio, Mariottini & Giacobbe, 2016 and J. superum sp. nov. for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12CFFE7E1D7FCA4D10BF913.taxon	description	SSH V – Fig. 6 P – R	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12CFFE7E1D7FCA4D10BF913.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, 5 – 6 spaced spiral cords, elevated, granulose, wide interspaces like the cords, with strong prosocline lamellae, peripheral cord very evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12CFFE7E1D7FCA4D10BF913.taxon	etymology	Etymology The species is named after our friend Frank Swinnen, for his help in finding specimens of Jujubinus.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12CFFE7E1D7FCA4D10BF913.taxon	materials_examined	Type material Holotype CANARY ISLANDS • lv (height 6.5 mm, width 4.8 mm); Gran Canaria, Arinaga; 27 ° 51 ′ 21.24 ″ N, 15 ° 23 ′ 46.61 ″ W; 25 Jan. 2016; low tide, amidst weeds, on rocky plateau with small stones; F. Swinnen and Martin Coysman leg.; used for DNA sequencing; MNHN-IM- 2000 - 40309 (ex BAU 2944.1). Paratypes CANARY ISLANDS • 1 dd; same data as for holotype; MNHN-IM- 2000 - 403010 (paratype 1) • 1 dd; same data as for holotype; MCZR- 00094 / P (paratype 2) • 16 lv; same data as for holotype; used for DNA sequencing; BAU 2944.2 to BAU 2944.5, BAU 2945.1 to BAU 2945.5, BAU 2946.1, BAU 2947.1 to BAU 2947.4, BAU 2948.1, BAU 2948.2 (paratypes 3 – 18) • 8 lv; same data as for holotype; CS-PM (paratypes 19 – 26) • 4 lv; same data as for holotype; FS (paratypes 27 – 30). Other material examined CANARY ISLANDS • 8 dd; Tenerife, Punta de Teno; on stones at low tide; CS-PM • 30 dd; Gran Canaria, Meloneras; low tide; CS-PM • 5 dd; unprecised locality; “ Canarie! D. Tomlin 1919 ”; MCZR-M- 11761. MADEIRA • 20 dd; unprecised locality; “ Madera ”, ex Watson collection, labelled as “ Mirulinus watsoni ”; MCZR-M- 11715 • 5 dd.; unprecised locality; ex Watson collection; MCZR-M- 11715 • 3 dd; unprecised locality; “ Madera ”, ex Nobre collection; MCZR-M- 11715. Description (in parentheses data of the holotype) SHELL. Shell of medium size for the genus, height 5.2 – 6.7 (6.5) mm, width 4.1 – 4.8 (4.8) mm, height / width ratio 1.20 – 1.35, conical, shiny in type material. Protoconch of ca 1 whorl 0.23 – 0.24 (0.24) mm in width, sculptured by one spiral thread. Teleoconch of 5.5 – 6.0 (5.5) slightly convex whorls. Sculpture of 4 – 6 (6) spaced abapical spiral cords, smooth or slightly grainy, of about the same strength, and a very evident peripheral cord, slightly shouldered, formed by 4 – 6 (4) merged cordlets. Fine prosocline growth striae and lamellae, particularly in the interspaces. Suture incised. Base convex and rounded, with 6 – 7 (7) regularly spaced, flat basal spiral cords; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the lower portion. Interior of the aperture whitish, greyish-greenish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish, remaining whorls and base bright greenish cream or greenish whitish; spiral cords with alternating spots, whitish or reddish or blackish sutural and peripheral cord articulated in black and white. SOFT PARTS. Observed in alcohol preserved specimens, no dark terminal strip observed on the creamy band of the snout.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12CFFE7E1D7FCA4D10BF913.taxon	distribution	Distribution Canary Islands, all living specimens collected in the intertidal. The historical records from Madeira in the Monterosato collection need confirmation.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12CFFE7E1D7FCA4D10BF913.taxon	discussion	Remarks Jujubinus swinneni sp. nov. has been reported by Segers et al. (2009: 69, pl. 6 fig. 5) under the name Jujubinus vexationis, a distinct species probably endemic to Madeira. In the Monterosato collection are kept three vials, containing a total of 28 shells, of J. vexationis (probably all from Madeira) labelled “ J. watsoni ”, “ Mirulinus watsoni ”, and “ Trochus striatus ”, respectively. It has also been confused with J. gravinae (e. g., Rolán & Swinnen 2009: figs 33 – 39; Hernández et al. 2011: 428, fig. 14 a – e). Poppe & Goto (1991) reported but did not figure J. gravinae from the Canary Islands (where it is actually absent) probably referring to J. swinneni. The confusion may have been caused by the colouration of J. swinneni that somehow mimics the sutural and peripheral sculpture of J. gravinae (see below). Jujubinus swinneni differs from J. vexationis in its less slender shell with a wider base and for the sculpture of regularly spaced cords similar in size, versus the more numerous and irregularly spaced ones of J. vexationis; additionally, the characteristic pattern with dull colours of the latter allows an easy separation. Jujubinus errinae (endemic to the Messina Strait) also presents a tuberculate sutural cord in the first teleoconch whorls, but with different colouration (spiral cords streaked with red) and marked sculpture (with strong prosocline lamellae in the interspaces) (Smriglio et al. 2016; Giacobbe & Renda 2019). See under J. gravinae for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12FFFE1E248F96ED2E7FBAE.taxon	diagnosis	Diagnosis Shell of medium size for the genus, thick, elevated, moderately slender, with 4 spaced spiral cords, elevated and granulose, broad interspaces with strong prosocline lamellae, conspicuous and evident peripheral cord, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12FFFE1E248F96ED2E7FBAE.taxon	materials_examined	Type material Lectotype FRANCE • dd; Paulilles; MCZR 00218 / L. Paralectotypes CORSICA • 20 dd; Pietranera; “ J. baudoni var. incomparabilis ”; MCZR 00219 / 176 to MCZR 00219 / 195. FRANCE • 133 dd; same data as for holotype; MCZR 00219 / 2 to MCZR 00219 / 133 • 30 dd; Roussillon; MCZR 00219 / 134 to MCZR 00219 / 163 • 12 dd; Département de l’Aude, Pyrénées Orientales; MCZR 00219 / 164 to MCZR 00219 / 175. Other material examined CORSICA • 6 lv; Calcataggio, Tour d’Ancone; 42 ° 02 ′ 36 ″ N, 08 ° 43 ′ 15 ″ E; 15 m depth; 28 Aug. 2012; used for DNA sequencing; BAU 3762, 3763 3765 to 3768 • 2 lv; Tour D’Omigna; 42 ° 08 ′ 42.3 ″ N, 8 ° 33 ′ 23.2 ″ E; 15 m depth; 27 Aug. 2012; used for DNA sequencing; BAU 3769, 3770. • 4 lv; S of Lavezzi Is., CORSICABENTHOS stn CS 49; 41 ° 19 ′ N, 9 ° 15.8 ′ E; 30 – 33 m depth; 19 Oct. 2020; used for DNA sequencing; MNHN-IM- 2000 - 13569 to 13571, 13584 • 3 lv; Large Punta Rossa, CORSICABENTHOS stn CB 44; 42 ° 25.5 ′ N, 8 ° 36.2 ′ E; 23 – 32 m depth; 26 May 2021; used for DNA sequencing; MNHN- IM- 2000 - 17991, 17994, 17997 • 1 lv; CORSICABENTHOS stn CS 83; 42 ° 21.2 ′ N, 8 ° 32.1 ′ E; 52 m depth; 21 May 2021; used for DNA sequencing; MNHN-IM- 2000 - 17889 • 1 lv; Capu Rossu, CORSICABENTHOS stn CB 87; 42 ° 25.5 ′ N, 8 ° 36.2 ′ E; 23 – 32 m depth; 26 May. 2021; used for DNA sequencing; MNHN- IM- 2000 - 17626 • 3 lv; Danger du Toro; 41 ° 30.5 ′ N, 9 ° 23.6 ′ E; 13 – 20 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12262 to 12264 • 3 lv; L’Arche; 41 ° 31.7 ′ N, 9 ° 24.1 ′ E; 32 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12787, 12789, 12819 • 1 lv; Tête de Cheval; 41 ° 20.3 ′ N, 9 ° 14.5 ′ E; 20 – 30 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13462 • 1 lv; Capu Rossu, 42 ° 14.2 ′ N, 8 ° 31.9 ′ E; 56 m depth; used for DNA sequencing; MNHN-IM- 2019 - 18109 • 2 lv; South of Garganellu Is.; 42 ° 21.9 ′ N, 8 ° 32.3 ′ E; 3 – 51 m depth; used for DNA sequencing; MNHN-IM- 2019 - 18141, 18165 • 1 lv; Punta Vecchia-Cala Genovese; 42 ° 59.8 ′ N, 9 ° 27.2 ′ E; 11 m depth; used for DNA sequencing; MNHN- IM- 2019 - 4683 • 1 lv; Sec de Nonza; 42 ° 47.2 ′ N, 9 ° 19.8 ′ E; 37 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5663 • 1 lv; Punta di Canelle; 42 ° 49.8 ′ N, 9 ° 18.4 ′ E; 15 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5863 • 2 lv; Centuri depths; 42 ° 59.3 ′ N, 9 ° 17.8 ′ E; 21 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5899, 5902. SARDINIA • 3 dd; Capo Caccia; 25 m depth; CS-PM • 59 dd; Caprera I.; 25 m depth; CS-PM • 91 dd; La Maddalena I.; 25 m depth; CS-PM • 62 dd; Spargi I.; 20 m depth; CS-PM. SPAIN • 1 dd; Cadaqués; 4 m depth; CS-PM.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12FFFE1E248F96ED2E7FBAE.taxon	description	Description SHELL. Shell of medium size for the genus, height 6.5 – 7.5 mm, width 4.8 – 5.8 mm, height / width ratio 1.28 – 1.35, conical, very robust. Protoconch seemingly smooth, of ca 1 whorl, 0.28 – 0.29 mm in width. Teleoconch of 5.5 – 6.5 slightly convex whorls. Sculpture of 4 – 5 strong very granulose, regularly spaced spiral cords of the same strength, and an evident broad peripheral cord, formed by 2 merged granulose cordlets; suture well-marked. Interspaces with strong, regular, prosocline lamellae. Base convex, with 6 – 8 concentric reddish and white, regularly spaced spiral cords, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, pinkish or greenish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls glossy red or pinkish; remaining whorls and base with alternating whitish and reddish spirals and irregular blotches. SOFT PARTS. Observed in alcohol preserved specimens, a dark terminal strip absent on the creamy band of the snout.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12FFFE1E248F96ED2E7FBAE.taxon	distribution	Distribution Jujubinus baudoni is limited to the northern and western Mediterranean Sea, from southern Spain to Corsica and Sardinia (Scaperrotta et al. 2011; Spanu 2011; Mariottini et al. 2013), with living specimens collected at depths of 15 – 50 m. As already reported by Mariottini et al. (2013), the specimens in the Monterosato collection (MCZR 11761) from Azores labelled as “ Calliostoma baudoni ” actually are J. pseudogravinae.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12FFFE1E248F96ED2E7FBAE.taxon	discussion	Remarks Little known species with an unmistakable sculpture. Three major colour patterns were so far known: one with prosocline reddish-brown and milky-white flammulae; the second, with emerald-green background, bright orange and / or red flammulae, milky-white prosocline stripes, and white spots on the basal cord (the variety incomparabilis); and the third, monochrome brownish-green or with prosocline faint whitish flammules (Mariottini et al. 2013). Additionally, we report here peculiar specimens from Caprera Island, with protoconch and initial teleoconch whorls glossy red, greenish base, and small whitish flammulae on the spiral cords.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF129FFE3E226FB01D2C6FC1A.taxon	diagnosis	Diagnosis Shell of large size for the genus, solid, several closely set spiral cords, not elevated, barely granulose, narrow interspaces with closely set and numerous prosocline lamellae and growth lines, peripheral cord not evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF129FFE3E226FB01D2C6FC1A.taxon	materials_examined	Type material Lectotypes of Trochus striatus var. e lenchoides Issel, 1878 TURKEY • 2 dd; Cape Barbieri; 20 m depth; “ Barbieri (m. 20) ” “ Isola Barbieri / 20 m / (Issel)! ”; MCZR- M- 11698 - L (designated by Curini-Galletti 1982 a). Paralectotype of Trochus striatus var. e lenchoides Issel, 1878 TURKEY • 1 dd; same data as for lectotype; MCZR-M- 11698 - P. Syntype of Jujubinus aegeensis Nordsieck, 1973 GREECE • 1 dd; Karpathos Is. (type locality); SMF 341823. Syntype of Jujubinus elenchoides orientalis Nordsieck, 1973 GREECE • 1 dd; Karpathos Is. (type locality); SMF 341821. Other material examined CYPRUS • 1 lv; Cape Greco; 34.9860 ° N, 34.0786 ° N; 30 m depth; 7 May 2022; on Posidonia oceanica; used for DNA sequencing, CYP 082; SZN-MOL 0054. GREECE • 4 lv; Astypalea Is., Analipsi; 36.575946 ° N, 26.393070 ° E; 0 – 1 m depth; 28 – 30 Jun. 2017; on Posidonia oceanica; used for DNA sequencing; BAU 3040.1 to 3040.4 • 2 lv; same data as for prececeding; used for DNA sequencing; BAU 3041.1, 3041.2 • 4 lv; same data as for prececeding; used for DNA sequencing; BAU 3044.1 to 3044.4 • 1 lv; same data as for prececeding; used for DNA sequencing; BAU 3044.6 • 3 dd; Kythnos Is.; 6 m depth; CS-PM • 12 dd; Astypalea Is., Analipsi; 5 m depth; on Posidonia oceanica; LT. MALTA • 1 dd; labelled “ coste / Malta ”; MCZR-M- 11703. TUNISIA • 2 dd; Gulf of Gabès; labelled “ Gabes Pallary ”; MCZR-M- 11712. TURKEY • 10 dd; Bozcaada Island; 8 m depth; CS-PM • 9 dd; Datça; 36 ° 42 ′ 16.6 ″ N, 27 ° 40 ′ 52.3 ″ E; 15 m depth; on Posidonia oceanica; LT • 2 dd; Datça harbour; 36 ° 43 ′ 20.9 ″ N, 27 ° 41 ′ 22.1 ″ E; 8 m depth; LT. UNPRECISED LOCALITIES • 15 dd and fragments; Mediterranean coasts of Northern Africa; labelled “ C. d’Afr. ” (Appolloni et al. 2018); MCZR-M- 11682 • 10 dd; Mediterranean coasts of Northern Africa; labelled “ J. (Zizyphinus) sartorii, Aradas. C. di Barberia! ” plus “ J. sartorii, Ar. Coste d’Africa ”; MCZR-M- 11758 • 4 dd; Adriatic? [probably wrong]; “ Trochus striatus Gm. Adr. Stoss. ”; MCZR-M- 11778 • 8 dd; Northern Africa; labelled “ J. sericeus nelle spugne ”; MCZR-M- 11683. Description (in parentheses data of the lectotype) SHELL. Shell of large size for the genus, height 7.4 – 13.0 (11.7) mm, width 5.3 – 7.5 (7.3) mm, height / width ratio 1.43 – 1.60, conical, shiny but opaque in type material. Protoconch, of ca 1.1 whorls, 0.23 – 0.24 mm in width, sculptured by sparse micropustules. Teleoconch of 6.5 – 7.5 (7.5) slightly convex whorls. Sculpture of 12 – 16 (14) closely set abapical spiral cords of about the same strength, and a not evident peripheral cord, slightly shouldered, formed by 2 – 3 (2) merged cordlets. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae and lamellae, irregularly set. Base convex and rounded, with 11 – 13 (12) regularly spaced, basal spiral flat cords, arranged in pairs; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright whitish greyish; remaining whorls and base dark background colour, black, brownish, reddish, with alternating sagittal bands, whitish or reddish or pinkish, which are highlighted depending on the background colour. SOFT PARTS. Epipodial tentacles whitish, densely covered by red speckles. Foot background whitish mottled with dark spots, sole white. Cephalic tentacles whitish densely covered by dark speckles, with a dorsal dark stripe. Eye stalks brownish dorsally, whitish ventrally; eyes black surrounded by a brown blotch, encircled by white. Snout light or dark brown with a creamy terminal band.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF129FFE3E226FB01D2C6FC1A.taxon	distribution	Distribution Eastern Mediterranean Sea, from the Aegean Sea (Manousis 2012) to eastern Turkey (this work), and Cyprus (Tornaritis 1987, as “ Jujubinus striatus Linne ”), living specimens in very shallow waters (0 – 1 m) occasionally down to a depth of 30 m. Also reported in the northern African coasts, from Morocco to Libya, (Ghisotti & Melone 1975) and Malta (MCZR-M- 11703 collection): while we are able to confirm its presence in the eastern Mediterranean Sea, the records from Malta and North Africa are almost all based on labels of historical collections and need further checking.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF129FFE3E226FB01D2C6FC1A.taxon	discussion	Remarks Described by Issel (1878) as Trochus striatus var. elenchoides, it may be argued whether it is to be considered as infrasubspecific or subspecific (ICZN 1999: art. 45.6.4); in any case, it was elevated to specific rank at first by Monterosato (1884: 47), which would make it available from its original introduction by Issel (ICZN 1999: art. 45.6.4.1). Issel (1878: 436) reported the locality “ Barbieri (m. 20) ” which is actually not an island (as suggested by Monterosato’s label, see also Curini-Galletti 1982 a: 219) but rather the small promontory (Capo Barbieri) in correspondence with the ancient locality of Rhoiteion (Troades, northern Anatolia). Jujubinus elenchoides differs from J. ruscurianus in its narrower interspaces and more numerous spiral cords (4 – 5 in ruscurianus vs 12 – 15 finer in elenchoides), and from J. vulgaris, J. exasperatus and J. pseudogravinae in its smooth, non-tuberculated and more numerous spiral cords. It also differs from J. swinneni sp. nov., J. vexationis, and J. errinae in its colouration and the larger size and, from the latter two species, for the lack of tubercles on the peripheral cord in the first teleoconch whorls. See under J. striatus, J. depictus, J. fraterculus, J. superum sp. nov., J. inopinatus sp. nov., J. kyrnos sp. nov., J. sartorii, J. smaragdinus, J. angustibasis sp. nov., J. gofasi sp. nov., for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12BFFE2E1D5FC75D135FA8F.taxon	description	SSH VIII – Fig. 7 A – C	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12BFFE2E1D5FC75D135FA8F.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, several closely set spiral cords, not elevated, narrow interspaces with closely set and numerous prosocline lamellae and growth lines, peripheral cord not evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12BFFE2E1D5FC75D135FA8F.taxon	etymology	Etymology After the ancient Latin name of the Adriatic Sea (Mare Superum), used as a noun in apposition.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12BFFE2E1D5FC75D135FA8F.taxon	materials_examined	Type material Holotype SLOVENIJA • lv; Izola; 45 ° 32 ′ 51.4 ″ N, 13 ° 42 ′ 12.9 ″ E; 4 m depth; on Posidonia meadow; used for DNA sequencing; MNHN-IM- 2000 - 40311 (ex BAU 2077.1). Paratypes SLOVENIJA • 2 lv; Izola; 45 ° 32 ′ 51.4 ″ N, 13 ° 42 ′ 12.9 ″ E; 4 m depth; on Posidonia meadow; used for DNA sequencing; BAU 2077.2, BAU 2077.3 (paratypes 1 – 2) • 2 lv; same data as for preceding; RM 3 - Jis 1, RM 3 - Jis 2 (paratypes 3 – 4) • 7 lv; same data as for preceding; CS-PM (paratypes 5 – 11). Description (in parentheses data of the holotype) SHELL. Shell conical, of medium size for the genus, height 6.0 – 6.5 (6.0) mm, width 4.7 – 4.9 (4.7) mm, height / width ratio 1.27 – 1.34, conical, solid, shiny. Protoconch seemingly smooth, of ca 1 whorl, 0.27 – 0.28 (0.28) mm in width. Teleoconch of 5.5 – 6.5 (5.5) slightly convex whorls. Sculpture of 5 – 6 (5) closely set abapical spiral cords of approximately the same strength, regularly spaced, and a not evident peripheral cord, slightly shouldered, formed by 2 (2) merged cordlets. Suture not very evident. Teleoconch surface covered by barely visible prosocline growth striae irregularly set and arranged and regularly lamellae in the interspaces. Base convex and rounded, with 7 – 8 (7) regularly spaced, basal spiral flat cords regularly arranged and with wide interspaces; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish-greyish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright whitish-greyish; remaining whorls and base dark greyish-blackish background colour, with alternating black and white sagittal stripes of irregular size. SOFT PARTS. Epipodial tentacles whitish translucent with sparse scattered very tiny white speckles. Foot background creamy or brownish with darker brown blotches and covered by sparse white speckles; sole white. Cephalic tentacles translucent with a dorsal white stripe and irregularly scattered white speckles. Eye stalks whitish or creamy dorsally, brown ventrally; eyes black. Snout light or dark brown with a creamy terminal band.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12BFFE2E1D5FC75D135FA8F.taxon	distribution	Distribution Currently known only for the Istrian peninsula (Slovenija) in shallow waters (4 m deep) on Posidonia oceanica (L.) Delile meadows.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12BFFE2E1D5FC75D135FA8F.taxon	discussion	Remarks Jujubinus superum sp. nov. differs from J. vulgaris, J. exasperatus and J. pseudogravinae in its non-tuberculated spiral cords. It differs from J. swinneni sp. nov. in its more slender outline and the different colouration, always with a dark greyish-blackish background. From J. vexationis, and J. errinae it differs in its larger size, in the lack of tubercles on the peripheral cord in the first teleoconch whorls, and in the colouration (dark greyish-blackish background with sagittal stripes in J. superum, cream background with alternating pinkish-brownish-reddish longitudinal and opisthocline flammules in J. vexationis and reddish-creamy, with spiral cords and basal cord red interrupted by short white spots in J. errinae). From J. elenchoides it differs in having fewer but stronger spiral cords (5 – 6 in J. superum vs 12 – 15 weaker cords in J. elenchoides). See under J. striatus, J. depictus, J. fraterculus, J. dianiensis sp. nov., J. inopinatus sp. nov., J. kyrnos sp. nov., J. sartorii, J. smaragdinus, J. angustibasis sp. nov., J. gofasi sp. nov., for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12AFFDCE24BFAE1D2C6FA9A.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, 4 – 5 spaced spiral cords, elevated, broad interspaces with numerous prosocline growth lines, peripheral cord evident and rippled, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12AFFDCE24BFAE1D2C6FA9A.taxon	materials_examined	Type material Syntypes FRANCE • 4 dd; Cannes, Cap Croisette (type locality); ex Dautzenberg collection; MCZR-M- 11689 / S • other syntypes; Cannes, Cap Croisette; Dautzenberg collection; RBINS I. G. 10591. Other material examined CORSICA • 4 dd; Bastia; Caziot leg., 1896; MCZR-M- 11689 • 5 dd; Corsica; Del Prete leg.; MCZR-M- 11689 • 19 dd; L’Île Rousse, “ Les Olivieres ”; 42 ° 38 ′ 01.4 ″ N, 8 ° 57 ′ 03.3 ″ E; camping shore, beached; LT • 1 lv; Calcataggio, Tour d’Ancone; 42 ° 02 ′ 36 ″ N, 8 ° 43 ′ 15 ″ E; 15 m depth; 28 Aug. 2012; used for DNA sequencing; BAU 3764 • 3 lv; Lavezzi Is., 41 ° 20 ′ N, 9 ° 15.6 ′ E; 0 – 2 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13574, 13598, 13600 • 2 lv; Punta Di Capineru; 41 ° 27.4 ′ N, 9 ° 02.4 ′ E; 0 – 1 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13679, 13690 • 3 lv; Cargèse Port; 42 ° 07.9 ′ N, 8 ° 35.8 ′ E; 7 m depth; used for DNA sequencing; MNHN-IM- 2019 - 16793, 16822, 16852. CROATIA • 9 dd; Brač; CS-PM. FRANCE • 4 dd; La Ciotat; MCZR-M- 11689 • 4 dd; Cannes; MCZR-M- 11689 • 1 dd; Roussillon; MCZR-M- 117747. ITALY • 5 dd; Livorno, Ardenza; MCZR-M- 11684 • 1 dd; Elba I., Porto Azzurro, Naregno; bioclastic sediment; LT • 15 lv; Giannutri I.; Cystoseira; CS-PM • 62 dd; Giannutri I.; 15 m depth; bioclastic sediment; CS-PM • 3 lv; Giannutri I.; 42 ° 15 ′ 7.05 ″ N, 11 ° 05 ′ 31.51 ″ E; used for DNA sequencing; RM 3 - 174 - Jgra 1, RM 3 - 175 - Jgra 2, RM 3 - 176 - Jgra 3 • 130 lv; Giannutri I.; 11 – 25 m depth; on algae; CS-PM • 1 dd; Argentario; 30 m depth; MO • 5 dd; Civitavecchia, Sant’Agostino; beached; CS-PM • 5 lv; Civitavecchia, Sant’Agostino; on Posidonia; CS-PM • 3 dd; Civitavecchia; beached; CS-PM • 3 dd; Palo Laziale; MO • 22 lv; Ventotene I., Cala Rossano; 40 ° 48 ′ 00.1 ″ N, 13 ° 25 ′ 53.8 ″ E; 1.5 – 3 m depth; on brown algae; LT • 3 dd; Taranto; Abate Gravina legit; MCZR-M- 11689. SARDINIA • 5 dd; La Maddalena I.; MO • 1 dd; Asinara I.; 25 m depth; MO • 13 dd; Stintino, Roccaruja; beached; CS-PM • 6 dd; Alghero; MCZR-M- 11689 • 3 dd; Capocaccia; beached; CS-PM • 32 dd; Gulf of Arzachena; CS-PM • 2 lv; Baccu Mandara; 39 ° 09 ′ 27.8 ″ N, 9 ° 23 ′ 06.8 ″ E; 1 – 3 m depth; amidst algae, used for DNA sequencing; BAU 2078.1, BAU 2078.3 • 9 dd; Bosa Marina; beached; CS-PM • 4 dd; Siniscola, Santa Lucia; 7 m depth; on brown algae; LT • 14 dd; La Maddalena I.; 25 m depth; bioclastic sediment; CS-PM • 6 dd; Gulg of Arzachena; beached; CS-PM. SICILY • 3 lv; Ognina, Catania, north of the port; 37 ° 31 ′ 55.4 ″ N, 15 ° 07 ′ 14.3 ″ E; 0.5 – 3 m depth; on brown algae; LT • 7 lv; Catania, Stazzo; 37 ° 39 ′ 24.0 ″ N, 15 ° 11 ′ 59.0 ″ E; 0.5 – 4 m depth; on brown algae; LT • 10 lv; Capo San Vito, Macari; 38 ° 08 ′ 22.8 ″ N, 12 ° 44 ′ 14.2 ″ E; 1 – 4 m depth; on brown algae; LT • 6 lv; Favignana I.; 4 m depth; on brown algae; LT • 25 dd plus, 4 dd; Palermo; MCZR-M- 11689 • 8 dd; Messina; Sulliotti leg.; MCZR-M- 11689 • 6 dd; Acitrezza; MO • 24 dd; Ognina; MCZR-M- 11684 • 4 dd; Sciacca; MCZR-M- 11689 • 90 dd; Favignana I.; CS-PM • 5 dd; Lampedusa Cala Maluc; 8 m depth; CS-PM • 22 dd; Pantelleria; MCZR-M- 11689. SPAIN • 1 dd; Valencia; MCZR-M- 11689.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12AFFDCE24BFAE1D2C6FA9A.taxon	description	Description SHELL. Shell of medium size for the genus, height 5.1 – 6.5 (6.3) mm, width 4.6 – 5.2 mm, height / width ratio 1.12 – 1.31, conical, not very bright. Protoconch, of ca 1 whorl, 0.24 – 0.25 mm in width, sculptured by one spiral thread. Teleoconch of 4.5 – 6.0 slightly convex whorls. Sculpture of 4 – 5 flat, closely set abapical spiral cords of about the same strength, a rippled peripheral cord formed by 4 – 5 merged cordlets, and 6 – 7 regularly spaced, basal flat spiral cords. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish or pinkish nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish-pinkish; remaining whorls and base with cream or whitish background, with brownish-reddish flammules of different size, peripheral cord white dotted. SOFT PARTS. Epipodial tentacles translucent, covered by scattered very tiny white speckles. Foot background creamy or reddish mottled with brownish or reddish and white blotches; sole whitish or creamy. Cephalic tentacles translucent, with a dorsal white stripe. Eye stalks brownish dorsally, whitish ventrally; eyes black encircled by white. Snout brownish to reddish with a creamy terminal band.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12AFFDCE24BFAE1D2C6FA9A.taxon	distribution	Distribution Mediterranean Sea, from the Adriatic to the Alboran Sea, in 0.5 – 25 m depth; not reported for the eastern basin by Ghisotti & Melone (1975), but recently and erroneously reported for the Aegean Sea by Manousis (2012: 92, first unnumbered figure) who figured a juvenile specimen of the J. striatus complex. Atlantic, from the Portuguese coasts to Dakar, Senegal (Cretella 1992); the record under this name from the Canary Islands (Rolán & Swinnen 2009: figs 33 – 39; Hernández et al. 2011: 428, fig. 14 a – e), actually represents Jujubinus swinneni sp. nov. (see under this species).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF12AFFDCE24BFAE1D2C6FA9A.taxon	discussion	Remarks Jujubinus gravinae has a very peculiar sculpture with the peripheral cord regularly and markedly crenulated which makes it easily diagnosed – for instance – from J. vexationis and J. errinae that exhibit this crenulation only on the first 2 – 3 teleoconch whorls. Jujubinus swinneni sp. nov., similar in the general aspect, can be easily diagnosed by the complete lack of tubercles on the peripheral cord and for the different colouration, bright greenish cream or greenish whitish, with alternating whitish or reddish or blackish spots on the peripheral cord vs cream or whitish background, with brownish-reddish flammules of J. gravinae. Jujubinus ruscurianus, J. elenchoides and J. superum sp. nov. lack any sutural and peripheral crenulation on all teleoconch whorls. Jujubinus gravinae differs from J. vulgaris, J. exasperatus and J. pseudogravinae in its non-tuberculated spiral cords. See under J. striatus, J. depictus, J. fraterculus, J. dianiensis sp. nov., J. inopinatus sp. nov., J. kyrnos sp. nov., J. sartorii, J. smaragdinus, J. angustibasis sp. nov., J. gofasi sp. nov., J. tingitanus, for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF114FFDEE1A1FAF5D04DFDE1.taxon	description	SSH X – Fig. 7 G – I	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF114FFDEE1A1FAF5D04DFDE1.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, 5 – 7 spaced and dimensionally regular spiral cords, elevated, broad interspaces with strong prosocline lamellae and fine growth lines, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF114FFDEE1A1FAF5D04DFDE1.taxon	etymology	Etymology The species is named from one of the names of Corsica in ancient Greek, ‘ Κύρνος’ (= ‘ Kyrnos’), used as a noun in apposition.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF114FFDEE1A1FAF5D04DFDE1.taxon	materials_examined	Type material Holotype CORSICA • lv (7.1 (H) × 5.0 (W) mm); Capo Rosso, Torra di Turhju (type locality), CORSICABENTHOS stn CB 32; 42 ° 14.4 ′ N, 8 ° 33.1 ′ E; 14 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17191. Paratypes CORSICA • 1 lv (6.5 (H) × 4.9 (W) mm); Punta Muchillina, CORSICABENTHOS stn CB 40; 42 ° 20.1 ′ N, 8 ° 33.3 ′ E; 10 m depth; used for DNA sequencing; MNHN-IM- 2019 - 18126 • 1 lv (9.8 (H) × 6.0 (W) mm); Le Sphinx, CORSICABENTHOS stn CF 02; 42 ° 43.3 ′ N, 9 ° 14.9 ′ E; 17 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5397 • 1 lv; North of Gargalo Is., CORSICABENTHOS stn CB 31; 42 ° 22.3 ′ N, 8 ° 32.4 ′ E; 26 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17184 • 1 lv; same data as for preceding; used for DNA sequencing; MNHN-IM- 2019 - 17269 • 1 lv; same data as for preceding; used for DNA sequencing; MNHN-IM- 2019 - 17281 • 1 lv; Punta di Ficaghjola, CORSICABENTHOS stn CB 33; 42 ° 15.3 ′ N, 8 ° 37.3 ′ E; 21 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17324. Other material examined SPAIN • 1 lv; Almería, Cabo de Gata, Playazo de Rodalquilar; 36 ° 51 ′ 28 ″ N, 2 ° 00 ′ 06 ″ W; 10 m depth; subadult, used for DNA sequencing; BAU 2858.2 • 1 lv; Murcia, Cabo de Palos; 37 ° 38 ′ 08.0 ″ N, 0 ° 41 ′ 30.4 ″ W; subadult, used for DNA sequencing; MNHN-IM- 2013 - 32869. Description (in parentheses data of the holotype) SHELL. Shell of medium size for the genus, height 3.7 – 7.2 (7.1) mm, width 3.2 – 5.0 (5.0) mm, height / width ratio 1.16 – 1.44, conical, cyrtoconoid outline, quite robust, not very bright. Protoconch seemingly smooth, of ca 1 whorl, 0.28 – 0.29 (0.29) mm in width. Teleoconch of 7 – 7.5 flat whorls. Sculpture of 5 – 7 flat, spiral cords regularly spaced of about the same strength, with wide interspaces sculptured with strong prosocline lamella, the 2 peripheral ones (bigger) forming the evident peripheral cord, 6 – 7 closely spaced flat basal cords. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae rather regularly set. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, columellar callus thinner in the basal portion. Aperture whitish or pearly-greyishgreenish nacreous. COLOURATION. Colour of protoconch and first teleoconch whorls whitish-greyish or reddish white, remaining whorls reddish or blackish grey, with irregularly sparse whitish flammules of various sizes; peripheral cord reddish or grayish and slightly dotted with white in the first whorls (more evidently in sub-adult specimens). SOFT PARTS. Epipodial tentacles whitish, densely covered by tiny purplish speckles interconnected to create an irregular network. Foot background reddish or creamy, densely mottled with brown or reddish-brown irregularly shaped blotches; sole white. Cephalic tentacles whitish, with a dorsal purplish stripe and covered by small purplish speckles. Eye stalks reddish or brownish; eyes black. Snout dark brown, creamy terminal band with a thin dark terminal strip.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF114FFDEE1A1FAF5D04DFDE1.taxon	distribution	Distribution Corsica, at depths of 10 – 26 m. Additional material, here conservatively considered as conspecific, from Cabo de Gata and Cabo de Palos, in southern Spain.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF114FFDEE1A1FAF5D04DFDE1.taxon	discussion	Remarks This species is diagnosed by the rather heavy shell with wide and regular but shallow spiral interspaces, and by the not very thick columella with the callous columellar tooth not prominent. It differs from J. vulgaris, J. exasperatus and J. pseudogravinae by the non-tuberculated spiral cords; from J. gravinae, J. vexationis and J. errinae by the absence of the typical tubercles on the peripheral cord and by the different colouration; from J. ruscurianus by the more slender outline and the different sculpture with narrower and less incised spaces. It differs from J. swinneni sp. nov. by the slender outline, the different colouration and the absence of sutural crenulation on the whorls of the teleoconch. It differs from J. elenchoides by its smaller size and by the sculpture formed by 5 – 7 spiral cords, vs 12 – 16 in J. elenchoides. It differs from J. superum sp. nov. in its larger size, its different colouration (reddish and / or whitish, with whitish and greyish flammules in J. kyrnos sp. nov. vs more uniform background with characteristic oblique black and whitish stripes in J. superum) and the sculpture formed by a smaller number of spiral cords (6 – 7 in J. kyrnos vs 12 – 16 in J. superum). Jujubinus kyrnos differs from J. elenchoides in its base with an acute and non-rounded angle and in the evident spiral cords (6.7 in J. kyrnos vs 12 – 15 in J. elenchoides). See under J. depictus, J. sartorii, J. fraterculus, J. gofasi sp. nov., J. dianiensis sp. nov., J. striatus, J. smaragdinus, J. inopinatus sp. nov., Jujubinus angustibasis sp. nov., J. dispar and J. tingitanus for comparative comments.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF116FFD9E1A5FDFBD60EF9A8.taxon	description	SSH XI – Fig. 7 J – L	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF116FFD9E1A5FDFBD60EF9A8.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, 5 – 6 spaced spiral cords, elevated, granulose, wide interspaces like the cords, with strong prosocline lamellae, peripheral cord very evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF116FFD9E1A5FDFBD60EF9A8.taxon	etymology	Etymology The species is named after Serge Gofas, for his great contribution to malacology.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF116FFD9E1A5FDFBD60EF9A8.taxon	materials_examined	Type material Holotype SPAIN • lv (6.8 (H) × 5.5 (W) mm); Málaga (type locality); 2 – 5 m depth; used for DNA sequencing; MNHN-IM- 2000 - 40312 (ex BAU 2075.1). Paratypes SPAIN • 1 lv; same data as for holotype; used for DNA sequencing; MCZR-M-TYPE 00095 (ex BAU 2075.3) (paratype 1) • 3 lv; same data as for holotype; BAU 2066.3 (paratype 2), BAU 2066.5 (paratype 3), BAU 2068.2 (paratype 4) • 5 lv; same data as for holotype; MO (paratypes 5 – 9) • 10 lv; same data as for holotype; CS-PM (paratypes 10 – 19) • 6 lv; same data as for holotype; JMP (paratypes 20 – 25) • 3 lv; Almería; 20 – 30 m depth; trawled; used for DNA sequencing; RM 3 - Jst 7 to RM 3 - Jst 9 (paratypes 26 – 28) • 4 lv; Almería; CS-PM (paratypes 29 – 32). Other material examined FRANCE • 10 lv; Brittany, Ploubazlanec, Baie de Launay; 48 ° 48 ′ 5 ″ N, 3 ° 00 ′ 10 ″ W; intertidal; used for DNA sequencing; BAU 2659.1 to 2659.5, BAU 2662.1 to 2662.5. GREAT BRITAIN • 2 dd; Jersey; “ J. striatus Jersey England d. Chaster. 95 ”; MCZR-M- 11680. MOROCCO • 31 dd; Tangeri; “ Tanger Jujub. Depictus De dd. Tangeri (M. Pallary) ”; MCZR-M- 11681 • 5 dd; Tangeri; “ Tanger J. depictus De dd. Tangeri ”; MCZR-M- 11694. Description (in parentheses data of the holotype) SHELL. Shell of medium size for the genus, height 6.4 – 8.8 (5.8) mm, width 4.9 – 6.8 (5.8) mm, height / width 1.21 – 1.38 conical, solid, slightly cyrtoconoid, rather shiny. Protoconch seemingly smooth, of ca 15 whorl, 0.25 – 0.26 (0.25) mm in width. Teleoconch of 6.5 (5.5) slightly convex whorls. Sculpture of 5 – 6 (6) strong, closely set abapical spiral cords of about the same strength, including the 2 – 3 (3) peripheral ones forming the strong peripheral cord, and 6 – 7 (7) regularly spaced, basal weaker spiral cords and flat. Interspaces with thick and strong regularly set prosocline lamellae. Suture well incised. Teleoconch surface covered by barely visible prosocline growth striae, rather regular. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with columellar callus thickened in the middle portion. Interior of the aperture whitish / greyish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish or greyish; remaining whorls and base cream or blackish background with alternating large blackish / whitish blotches. SOFT PARTS. Observed in alcohol preserved specimens, dark terminal strip on the creamy band of the snout.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF116FFD9E1A5FDFBD60EF9A8.taxon	distribution	Distribution Ranging from the Mediterranean of southern Spain to the Atlantic, southward along the coasts of Morocco and northward to Brittany (France), and Jersey Is. (UK), living specimens from the intertidal to 5 m deep.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF116FFD9E1A5FDFBD60EF9A8.taxon	discussion	Remarks In some specimens the background colour is blackish with alternating large whitish blotches. It differs from J. exasperatus, J. pseudogravinae and J. gravinae in the lack of tubercles on the spiral cords; from J. vulgaris in the more numerous spiral cords (5 – 6 in J. gofasi sp. nov. vs 3 in J. vulgaris) and the colouration (cream or blackish with alternating large blackish-whitish blotches in J. gofasi vs white or whitish background with reddish brown in J. vulgaris); from J. vexationis (Madeira) in its larger size and the different sculpture (7 – 8 weak spiral cords, with slight undulations of the peripheral cord in the first 2 – 3 teleoconch whorls, broad interspaces with fine prosocline lines in J. vexationis vs 6 strong spiral cords and narrow adorned with strong prosocline lamellae in J. gofasi). From J. swinneni sp. nov. (Canary Islands) it differs in its stronger sculpture (with the prosocline lamellae crossing the spiral cords, whereas fine growth lines and lamellae are present mostly in the interspaces in J. swinneni). It differs from J. depictus in the more numerous spiral cords (3 – 4 in J. depictus vs 5 – 6 in J. gofasi sp. nov.) and weaker sculpture; from J. sartorii in the spiral cords with narrower interspaces and with more marked axial lamellae; from J. kyrnos sp. nov. in its large size, the colouration (cream or whitish background, with blackish and greyish flammules in J. kyrnos vs darker background with alternating large blackish blotches in J. gofasi), and the more numerous spiral cords (3 in J. kyrnos vs 5 – 6 in J. gofasi); from J. ruscurianus in its stronger sculpture with narrower interspaces with more marked lamellae; from J. elenchoides in its smaller size, and the fewer cords (12 – 16 spiral cords in J. elenchoides vs 5 – 6 in J. gofasi); from J. fraterculus in its weaker sculpture with less marked prosocline lamellae in the interspaces; from J. superum sp. nov. in its colouration (dark greyish-blackish background in J. superum vs cream or blackish background with alternating large blackish blotches in J. gofasi) the larger size and the more marked sculpture with (vs very fine lamellae in the interspaces of J. superum).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF111FFDAE26CF902D121FAD8.taxon	description	SSH XII – Fig. 7 M – O	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF111FFDAE26CF902D121FAD8.taxon	diagnosis	Diagnosis Shell of large size for the genus, solid, 5 – 6 spaced spiral cords, elevated, granulose, interspaces twice as broad as the cords, with strong prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF111FFDAE26CF902D121FAD8.taxon	materials_examined	Type material Syntypes SICILY • 1 dd; Acirezza (type locality); University of Catania, Aradas collection • 2 dd; Acitrezza; Civico Museo di Storia Naturale, Milano, Mo 28763, Mo 28771. Other material examined CORSICA • 2 lv; Sec St Antonio, CORSICABENTHOS stn CSXX; 41 ° 20.4 ′ N, 9 ° 10.9 ′ E; 43 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13658, 13659 • 2 lv; off South coasts, CORSICABENTHOS stn CSXX; 41 ° 20.4 ′ N, 9 ° 10.8 ′ E; 42 m depth; used for DNA sequencing; MNHN-IM- 2019 - 14585, 14586. CROATIA • 4 dd; Velirat; “ Jujubinus goniobasis Coen, 1937 fide Monterosato ”; MCZR-M- 11728. CYPRUS • 14 dd; Kerinia; (13 dd + 1 dd form concinnus) “ Kerinia (Cipro) Lm. 1899 + vr. Kerinia copia a Coen ”; MCZR-M- 11707. FRANCE • 6 dd; coasts of the Pyrénées-Orientales; “ Roussillon ”; MCZR-M- 11774. GREECE • 4 dd; Preveza; “ Spiaggia di Prevesa, Jonio ”; MCZR-M- 11735. MALTA • 17 dd; unprecised locality; as “ Mirulinus rudi-striatus, Monts. Malta! ”; MCZR-M- 11702 • 24 dd; unprecised locality; “ Malta ”; MCZR-M- 11732 • 1 dd; unprecised locality; “ Malta ”; MCZR-M- 11758 • 2 lv; Gozo Is., Xwejni Bay; 36 ° 04 ′ 48.62 ″ N, 14 ° 14 ′ 50.80 ″ E; 10 m depth; on P. oceanica, used for DNA sequencing; BAU 2676.1, BAU 2676.2. SICILY • 12 dd; Palermo, Arenella; “ Arenella ”; MCZR-M- 25224 • 2 dd; Lampedusa; “ Lampedusa ”; MCZR-M- 11766 • 11 dd; Palermo, Romagnolo; “ Romagnolo (coraliné) 12 giug. 1907 ”; MCZR-M- 11756 • 8 dd; Palermo, Romagnolo; “ vr. Romagnoli monts copia a Cossmann 1913 ”; MCZR-M- 11707.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF111FFDAE26CF902D121FAD8.taxon	description	Description SHELL. Shell of large size for the genus, height 6.5 – 10.3 mm, width 5.6 – 7.1 mm, height / width ratio 1.34 – 1.45, conical, light. Protoconch seemingly smooth, of ca 1 whorl, 0.34 – 0.35 mm in width. Teleoconch of 6.5 – 7.0 slightly convex whorls. Sculpture of 5 – 6 flat, closely set abapical spiral cords of about the same strength, including the 2 – 3 peripheral ones forming the peripheral cord, slightly shouldered, and 8 – 9 regularly spaced, basal spiral cords large and flat. Suture well incised. Teleoconch surface covered by barely visible prosocline growth striae irregularly set and interspaces adorned with strong prosocline lamellae. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with columellar callus thickened in the median portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright whitish- pinkish; remaining whorls and base cream or whitish striated by fine brownish-reddish opisthocline or orthocline lines. Málaga, Spain. M – O. J. sartorii Aradas & Maggiore, 1840, RUC, syntype of Trochus sartorii, height 10.2 mm. Acitrezza, Sicily, Italy. P – R. J. fraterculus Monterosato, 1889, CS-PM, height 5.4 mm, Borj El Hsar (34 ° 42 ′ 38 ″ N, 11 ° 09 ′ 04 ″ E), Kerkennah Islands, Tunisia. S – U. J. depictus (Deshayes, 1835), CS-PM, height 8.2 mm, Giannutri Island (42 ° 15 ′ 07.05 ″ N, 11 ° 05 ′ 31.51 ″ E), Tuscany, Italy, on P. oceanica, 20 m depth. V – X. J. ruscurianus (Weinkauff, 1868), BAU 2858.1, height 4.5 mm, Playazo de Rodalquilar, Spain. Scale bars = 2 mm. SOFT PARTS. Epipodial tentacles whitish, densely covered by tiny purplish speckles interconnected to create an irregular network and sparingly white spots. Foot background reddish-creamy or reddish-brown mottled with darker blotches and basal white strips; sole white. Cephalic tentacles whitish, with a dorsal purplish stripe and densely covered by small purplish speckles. Eye stalks red-brownish; eyes black encircled by dark brown with a ventral white spot. Snout reddish, creamy terminal band with a thin dark terminal strip.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF111FFDAE26CF902D121FAD8.taxon	distribution	Distribution Sicily (Aradas & Maggiore 1840), Lampedusa, Cyprus and Malta; Corsica; Puntebianche (Veli Rat) Croatia (Coen 1937); living specimens at depths of 10 – 42 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF111FFDAE26CF902D121FAD8.taxon	discussion	Remarks In the Monterosato collection, one vial (MCZR-M- 11728) is preserved containing 4 specimens with the label handwritten by Monterosato reading “ goniobasis Mont. Velirat! ”, which clearly refers to the taxon subsequently described as J. goniobasis Coen, 1937. Despite the slightly more elevated form, the sculpture is very similar to that of J. sartorii, and we consider the two taxa as synonyms. Jujubinus sartorii differs from J. exasperatus, J. pseudogravinae, J. gravinae and J. dianiensis sp. nov. in the complete or partial absence of tubercles on the spiral cords; from J. vexationis (Madeira) and J. swinneni sp. nov. (Canary Islands) in its stronger sculptures; from J. superum sp. nov. and J. vulgaris in its stouter outline, and stronger sculpture; from J. ruscurianus (Western Mediterranean) in its usually larger size and stronger sculpture (strong lamellae in the interspaces in J. sartorii vs fine prosocline lamellae in J. ruscurianus); from J. elenchoides in its smaller size and a fewer and stronger spiral cords; from J. kyrnos sp. nov. in its more slender shape and stronger sculpture (strong lamellae in the interspaces in J. sartorii vs fine prosocline lamellae in J. ruscurianus); from J. depictus mostly in the outline (more cyrtoconoid in J. depictus) colouration (bright whitish-pinkish in J. sartorii vs reddish, reddish brown or whitish-reddish background with white blotches in J. depictus). See under J. fraterculus, J. gofasi sp. nov., J. dianiensis, J. striatus, J. smaragdinus, J. inopinatus sp. nov., J. angustibasis sp. nov., J. dispar, for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF112FFD5E247FAB3D25FF871.taxon	description	SSH XIII – Fig. 7 P – R	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF112FFD5E247FAB3D25FF871.taxon	diagnosis	Diagnosis Shell of large size for the genus, not very solid, 4 – 5 elevated, flat, equidistant spiral cords, interspaces with fine prosocline lamellae, peripheral cord very evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF112FFD5E247FAB3D25FF871.taxon	materials_examined	Type material Syntypes TUNISIA • 8 dd; Sfax; “ Monotrochus fraterculus Monts Spugne di Sfax! ”; MCZR-M- 11705 / S • 26 dd; Sfax; “ M. fraterculus Monts (Zyziph.), Boll. Malacologico Italiano 1880 p. 220 spugne di Sfax ” MCZR- M- 11705 / S • 2 dd; Gulf of Gabès; “ Golfo di Gabes ”; MCZR-M- 11705 / S • 6 dd; “ Sfax ”; MCZR-M- 11705 / S • 13 dd; Mediterranean coasts of Northern Africa [unprecised locality]; “ Africa ”; MCZR-M- 11758 / S • 8 dd; Mediterranean coasts of Northern Africa [unprecised locality]; labelled “ C. d’Afr. ” (Appolloni et al. 2018); MCZR-M- 11758 / S. Other material examined TUNISIA • 1 dd; Sfax; “ smaragd. vr. Altior Mts Sfax, spugne ”; MCZR-M- 11761 • 34 dd; N of Zarzis; 33 ° 36 ′ 28 ″ N, 11 ° 04 ′ 04 ″ E; 0 – 7 m depth; MO • 3 dd; Djerba I.; 0.5 m depth; lagoon bioclastic sediment; LT • 1 dd; Djerba I.; beached; MO • 3 sh; Kerkennah; beached; MO • 4 dd; Kerkennah, Borj El Hsar Is.; beached; CS-PM • 150 dd; Kerkennah, Sidi Yousef; beached; CS-PM • 4 dd; Kerkennah, West of Gharbi I., Sidi Youssef; 0.5 m depth; LT • 2 lv; Kerkennah, Gharbi I., Sidi Youssef; 34 ° 45 ′ 50 ″ N, 11 ° 19 ′ 55 ″ E; 0 – 1 m depth; used for DNA sequencing; BAU 2314.1 – 2 • 1 lv; Kerkennah, Chergui I.; 34 ° 43 ′ 21 ″ N, 11 ° 09 ′ 21 ″ E; 0 – 2 m depth; used for DNA sequencing; BAU 2311.1 • 1 lv; Kerkennah, Chergui I.; 34 ° 43 ′ 21 ″ N, 11 ° 09 ′ 21 ″ E; 0 – 2 m depth; used for DNA sequencing; BAU 2312.1 • 4 dd; unprecised locality; “ Jujubinus sp. C. di Barberia ”; MCZR-M- 11762.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF112FFD5E247FAB3D25FF871.taxon	description	Description SHELL. Shell of large size for the genus, height 10.0 – 11.0 mm, width 7.5 – 8.0 mm, height / width ratio 1.34 – 1.38 conical, light. Protoconch seemingly smooth, of ca 0.8 whorls, 0.25 – 0.26 mm in width. Teleoconch of 5.5 – 6.5 slightly convex whorls. Sculpture of 4 – 5 flat, closely set abapical spiral cords of about the same strength, including the 2 peripheral ones forming the peripheral cord, slightly shouldered, and 6 – 7 regularly spaced, basal spiral cords large and flat. Spiral interspaces with thick and strong prosocline lamellae regularly set. Suture incised. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish or greyish; remaining whorls and base cream or brownish with alternating notches of colour brownish-reddish SOFT PARTS. Observed in alcohol preserved specimens, no dark terminal strip is present on the creamy band of the snout.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF112FFD5E247FAB3D25FF871.taxon	distribution	Distribution Kerkennah Islands (Gulf of Gabès, Tunisia), as a variety of J. unidentatus (Ghisotti & Melone 1975). Specimens reported under this name by Scuderi & Terlizzi (2012) from Apulia belong to another species. Currently J. fraterculus is known only from the Gulf of Gabès, Tunisia, and it can be considered endemic to that area; living specimens at depths of 0 – 2 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF112FFD5E247FAB3D25FF871.taxon	discussion	Remarks Jujubinus fraterculus differs from J. exasperatus, J. pseudogravinae, J. gravinae and J. dianiensis sp. nov. in the complete or partial absence of tubercles on the spiral cords; from J. vexationis (Madeira), J. swinneni sp. nov. (Canary Islands) and J. ruscurianus (western Mediterranean), in the more slender outline and stronger sculpture (with marked striae in the spiral interspaces in J. fraterculus vs thin prosocline striae in J. swinneni and J. ruscurianus); from J. superum sp. nov. and J. vulgaris in its stouter outline, and more marked sculpture; from J. elenchoides in its smaller size, and the fewer and slightly more marked spiral cords; from J. kyrnos sp. nov., in its more slender shape and more marked prosocline lamellae in the interspaces; from J. depictus) in its more slender outline (vs a more cyrtoconoid one in J. depictus) and in the colouration (whitish or greyish or brownish in J. fraterculus vs wine reddish, reddish brown or whitish-reddish background with white blotches in J. depictus); from J. sartorii in its smaller size with narrower base and more angled periphery, and in the colouration (whitish or greyish or brownish of J. fraterculus vs bright whitish-pinkish J. sartorii). See under J. gofasi sp. nov., J. dianiensis, J. striatus, J. smaragdinus, J. inopinatus sp. nov., J. angustibasis sp. nov., J. dispar, for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11CFFD6E25EFE90D18FFD53.taxon	description	SSH XIV – Fig. 7 S – U	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11CFFD6E25EFE90D18FFD53.taxon	diagnosis	Diagnosis Shell of large size for the genus, solid, 3 – 4 spaced and raised spiral cords, broad interspaces with very strong prosocline lamellae and fine growth lines, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11CFFD6E25EFE90D18FFD53.taxon	materials_examined	Type material Syntypes GREECE • 1 dd; Peloponnese; “ Morea ” (type locality); MCZR-M- 11682 / S ex collection Deshayes • 8 dd; Peloponnese; “ Morea ”; MNHN-IM- 2.000 - 31.122. Other material examined CORSICA • 4 lv; Saint Florent; 42 ° 42.3 ′ N, 9 ° 19.2 ′ E; 9 m depth; used for DNA sequencing; MNHN- IM- 2019 - 5498, 5671, 5688, 5903 • 1 lv; Sec de Giraglia; 43 ° 01.7 ′ N, 9 ° 24.4 ′ E; 21 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6378 • 1 lv; Centuri depths; 42 ° 59.5 ′ N, 9 ° 17.5 ′ E; 36 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6379 • 4 lv; North Cap Corse; 43 ° 02.6 ′ N, 9 ° 19.9 ′ E; 60 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6380, 6381, 6429, 12734 • 1 lv; North-West Cap Corse; 42 ° 59.6 ′ N, 9 ° 17.6 ′ E; 41 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12745 • 2 lv; Golfu di Sant’Amanza; 41 ° 25.2 ′ N, 9 ° 14 ′ E; 24 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12840, 12841 • 3 lv; Baie de Figari; 41 ° 28.3 ′ N, 9 ° 04 ′ E; 0 – 1 m depth; used for DNA sequencing; MNHN- IM- 2019 - 12852, 14030, 16774 • 1 lv; Sec St Antonio; 41 ° 20.4 ′ N, 9 ° 10.9 ′ E; 43 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17049 • 1 lv; between Gargalo Is. and Garganellu Is.; 42 ° 22 ′ N, 8 ° 32.5 ′ E; 28 – 30 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17620 • 1 lv; Punta Scandola; 42 ° 20 ′ N, 8 ° 33.6 ′ E; 12 – 17 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17621 • 1 lv; Punta Muchillina; 42 ° 20.1 ′ N, 8 ° 33.3 ′ E; 10 m depth; used for DNA sequencing; MNHN-IM- 2019 - 18474. CROATIA • 1 dd; unprecised locality; “ Dalm: ”; MCZR-M- 11740. FRANCE • 13 dd; Cannes; “ Jujubinus heilmanni Monts Cannes! ms ” plus “ Cannes 1885 ”; MCZR-M- 11708 • 1 dd; Bandol; “ Bandol D. Chatelet ”; MCZR-M- 11735. GREECE • 3 dd; Crete, Kanari; “ Kanari-Creta Maltzan ”; MCZR-M- 11735. ITALY • 7 dd; Elba I., Lacona; 10 m depth; bioclastic sediment; CS-PM • 28 dd; Elba I., Porto Azzurro, Naregno; 3 m depth; bioclastic sediment; LT • 2 lv; Elba I., Fetovaia; 42 ° 43 ′ 58.1 ″ N, 10 ° 09 ′ 13.1 ″ E; used for DNA sequencing; MNHN-IM- 2013 - 32668, 32670 • 2 lv; Elba Is., Fetovaia; 42 ° 43 ′ 58.1 ″ N, 10 ° 09 ′ 13.1 ″ E; used for DNA sequencing; MNHN-IM- 2013 - 5495, 5496 • 21 lv; Giannutri I.; 15 m depth; Posidonia; CS-PM • 5 lv; Giannutri I.; 42 ° 15 ′ 7.05 ″ N, 11 ° 5 ′ 31.51 ″ E; used for DNA sequencing; BAU 1793.2, 1793.4, RM 3 - Jco 1 to RM 3 - Jco 3 • 1 lv; Giannutri Is., Punta Secca, stn. GMM 2; 42 ° 15 ′ 41 ″ N, 11 ° 06 ′ 24 ″ E; 12 – 13 m depth; used for DNA sequencing; BAU 2668.2 • 2 lv; Giglio Is., Italy; 42 ° 22 ′ 7.82 ″ N, 10 ° 55 ′ 5.11 ″ E; used for DNA sequencing; RM 3 - Jco 4, RM 3 - Jco 5 • 4 dd; Santa Marinella; 15 – 20 m depth; fishing nets with Posidonia residues; LT • 27 dd; Torre Valdaliga; 25 m depth; bioclastic sediment; CS-PM • 4 dd; Marina di Camerota; 25 m depth; bioclastic sediment; CS-PM • 29 dd; La Maddalena Island; 25 m depth; bioclastic sediment; CS-PM • 37 dd; Golfo d’Arzachena; beached; CS-PM • 3 dd; Argentario; 30 m depth; MO • 2 dd; Giglio Is.; “ Isola del Giglio 1917 - Di Gestro ”; MCZR-M- 11761 • 50 dd; Giannutri Is.; 30 m depth; CS-PM • 8 dd; Capo Linaro; 35 m depth; MO • 15 dd; Tor Paterno MPA; 23 – 33 m depth; MO • 1 dd; Napoli; “ Jujubinus concinnus var. rubra Porro Napoli ”; MCZR-M- 11711 • 10 dd; Napoli; “ Mixtus. Monts Napoli-Bacoli (Praus) 900 ”; MCZR-M- 11702 • 3 dd; Napoli; “ Napoli ”; MCZR-M- 11758 • 10 dd; Napoli; “ J. litoralis Brusina Napoli! ”; MCZR-M- 11758 • 8 dd; Napoli; “ M. crassus M. l, Napoli ”; MCZR-M- 11740 • 2 dd; Capri Is.; “ Napoli Capri Z. mullip. e ”; MCZR-M- 11758 •. 4 dd; Capo Palinuro Occhi cave; 12 m depth; MO • 5 dd; Brindisi; 3 m depth; algae; LT. SICILY • 55 dd; Palermo; “ Pal. poco profondo ”; MCZR-M- 11746 • 2 dd; Salina Is.; 35 m depth; MO • 1 dd; Siracusa; “ Siracusa ”; MCZR-M- 11735 • 4 dd; Isola delle Correnti; 18 m depth; MO • 2 dd; Marettimo Is.; 20 m depth; MO • 14 dd; Levanzo Is., Punta Altarella; 37 ° 59 ′ N, 12 ° 21 ′ E; 25 m depth; MO • 1 dd; Favignana; 5 m depth; on Posidonia oceanica; LT • 8 dd; Lampedusa Is.; “ Juj. rectoconicus. M. Lampedusa ”; MCZR-M- 11740 • 5 dd; Pantelleria Is.; 30 m depth; MO • 6 dd; Pantelleria Bank; 37 ° 10 ′ N, 12 ° 05 ′ E; 20 – 50 m depth; MO.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11CFFD6E25EFE90D18FFD53.taxon	description	Description SHELL. Shell of large size for the genus, height 7.9 – 9.7 mm, width 5.3 – 6.1 mm, height / width ratio 1.36 – 1.75, conical, robust, slightly cyrtoconoid, not very bright. Protoconch seemingly smooth, of ca 1 whorl, 0.29 – 0.30 mm in width. Teleoconch of 7 – 7.5 flat and inclined spirals whorls. Sculpture of 5 – 6 spiral cords of about the same strength, separated by deep and wide interspaces regularly spaced with wide interspaces, including the 2 – 3 peripheral ones forming the peripheral cord. Whole teleoconch surface strongly sculptured by thick prosocline lamellae, regularly spaced. Base convex adorned with 7 – 8 spiral cords with wide interspaces sculptured by strong prosocline lamellae, umbilicus closed also in juveniles, covered with a whitish callus. Suture incised but not very evident. Aperture quadrangular, with the columellar callus not very thick in the basal portion. Interior of the aperture whitish or greenish-greyish nacreous. COLOURATION. Colour of the shell variable, reddish, reddish-brown or whitish-reddish background with sparse white blotches, occasionally with broad vertical whitish-beige oblique bands; base reddish, with blackish and whitish flammulae of different size, peripheral cord spotted with red, black and white. SOFT PARTS. Epipodial tentacles whitish, densely covered by tiny purplish speckles interconnected to create an irregular network. Foot background reddish or reddish-brown mottled with longitudinal darker stripes; sole white. Cephalic tentacles whitish, with a dorsal purplish stripe and densely covered by small purplish speckles. Eye stalks reddish or brownish; eyes black encircled by dark brown with a ventral white spot. Snout light or dark brown, creamy terminal band with a thick dark terminal strip.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11CFFD6E25EFE90D18FFD53.taxon	distribution	Distribution Mediterranean, from southern France, Italy, Sicily, Croatia and Greece; living specimens at depths of 0 – 45 m, occasionally down to 60 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11CFFD6E25EFE90D18FFD53.taxon	discussion	Remarks Despite having very seldom been reported, this is not a rare species. Ghisotti & Melone (1975: 170, fig.) treated it as a subspecies of J. striatus (“ Jujubinus striatus (L.), es. variegato tendente alla ssp. depictus ”), and in the same work they referred it to two forms with no taxonomic value: “ J. striatus depictus (Desh.) forma magnificus Coen coll. Coen, da Palermo ” and “ J. striatus depictus (Desh.) forma concinnus MTS coll. Coen, da Napoli ” (1975: 173, fig.). This is not Trochus concinnus Philippi, 1846 (currently Turbo). The actual range may be wider than currently known, simply due to misidentification with other taxa (e. g., Jujubinus striatus and related species: see below). It differs from J. exasperatus in the absence of tubercles on the spiral cords and in its colour pattern usually dominated by brownish with less redreddish, whereas red is largely dominant in most J. exasperatus populations. Jujubinus pseudogravinae is also similar to J. depictus, but the latter differs in its smaller average size, the more marked and regular sculpture, with spiral cords and regular interspaces vs a sculpture with spiral cords and interspaces of different sizes in J. pseudogravinae. It differs from J. ruscurianus (Western Mediterranean), J. vexationis (Madeira) and J. swinneni sp. nov. (Canary Islands) in its stronger sculpture and larger size. From J. elenchoides it differs in its smaller size, the fewer spiral cords (12 – 16 in elenchoides vs 5 – 6 in depictus) and the stronger sculpture; from J. gravinae, in the different sculpture, with tuberculous peripheral cord, never present in J. depictus. From J. superum sp. nov. (northern Adriatic), it differs in the colouration (dark greyish-blackish background colour with irregular black and white sagittal stripes in J. superum vs reddish, reddish brown or whitish-reddish background with white blotches in depictus). From J. kyrnos sp. nov., it differs in its scuplture (spaced and smooth abapical spiral cords in J. depictus vs weaker sculpture of flat and closely set abapical spiral cords in J. kyrnos). See under J. sartorii, J. fraterculus, J. gofasi sp. nov., J. dianiensis sp. nov., J. striatus, J. smaragdinus, J. inopinatus sp. nov., J. angustibasis sp. nov., and J. dispar, for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11EFFD1E243FD2BD709F808.taxon	description	SSH XV – Fig. 7 V – X	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11EFFD1E243FD2BD709F808.taxon	diagnosis	Diagnosis Shell of medium size for the genus, rather solid, 4 – 5 spaced spiral cords, elevated, broad interspaces with thick prosocline lamellae and growth lines, peripheral cord evident, high height / width ratio, colour red.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11EFFD1E243FD2BD709F808.taxon	materials_examined	Type material Syntypes of Trochus ruscurianus Weinkauff, 1868 ALGERIA • 3 dd; Algier (type locality); MCZR-M- 11780 / S, ex Weinkauff. Holotype of Jujubinus guanchus Curini-Galletti, 1985 CANARY ISLANDS • dd; Lanzarote Is., harbour of Arrecife (type locality); 28 ° 57 ′ N, 13 ° 32 ′ W; USNM 784714. Paratypes of Jujubinus guanchus Curini-Galletti, 1985 CANARY ISLANDS • 1 dd; same data as for holotype; USNM 784715 (paratype D) • 2 dd; same data as for holotype; RBINS 388 - 26439 (paratypes A, C). Other paratypes in the collections G. T. Poppe (Mactan) and M. Curini-Galletti (Sassari). Other material examined ALGERIA • 3 dd; Oran; labelled “ Calliostoma ruscurianus Weinkauff. Oran, récoltes Paul Pallary ”; MCZR-M- 11780 • 2 dd; Oran; ex Pallary collection; MCZR-M- 11779 • 14 dd; Alger; labelled “ Porto di Algeri (Lhotellier) ” ex Pallary collection; MCZR-M- 11780 • 4 dd; Alger; labelled “ Troruscurianus Weink-Algieri ”; MCZR-M- 11780 • 1 dd; Alger; labelled “ T. ruscurianus, Alger! Weink. ”; MCZR-M- 11780. CANARY ISLANDS • 1 lv; Lanzarote I., Puerto del Carmen, Playa de Matagorda; 28 ° 55 ′ 55.59 ″ N, 13 ° 36 ′ 46.62 ″ W; used for DNA sequencing, corresponding to J. guanchus; BAU 3771 – 3773 • 4 dd; Lanzarote Is., Arrecife; 2 m depth; MO. MOROCCO • 100 dd; Al Hoceima; beached; young and adult shells; CS-PM • 1 dd; Ceuta; beached; CS-PM • 4 lv; Torres de Alcalá, Cala Iris; 1 – 4 m depth; on brown algae; LT. SPAIN • 3 dd; Getares, Punta Carnero; beached; CS-PM • 51 dd; Algeciras, Getares; 20 m depth; washing of algae; LT • 6 lv; Getares, 4 km S of Algeciras; 20 m depth; washing of algae; LT • dd; Málaga; 25 m depth; CS-PM • 1 dd; Málaga; 2 m depth; CS-PM • 3 dd; Murcia, Águilas; MO • 3 dd; Málaga; 3 m depth; CS-PM • 2 dd; Cadiz; 2 m depth; CS-PM • 1 lv; Málaga; used for DNA sequencing; BAU 2076 • 5 lv; Málaga, Benalmádena; 36 ° 35.3 ′ N, 4 ° 31.7 ′ W; used for DNA sequencing; BAU 2664.1 to 2664.5 • 1 lv; Playazo de Rodalquilar; 36 ° 51 ′ 28 ″ N, 2 ° 00 ′ 06 ″ W; used for DNA sequencing; BAU 2858.1.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11EFFD1E243FD2BD709F808.taxon	description	Description SHELL. Shell of medium size for the genus, height 4.3 – 5.7 mm, width 3.7 – 4.7 mm, height / width ratio 1.16 – 1.30, conical, shiny in type material. Protoconch seemingly smooth, of ca 1 whorl 0.26 – 0.27 mm in width. Teleoconch of 5.5 – 6.0 slightly convex whorls. Sculpture of 5 – 6 spaced abapical spiral cords, smooth, of about the same strength, including the 2 – 3 peripheral ones forming the peripheral cord, and 7 – 8 regularly spaced, basal spiral flat cords. Suture incised. Teleoconch surface covered by thick, yet barely visible prosocline growth lamellae, irregular set. Base convex and rounded, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the lower portion. Interior of the aperture greenish whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish reddish or pinkish, remaining whorls and base bright reddish whitish or creamy white; spiral cords with alternating flammules, cream or reddish or yellowish, with yellow stripes in the interspaces. Sutural and peripheral cord articulated in red and cream or white. SOFT PARTS. Observed in alcohol preserved specimens, no dark terminal strip is present on the creamy band of the snout.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11EFFD1E243FD2BD709F808.taxon	distribution	Distribution Western Mediterranean, Alboran Sea until Almería, southern Spain and Algeria (Weinkauff 1867; Curini- Galleti & Palazzi 1980; Curini-Galletti 1982 b; Sánchez-Moyano et al. 2000); living specimens at depths of 1 – 20 m. In the neighbouring Atlantic from the Berlenga Islands and South Portugal to Morocco. Canary Islands (Pallary 1920; Cretella 1993; Gofas et al. 2011; Alf et al. 2020) probably referring to the morphotype described as J. guanchus.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11EFFD1E243FD2BD709F808.taxon	discussion	Remarks This is a morphologically very characteristic and common species in the western Mediterranean and neighbouring Atlantic. Morphologically, it can be diagnosed from J. exasperatus, J. pseudogravinae, J. gravinae, J. errinae and J. vexationis by the lack of tubercles or undulations on sutural cords and basal cords and for the strong spiral cords regularly spaced by deep interspaces. It differs from J. striatus, J. depictus, J. fraterculus, J. superum sp. nov., J. inopinatus sp. nov., J. kyrnos sp. nov., J. sartorii, J. smaragdinus, J. angustibasis sp. nov., J. gofasi sp. nov., and J. superum in the less marked sculpture with apparently smooth interspaces due to the very weak prosocline growth lines. The specimens in the Monterosato collection, with the label autograph by Weinkauff, are likely syntypes, and were the only available source to confirm the actual identity of this taxon (the original Weinkauff collection having largely been dispersed: Curini-Galletti & Palazzi 1980). Curini-Galletti (1985) described J. guanchus from the Canary Islands, based on a particular morphotype with ca six teleoconch whorls separated by incised sutures, and a sculpture of four smooth spiral cords crossed by numerous very fine markedly prosocline growth lines; the background colour is shiny olive-green, with white blotches arranged in irregular longitudinal band across the whorls and the peripheral cord, the basal ridges with regularly white dotted. We have assayed three specimens from Lanzarote, morphologically corresponding to typical J. guanchus, which proved to be genetically indistinguishable from Western Mediterranean typical J. ruscurianus. Jujubinus poppei, has been described on syntopic samples from Lanzarote, differing from J. guanchus in the different number of spiral cords (6 – 7 in J. poppei vs 4 in J. guanchus). See under J. gravinae, and J. superum sp. nov. for detailed comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	description	urn: lsid: zoobank. org: act: 30 C 0 A 441 - 53 A 9 - 4266 - A 5 CF-B 2 B 44133479 C SSH XVI – Fig. 8 A – C	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	diagnosis	Diagnosis Shell of medium size for the genus, not very solid, 6 – 7 spaced spiral cords, elevated, narrow interspaces with prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	etymology	Etymology The species name is after Dianium, the ancient name of Giannutri Island.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	materials_examined	Type material Holotype ITALY • lv; Giannutri I., Punta San Francesco (type locality); 42 ° 15 ′ 14 ″ N, 11 ° 06 ′ 13 ″ E; 10 – 15 m depth; used for DNA sequencing; MNHN-IM- 2000 - 30318 (ex BAU 2667.5). Paratypes ITALY • 1 lv; same data as for holotype; paratype 1, MNHN-IM- 2000 - 30319 • 2 lv; same data as for holotype; paratypes 2 – 3, MZB- 60155, MZB- 60156 • 1 lv; same data as for holotype; paratype 4, BAU 2359.6 • 2 lv; same data as for holotype; paratypes 6 – 7, 12, MCZR-M-TYPE 00097 • 34 lv; same data as for holotype; paratypes 5, 8 – 11,13 – 40, CS-PM • 3 lv; Giannutri Is., Italy; 42 ° 15 ′ 7.05 ″ N, 11 ° 05 ′ 31.51 ″ E; 10 – 15 m depth; used for DNA sequencing; RM 3 - Jdia 1 to RM 3 - Jdia 3. Other material examined CORSICA • 1 lv; Tour de Sponsaglia; 41 ° 29.2 ′ N, 9 ° 17.2 ′ E; 16 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12258 • 1 lv; same data as for preceding; used for DNA sequencing; MNHN- IM- 2019 - 12259 • 1 lv; Punta di U Capricciolu; 41 ° 25.7 ′ N, 9 ° 15.9 ′ E; 5 – 25 m depth; used for DNA sequencing; MNHN-IM- 2019 - 14245. ITALY • 6 lv; Giglio Island; 10 m depth; Cystoseira; CS-PM • 277 lv and dd; Giannutri I.; 15 m depth; Posidonia and Cystoseira; CS-PM • 17 dd; Giannutri I.; 18 m depth; bioclastic sediment; CS-PM • 10 lv; Giannutri I., Italy; 42 ° 15 ′ 7.05 ″ N, 11 ° 5 ′ 31.51 ″ E; used for DNA sequencing; BAU 2359.1 to 2359.10 • 5 lv; Giannutri I., Punta Secca, stn GMM 2; 42 ° 15 ′ 41 ″ N, 11 ° 06 ′ 24 ″ E; 12 – 13 m depth; used for DNA sequencing; BAU 2667.1 to 2667.5 • 3 lv; same data as for preceding; used for DNA sequencing; BAU 2669.1 to 2669.3 • 2 lv; same data as for preceding; used for DNA sequencing; BAU 2671.1, 2671.2 • 1 lv; same data as for preceding; used for DNA sequencing; BAU 2672 • Giannutri Is., ‘ Le Cerniette’, stn GMM 1; 42 ° 15 ′ 10 ″ N, 11 ° 05 ′ 32 ″ E; 10 – 17 m depth; used for DNA sequencing; BAU 3653 • Giannutri Is.; 42 ° 15 ′ 7.05 ″ N, 11 ° 5 ′ 31.51 ″ E; 10 – 15 m depth; used for DNA sequencing; RM 3 - Jgi 1, RM 3 - Jgi 2.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	description	Description (in parentheses data of the holotype) SHELL. Shell of medium size for the genus, height 5.4 – 8.0 (7.5) mm, width 3.7 – 5.5 (5.0) mm, height / width ratio 1.35 – 1.51, conical, not very solid, not very bright. Protoconch, of ca 1 whorl, width 0.24 – 0.25 (0.24) mm, sculptured by two spiral threads. Teleoconch of 5.5 – 6.5 (6.5) slightly convex whorls. Sculpture of 6 – 7 (6) prominent spiral cords of equal strength, interspaces well incised and slightly shouldered, and 6 – 7 narrow basal cords, regularly spaced, with interspaces wider than the cords. Peripheral cord pronounced, rounded and divided by a furrow, which produces four / five very light and parallel cords of the same size. Suture evident and well incised. Teleoconch surface covered by thick prosocline growth striae, irregularly set. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, columellar callus thickened in the middle portion, internally whitish or pinkish nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish-pinkish; remaining whorls and base with predominantly blackish brown background, with whitish blotches anteriorly bordered with dark brown line. SOFT PARTS. Epipodial tentacles rosy or creamy, covered by scattered very tiny white. Foot background creamy or reddish mottled with brownish, reddish or white blotches; sole white. Cephalic tentacles whitish or purplish, with a dorsal whitish stripe and covered by small light white speckles. Eye stalks brownish dorsally, whitish ventrally; eyes black encircled by white. Snout brownish to reddish, a creamy terminal band with a thin dark terminal stripe.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	distribution	Distribution Currently known only from the type locality (Giannutri Island) and the neighbouring Giglio Island, and from Corsica; living specimens at depths of 10 – 18 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF118FFD3E1CDFE90D65BFA50.taxon	discussion	Remarks A small proportion of specimens shows a reversed colouration, with whitish background with axial irregular reddish-brown blotches, very rare reddish specimens. There is a percentage of 10 % of specimens with the peripheral cord tuberculate until the last whorl. This is among the species of Jujubinus with the most restricted known range (along with J. catenatus Ardovini, 2006, J. errinae, J. curinii Bogi & Campani, 2006, J. trilloi Smriglio, Di Giulio & Mariottini, 2014, J. eleonorae Smriglio, Di Giulio & Mariottini, 2014, J. roseus Smriglio, Mariottini & Oliverio, 2026, and J. alboranensis Smriglio, Mariottini & Oliverio, 2015). This species can be confused with Jujubinus gravinae, which co-occurs at the type locality and has a very similar teleoconch colouration. However, J. dianiensis sp. nov. differs in its slenderer outline and the peripheral cord with tubercles only on the first whorls vs tubercles on the peripheral cord of all whorls in J. gravinae. Jujubinus dianiensis differs from Jujubinus angustibasis sp. nov., J. kyrnos sp. nov., J. superum sp. nov., J. gofasi sp. nov., J. elenchoides Monterosato, 1884, J. ruscurianus, J. vulgaris, J. depictus, J. sartorii, and J. fraterculus in the tubercles on the peripheral cord of the early whorls, absent in the other species. It also differs from J. exasperatus and J. pseudogravinae in the lack of tubercles on the spiral cords; from J. vexationis (Madeira) and from J. swinneni sp. nov. (Canary Islands) in the lack of the spiral cords on the peripheral cord and for the narrower base.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	description	urn: lsid: zoobank. org: act: B 77 E 0260 - 2 AD 4 - 42 A 8 - 8 A 7 E-BDAEA 7 F 3 A 4 DF SSH XVII – Fig. 8 D – F	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	diagnosis	Diagnosis Shell of large size for the genus, elevated and robust, with dense, raised and non-granulose spiral cords, wide interspaces, sculptured by weak, minute and prosocline lamellae; distinct peripheral cord, narrow base, umbilicus closed, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	etymology	Etymology The species name is from the Latin ‘ angustus ’ (= ‘ narrow’) and ‘ basis ’ (= ‘ base’), referring to the narrow base.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	materials_examined	Type material Holotype SICILY • lv; Costa Saracena-Castelluccio (type locality); 37 ° 18 ′ 33.68 ″ N, 15 ° 07 ′ 58.08 ″ E; 5 m depth; on P. oceanica; used for DNA sequencing; MNHN-IM- 2000 - 40314 (ex RM 3 - Jst 4). Paratypes SICILY • 2 lv; same data as for holotype; used for DNA sequencing; RM 3 - Jst 5, RM 3 - Jst 6. Other material examined CORSICA • 1 dd; Bastia; “ Bastia (Caziot) ”; MCZR-M- 11761 • 3 dd; Bastia; “ JuJ. di Bastia ”; MCZR-M- 11738. CROATIA • 27 dd; Zara; “ Zizyphinus parvulus Brus. non Phil. Zara ”; MCZR-M- 11702 • 6 dd; Velirat; “ Pictus Monts. Velirat! ”; MCZR-M- 11754 • 79 dd plus, 17 dd; Velirat; “ J. turritus I più Velirat Dal. Copia a Tomlin ed altri e al Dr. Bavay 1916 ” “ varie forme Velirat!! 1910 ”; MCZR-M- 11778 • 1 dd; Molat; “ Melada (Klec.) ”; MCZR-M- 11748 • 1 dd; Velirat; “ Velirat! 1909 ”; MCZR-M- 11761 • 2 dd; Velirat; “ Velirat !! (Dalm.) 1910 ”; MCZR-M- 1177. FRANCE • 2 dd; Nice; “ Nice (Caziot) ”; MCZR-M- 11694. GREECE • 5 dd; Athens Varkiza; 12 m depth; bioclastic sediment; LT • 2 dd; Paros; “ Paros vr. Kviegis ”; MCZR-M- 11738 • 3 dd; Mt. Athos; “ Mt Athos ”; MCZR-M- 11738 • 3 dd; Crete, Kanari; “ Kanari-Creta- Maltzan ”; MCZR-M- 11738 • 2 lv; Vai, Astypalea Is., Greece; 36 ° 35 ′ 13 ″ N, 026 ° 24 ′ 10 ″ E; 1 – 6 m depth; 24 Jun. 2017; under rocks, used for DNA sequencing; BAU 1375.1, 1375.2 • 4 lv; Vai, Astypalea Is., Greece; 36 ° 35 ′ 13 ″ N, 026 ° 24 ′ 10 ″ E; 1 – 6 m depth; 24 Jun. 2017; under rocks, used for DNA sequencing; BAU 3037.1 to 3037.4. ITALY • 4 dd; Genova; “ Zampa – Genova Bofill ”; MCZR-M- 11681 • 2 dd; Torre Valdaliga; 20 m depth; bioclastic sediment; CS-PM • 1 dd; Civitavecchia; LT • 2 dd; Ponza Is.; 3 m depth; LT • 1 dd; Anzio harbour; “ Porto D’Anzio ”; MCZR-M- 11761 • 12 dd; Napoli; “ Napoli ”; MCZR-M- 11681 • 2 dd; Napoli; “ Napoli ”; MCZR-M- 11681 • 9 dd; Napoli; “ Napoli (Praus) ”; MCZR-M- 11723 • 1 dd; Napoli; “ JuJ. concinnus vr. Rubra Napoli (Praus) 1906 ”; MCZR-M- 11711 • 3 dd; Napoli; “ Nap. ”; MCZR-M- 11748 • 4 dd; Napoli; “ Napoli ”; MCZR-M- 11758 • 2 dd; Napoli?; “ carminum Napoli? ”; MCZR-M- 11760 • 1 dd; Napoli; “ Napoli ”; MCZR-M- 11770 • 24 dd; Capri I.; “ Capri! (Bellini) ”; MCZR-M- 11681 • 3 dd; Capri I.; “ Capri ”; MCZR-M- 11723 • 102 dd; Le Castella; 25 m depth; bioclastic sediment; CS-PM • 1 dd; Tricase; “ Tricase (Adriatico) ”; MCZR-M- 11735 • 43 dd; Monopoli; beached; CS-PM • 8 dd; Brindisi; 4 m depth; bioclastic sediment; LT • 2 dd; Tricase Marina; 3 m depth; bioclastic sediment; CS-PM • 2 dd; Porto Cesareo; 3 m depth; bioclastic sediment; CS-PM • 1 lv; Lecce, S. Isidoro; used for DNA sequencing; BAU 1679.3 • 3 lv; Torre Colimena; 40 ° 17 ′ 40 ″ N, 017 ° 45 ′ 20 ″ E; 2 – 7 m depth; Aug. 2007; under rocks; used for DNA sequencing; BAU 1726.2, 1726.4, 1726.5 • 3 lv; Punta Prosciutto; 40 ° 17 ′ 37 ″ N, 17 ° 45 ′ 47 ″ E; used for DNA sequencing; BAU 2658.1, 2658.4, 2658.6 • 1 lv; Nardò, Santa Maria al Bagno; used for DNA sequencing; BAU 3051.1 • 1 lv; Otranto, Punta Remiti; used for DNA sequencing; BAU 3050.1. SARDINIA • 23 dd; unprecised locality; “ striatus L. Sardegna ”; MCZR-M- 11685 • 1 dd; unprecised locality; “ Sard. a ”; MCZR-M- 11712 • 1 dd; unprecised locality; “ Sardegna !! ”; MCZR-M- 11758. SICILY • 8 dd; Palermo; “ Palermo ”; MCZR-M- 11694 • 10 dd; Palermo; “ Jujubinus depictus Desh = sartori = gravesi vr. Palermo ”; MCZR-M- 11681 • 2 dd; Palermo; “ Palermo ”; MCZR-M- 25359 • 34 dd; Palermo; “ Mirulinus elenchoides Monts Palermo ” [including 3 dd of Jujubinus exasperatus in the same vial], ex coll. Beltrani; MCZR-M- 25293 • 24 dd; Palermo; “ Mirulinus elenchoides Monts Palermo ”; MCZR-M- 25293 • 47 dd; Palermo, Arenella; “ Mirulinus finitimus, M. Arenella! ”; MCZR-M- 11714 • 7 dd; Palermo; “ Paler. ”; MCZR-M- 11759 • 1 dd; Palermo; “ Pal! Feb. 1916 ”; MCZR-M- 11759 • 18 dd; Palermo, Aspra; “ J. depictus Pal. All’Aspra giug. 85 ”; MCZR-M- 11759 • 17 dd; Palermo, harbour; “ Jujubinus depictus De dd. Var. ex col. Rada Palermo. Beltrani ” [2 dd of Jujubinus exasperatus in the same vial]; MCZR-M- 11759 • 92 dd; Palermo; “ Palermo! J. multistriatus Monts. Inedito ”; MCZR-M- 11772 • 24 dd; Palermo; “ J. multistriatus Monts pallescens assieme al tipo Palermo ”; MCZR-M- 11772 • 18 dd; Palermo; “ Palermo ”; MCZR-M- 11749 • 18 dd; Palermo; “ JuJ. Depictus di Romagnolo D. Roccand 1903 ”; MCZR-M- 11749 • 15 dd; Palermo; “ var. fulvo-picta Paler. ” in a single vial together with 3 shells of J. striatus; MCZR-M- 11749 • 1 dd; Palermo; “ Pal. ”; MCZR-M- 11757 • 20 dd; Palermo harbour; “ J. depictus porto di Palermo ”; MCZR-M- 11681 • 1 dd; Ustica Is.; “ Ustica! 1918 ”; MCZR-M- 11759 • 3 lv; San Vito Lo Capo, Sicily; used for DNA sequencing; BAU 2074.1 to 2074.3 • 27 dd; Favignana Is.; beached; CS-PM • 3 dd; Lipari Is.; “ Lipari ” [plus 6 dd with no data]; MCZR-M- 11682 • 1 lv; Castel di Tusa; 3 m depth; on Posidonia oceanica; CS-PM • 12 dd; Messina, Ognina; “ Ognina Brug ”; MCZR-M- 11684 • 21 dd; Messina; “ J. depictus vr funerea Sull. Boll. Mal. Ital. 1889 Messina ”; MCZR-M- 11681 • 10 lv; Costa Saracena-Castelluccio; 37 ° 18 ′ 33.68 ″ N, 15 ° 07 ′ 58.08 ″ E; 5 m depth; on Posidonia oceanica; CS-PM • 5 dd; Messina, Ognina; “ Ognina ”; MCZR-M- 11684 • 19 dd; Trapani; “ Trapani ”; MCZR-M- 11684 • 1 dd; Trapani; “ Trap? ”; MCZR-M- 11684 • 5 dd; Catania, Ognina, N of port; 37 ° 31 ′ 55.4 ″ N, 15 ° 07 ′ 14.3 ″ E; 0.5 – 3 m depth; on brown algae; LT • 9 dd; Ognina; fishing nets; LT • 29 dd; Gulf of Catania; fishing nets; LT • 1 dd; Gulf of Catania; fishing nets; LT • 2 dd; Messina; “ Messina ”; MCZR-M- 11756 • 2 dd; Messina; “ Messina (Sulliotti) ”; MCZR-M- 11759 • 9 dd; Trapani; “ Trapa [ni] ”; MCZR-M- 11759 [+ 1 dd of Jujubinus gravinae in the same vial] • 1 dd; Ognina; “ J. pulchellus Ognina! M. ”; MCZR-M- 11759 • 4 dd; Sciacca; “ Sciacca ”; MCZR-M- 11772 • 8 dd; Messina; “ Zizyphinus incertus Monts Messina! ” [including 2 dd labelled “ D. Praus Nap? ”, and 3 dd of Jujubinus exasperatus in the same vial]; MCZR-M- 11776 • 13 dd; Trapani; “ Trapani ”; MCZR-M- 11758, MT • 11 dd; Messina; “ Messina! funereus, Sull ”; MCZR-M- 11711 • 1 dd; Messina; “ J. armillatus. Messina Monts. Jujubinus Renieri ms. = depictus, vr elegans Sull. 1889, p. 69 ”; MCZR-M- 11711 • 1 dd; Messina, Santa Romina; “ J. funereus, Sull. S. Romina Messina Bull. Mal. Ital. 1889, p. 69 come J. depictus vr- funereus ”; MCZR-M- 11711 • 1 dd; Magnisi; “ Magnisi ”; MCZR-M- 11770 • 7 dd; unprecised locality; “ Sicilia ”; MCZR-M- 11770. SPAIN • 1 dd; Almería; “ Almeria Spain Tr. Smaragdinus? ”; MCZR-M- 11760 • 17 dd; Menorca Is., Mahón; “ Mahon ”; MCZR-M- 11694.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	description	Description (in parentheses data of the holotype) SHELL. Shell of large size for the genus, height 6.3 – 10.2 (7.6) mm, width 4.6 – 7.2 (4.8) mm, height / width ratio 1.36 – 1.58 (1.58), conical, light. Protoconch of ca 1 whorl, 0.24 – 0.25 (0.24) mm in width, sculptured by two spiral threads. Teleoconch of 6.5 – 7.5 (7.0) slightly convex whorls. Sculpture of 4 – 7 (7) flat, closely set abapical spiral cords of about the same strength, 2 – 3 (3) peripheral ones forming the peripheral cord, slightly stepped, and 5 – 6 (6) regularly spaced, basal spiral cords large and flat. Suture little incised. Teleoconch surface covered by barely visible prosocline irregularly set growth striae, and small but evident prosocline lamellae in the interspaces. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, greyish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright greyish or brownish or greenish; remaining whorls and base with alternating brownish-greyish and greenish colour blotches, with scattered small whitish patches on the spiral cords. SOFT PARTS. Epipodial tentacles translucent, densely covered by scattered very tiny black and white speckles. Foot background creamy mottled with brown or reddish irregularly-shaped blotches, and covered by white speckles; sole white. Cephalic tentacles translucent, with two dorsal brown stripes and densely covered by small whitish and brownish speckles. Eye stalks brownish dorsally, creamy ventrally; eyes black. Snout dark brown, a creamy terminal band with a thin dark terminal stripe.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	distribution	Distribution Almost the entire Mediterranean Sea, from Spain to Greece; living specimens at depths of 1 – 7 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF11BFFCCE1C3FA2BD75BFBEA.taxon	discussion	Remarks It differs from J. exasperatus and J. pseudogravinae in the lack of tubercles on the spiral cords; from J. gravinae, J. dianiensis sp. nov. and J. vexationis in the absence of tubercles or undulations on the peripheral cord; from J. swinneni sp. nov. (from Canary Islands) in its larger size and different colouration (whorls and base with alternating brownish-greyish-greenish colour patches on the spiral cords in J. angustibasis sp. nov. vs alternating whitish or reddish or blackish spots on spiral cords in J. swinneni); from J. depictus, J. sartorii, J. fraterculus, J. gofasi sp. nov., and J. smaragdinus in its weaker sculpture and barely evident prosocline lamellae in the interspaces; from J. superum sp. nov. and J. kyrnos sp. nov. in its larger size and different colouration (brownish-greyish-greenish colour patches on the spiral cords in J. angustibasis vs dark greyish-blackish background in J. superum and whitish greyish or beige in J. kyrnos); from J. ruscurianus in its more slender outline and less marked and more spaced spiral cords; from J. elenchoides in its fewer spiral cords; from J. inopinatus sp. nov. in its less slender outline and narrower spiral interspaces; from J. striatus in its narrower base; from J. vulgaris in its more numerous spiral cords (4 – 6 in J. angustibasis vs 3 in J. vulgaris).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	description	urn: lsid: zoobank. org: act: C 964 DC 4 E-B 3 DB- 443 F- 9 FA 6 - A 680933 ADF 98 SSH XVIII – Fig. 8 G – I	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, slender, 5 spaced and elevated spiral cords, broad interspaces with fine prosocline lamellae and growth lines, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	etymology	Etymology The name inopinatus is from the Latin: ‘ unexpected’.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	materials_examined	Type material Holotype SICILY • lv; San Vito Lo Capo (type locality); used for DNA sequencing; MNHN-IM- 2000 - 40313 (ex BAU 1680). Other material examined ITALY • 1 dd; Napoli; labelled “ Nap! ”; MCZR-M- 00092.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	description	Description (in parentheses data of the holotype) SHELL. Shell of medium size for the genus, height 9.0 – 9.1 (9) mm, width 5.7 – 6.0 (5.7) mm, height / width ratio 1.51 – 1.58 (1.58). Protoconch seemingly smooth, of ca 1 whorl, ca 0.29 mm in width. Teleoconch of 6.5 – 7.5 (7.5) slightly convex whorls. Sculpture of 5 not very strong and not granulose spiral cords of about the same strength and regularly spaced, including the 2 peripheral ones, forming the broad peripheral cord; well-marked suture. Interspaces with fine and dense prosocline lamellae regularly set. Base convex, with 5 – 6 concentric black and white spiral cords regularly spaced, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls glossy pinkish with whitish blotches, with spiral cords articulated in black and white. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	distribution	Distribution Species currently known only from the type locality, San Vito Lo Capo, Sicily, and Gulf of Naples, Italy: the holotype has been collected at unprecised depth, but in very shallow waters.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF104FFCFE1C9FBC5D01DFB00.taxon	discussion	Remarks Jujubinus inopinatus sp. nov. is currently known from only two specimens. It is very similar to J. angustibasis sp. nov., but the molecular analysis (on the holotype) warrants very confidently the separation. The paratype was found in the Monterosato collection (MCZR-M- 11760) and it is almost identical in shape and colouration to the holotype; it is possible that other specimens exist in public or private collections, misidentified as ‘ J. striatus ’ (with reference to J. angustibasis). It differs from J. exasperatus and J. pseudogravinae in the lack of tubercles on the spiral cords; from J. gravinae, J. dianiensis sp. nov. and J. vexationis in the absence of tubercles or undulations on the peripheral cord; from J. striatus in its more slender outline and in the more spaced spiral cords and finer basal cords; from J. elenchoides in its smaller size and the fewer spiral cords; from J. superum sp. nov. and J. kyrnos sp. nov., in its larger size and the reddish colour (absent in the other two species); from J. depictus, J. sartorii, J. fraterculus, J. gofasi sp. nov. and J. smaragdinus in its weaker sculpture with finer prosocline lamellae in the interspaces; from J. ruscurianus in the more slender outline and the fewer, more spaced and less marked spiral cords; from J. vulgaris in the more numerous spiral cords (5 vs normally 3 in J. vulgaris) with narrower interspaces.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF107FFC8E227FA9BD775FCFD.taxon	description	SSH XIX – Fig. 8 J – L	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF107FFC8E227FA9BD775FCFD.taxon	diagnosis	Diagnosis Shell of large size for the genus, very solid, 5 – 6 well spaced spiral cords, elevated, deep interspaces with internally strong prosocline spaced apart lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF107FFC8E227FA9BD775FCFD.taxon	materials_examined	Type material Lectotype of Trochus striatus Linnaeus, 1758 MEDITERRANEAN • dd; Mediterranean (type locality); lectotype here designated; LSL. 508, The Linnean Society. Paralectotypes of Trochus striatus Linnaeus, 1758 MEDITERRANEAN • 6 dd; Mediterranean; Peter Dance’s label PZ 0010802; LSL. 508, The Linnean Society. Syntypes of Jujubinus aequistriatus Monterosato, 1884 SICILY • 45 dd; Palermo; NA.; labelled “ Tr. (Jujub.) aequistriatus Palermo! ”; MCZR-M- 11688 / S • 8 dd; Palermo, Mondello; NA.; labelled “ J. aequistriatus Monts detrito Mondello ” • 8 dd; Palermo, Mondello; NA.; labelled “ J. Aequistriatus Mont. Detrito Mondello ”; MCZR-M- 11688 • 25 dd; Palermo, Arenella; NA.; labelled “ aequistriatus di Arenella, 1918 assieme al littoralis ”; MCZR-M- 11521 • 3 dd; Palermo and Trapani; NA.; labelled “ Paler Trapa ”; MCZR-M- 11694. Other material examined ALGERIA • 4 dd; Annaba; “ Bône-Alg. Hangenmuller ”; MCZR-M- 11682 • 5 dd; Algeri; “ Algeri (Joly) ”; MCZR-M- 11682 • 7 dd; Oran; “ attenuans Oran e Argelia Pallary ”; MCZR-M- 11682 • 2 dd; unprecised locality; “ coll Weink. ”; MCZR-M- 11682. CROATIA • 1 dd; Brac I.; “ Dalm. Bras ”; MCZR-M- 11682. FRANCE • 13 dd; Brest; “ Tr. Striatus L. Brest dedit Petit ”; MCZR-M- 11680 • 25 dd; Grand Camp- Maisy; “ Mi. striatus L. Grand Camp-Calvados, dedit Chapar ”; MCZR-M- 11680 • 1 dd; Arcachon; “ JuJ. Striatus, Auct. Arcachon! Atl-Franc ”; MCZR-M- 11680 • 2 dd; La Ciotat; “ La Ciotat!. Bartesago ”; MCZR-M- 11712 • 3 dd; Bandol; “ Bandol 1908 ”; MCZR-M- 11735 • 4 dd; La Ciotat; “ La Ciotat ”; MCZR-M- 11743 • 7 dd; Port Vendres; “ Trochus striatus Lin. Port Vendre ”; MCZR-M- 11776 • 2 dd; Toulon; “ Toulon (Caziot) ”; MCZR-M- 11775 • 12 dd; unidentified locality; “ Portane Pauls littoral ”, ex coll Pallary; MCZR-M- 11775. GREAT BRITAIN • 3 dd; unprecised locality; “ England ”; MCZR-M- 11694 • 6 dd; Falmouth; “ Trochus striatus L. Riants Falmouth ”; MCZR-M- 11756 • 6 dd; Devonshire; “ Trochus striatus L. Jorbay Devonshire ” plus “ Mi. striatus L. Jorbay (Devon.) England dedit Venticuss ”; MCZR-M- 11757 • 5 dd; Guernsey I.; “ Is Guernsay ”; MCZR-M- 11694. GREECE • 1 dd; Astypalea I.; 10 m depth; fishing nets; LT • 28 dd; Astypalea I., near Maltezana; 10 m depth; on Posidonia oceanica; LT. ITALY • 2 dd; Castiglioncello; “ Castiglioncello ”; MCZR-M- 11757 • 6 dd; Terracina; “ Terracina ”; MCZR-M- 11748 • 2 dd; Anzio; “ Porto d’Anzio. Da Praus col nome di mixtus sine loco! D. R. ”; MCZR-M- 11682 • 1 dd; Anzio, harbour; “ Porto d’Anzio ”; MCZR-M- 11735 • 1 dd; Napoli; “ hieroglyphus. Monts. Napoli! ”; MCZR-M- 11748 • 1 dd; Napoli; “ propinquus Monts desiderato Napoli ”; MCZR-M- 11749 • 1 dd; Napoli; “ Napoli ”; MCZR-M- 25224 • 1 dd; Chioggia; “ Chioggia Chiamenti ”; MCZR-M- 11758 • 1 dd; Taranto?; “ Taranto? (D. Praus) ”; MCZR-M- 11758. LYBIA • 1 dd; Tripoli; “ Tripolit Marsa al Kmescia ”; MCZR-M- 11757. MALTA • 1 dd; unprecised locality; “ Malta ”; MCZR-M- 11682 • 1 dd; unprecised locality; “ Malta? ”; MCZR-M- 11720 • 2 dd; unprecised locality; “ Malta ”; MCZR-M- 11735 • 1 dd; unprecised locality; “ ubi? Malta? ”; MCZR-M- 11758 • 2 lv; Gozo Is., Xwejni Bay; 36 ° 04 ′ 48.62 ″ N, 14 ° 14 ′ 50.80 ″ E; 10 m depth; on P. oceanica, used for DNA sequencing; BAU 2675.1 – 2. MOROCCO • 1 dd; Tanger; “ Tanger ”; MCZR-M- 11758 • 9 dd; Casablanca; “ J. strigosus (Gm) Ph Casa Blanca Marocco ”; MCZR-M- 11715. PORTUGAL • 5 dd; Setubal, Portinho de Arrabida; fishing boat; CS-PM. SARDINIA • 6 dd; Cagliari; “ Cagliari Fra Piero 1892 ”; MCZR-M- 11682 • 49 dd; San Pietro I.; “ JuJ. microconus copia a Pallary S. Pietro-Sard. Hangenmuller ”; MCZR-M- 11712 • 8 dd; Golfo d’Arzachena; beached; CS-PM • 1 dd; unprecised locality; “ Sardegna ”; MCZR-M- 11712 • 9 dd; unprecised locality; “ Sard! De Ribbis 1912 ”; MCZR-M- 11743 • 1 dd; unprecised locality; “ Sardegna ”; MCZR-M- 11748 • 1 dd; unprecised locality; “ (Praus) Sardegna ”; MCZR-M- 11749 • 1 dd; unprecised locality; “ Sardegna Chafers ”; MCZR-M- 11758 • 1 dd; unprecised locality; “ Sardegna Mme Bibbin ”; MCZR-M- 11758 • 1 dd; unprecised locality; “ Sardegna ”; MCZR-M- 11770. SICILY • 12 dd; Palermo; “ Pal. Euggeus 1912 ”; MCZR-M- 11684 • 4 dd; Palermo; “ Paler ”; MCZR-M- 11684 • 1 dd; Palermo; “ Pal. ”; MCZR-M- 11684 • 2 dd; Palermo; “ Prominulus Pal. Littorale ”; MCZR-M- 11684 • 7 dd; Palermo; “ J. rotulatus Palermo ”; MCZR-M- 11711 • 40 dd; Palermo; “ JuJ. Di Pal. Nel Porto. 89 ”; MCZR-M- 11711 • 5 dd; Palermo; “ Palermo ”; MCZR-M- 11715 • 4 dd; Palermo; “ P ”, written on the cork stopper of the vial; MCZR-M- 11735 • 3 dd; Palermo; “ var. fusco-picta Paler. ” [in a single vial together with 15 shells of J. angustibasis]; MCZR-M- 11749 • 6 dd; Palermo; “ P ”, written on the cork stopper of the vial; MCZR-M- 11735 • 4 dd; Palermo; “ nel Porto di Palermo ”, freak; MCZR-M- 11761 • 14 dd; Palermo; “ Porto di Pal! ”; MCZR-M- 25359 • 9 dd; Palermo, harbour; “ Porto di Palermo ”; MCZR-M- 11735 • 5 lv; Palermo, Barcarello; 3 m depth; used for DNA sequencing; BAU 2069.1, 2069.3 to 2069.6 • 1 dd; Palermo, Romagnolo; “ Romagnolo det. Beltrani, 1918 ”; MCZR-M- 11749 • 1 dd; Palermo?; “ Trochus unidentatus var- Philippi Palermo? Da Benoit ”; MCZR-M- 11761 • 1 dd; Palermo, Punta Raisi; 38 ° 10 ′ 47.0 ″ N, 13 ° 05 ′ 21.1 ″ E; beached; LT • 3 dd; Trapani; “ Trap ”; MCZR-M- 11684 • 3 dd; Trapani; “ Trapa ”; MCZR-M- 11684 • 12 dd; Favignana I.; beached; CS-PM • 5 dd; Favignana I.; beached; CS-PM • 1 dd; Mazzara del Vallo?; “ Mazzara? ”; MCZR-M- 11746 • 1 dd; Sciacca; “ pulchrae-colorata Sciacca ”; MCZR-M- 11756 • 3 lv; Castellammare del Golfo; 38 ° 01 ′ 48.05 ″ N, 12 ° 52 ’ 58.32 ″ E; 2 m depth; used for DNA sequencing; RM 3 - Jst 1 to RM 3 - Jst 3 • 8 dd; Ognina; “ Trap. 2 Ognina ”; MCZR-M- 11756 • 3 dd; Messina; “ J. renieri, Messina lo stesso alle Baleari ”; MCZR-M- 11758 • 2 lv; Mezzoforno-Cefalù; 5 m depth; Posidonia; CS-PM • 1 dd; Lampedusa; “ Lampedusa ”; MCZR-M- 11746 • 2 dd; Lampedusa; “ Lampedusa ”; MCZR-M- 11749. SPAIN • 4 dd; Algeciras; “ Algeciras (Kobelt) anche di Gibilterra (Tomlin) ”; MCZR-M- 11682 • 43 dd; Getares, 4 km S of Algeciras; 20 m depth; washing of algae; LT • 2 dd; Gibraltar; “ Gibilterra (Ponsonby) ”; MCZR-M- 11680 • 3 dd; Gibraltar; “ Gibilterra ”; MCZR-M- 11681.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF107FFC8E227FA9BD775FCFD.taxon	description	Description (in parentheses data of the lectotype) SHELL. Shell of large size for the genus, height 7.0 – 12.3 (6.8) mm, width 6.1 – 8.2 (5.3) mm, height / width ratio 1.35 – 1.51, conical, very solid. Protoconch seemingly smooth, of ca 1 whorl, 0.31 – 0.32 μm in width. Teleoconch of 7.5 (7.5) slightly convex whorls. Sculpture of 5 – 7 (6) flat, closely set abapical spiral cords of about the same strength, including the 2 – 3 (3) peripheral ones forming the peripheral cord, slightly shouldered, and 8 – 9 (9) regularly spaced, basal spiral cords narrow and marked. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae, and strong prosocline lamellae. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright pinkish-cream; remaining whorls and base cream or hazelnut colour striated by fine brownish-reddish opisthocline or orthocline lines. SOFT PARTS. Epipodial tentacles translucent, densely covered by very tiny white speckles. Foot background creamy mottled with brown or reddish irregularly-shaped blotches, and covered by white and brownish speckles; sole white. Cephalic tentacles translucent, with two dorsal brown stripes and covered by small whitish and brownish speckles. Eye stalks brownish dorsally, creamy ventrally; eyes black. Snout dark brown, a creamy terminal band with a thin dark terminal stripe.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF107FFC8E227FA9BD775FCFD.taxon	distribution	Distribution Atlantic, from Great Britain to Spain, and across the entire Mediterranean Sea; living specimens at depths of 2 – 10 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF107FFC8E227FA9BD775FCFD.taxon	discussion	Remarks The binomen Jujubinus striatus has been widely used in the past for specimens belonging to a number of species (several cryptic), not all of them phylogenetically related. We restrict here the concept of Trochus striatus to the broad-based specimens in the series of Linnean original syntypes because the narrow-base forms are a complex of different species, as evidenced by the present work. We note that a single shell of the Linnean type series is a specimen of J. angustibasis sp. nov. Therefore, we have selected one of the broad-based specimens as lectotype of Trochus striatus, to stabilize the use of the name. Jujubinus striatus differs from J. vexationis, J. swinneni sp. nov., J. elenchoides, J. superum sp. nov., J. gofasi sp. nov., J. kyrnos sp. nov., J. dianiensis sp. nov., and J. fraterculus in its broad base and the less slender outlline; from J. depictus, J. ruscurianus, and J. sartorii in the more regular and less marked sculpture; from J. gravinae in its larger size and the peripheral cord not crenulated.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF100FFC5E26DFCD0D259F9A2.taxon	diagnosis	Diagnosis Shell of medium size for the genus, elevated, solid, 5 – 7 well spaced spiral cords with not very deep interspaces with internally numerous prosocline lines and lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF100FFC5E26DFCD0D259F9A2.taxon	materials_examined	Type material Trochus smaragdinus (lot MCZR-M- 11700, label in the box: “ T. (Zizyphinus) striatus, Lin. var. smaragdina, Monts Boll. M. Ital. 1880, p. 220. C. D’Afr.! ”). Lectotype of Trochus smaragdinus Monterosato, 1880 TUNISIA • dd; Sfax; labelled “ Mirulinus smaragdinus Monts. Sfax ”; lectotype here designated; MCZR- M- 11700 / L Paralectotypes of Trochus smaragdinus Monterosato, 1880 TUNISIA • 6 dd; labelled “ Mirulinus smaragdinus Monts. Sfax ”; MCZR-M- 11700 / P • 154 dd; unprecised locality; “ Jujubinus smaragdinus C. d’Africa ”; MCZR-M- 11700 / P • 4 dd; unprecised locality; labelled “ Jujubinus smaragdinus Monts. Coste d’Africa ”. Syntypes of Jujubinus delpreteanus Sulliotti, 1889 The original material in the Sulliotti collection is lost (M. Tancredi, Museo Civico di Storia Naturale “ Giacomo Doria ”, Genova, Italy, pers. com., 12 Dec. 2018). SICILY • 5 dd; Messina, Lake Faro; labelled “ Faro di Messina ”; MCZR-M- 25224 - S, ex Sulliotti collection. Other material examined Morphotype smaragdinus TUNISIA • 77 dd; Kerkennah, Borj El Hsar; beached; CS-PM • 5 dd; Kerkennah, West of Gharbi Island, Sidi Youssef; 0.5 m depth; lagoon bioclastic sediment; LT • 3 dd; Djerba I.; beached; LT • 1 dd; Djerba I.; 0.5 m depth; lagoon bioclastic sediment; LT • 2 lv; Djerba I., Borj Jellij; 33 ° 53.05 ′ N, 10 ° 43.58 ′ E; 0 – 1 m depth; used for DNA sequencing; MNHN-IM- 2013 - 31801 - 2 • 1 lv; Djerba I., Sidi Jmour; 33 ° 49.88 ′ N, 10 ° 44.83 ′ E; 0 – 1 m depth; used for DNA sequencing; MNHN-IM- 2013 - 31891 • 4 lv; Djerba I., Sidi Jmour; 33 ° 49.88 ′ N, 10 ° 44.83 ′ E; 0 – 1 m depth; used for DNA sequencing; MNHN-IM- 2013 - 31988, 31998, 32000, 32113 • 2 lv; Djerba I., Sidi Solimen; 34 ° 02 ′ N, 11 ° 05 ′ E; 0 – 1 m depth; used for DNA sequencing; MNHN-IM- 2013 - 31920, 31923. Morphotype delpreteanus FRANCE • 13 dd; Toulon; France; labelled “ J. propinquus ”; MCZR-M-E 12 / 16 • 98 dd; Toulon; labelled “ J. propinquus Monts. Toulon inedito ”, “ Toulon (Vimont) ”; MCZR-M- 11727 • 82 dd; Toulon; labelled “ Mirulinus propinquus. M. Toulon! ”; MCZR-M- 11727. SICILY • 13 dd; Messina, Lake Faro; 0.2 m depth; amidst Cystoseira; CS-PM • 2 dd; Messina, Lake Ganzirri; CS-PM • 6 lv; Messina, Lake Ganzirri; 38 ° 15 ′ 44.5 ″ N, 15 ° 37 ′ 32.0 ″ E; 0.5 – 1 m depth; used for DNA sequencing; BAU 2943.1 to BAU 2943.6 • 2 dd; Messina, Lakes (Faro and / or Ganzirri); labelled “ Trochus pumilio ”; MCZR-M- 11269 • 2 dd; Messina, Lake Ganzirri; 2 m depth; LT • 20 lv; Lake Ganzirri; Messina; CS-PM. Description (in parentheses data of the lectotype) Morphotype smaragdinus SHELL. Shell of medium size for the genus, height 6.8 – 8.0 (6.8) mm, width 4.0 – 4.5 (4.0) mm, height / width ratio 1.70 – 1.80, conical, light. Protoconch seemingly smooth, of ca 1 whorl, 0.20 – 0.21 μm in width. Teleoconch of 5.5 – 6.5 (6.5) slightly convex whorls. Sculpture of 8 – 10 (8) flat, closely set abapical spiral cords of about the same strength, including the 2 – 3 (2) peripheral ones forming the peripheral cord, slightly shouldered, and 8 – 10 (8) regularly spaced, basal spiral cords large and flat. Suture well incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish-greenish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish-greenish; remaining whorls and base greenish adorned with alternating yellowish and white patches on the spiral cords. SOFT PARTS. Unknown for the open sea morphotype. Morphotype delpreteanus Epipodial tentacles translucent, densely covered by scattered tiny black (some white) speckles. Foot background creamy mottled with irregularly-shaped brown blotches (sometimes dotted), and covered by white speckles; sole white. Cephalic tentacles translucent, with a dorsal irregular and interrupted white stripe, and covered by small whitish and brownish speckles. Eye stalks brownish dorsally, creamy ventrally; eyes black. Snout dark brown, a creamy terminal band with a thin dark terminal stripe.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF100FFC5E26DFCD0D259F9A2.taxon	distribution	Distribution Jujubinus smaragdinus as genetically tested is known with certainty from the Gulf of Gabès, Tunisia (Monterosato 1880), with living specimens at depths of 0 – 1 m. We have examined samples morphologically very similar to the Tunisian material, from the Aegean and Ionian seas (see also Scuderi & Terlizzi 2012: 39, pl. 3 fig. 6, from Apulian Ionian Sea, as ‘ J. fraterculus ’), but these need to be tested genetically. The morphotype delpreteanus occurs in the coastal lakes of Pantano Piccolo and Ganzirri (Messina, Sicily) (Sulliotti 1889; Curini-Galletti & Palazzi 1979, as Trochus pumilio Philippi, 1844; Giacobbe 2012; Scuderi et al. 2019), at depths of 0.5 – 2 m. Specimens in the lot MCZR-M- 11727, from “ Toulon ”, France (possibly the Hyères salines, including the Étang de Pesquiers) are morphologically strikingly similar to the Sicilian delpreteanus morphotype.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF100FFC5E26DFCD0D259F9A2.taxon	discussion	Remarks We have tested Sicilian samples from a coastal lake environment, corresponding to the nominal taxon J. delpreteanus characterised by an impoverished sculpture, with denser but weaker spiral cords and a dark blackish-brown monochrome colouration. Given the relatively low genetic divergence between the Tunisian open sea form corresponding to Trochus smaragdinus, and the Sicilian coastal lake samples corresponding to J. delpreteanus (<3 %), and despite the striking morphological differences, we conservatively consider them as belonging to a single species. Trochus pumilio has been considered either as a synonym of Gibbula adriatica (Philippi, 1844) by Ghisotti & Melone (1975) or as a subspecies of Jujubinus striatus (J. striatus pumilio) by Curini-Galletti & Palazzi (1979), identical with J. delpreteanus; we agree with the latter identification. However, Trochus pumilio is a junior homonym of Trochus pumilio Dillwyn, 1817, currently accepted as Obeles pumilio (Dillwyn, 1817) (Pulmonata, Geomitridae). The open sea form of J. smaragdinus differs from J. striatus in its typical greenish-yelowish-whitish colour pattern, the smaller size, the more evident prosocline lamellae in the interspaces. It differs from J. exasperatus, J. pseudogravinae, J. vulgaris, J. gravinae, and J. dianiensis sp. nov. in its less marked sculpture in all teleoconch whorls; from J. depictus, J. sartorii, J. ruscurianus in the weaker and more regular sculpture with barely evident prosocline lamellae; from J. elenchoides in its smaller size and the fewer spiral cords; from J. superum sp. nov. in the different colouration (dark greyish-blackish in J. superum, greenish with whitish spots in J. smaragdinus). Colouration is also diagnostic with J. kyrnos sp. nov., J. fraterculus, J. gofasi sp. nov., and J. vexationis (Madeira). See under J. delpreteanus, J. inopinatus sp. nov., J. angustibasis sp. nov., and J. dispar for comparative remarks.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10DFFC7E222F93DD119FE6C.taxon	diagnosis	Diagnosis Shell small for the genus, with scalariform body whorls, hint of umbilical fissure, low height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10DFFC7E222F93DD119FE6C.taxon	materials_examined	Type material Syntypes MOROCCO • 1 syntype; Tanger (type locality); MCZR-M- 30081 / S • 1 syntype; Tanger; MNHN- IM- 2.000 - 32.439. Other material examined MOROCCO • 79 dd; Al Hoceima, Cala Iris; 3 m depth; CS-PM. SPAIN • 3 dd; Algeciras; 10 m depth; CS-PM • 1 lv; Málaga; 2 m depth; under rocks; CS-PM • 4 lv; Getares; 0 – 1 m depth; washing sea-weeds; MO.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10DFFC7E222F93DD119FE6C.taxon	description	Description SHELL. Shell small for the genus, height 2.3 – 4.0 mm, width 2.4 – 3.8 mm, height / width ratio 0.95 – 1.05, body with stepped whorls, angled shoulder. Protoconch seemingly smooth, of ca 1 whorl, 0.21 – 0.22 mm in width. Teleoconch of 3.5 – 4.0 convex whorls. Sculpture of 9 abapical spiral cords of various sizes; 3 – 4 smaller on the shoulder, 2 strongly keeled cords (the abapical peripheral) with a large interspace sculptured by two tiny cords, regularly spaced. Suture scarcely incised. Teleoconch covered by barely visible, wavy, regularly spaced prosocline lamellae, more evident in the wide interspace between the two large, keeled cords. Base convex, with 7 spiral, regularly spaced cords, with dense prosocline lamellae in interspaces, umbilicus narrowly open, white. Aperture quadrangular, internally whitish, nacreous, iridescent in fresh specimens. COLOURATION. Colour of protoconch and teleoconch whorls whitish or light beige, opaque, with small (occasionally large) darker spots and blotches (brown or red). SOFT PARTS. Epipodial tentacles translucent, whitish, covered by scattered tiny white speckles. Foot background creamy mottled with irregularly-shaped brown to blackish lateral bands (sometimes dotted), and covered by white or yellowish speckles; sole whitish. Cephalic tentacles translucent whitish, with a white or yellow stripe. Eye stalks creamy ventrally; eyes black. Snout variable, whitish to dark brown or black.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10DFFC7E222F93DD119FE6C.taxon	distribution	Distribution Western Mediterranean and Ibero-Moroccan Gulf. From southern Spain (Málaga), and North African coast (Algeria and Morocco), across the Strait of Gibraltar area (Jeffreys 1883; Monterosato 1890; Pallary 1902, 1920; Schirò 1971; Ghisotti & Melone 1972; Verduin 1979; and as Gibbula ditropis: Manzoni 1871; Van Aartsen et al. 1984; Scaperrotta et al. 2010; Gofas et al. 2011); living specimens at depths of 0 – 1 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10DFFC7E222F93DD119FE6C.taxon	discussion	Remarks This species has been almost invariably considered as a member of the genus Gibbula (s. lat.) until it was recently reassigned to Jujubinus based on molecular phylogenetics (Uribe et al. 2017), which is confirmed here. We highlight that juveniles of J. tingitanus and J. gravinae show very similar outlines, in turn very different from, e. g., Steromphala racketti (Payraudeau, 1826), suggesting that J. tingitanus with its peculiar adult morphology, may represent a case of pedomorphosis (Fig. 9 A – F). Monterosato (1890: 144) reported a single very small specimen from Sicily, off the coast of Palermo, identified by Jeffreys as Gibbula ditropis. Despite an accurate search at MCZR in the Monterosato collection, this specimen could not be found, and we keep this record as doubtful. Trochus ditropis S. Wood, 1848, is a very similar, but distinct fossil species from the Pliocene (Coralline and Red Crag) of England (see also Verduin 1979).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10FFFC6E256FE4FD0DAFC1A.taxon	diagnosis	Diagnosis Shell of medium size for the genus, with strong spiral cords, high and very grainy, broad interspaces with strong prosocline lamellae, distinct peripheral cord, wide base, umbilicus closed, low height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10FFFC6E256FE4FD0DAFC1A.taxon	materials_examined	Type material Holotype Not found. Paratypes MOROCCO • 36 dd; Morocco, Tangeri (type locality); MCZR-M- 11751 / P. Other material examined SPAIN • 9 dd; Málaga; 60 m depth; trawled; CS-PM • 1 dd; Málaga; 80 m depth; CS-PM • 34 lv; Málaga; 20 – 30 m depth; CS-PM • 4 lv; Málaga; 20 – 30 m depth; used for DNA sequencing; BAU- 2079.1 to BAU- 2079.4 • 2 dd; Málaga Bay; 100 – 150 m depth; fishing nets; LT • 4 dd; Getares; 15 m depth; greyish-green colour form; CS-PM • 6 dd; Algeciras; beached; MO. MOROCCO • 4 dd; Ceuta, El Nacho; MO.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10FFFC6E256FE4FD0DAFC1A.taxon	description	Description SHELL. Shell of medium size for the genus, height 4.9 – 5.7 mm, width 3.9 – 5.4 mm, height / width ratio 1.12 – 1.25, conical, light. Protoconch seemingly smooth, of ca 1 whorl, 0.35 – 0.36 mm in width. Teleoconch of 4.5 – 5.5 (5.5) slightly convex whorls. Sculpture of 4 – 5 (5) prominent, closely set abapical spiral cords of about the same strength, including the 2 – 3 (2) peripheral ones forming the peripheral cord, slightly shouldered, and 4 – 5 (4) regularly spaced, basal spiral cords large and flat. Suture incised. Teleoconch surface covered by strong prosocline growth striae, irregularly set. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, or pinkish nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright red; remaining whorls and base with alternating brownish-reddish adorned by white-greyish patch of different size (also in the holotype). SOFT PARTS. Unknown.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10FFFC6E256FE4FD0DAFC1A.taxon	distribution	Distribution Strait of Gibraltar (Curini-Galletti 1985; Gofas et al. 2011), Morocco and Mauritania coasts (Curini-Galletti 1982 b, 1985; Cretella 1993) Western Mediterranean, Alboran Sea, Spanish coasts; living specimens at depths of 20 – 30 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10FFFC6E256FE4FD0DAFC1A.taxon	discussion	Remarks It differs from J. exasperatus, J. pseudogravinae, J. gravinae, J. dianiensis sp. nov., and J. vexationis (Madeira) in the absence of tubercles or undulations on the peripheral cord; from J. swinneni sp. nov. (Canary Islands) in its colouration (brownish-reddish with white-greyish blotches of varying size in J. dispar vs bright greenish-cream or greenish-whitish with alternating whitish or reddish or blackish spots on spiral cords in J. swinneni). Jujubinus depictus, J. sartorii, J. fraterculus, J. gofasi sp. nov., and J. smaragdinus all have evident sculptures and prosocline lamellae in the interspaces similar to J. dispar; however, the sculpture is more marked in J. dispar, with the spiral cords crossed by very sharp prosocline lamellae giving a wrinkled appearance to the entire teleoconch, including the base. It differs from J. superum sp. nov., J. kyrnos sp. nov., and J. angustibasis sp. nov. in its larger size and stronger sculpture, and in the different colouration (brownish-reddish with white-greyish patches of different size in J. dispar vs dark greyish-blackish background in J. superum, whitish greyish or beige in J. kyrnos, brownish-greyish-greenish colour patches on the spiral cords in J. angustibasis; from J. ruscurianus in its more slender outline, the fewer, less evident and more spaced spiral cords, and the more marked prosocline lamellae (vs fine very close streaks in J. ruscurianus); from J. elenchoides in the fewer spiral cords (4 – 5 vs 12 – 16 in J. elenchoides) and the marked prosocline lamellae (almost lacking in J. elenchoides); from J. inopinatus sp. nov. in its less slender outline, narrower interspaces with marked prosocline lamellae (almost lacking in J. inopinatus); from J. striatus (Linnaeus, 1758) and J. vulgaris (Risso, 1826) in the stronger sculpture and less slender outline.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10EFFC3E229FC75D049FE67.taxon	diagnosis	Diagnosis Shell of medium size for the genus, not very high, strong, sparse, raised and very grainy spiral cords, wide interspaces, and very sculptured by prosocline lamellae, distinct peripheral cord, narrow base, umbilicus closed, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10EFFC3E229FC75D049FE67.taxon	materials_examined	Type material Syntypes of Trochus montagui Wood, 1828 IRELAND • 5 dd; Ireland (type locality); J. E. Gray collection; NHMUK 1968623, Trochus parvulus Philippi, 1844 Type material (from Italy, Reggio Calabria, Pezzo) lost (Óscar Alfredo Gálvez Herrera, Museo Nacional de Historia Natural, Santiago de Chile, Chile, pers. com., 12 Feb. 2020; Christine Zorn, Museum für Naturkunde, Berlin, Germany, pers. com., 26 Feb. 2020). Other material examined CORSICA • 5 dd; Bastia; Caziot leg.; MCZR-M-E 12 - 11715 • 14 dd; Capo Corso; 95 – 105 m depth; MO • 4 dd; Capo Corso; 60 m depth; residuals of fishing boats; CS-PM • 3 lv; Gulf de Saint-Florent; 42 ° 44.1 ′ N, 9 ° 19.3 ′ E; 60 m depth; used for DNA sequencing; MNHN-IM- 2019 - 5675, 5676, 5678 • 1 lv; North Cap Corse; 42 ° 59.8 ′ N, 9 ° 18.6 ′ E; 95 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6259 • 1 lv; East of Bastia; 42 ° 41.2 ′ N, 9 ° 27.7 ′ E; 50 m depth; used for DNA sequencing; MNHN-IM- 2019 - 6402 • 2 lv; Large Punta di u Capicciolu; 41 ° 24.3 ′ N, 9 ° 16.9 ′ E; 71 – 71 m depth; used for DNA sequencing; MNHN- IM- 2019 - 12120, 12423 • 1 lv; South of Gulf de Porto Novo; 41 ° 29.7 ′ N, 9 ° 17.8 ′ E; 48 – 51 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12694 12695 • 1 lv; Large Porto-Vecchio; 41 ° 32.8 ′ N, 9 ° 27.2 ′ E; 89 – 90 m depth; used for DNA sequencing; MNHN-IM- 2019 - 12797 • 2 lv; off South coasts; 41 ° 21 ′ N, 9 ° 10.6 ′ E; 73 – 73 m depth; used for DNA sequencing; MNHN-IM- 2019 - 13293, 13294 • 4 lv; Gulf de Girolata; 42 ° 20.8 ′ N, 8 ° 35.8 ′ E; 45 – 45 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17224, 17356, 17357, 17359 • 1 lv; between Capo Cavallo and Capu di a Mursetta; 42 ° 29.3 ′ N, 8 ° 38.5 ′ E; 90 – 91 m depth; used for DNA sequencing; MNHN-IM- 2019 - 17968. GREAT BRITAIN • 5 dd; Aberdeen; labelled “ Trochus montacuti ”; MCZR-M- 11778 • 11 dd; Shetlands; “ Shetland, Mc’Andrew ”; MCZR-M-E 12 / 14. ITALY • 43 dd; Capo D’Anzio, 3 miles off; 40 m depth; bioclastic sediment; CS-PM • 4 dd; Civitavecchia; 40 m depth; fishing boat; CS-PM • 2 dd; San Felice Circeo (LT); 40 m depth; dredging; LT • 2 dd; Ventotene Island; 50 m depth; fishing boat; CS-PM • 1 dd; Livorno; residuals of fishing boats; CS-PM • 1 dd; Capraia Is.; 100 m depth; residuals of fishing boat; CS-PM • 1 dd; Montalto di Castro; 57 m depth; CS-PM • 4 dd; Fiumicino; 160 m depth; residuals of fishing boats; CS-PM • 125 dd; Anzio; 50 m depth; CS-PM • 1 dd; Cape Circeo; 35 m depth; CS-PM • 2 dd; Capri Is.; MCZR-M-E 12 - 11716 • 1 dd; Maratea; 80 m depth; residuals of fishing boats; CS-PM. SARDINIA • 1 dd; Caprera Is., Punta Crucitta; 78 m depth; CS-PM • 3 dd; La Maddalena Island; 60 m depth; dredging; LT. SICILY • 3 dd; Palermo, Gulf of Carini; 120 m depth; residuals of fishing boats; CS-PM • 16 dd; Pantelleria Is.; 42 m depth; MO. SPAIN • 6 dd; Getares (Algeciras); 20 m depth; washing of algae; LT • 4 dd; Málaga; 90 m depth; trawled; CS-PM • 3 lv; Málaga; 90 m depth; trawled, used for DNA sequencing; BAU- 2072.1 to BAU- 2072.3 • 2 lv; Málaga Bay; 80 – 90 m depth; fishing nets, used for DNA sequencing; BAU- 2073.1, BAU- 2073.2 • 1 dd; Málaga Bay; 80 – 90 m depth; fishing nets; LT • 1 dd; Baleares, Majorca Is.; residuals of fishing boats; CS-PM • 3 dd; Alboran Sea; 100 – 150 m depth; fishing nets; LT.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10EFFC3E229FC75D049FE67.taxon	description	Description SHELL. Shell of medium size for the genus, height 5.5 – 7.1 mm, width 4.2 – 5.1 mm, height / width ratio 1.30 – 1.38, conical, slightly shiny. Protoconch seemingly smooth of ca 1 whorl, 0.21 – 0.22 mm in width. Teleoconch of 4.5 – 6.5 slightly convex whorls. Sculpture of 5 – 6 closely set, slightly shouldered spiral cords of about the same strength, including the 2 peripheral ones forming the peripheral cord, and 7 – 8 regularly spaced, very evident, basal spiral cords. Suture well incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set and by strong prosocline lamellae in spiral interspaces. Base convex, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish brownish; remaining whorls and base cream or hazelnut colour with axial wide lines brownish-reddish spotted and of different size. SOFT PARTS. Unknown.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10EFFC3E229FC75D049FE67.taxon	distribution	Distribution The Mediterranean Sea (e. g., Nordsieck 1968; Ghisotti & Melone 1975; Terreni 1981; Curini-Galletti & Palazzi 1982; Poppe & Goto 1991; Barash & Danin 1992; Giannuzzi-Savelli et al. 1994; Brunet Navarro & Capdevila 2005; Scaperrotta et al. 2010; Gofas et al. 2011; Manousis 2012; Smriglio et al. 2019), and the northeastern Atlantic Ocean from Great Britain and Ireland to the Canary Islands (Jeffreys 1883; Wood 1828; Fretter & Graham 1962; Nordsieck 1968; Rolán 1983; Poppe & Goto 1991; Gofas et al. 2011; Smriglio et al. 2019), Madeira (Segers et al. 2009) and the Canary Islands (Hernández et al. 2011); living specimens at depths of 50 – 95 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10EFFC3E229FC75D049FE67.taxon	discussion	Remarks We postpone a decision on the generic classification of this species, pending a study with a wider taxonomic coverage, including more species that are morphologically similar (like e. g., Jujubinus tumidulus (Aradas, 1846), Clelandella artilesi, C. azorica Gofas, 2005, C. dautzenbergi, C. madeirensis, C. myriamae Gofas, 2005, C. perforata Gofas, 2005).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10BFFC3E221FE7AD1D0F809.taxon	diagnosis	Diagnosis Shell of medium size for the genus, solid, elevated; sculpture of 4 – 5 very spaced spiral cords, raised and very granulated, broad interspaces with strong prosocline lamellae, peripheral cord evident.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10BFFC3E221FE7AD1D0F809.taxon	materials_examined	Type material Holotype SICILY • dd; off Mazara del Vallo; 45 – 70 m depth; Museo Malacologico Piceno, Cupra Marittima. Paratypes SICILY • 1 dd; same data as for holotype; Museo Malacologico Piceno, Cupra Marittima (paratype 1) • 2 dd; same data as for holotype; collection Roberto Ardovini, Rome (paratypes 2 – 3). Material examined STRAIT OF SICILY • 3 dd; unprecised locality in the Strait of Sicily; ca 90 m depth; CS-PM. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, of medium size for the genus, height 6.0 – 6.7 (6.3) mm, width 4.1 – 4.9 (4.9) mm. Protoconch seemingly smooth, of ca 1 whorl, ca 0.2 mm in width. Teleoconch of 6 – 7 very convex whorls. Sculpture of 4 strong, granulose spiral cords, interspaces almost as wide as cords with very strong prosocline lamellae, and a a larger and evident peripheral cord. Suture barely visible. Base convex, with 7 – 8 regularly spaced basal cords, interspaces crossed by marked prosocline lamellae. Aperture quadrangular, with white columellar callus thickened in the central portion; interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch milky white; teleoconch with first three whorls reddish or reddish pink, remaining whorls and base uniformly yellowish greenish with spiral cords reddish brown. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10BFFC3E221FE7AD1D0F809.taxon	distribution	Distribution Currently only known from the Strait of Sicily, Mediterranean Sea, empty shells at depths of 45 – 90 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10BFFC3E221FE7AD1D0F809.taxon	discussion	Remarks Jujubinus catenatus is clearly diagnosed from species with a similar strongly ‘ granulose’ sculpture [like Jujubinus montagui (W. Wood, 1828), J. tumidulus, or J. errinae] by its ‘ pear-shaped’ outline. From J. montagui it also differs in the less robust spiral cords sculptured by smaller and more spaced tubercles. From J. tumidulus it also differs in its larger size (6.0 – 6.7 mm vs 3 – 4.5 mm in J. tumidulus), and a less dense and weaker tubercles on the spirals.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10AFFBDE1A0FE90D78BFE50.taxon	diagnosis	Diagnosis Shell of medium size for the genus, slightly turriculate; sculpture 5 spiral cords, with incised interspaces; strong prosocline lamellae between spiral cords.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10AFFBDE1A0FE90D78BFE50.taxon	materials_examined	Type material Holotype SICILY • dd; Strait of Messina; stn POP 95 - DG 04; 38 ° 14 ′ 45 ″ N, 15 ° 37 ′ 36 ″ E; 100 m depth; MZB- 60155. Paratypes SICILY • 8 dd; same data as for holotype; Benthic Ecology Lab. Messina University (paratypes A – H) • 1 dd; Strait of Messina; stn POP 95 - DG 001; 38 ° 14 ′ 45 ″ N, 15 ° 37 ′ 28 ″ E; 90 – 115 m depth; Benthic Ecology Lab. Messina University (paratype I) • 1 dd; same data as for preceding; CWR (paratype M) • 1 dd; Rada Paradiso, stn PIC 02 - St 2; 38 ° 13 ′ 27 ″ N, 15 ° 36 ′ 02 ″ E; 90 – 115 m depth; Benthic Ecology Lab. Messina University (paratype L) • 5 dd; off Borgo Chianalea (Messina); 38 ° 15 ′ 36 ″ N, 15 ° 43 ′ 08 ″ E; 40 – 50 m depth; CS-PM (paratypes N – R) • 3 dd; same data as for preceding; coll. Giuseppe Notaristefano, Messina (paratypes S – U) • 1 dd; same data as for preceding; coll. Rafael Mesa, Lanzarote (paratype V) • 1 dd; same data as for preceding; coll. Bruno Amati, Roma (paratype X) • 1 dd; same data as for preceding; coll. Ermanno Quaggiotto, Logare (paratype Y). Description (in parentheses data of the holotype) SHELL. Shell conical, of medium size for the genus, height 4.5 – 6.0 (4.9) mm, width 3.8 – 4.7 (4.0) mm, slightly shiny. Protoconch seemingly smooth, of ca 1 whorl, 0.26 – 0.29 (0.28) mm in width. Teleoconch of 4 – 4.5 (4.5) slightly convex whorls. Sculpture of 5 strongly granulose, closely set, abapical spiral cords of about the same strength, and a broader peripheral cord formed by 2 merged cordlets; peripheral cord rippled on first two whorls, almost smoot on remaining whorls. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set. And very evident lamellae in the spiral interspaces. Base convex, with 6 regularly spaced, narrow basal spiral cords; umbilicus closed and covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion; interior whitish nacreous. COLOURATION. Colour of protoconch whitish, teleoconch spire and base pinkish-creamy, with red spiral cords interrupted by short white spots. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10AFFBDE1A0FE90D78BFE50.taxon	distribution	Distribution Currently only known from the Strait of Messina and Ustica Is. (Giacobbe & Renda 2019). Empty shells at depths of 90 – 115 m, but the record from a bank off Ustica Is. (Giacobbe & Renda 2019) in a sample suggests an origin in a shallower kelp-related habitat.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF10AFFBDE1A0FE90D78BFE50.taxon	discussion	Remarks Jujubinus catenatus differs from J. errinae in the neatly convex whorls (yielding the typical ‘ pearl-like’ outline), the more widely spaced and more granulose spiral cords, and the continuous reddish spiral lines vs the typical reddish spiral lines interrupted by short white spots of J. errinae. Jujubinus montagui differs in its higher height / width ratio, the sculpture of less dense and larger tubercles, and the colouration lacking the interrupted red lines. Jujubinus tumidulus differs in its smaller size, the lower height / width ratio, the slightly weaker sculpture. None of the three species has the initial teleoconch whorls so markedly rippled as J. errinae, at variance with the remaining whorls.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF175FFBCE272FE2BD0BDFE9A.taxon	diagnosis	Diagnosis Shell of small size for the genus, solid; sculpture of 3 – 4 spaced spiral cords, raised, broad interspaces with dense prosocline growth lines; peripheral cord evident and rippled, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF175FFBCE272FE2BD0BDFE9A.taxon	materials_examined	Type material Holotype GREECE • dd; Karpathos Is.; SMF 341874. Paratypes GREECE • unprecised number dd; Karpathos Is.; SMF. Other material examined GREECE • 3 dd; Petroch ṓri; 0.5 m depth; CS-PM. Description (in parentheses data of the holotype) SHELL. Shell of small size for the genus, conical, height 3.9 – 4.9 (5.0) mm, width 3.6 – 4.8 (4.0) mm, height / width ratio 1.0 – 1.25 (1.25), rather shiny. Protoconch smooth, of ca 1.0 whorls, ca 0.2 mm in width. Teleoconch of 4.5 – 5.0 slightly convex whorls. Sculpture of 3 – 5 flat abapical spiral cords of about the same strength, and a broad, evident and rippled peripheral cord formed by 4 – 5 cordlets. Suture incised. Teleoconch surface covered by barely visible, irregularly set prosocline growth striae. Base convex, with 6 – 7 regularly spaced, basal flat spiral cords; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, yellowish or pinkish nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls whitish; remaining whorls and base of cream or whitish background, with brownish-reddish-whitish flammules, of variable size on the basal flat spiral cords, peripheral cord with alternating whitish-yellowish and brownish-red spots. SOFT PARTS. Unknown.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF175FFBCE272FE2BD0BDFE9A.taxon	distribution	Distribution Currently known only from the Aegean Sea, empty shells in shallow waters (0.5 m deep, our material).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF175FFBCE272FE2BD0BDFE9A.taxon	discussion	Remarks Jujubinus karpathoensis recalls J. gravinae with which it shares the general aspect, and in particular the evident and rippled peripheral cord; the two taxa are clearly related, and we cannot exclude that they are synonyms. It is not common in the Aegean Sea, yet specimens under this names are frequently found in the collections also from other Mediterranean areas. We assume that all records of J. karpathoensis from outside the Aegean Sea are misidentifications. Two other species have a similarly rippled peripheral cord, namely J. vexationis and J. errinae; both exhibit the rippled cord only on the first 2 – 3 teleoconch whorls, while J. karpathoensis (like J. gravinae) on all teleoconch whorls.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF174FFBEE223FEF3D00FFD48.taxon	diagnosis	Diagnosis Shell cyrtoconoid, of small size for the genus, elevated, thick; sculpture of 4 – 5 spaced spiral cords, raised and granulose, broad interspaces with dense prosocline lamellae, peripheral cord poorly defined, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF174FFBEE223FEF3D00FFD48.taxon	materials_examined	Type material Not yet found. Probably the four syntypes (Aradas 1846: 165) are to be searched in the Aradas collection, which is stored partly in the Natural History Museum of Milan, and partly in the Museum of Zoology of Catania. Other material examined ITALY • 40 dd; Scilla; 42 m depth; CS-PM. SICILY • 2 dd; Palermo; “ Jujubinus tumidulus (Aradas) Palermo ”, ex Bellini collection; MCZR • 123 dd; Palermo; “ P ”; MCZR-M- 11713, MTS • 13 dd; San Vito Lo Capo; “ S. vito ”; MCZR-ME 12 / 4, MTS • 11 dd; Linosa Is., Punta Calcarella; 36 m depth; CS-PM • 50 dd; Linosa Island, Sicchitella Due Ceppi; 28 m depth; CS-PM • 20 dd; Linosa Is.; 50 – 60 m depth; MO • 15 dd; Pantelleria Is., stn UR 92 # 2; 36 ° 49 ′ 24 ″ N, 12 ° 00 ′ 53 ″ E; 53 m depth; MO • 3 dd; Marettimo Is.; 80 m depth; MO • 11 dd; Marettimo Is.; 45 m depth; CS-PM • 200 + dd; Lampedusa Is.; 70 – 80 m depth; residuals of trawlers; CS-PM • 3 dd; Lampedusa Is.; 92 m depth; MO. UNPRECISED LOCALITY (probably Sicily) • 1 dd; “ Trochus tumidulus Aradas Pliocen. Pleistocenico ”; MCZR. Description (holotype data not available) SHELL. Shell solid, cyrtoconoid, of small size for the genus, height 3 – 4.5 mm, width 2.5 – 3.3 mm. Protoconch seemingly smooth, of ca 1 whorl, ca 0.2 mm in width. Teleoconch of 4.5 – 5 regularly convex whorls. Sculpture of 5 – 6 finely tuberculated spiral cords, interspaces twice as wide as the spirals, and 2 slightly smalle cords merged to form a non-prominent, broader peripheral cord. Suture incised. Teleoconch surface covered by fine prosocline growth lines, interspaces crossed by dense prosocline lamellae. Base convex, with 6 – 7 flat, granulose basal cords; umbilicus closed; covered by a whitish callus. Aperture quadrangular, with columella callus only barely thickend in the middle portion. Interior of the aperture whitish nacreous. COLOURATION. Protoconch whitish. Colouration variable, teleoconch of whitish, yellowish or pinkish background, with irregularly arranged darker or lighter spots and blotches, sometimes with tubercles on the spirals darker, usually with alternating white and darker blotches on the peripheral cord. Completely dark or light monochrome specimens are also known. Base generally the same colour as the protoconch with spirals dotted of reddish or brownish. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF174FFBEE223FEF3D00FFD48.taxon	distribution	Distribution Described as a fossil from Gravitelli (Messina, Sicily, probably Pliocene) by Aradas (1846) and reported as Recent from the port of Messina, Lipari and Palermo (Sicily) by Curini-Galletti & Palazzi (1982), from the Gulf of Carini (Palermo) by Scaperrotta et al. (2009) and from Ganzirri, Messina by Scuderi et al. (2019), here recorded from Marettimo, Linosa and Lampedusa, empty shells at depths of 30 – 100 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF174FFBEE223FEF3D00FFD48.taxon	discussion	Remarks Although never recorded from outside the Mediterranean, a specimen from the Canary Islands figured under the name “ Jujubinus vexationis ” by Hernández et al. (2011: fig. 15 j – k) is very similar to J. tumidulus. It is an easily recognizable species, resembling a small J. montagui, from which it differs in the more neatly cyrtoconoid outline and the smaller size.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF176FFB9E267FD23D0D4FD92.taxon	diagnosis	Diagnosis Shell of medium size for the genus, elevated, thick; sculpture of 5 – 6 spaced spiral cordlets, raised and scarcelu scultured, broad interspaces with weak prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF176FFB9E267FD23D0D4FD92.taxon	materials_examined	Type material Syntypes SICILY • unprecised number dd; Messina; Coen collection (n. 6527), Hebrew University of Jerusalem (Israel). Other material examined SICILY • 14 dd; Messina; MCZR-M- 11471 • 114 dd; S. Reniero, Messina; MCZR-M- 11471 • 2 dd; Messina; CS-PM. Description (holotype data not available) SHELL. Shell conic, solid, with medium size for the genus, height 3.5 – 5.4 mm, width 3.0 – 4.5 mm. Protoconch seemingly smooth, of ca 1 whorl, ca 0.2 mm in width. Teleoconch of 4.5 – 5.5 convex whorls. Sculpture of 5 – 6 spiral cords of almost equal size (sometimes formed by two merged cordlets), and a broader, not very prominent peripheral cord. Interspaces narrow, sculptured by weak prosocline lamellae. Suture not incised. Base flat or barely convex, with 5 – 6 flat basal spiral cords. Umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the lower third. COLOURATION. Interior of aperture whitish nacreous. Colour of the protoconch whitish. Teleoconch of greyish-brown background, with small whitish spots on spiral cords, iridescent in fresh specimens SOFT PARTS. Epipodial tentacles greyish, with white knob-shaped sense organ at the base. Foot background creamy, with brown small dashes and large spots arranged in two main rows and one of smaller spots, alternated to white ones. Cephalic tentacles greyish, with thin dark little lines and small white dots. Eye stalks brownish dorsally, whitish ventrally, eyes black encircled by a creamy crown. Snout dark-brown, a whitish terminal band with a thin dark terminal stripe (Scuderi 2023).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF176FFB9E267FD23D0D4FD92.taxon	distribution	Distribution Currently only known from the eastern coasts of Sicily, in very shallow waters (Scuderi et al. 2019; Scuderi 2023).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF176FFB9E267FD23D0D4FD92.taxon	discussion	Remarks Jujubinus seguenzae is similar to J. striatus and J. angustibasis sp. nov. (with the latter is also found sympatrically) but is constantly smaller than both species but its size (3.5 – 5.4 mm vs 6.1 – 8.2 mm of J. striatus and 4.6 – 7.2 mm of J. angustibasis), and the microsculture of prosocline lamellae is rather weak in J. seguenzae vs strong in J. striatus and J. angustibasis.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF171FFB8E24FFDEDD09EFE51.taxon	diagnosis	Diagnosis Shell of small size for the genus, elevated, thick; sculpture of 5 – 6 spaced spiral cords, raised and nodulose, with smaller interspaces and prosocline irregular striae; peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF171FFB8E24FFDEDD09EFE51.taxon	materials_examined	Type material Holotype SÃO TOMÉ • dd; Minerio Reef; 00 ° 23 ′ 016 ″ N, 6 ° 46 ′ 228 ″ E; 43 m depth; MNCN 15.05 / 47563. Paratypes SÃO TOMÉ • 1 dd; same data as for holotype; MNHN • 1 dd; same data as for holotype; MHNS • 1 dd; same data as for holotype; BMNH • 1 dd; same data as for holotype; collection Sandro Gori, Livorno. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, of small size for the genus, height 4 – 4.9 (4.4) mm, width 3.0 – 3.4 (3.0) mm. Protoconch seemingly smooth, of ca 1 whorl, 0.15 – 0.20 mm in width. Teleoconch of 6 – 6.5 (6) flat and rapidly increasing whorls. Sculpture of 4 barely nodulose cords separated by smaller interspaces on first teleoconch whorl; 5 – 6 nodulose cords on following whorls, and a larger and prominent, nodulose, peripheral cord. Microsculpture of very prosocline irregular striae between the nodulose cords. Base convex, with 7 – 9 spiral cords with interspaces of similar size. Aperture broadly quadrangular, rounded anteriorly, with a columellar callus on the lower third. Interior of the aperture white, nacreous. COLURATION. Colour of protoconch milky white; teleoconch red, with sparse white spots, larger on the peripheral cord; alternatig red and white segments on the spirals of the base. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF171FFB8E24FFDEDD09EFE51.taxon	distribution	Distribution Only known from the type locality, Minerio Reef, São Tomé; empty shells at a depth of 43 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF171FFB8E24FFDEDD09EFE51.taxon	discussion	Remarks Jujubinus augustoi is very similar to J. pseudogravinae, in their granulose spirals and the colouration dominated by red on the teleoconch, and the protoconch white. It differs in its smaller size, smaller protoconch, and more numerous spiral cords Jujubinus augustoi is also similar to J. exasperatus, with which it was originally compared; it differs in the smaller size, the larger nodules on the spirals, the fewer spirals on the base. It differs from J. gravinae in its smaller size and smaller protoconch, and the less nodulose spiral cords with more regular and stronger prosocline lamellae in the interspaces (Rolán & Gori 2009).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF170FFB8E20BFE2BD7CCF87C.taxon	diagnosis	Diagnosis Shell of medium size for the genus, elevated, thick; sculpture of 3 – 4 spaced spiral cordlets, raised and very granulated, broad interspaces with strong prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF170FFB8E20BFE2BD7CCF87C.taxon	materials_examined	Type material Holotype ANGOLA • dd; Baie de Lucira (Praia do Cesar); 13 ° 52 ′ S, 12 ° 30 ′ E; 10 m depth; MNHN-IM- 2000 - 30359. Paratypes ANGOLA • 1 dd; same data as for holotype; MNHN • 8 dd; Baie de Santa María; MNHN • 1 dd; same locality as for preceding; MNCN • 1 dd; same locality as for preceding; Instituto de Investigação Cientifica Tropical, Lisbon. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, of medium size for the genus, height 5.0 – 6.5 (6.5) mm, width 3.8 – 4.7 (4.7) mm, slightly shiny. Protoconch seemingly smooth, of ca 1 whorls, 0.28 mm in width. Teleoconch of 5.5 – 7 whorls, the first rather convex, the others slightly convex. Sculpture of 3 spiral cords on first whorl, four on the remaining ones, with interspaces twice as wide as the spiral cords, and a broader, weakly sculptured, peripheral cord, formed by 2 – 3 merged cordlets. Suture incised. Growth striae barely visible; interspaces sculptured by very strong, prosocline lamellae. Base flat (or barely concave), with 7 regularly spaced, basal spiral cords; umbilicus closed, covered with a whitish callus. Aperture quadrangular, with columellar callus in the lower third. Interior of aperture whitish nacreous. COLOURATION. Colour of the protoconch whitish. Teleoconch colour of red background, with broad vertical whitish flame-like blotches, extending over the peripheral cord. Base reddish, with basal cords slightly darker, interrupted by small white blotches. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF170FFB8E20BFE2BD7CCF87C.taxon	distribution	Distribution Currently only known from Angola (Gofas 1991) with empty shells at a depth of 10 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF170FFB8E20BFE2BD7CCF87C.taxon	discussion	Remarks Jujubinus fulgor differs from J. exasperatus in the different sculpture of less strongly tuberculated spiral cords, with wider interspaces, the less evident peripheral cord, and the slightly more convex outline. Jujubinus baud o ni is somehow similar, especially in the slightly convex outline and similar colouration, but is diagnosed by its stronger granulation of the spirals and the narrower interspaces.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBBE267FE90D647F952.taxon	diagnosis	Diagnosis Shell of small size for the genus, elevated, thick; sculpture of 3 – 5 well spaced and raised spiral cords, broad interspaces with strong prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBBE267FE90D647F952.taxon	materials_examined	Type material Holotype CANARY ISLANDS • dd; Punta del Lajial, El Hierro; MNCN 15.05 / 47568. Paratypes CANARY ISLANDS • 38 dd; same data as for holotype; MNHN • 1 dd; same data as for holotype; BMNH • 1 dd; same data as for holotype; MNHC • 2 dd; same data as for holotype; MHNS • 33 dd; same data as for holotype; FS • 1 dd; Santa Cruz de La Palma; coll. Ramón Gómez, La Palma • 2 dd; Tenerife, Puerto de la Cruz; coll. Ramón Gómez, La Palma • 20 dd; La Palma; coll. José María Hernández, Gáldar, Las Palmas • 10 dd; NW Gran Canaria; coll. José María Hernández, Gáldar, Las Palmas. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, shiny, of small size for the genus, height 3.0 – 5.0 (3.0) mm; width 2.7 – 4.4 (2.6) mm. Protoconch seemingly smooth, of ca 1 whorl, ca 0.20 mm in width. Teleoconch of 3.5 – 5.5 convex and whorls. Sculpture of 3 – 4 raised and stepped non-granulose spiral cords, the uppermost frequently stronger and more prominent, and a broad and prominent peripheral cord. Interspaces narrower than spiral cords, sculptured by dense prosocline growth lines. Base convex, with 6 – 7 (7) regularly spaced basal spiral cords, flat, smooth. Umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with columellar callus weakly thickened on lower third only in adults. COLOURATION. Interior of aperture whitish nacreous. Colour of the protoconch whitish; teleoconch with reddish, pink, brownish or greenish background, with white vertical blotches, often darker edges. Base with spirals of same colour as background, interrupted by white small blotches. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBBE267FE90D647F952.taxon	distribution	Distribution Only known from the Canary Islands, with empty shells at a depth of 5 – 27 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBBE267FE90D647F952.taxon	discussion	Remarks The sculpture of J. hernandezi is very peculiar, with its strong but smooth spiral cords (the uppermost frequently stronger and prominent), and a very prominent peripheral cord. This makes J. hernandezi easily diagnosable among the northeastern Atlantic species of the genus (Rolán & Swinnen 2009).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBAE276F92BD64EF9DE.taxon	diagnosis	Diagnosis Shell of small size for the genus, elevated, thick; sculpture of 4 – 5 well spaced and raised spiral cordls, the uppermost broader, interspaces with small prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBAE276F92BD64EF9DE.taxon	materials_examined	Type material Holotype CANARY ISLANDS • dd; Tenerife, Fañabé; 5 – 27 m depth; MNCN 15.05 / 47569. Paratypes CANARY ISLANDS • 1 dd; same data as for holotype; MNHN • 1 dd; same data as for holotype; BMNH • 1 dd; same data as for holotype; MNHC • 75 dd; same data as for holotype; MHNS • 5 dd; same data as for holotype; FS • 2 dd; same data as for holotype; coll. Ramón Gómez, La Palma • 2 dd; same data as for holotype; coll. Roland de Prins, Brussels • 5 dd; same data as for holotype; coll. José María Hernández, Gáldar, Las Palmas • 4 dd; same data as for holotype; coll. Juan Horro, Vigo. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, shiny, of small size for the genus, height 3.3 – 4.7 mm (4.7), width 2.4 – 3.4 (3.4) mm. Protoconch seemingly smooth, of ca 1.5 whorls, ca 0.30 mm in width. Teleoconch of 5.5 – 6.5 convex whorls. Sculpture of 5 – 7 (6) flat abapical cords, the uppermost broader, and a broader and slightly prominent peripheral cord. Suture not incised. Interspaces narrower than spiral cords, sculptured by dense prosocline growth lines. Base convex, with 6 – 7 (6) spiral cords; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus weakly thickened on lower third, only in adults. Interior of the aperture whitish nacreous. COLOURATION. Colour of protoconch white. Teleoconch of cream or reddish background, with irregular axial prosocline white stripes lined adaperturally with brown, producing white / brown blotches on the peripheral cord; basal cords with similar white blotches lined of brown, over the creamy or reddish background. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBAE276F92BD64EF9DE.taxon	distribution	Distribution Only known from the type locality, Tenerife, Canary Islands, with empty shells at depths of 5 – 27 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF173FFBAE276F92BD64EF9DE.taxon	discussion	Remarks The almost smooth spiral cords, the broader but not prominent uppermost cord, along with the typical colouration with white vertical stripes / blotches lined adaperturally of brown (similarly replicated on the base with white and brown blotches on the spirals), makes J. mabelae rather easily identifiable. The absence of strong prosocline lamellae in the interspaces and of tubercles on the spirals easily separates J. mabelae from most species with convex whorls (see under J. poppei for diagnostic characters).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF172FFB5E251F9B1D291F9B1.taxon	diagnosis	Diagnosis Shell of small size for the genus, cyrtoconoid; sculpture of 4 flat and smooth spiral cordlets, raised and not granulated, narrow interspaces with very fine prosocline lamellae, peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF172FFB5E251F9B1D291F9B1.taxon	materials_examined	Type material Holotype CANARY ISLANDS • dd; Lanzarote Is., harbour of Arrecife; 28 ° 57 ′ N, 13 ° 32 ′ W; USNM 784712. Paratypes CANARY ISLANDS • 1 dd; same data as for holotype; USNM 784713 (paratype I) • 2 dd; same data as for holotype; RBINS 387 - 26439 (paratypes A and H). Other paratypes in the collections G. T. Poppe (Mactan) and M. Curini-Galletti (Sassari). Other material examined CANARY ISLANDS • 4 dd; Lanzarote Is., Arrecife; 2 m depth; MO. Description (in parentheses data of the holotype) SHELL. Shell cyrtoconoid, solid, shiny, of small size for the genus, height 2.7 – 4.8 (3.4) mm, width 2.9 – 4.1 (2.9) mm. Protoconch seemingly smooth, of ca 1.5 whorls, ca 0.2 mm in width. Teleoconch with 4 – 5.5 (5) very convex whorls. Sculpture of 6 – 7 (7) spiral cords of about the same strength, and a broader and slightly prominent peripheral cord. Suture incised. Interspaces narrower than spirals; microsculpture of very fine, markedly prosocline lamellae. Base slightly convex, with 6 basal cords slightly smaller than interspaces; umbilicus closed and covered by a whitish callus, but umbilical chink barely visible in juveniles. Aperture quadrangular, with columellar callus barely thickened on lower third. Interior of aperture whitish-greyish nacreous. COLOURATION. Protoconch whitish. Teleoconch background colour brown on spire and base, sometimes with darker spirals and peripheral cord, and irregular white spots and blotches. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF172FFB5E251F9B1D291F9B1.taxon	distribution	Distribution Arrecife, Lanzarote, Canary Islands, with empty shells at a depth of 2 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF172FFB5E251F9B1D291F9B1.taxon	discussion	Remarks The small size, the almost smooth spiral cords, and the neatly cyrtoconoid outline make J. poppei easily distinguished from almost all other species of Jujubinus. It is similar to the nominal taxon Jujubinus guanchus (which is here recognised as a synonym of J. ruscurianus), described from the same area: J. poppei differs in its more neatly cyrtoconoid outline, and the weaker but more numerous spiral cords (6 – 7 vs 3 – 4 in J. guanchus and 5 – 6 in J. ruscurianus). Jujubinus tingitanus has a similarly Gibbula - like outline, but is easily diagnosed by its lower height / width ratio, and the strongly keeled shell with two major strong spiral cords.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17DFFB4E27FF90CD191FA32.taxon	diagnosis	Diagnosis Shell of medium size for the genus, elevated, thick; sculpture of 6 – 7 granulose spiral cords, interspaces with evident prosocline lamellae, rippled peripheral cord evident, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17DFFB4E27FF90CD191FA32.taxon	materials_examined	Type material Holotype CAPE VERDE • dd; Mordeira Bay, Sal Island; MNCN 15.05 / 44459. Paratypes CAPE VERDE • 1 dd; same data as for holotype; MNCN 15.05 / 44459 • 2 dd; same data as for holotype; MNHN • 2 dd; same data as for holotype; AMNH • 19 dd; same data as for holotype; coll. Emilio Rolán, Vigo. Other material examined CAPE VERDE • 1 dd; Maio Island; CS-PM. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, of medium size for the genus, height 4.5 – 6.0 (4.9) mm, width 3.4 – 4.8 (3.9) mm, shiny. Protoconch seemingly smooth, of ca 1 whorl, ca 0.30 mm in width. Teleoconch of 5 – 6 (6) convex whorls. Sculpture with 4 – 6 (6) narrow, finely granulose whorls, interspaces narrower than the spirals, and a prominent and rippled peripheral cord formed by 2 – 3 merged cordlets. Suture not incised. Growth lines barely visible, interspace of spirals crossed by thick, prosocline lamellae. Base convex, with 6 irregular spiral cords alternate with smaller cordlets; umbilicus closed, covered with a white callus. Aperture quadrangular, with the columellar callus slightly thickened in the lower third. Interior of aperture whitish nacreous. COLOURATION. Colour of the protoconch whitish. Teleoconch with variable colouration, background colour cream to light green, with scattered white spots and greenish-brown or light olive-green flammules, and red blotches on the peripheral cord; base with the same colour pattern with the basal cords colourful of alternating reddish-brownish and whitish. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17DFFB4E27FF90CD191FA32.taxon	distribution	Distribution Sal and Maio Islands, Cape Verde Archipelago, with empty shells at a depth of ca 10 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17DFFB4E27FF90CD191FA32.taxon	discussion	Remarks Jujubinus rubioi is rather similar to J. gravinae, especially for the rippled peripheral cord, but differs in its scabrous sculpture of prosocline lamellae. Jujubinus vexationis is also similar but its peripheral cord is evidently rippled only in the first 2 – 3 whorls.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17CFFB6E1C7F98DD093FACA.taxon	description	Species not assayed molecularly – Fig. 11 K – M	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17CFFB6E1C7F98DD093FACA.taxon	diagnosis	Diagnosis Small and little conical shell, with sculpture of incised spiral lines and strong prosocline lamellae between and over spiral cords, micropustules within interspaces	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17CFFB6E1C7F98DD093FACA.taxon	materials_examined	Type material Holotype CANARY ISLANDS • dd; La Gomera Is., San Sebastian, off Playa de la Cueva; 28 ° 06 ′ N, 17 ° 06 ′ W; 18 m depth; RBINS, MT. 3820 / I. G. 34082. Paratypes CANARY ISLANDS • 1 dd; same data as for holotype; MNHN-IM- 2014 - 7994 (paratype A) • 7 dd; same data as for holotype; CS-PM (paratypes B, M – S) • 10 dd; same data as for holotype; FS (paratypes C – L). Material examined CANARY ISLANDS • 3 dd juvs; same data as for holotype; FS. Description (in parentheses data of the holotype) SHELL. Shell conical, of small size for the genus, height 2.8 – 3.0 (2.8) mm, width 2.5 – 2.8 (2.8) mm, slightly shiny. Protoconch seemingly smooth, of about 1.5 whorls, ca 0.25 mm in width. Teleoconch of 4.5 slightly convex whorls, stepped shoulder on first two whorls. Sculpture of 6 regularly spaced abapical spiral cords, of about the same strength, and a strong basal cord (more pronounced on first whorls). Suture incised. Surface of the teleoconch completely covered by strong prosocline lamellae, regularly spaced, with dense micropustules in the interspaces. Base convex, with 6 – 7 (7) basal spiral cords, umbilicus closed and covered with a white callus. Aperture quadrangular, with the columellar callus slightly thickened in the middle portion. Interior of the aperture whitish nacreous. COLOURATION. Colour of protoconch whitish. Teleoconch yellowish-creamy, with brownish quadrangular spots on spire and base, brown blotches on basal cord. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17CFFB6E1C7F98DD093FACA.taxon	distribution	Distribution Currently only known from La Gomera Is., Canary Islands, empty shells in 18 m depth.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17CFFB6E1C7F98DD093FACA.taxon	discussion	Remarks Jujubinus silbogomerus is similar to J. poppei, from which differs in the strong prosocline lamellae (very fine in J. poppei) and the micropustules in the interspaces. Jujubinus montagui and J. tumidulus share a microsculpture of strong lamellae, but they lack the micropustules in the interspaces, and J. montagui is larger in size (height 5.5 – 7.1 mm vs 2.8 – 3.0 mm in J. silbogomerus).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17EFFB1E262FAA4D798FB65.taxon	diagnosis	Diagnosis Shell of medium size for the genus, elevated, thick; sculpture of 6 – 8, very flat spiral cordlets, separated by incised lines, and crossed by prosocline growth lines; peripheral cord not evident, prosocline lamellae between cords absent, high height / width ratio.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17EFFB1E262FAA4D798FB65.taxon	materials_examined	Type material Holotype CANARY ISLANDS • dd; Fuerteventura, Punta Jandía; 600 m depth; RBINS IG. 32400 MT. 2821. Paratypes CANARY ISLANDS • 1 dd; same data as for holotype; MNCN • 1 dd; same data as for holotype; coll. Roland de Prins, Brussels • 2 dd; same data as for holotype; FS • 1 dd; same data as for holotype; coll. Rafael Mesa, Lanzarote. Description (in parentheses data of the holotype) SHELL. Shell conical, solid, shiny, of medium size for the genus, height 6.0 – 8.5 (8.5) mm, width 4.8 – 6.2 (6.2) mm. Protoconch seemingly smooth, of ca 1 whorl, ca 0.2 mm in width. Teleoconch of 4.5 – 5.5 (5) flat or barely convex whorls. Sculpture of 9 – 10 narrow, not exactly equal in size, abapical spiral whorls, finely granulose in the first whorls, and a broad and not evident peripheral cord. Suture incised. Teleoconch surface covered by evident, dense, very prosocline growth lines. Base convex, with 10 – 12 irregular, flat spiral cords; umbilicus closed, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish-pinkish nacreous. COLOURATION. Colour of the protoconch white. Teleoconch of pink-reddish background, with scattered white spots and irregularly arranged whitish stripes of variable breadth; base with the same colour pattern. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17EFFB1E262FAA4D798FB65.taxon	distribution	Distribution Only known from the type locality, Fuerteventura, Canary Islands, with empty shells at a depth of 600 m (but see Remarks).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17EFFB1E262FAA4D798FB65.taxon	discussion	Remarks The known specimens of J. rafaelmesai (all empty shells) have been collected at a very high depth (600 m) compared with all the other species of the genus, and in our opinion, an origin of the shells from shallower habitats is very probable. The general habitus recalls that of the species in the J. curinii complex from which it differs in its larger size and the chromatic pattern: a relationship with this complex is worthy an investigation. Similarly, along with the peculiar chromatic pattern, the large size separates J. rafaelmesai from the other Canarian species devoid of evident tubercles or lamellae (6 – 8.5 mm vs 4.7 mm in J. mabelae, 3.5 mm in J. poppei, and 3.4 mm in J. guanchus (Curini-Galletti 1985 )).	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF179FFB0E1CAFB47D034FB87.taxon	conservation	Species not assayed molecularly – Fig. 12 A – C	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF179FFB0E1CAFB47D034FB87.taxon	diagnosis	Diagnosis Shell of medium size for the genus, light, slightly turriculate; sculpture of only incised spiral lines, prosocline lamellae between cords absent.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF179FFB0E1CAFB47D034FB87.taxon	materials_examined	Type material Holotype ALBORAN SEA • dd; Alborán Island; 35 ° 56 ′ 43.99 ″ N, 3 ° 02 ′ 03.73 ″ W; MNHN-IM- 2000 - 30131. Paratypes ALBORAN SEA • 1 dd; same data as for holotype; MNHN-IM- 2000 - 30132 (paratype 1) • 2 dd; same data as for holotype; MZB- 60153 (paratype 2), MZB- 60154 (paratype 3) • 2 dd; same data as for holotype; MCZR 00225 A (paratype 4), MCZR 00225 B (paratype 5) • 2 dd; same data as for holotype; MNCN 15.05 / 60160 (paratypes 6 – 7) • 13 dd; same data as for holotype; CS-PM (paratypes 8 – 20) • 102 dd; same data as for holotype; MO (paratypes 21 – 122). Other material examined ALBORAN SEA • 8 dd; same data as for holotype; CS-PM.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF179FFB0E1CAFB47D034FB87.taxon	description	Description (in parentheses data of the holotype) SHELL. Shell conical, slightly turriculate, of medium size for the genus, height 6.5 – 8.9 (6.8) mm, width 5.0 – 6.0 (5.3) mm, light. Protoconch seemingly smooth, of ca 1.1 whorl, 0.37 – 0.40 (0.39) mm in width. Teleoconch of 4.5 – 5.5 (4.5) slightly convex whorls. Sculpture of 8 – 10 (9) flat, weak and closely set abapical spiral cords of about the same strength, and a weak peripheral cord, slightly shouldered, formed by 2 – 3 (3) merged cordlets. Suture incised. Teleoconch surface covered by barely visible prosocline growth striae, irregularly set. Base convex, with 7 – 8 (8) regularly spaced, basal spiral cords, large and flat; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and early teleoconch whorls bright red; remaining whorls and base with alternating brownish-reddish and yellowish-orange spiral stripes (of different size in the holotype), with the upper part of the brownish-reddish stripes dotted with red. SOFT PARTS. Unknown.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF179FFB0E1CAFB47D034FB87.taxon	distribution	Distribution Known only from the type locality, Alborán Island, with empty shells at depths of 30 – 37 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF179FFB0E1CAFB47D034FB87.taxon	discussion	Remarks Jujubinus alboranensis is a member of the J. curini complex, and differs from most of the Atlantic and Mediterranean Jujubinus species by lacking the typical prosocline lamellae in the spiral furrows between cords, see under J. curinii, J. eleonorae, J. browningleeae Smriglio, Mariottini & Swinnen, 2018, J. roseus, and J. trilloi for diagnostic differences.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF178FFB3E1FFFB1AD283FC86.taxon	description	Species not assayed molecularly – Fig. 12 D – F	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF178FFB3E1FFFB1AD283FC86.taxon	diagnosis	Diagnosis Shell of medium size for the genus, slightly turriculate; sculpture of flat spiral cords delimited by narrow incised spiral lines, prosocline lamellae between cords absent.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF178FFB3E1FFFB1AD283FC86.taxon	materials_examined	Type material Holotype PORTUGAL • dd; Gorringe Ridge, 130 miles off Cape St Vincent; RBINS: I. G. 33787 / MT. 3687. Paratype PORTUGAL • 1 dd; same data as for holotype; FS.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF178FFB3E1FFFB1AD283FC86.taxon	description	Description of the holotype SHELL. Shell conical, slightly turriculate, of medium size for the genus, height 5.2 mm, width 3.8 mm. Protoconch seemingly smooth, of ca 1 whorls, 0.27 mm in width. Teleoconch of 5.5 slightly convex whorls. Sculpture consisting of 6 flat, regularly spaced, spiral cords, slightly shouldered, delimited by incised spiral lines, and a broader and evident basal cord. Suture well incised. Teleoconch surface entirely covered by evident prosocline growth striae. Base convex, with 7 basal spiral cords, of unequal size, large and flat; umbilicus closed and covered with a whitish callus. Aperture quadrangular, with the columellar callus thickened in the middle portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and earliest teleoconch whorls reddish, remaining whorls and base uniformly dark-brown with axial wavy and narrow yellow stripes. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF178FFB3E1FFFB1AD283FC86.taxon	distribution	Distribution Only known from the bank of the Gorringe Ridge, with empty shells at a depth of 30 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF178FFB3E1FFFB1AD283FC86.taxon	discussion	Remarks Jujubinus browningleeae is the first species of the Jujubinus curinii complex discovered in the Atlantic Ocean, all other species being exclusively Mediterranean. Jujubinus browningleeae differs from J. curinii in its almost non-stepper whorls, the fewer spiral cords, the broader peripheral cord, and the darker colouration. From J. alboranensis and J. trilloi it differs in the fewer spiral cords, the broader peripheral cord, and the darker colouration. From J. eleonorae it differs in the weaker spiral sculpture with narrower interspaces, and the darker colouration.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17BFFB2E247FC18D09EFCA0.taxon	diagnosis	Diagnosis Shell of small size for the genus, solid, with stepped spire; sculpture of only incised spiral lines, prosocline lamellae between cords absent.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17BFFB2E247FC18D09EFCA0.taxon	materials_examined	Type material Holotype ITALY • dd; Strait of Messina, Scilla; 38 ° 15 ′ 36 ″ N, 15 ° 43 ′ 08 ″ E; 42 m depth; Museo di Storia Naturale, Livorno (Malacology, V- 730). Paratypes ITALY • 1 dd; same data as for holotype; MZB- 45234 (unumbered paratype) • 1 dd; same data as for holotype; coll. Cesare Bogi, Livorno (paratype A) • 1 dd; same data as for holotype; coll. Enzo Campani, Livorno (paratype B) • 2 dd; same data as for holotype; coll. Francesco Ciriaco, Livorno (unnumbered paratypes). Other material examined ITALY • 200 dd; same data as for holotype; CS-PM. Description (in parentheses data of the holotype) SHELL. Solid shell of small size for the genus, height 3.9 – 4.6 (4.0) mm, width 2.9 – 3.0 (3.1) mm, conical, solid. Protoconch seemingly smooth of ca 1.1 whorls, ca 0.3 mm in width. Teleoconch of 5 - 6 (6) flat or barely convex whorls. Sculpture of 10 flat, weak, smooth spiral cords, merged into 5 pairs separated by incised spiral lines, and a peripheral pair of slightly broader cords. Suture incised, with one nonprotruding sutural cingulate. Teleoconch with shiny surface covered by barely visible prosocline growth striae, irregularly set. Base convex, with 9 – 10 flat, broad, spaced spiral cords, umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with the columellar callus slightly thickened in the central portion. Interior of the aperture whitish, nacreous. COLOURATION. Colour of protoconch and early teleoconch of shiny reddish-brown background, remaining whorls brownish (occasionally greenish), with small white blotches and spots. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17BFFB2E247FC18D09EFCA0.taxon	distribution	Distribution Only known from the Strait of Messina, with empty shells at a depth of 42 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17BFFB2E247FC18D09EFCA0.taxon	discussion	Remarks Jujubinus curinii was the first of this group of ‘ smooth’ species described for the Mediterranean Sea (Bogi & Campani 2006), all lacking the typical prosocline lamellae in the spiral furrows between cords. It differs from J. alboranensis in its smaller size (3.9 – 4.6 mm vs 6.5 – 8.9 mm in J. alboranensis), and the different colouration (shiny reddish-brown with whitish blotches and spots vs yellowish-orange narrow spirals with alternating bright red dots and spotsin J. alboranensis). Jujubinus curinii is apparently endemic to the Strait of Messina, J. alboranensis to the Alboran Sea.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17AFFADE1D7FC3BD0AEFC43.taxon	description	Species not assayed molecularly – Fig. 12 J – L	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17AFFADE1D7FC3BD0AEFC43.taxon	diagnosis	Diagnosis Shell of medium size for the genus, stout; flat spiral cords, strong spiral sculpture; basal cord evident; prosocline lamellae between cords absent.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17AFFADE1D7FC3BD0AEFC43.taxon	materials_examined	Type material Holotype STRAIT OF SICILY • dd; Skerki Bank; 37 ° 45 ′ 00 ″ N, 10 ° 50 ′ 00 ″ E; 37 m depth; MNHN-IM- 2000 - 25883. Paratypes STRAIT OF SICILY • 1 dd; same data as for holotype; MZB- 49668 (paratype 1) • 1 dd; same data as for holotype; MCZR 00227 (paratype 2) • 37 dd; same data as for holotype; CS-PM (paratypes 3 – 39) • 20 dd; same data as for holotype; EDS (paratypes 40 – 59) • 10 dd; same data as for holotype; MO (paratypes 60 – 69) • 10 dd; same data as for holotype; IN (paratypes 70 – 79) • 40 dd; same data as for holotype; PT (paratypes 80 – 119) • 2 dd; same data as for holotype; BA (paratypes 120 – 121). Other material examined STRAIT OF SICILY • 25 dd; same data as for holotype; MO • 606 dd; same data as for holotype; IN.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17AFFADE1D7FC3BD0AEFC43.taxon	description	Description (in parentheses data of the holotype) SHELL. Shell conical, of medium size for the genus, height 5.5 – 6 mm (5.7), width 4 – 4.5 (4.7) mm, rather solid. Protoconch seemingly smooth, of ca 1.1 whorls, 0.24 – 0.25 mm in width. Teleoconch of 4.5 – 5.5 slightly convex whorls. Sculpture of 6 – 7 strong abapical spiral cords of about the same strength, slightly shouldered, and a pronounced basal cord, rounded, formed by 2 cordlets separated by a furrow, the lower twice as wide as the upper one. Suture barely visible. Teleoconch surface covered by faint prosocline growth striae, in some specimens more evident and irregular. Base convex, with 4 – 5 flat spiral cords, regularly spaced, crossed by numerous growth striae; umbilicus closed also in juveniles, covered with a white callus. Aperture quadrangular, with a columellar callus thickened in the middle portion. Interior of the aperture whitish nacreous. COLOURATION. Colour of protoconch and early teleoconch whorls bright red, remaining whorls and base light green-brownish with 3 regularly spaced, thick yellowish-orange spiral lines covering a portion of the spiral cords, and 4 – 5 lines on the base. SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17AFFADE1D7FC3BD0AEFC43.taxon	distribution	Distribution Only known from the Skerki Bank, Strait of Sicily, with empty shells at a depth of 37 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF17AFFADE1D7FC3BD0AEFC43.taxon	discussion	Remarks The stronger spiral sculpture and the peculiar chromatic pattern (with bright orange spiral lines) are diagnostic with the other species of the Jujubinus curini complex. Juveniles can also be easily diagnosed: Jujubinus curinii has weaker spiral sculpture, is lighter in colouration and lacks the orange spiral lines; J. alboranensis has also a weaker spiral sculpture, has a bright red protoconch and early teleoconch whorls, but has always small white blotches on the peripheral cord.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF165FFACE1B3FC5ED703FD26.taxon	diagnosis	Diagnosis Shell of medium size for the genus, slightly turriculate; sculpture of flat spiral cords; prosocline lamellae between cords absent.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF165FFACE1B3FC5ED703FD26.taxon	materials_examined	Type material Holotype SPAIN • lv; off Alborán Island, Fauna Ibérica IV stn 315 B; 35 ° 58.00 ′ N, 02 ° 58.46 ‛ W; 37 m depth; Jul. 1996; infralittoral bottom; Fauna Ibérica IV leg.; MNCN 15.05 / 35835 H. Paratypes SPAIN • 5 lv, 5 dd (including seven juveniles); same data as for holotype; MNCN 15.05 / 35835 P.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF165FFACE1B3FC5ED703FD26.taxon	description	Description SHELL. Shell of medium size for the genus, height 5.5 – 8.9 mm, width 4.3 – 6.3 mm, conical, light. Protoconch of 0.85 whorls, 0.27 mm in width, sculptured by two spiral threads. Teleoconch of 4.5 – 6.5 slightly convex whorls with stepped shoulder. Sculpture of 6 – 10 flat, closely set abapical spiral cords of about the same strength, broader than interspaces, and a peripheral cord, not prominent, slightly shouldered and ratchet, formed by 2 – 3 merged cordlets. Suture incised. Teleoconch surface covered by faint prosocline growth striae. Base convex, with 13 – 14 (13) regularly spaced, basal broad and flat spiral cordlets of variable size; umbilicus closed also in juveniles, covered with a whitish-pinkish callus. Aperture quadrangular, with columellar callus thickened in lower third. Interior of the aperture whitish-pinkish nacreous. COLOURATION. Colour of protoconch and initial teleoconch whorls bright intense pink; remaining whorls and base pale pink, with brownish-reddish flexuous axial lines of different size and arrangement. SOFT PARTS (based on Peñas et al. 2006: fig. 424, as Jujubinus sp.). Epipodial tentacles light purple-reddish. Foot reddish, sole creamy yellowish. Cephalic tentacles purple-reddish, with a dorsal purple stripe. Eye stalks reddish, eyes black encircled by creamy-yellow. Snout reddish with a creamy terminal band. Cephalic lappets whitish.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF165FFACE1B3FC5ED703FD26.taxon	distribution	Distribution Currently known only from Alborán Island, with living specimens at depths of 10 – 34 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF165FFACE1B3FC5ED703FD26.taxon	discussion	Remarks This is the last described species in the J. curinii complex, currently under description elsewhere (Smriglio et al. 2026). It has been collected sympatrically with J. alboranensis, from which it differs in the more numerous, flat spiral cords (13 – 14 vs 6 in J. alboranensis) and the consistently different pink colouration. From J. trilloi it differs in its larger shell (8 mm vs rarely up to 6.5 mm in J. trilloi), the fewer, flat spiral cordlets (12 – 13 vs 6 – 8 in J. trilloi) and the different colouration.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF164FFAEE1ADFCB8D2A9FD9F.taxon	diagnosis	Diagnosis Shell of medium size for the genus, slightly turriculate; sculpture of only incised spiral lines, prosocline lamellae between cords absent.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF164FFAEE1ADFCB8D2A9FD9F.taxon	materials_examined	Type material Holotype STRAIT OF SICILY • dd; Talbot Bank; 37 ° 30 ′ 00 ″ N, 11 ° 40 ′ 00 ″ E; 25 m depth; MNHN-IM- 2010 - 25882. Paratypes STRAIT OF SICILY • 1 dd; same data as for holotype; MZB- 49669 (paratype 1) • 1 dd; same data as for holotype; MCZR 00226 (paratype 2) • 52 dd; same data as for holotype; CS-PM (paratypes 3 – 54) • 5 dd; same data as for holotype; MO (paratypes 55 – 59) • 30 dd; same data as for holotype; PT (paratypes 60 – 89). Description (in parentheses data of the holotype) SHELL. Shell conical, slightly turriculate, of medium size for the genus, height 5 – 6.5 (5.4) mm, width 4 – 4.5 (4.6) mm, solid. Protoconch seemingly smooth, of ca 1 whorl, 0.24 – 0.25 (0.25) mm in diameter. Teleoconch of 4.5 – 5.5 (4.5) flat or slightly convex whorls. Sculpture of 6 – 8 (8) regularly spaced, fine, incised spiral lines. Basal cord rounded, formed by 2 cordlets separated by a fine furrow, the upper one as wide as one third to one quarted the lower one. Suture distinct. Teleoconch surface covered by faint prosocline growth striae. Base convex, with 9 – 11 (11) concentric fine incised spiral lines, irregularly spaced and crossed by numerous growth striae. Umbilicus closed also in juveniles covered with a white callus. Aperture quadrangular with a columellar callus thickened in the middle portion. Interior of the aperture whitish, iridescent. COLOURATION. Colour of protoconch and early teleoconch whorls bright red, remaining whorls and base greenish-brownish, with axial opisthocline brown-reddish narrow lines. Three to five (4) narrow yellowish-orange spiral lines on the spire, 4 – 6 (4) on the base SOFT PARTS. Not known.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF164FFAEE1ADFCB8D2A9FD9F.taxon	distribution	Distribution Only known from the Talbot Bank, Strait of Sicily, with empty shells at a depth of 25 m.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
CB4C87EFF164FFAEE1ADFCB8D2A9FD9F.taxon	discussion	Remarks Jujubinus trilloi differs from J. curinii in the less stepped whorls, and the colouration with bright red protoconch and early teleoconch whorls, the remaining whorls and base greenish-brownish, with narrow axial opisthocline brown-reddish lines, and the yellowish-orange spiral lines on spire and base (vs shiny reddish-brown protoconch and early teleoconch whorls, with sparse whitish spots in J. curinii). Jujubinus alboranensis differs in its slightly less convex whorls, and the white blotches on the peripheral cord. Jujubinus eleonorae differs in its stronger sculpture, and in the spiral orange lines that are similar but thicker than in J. trilloi.	en	Mariottini, Paolo, Aissaoui, Chrifa, Chiappa, Giacomo, Giulio, Andrea Di, Fassio, Giulia, Furfaro, Giulia, Modica, Maria Vittoria, Pezzotti, Simone, Puillandre, Nicolas, Smriglio, Carlo, Oliverio, Marco (2026): Integrative taxonomy of northeastern Atlantic top-shells of the genus Jujubinus Monterosato, 1884 (Gastropoda, Trochidae, Cantharidinae). European Journal of Taxonomy 1044: 1-96, DOI: 10.5852/ejt.2026.1044.3223, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3223/14285
