taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C418D608FFB5FFBB8970F885025010A2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881526/files/figure.png	https://doi.org/10.5281/zenodo.18881526	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFB5FFBB8970F885025010A2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838950/files/figure.png	https://doi.org/10.5281/zenodo.18838950	FIG. 3. — Frullania beauverdii Steph.: A, portion of male plant showing androecia; B, portion of female plant showing terminal gynoecium; C, D, stem leaves; E, stylus;F-I, stem underleaves;J, perianth;K, perianth beak; L-N, perianth cross sections: M, medial portion; L, N, above and below the medial portion respectively). All from Hegewald & Hegewald 8717 (MO). Scale bars: A, B, J, L-N, 1.25 mm; C, D, F-I, 0.5 mm; E, K, 125 µm. Credits: Bryan Espinoza-Prieto.	FIG. 3. — Frullania beauverdii Steph.: A, portion of male plant showing androecia; B, portion of female plant showing terminal gynoecium; C, D, stem leaves; E, stylus;F-I, stem underleaves;J, perianth;K, perianth beak; L-N, perianth cross sections: M, medial portion; L, N, above and below the medial portion respectively). All from Hegewald & Hegewald 8717 (MO). Scale bars: A, B, J, L-N, 1.25 mm; C, D, F-I, 0.5 mm; E, K, 125 µm. Credits: Bryan Espinoza-Prieto.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFB8FFB68975F9B305B016B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838952/files/figure.png	https://doi.org/10.5281/zenodo.18838952	FIG. 4. — Distribution of Frullania cuencensis Taylor (diamonds) and F. pyricalycina Steph. (stars).	FIG. 4. — Distribution of Frullania cuencensis Taylor (diamonds) and F. pyricalycina Steph. (stars).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFBBFFB08B23FDB407761678.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838956/files/figure.png	https://doi.org/10.5281/zenodo.18838956	FIG. 5. — Distribution of Frullania holostipula S.Hatt. & D.G.Griffin (diamonds), F. pluricarinata Gottsche (stars) and F. winteri Steph. (triangles).	FIG. 5. — Distribution of Frullania holostipula S.Hatt. & D.G.Griffin (diamonds), F. pluricarinata Gottsche (stars) and F. winteri Steph. (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFBDFFB08955FEF7036410A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838956/files/figure.png	https://doi.org/10.5281/zenodo.18838956	FIG. 5. — Distribution of Frullania holostipula S.Hatt. & D.G.Griffin (diamonds), F. pluricarinata Gottsche (stars) and F. winteri Steph. (triangles).	FIG. 5. — Distribution of Frullania holostipula S.Hatt. & D.G.Griffin (diamonds), F. pluricarinata Gottsche (stars) and F. winteri Steph. (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFBDFFB08955FEF7036410A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881528/files/figure.png	https://doi.org/10.5281/zenodo.18881528	FIG. 6. — Frullania pluricarinata Gottsche: A, habit; B, C, stem leaves; D-H, styli; I-P, stem underleaves; Q, R, innermost gynoecial leaves; S, perianth; T, cross- section of perianth; U, androecium. A, F-H, O, P, from probable type (S-B237892); B-D, I-K, R-U, from Espinoza-Prieto & Ccanto 2160 (MO); C, E, L-N, Q, from Hegewald & Hegewald 8373 (MO). Scale bars: A, 1.25 mm; B, C, I-R, 0.25 mm; D-H, 125 µm; S-U, 0.5 mm. Credits: Bryan Espinoza-Prieto.	FIG. 6. — Frullania pluricarinata Gottsche: A, habit; B, C, stem leaves; D-H, styli; I-P, stem underleaves; Q, R, innermost gynoecial leaves; S, perianth; T, cross- section of perianth; U, androecium. A, F-H, O, P, from probable type (S-B237892); B-D, I-K, R-U, from Espinoza-Prieto & Ccanto 2160 (MO); C, E, L-N, Q, from Hegewald & Hegewald 8373 (MO). Scale bars: A, 1.25 mm; B, C, I-R, 0.25 mm; D-H, 125 µm; S-U, 0.5 mm. Credits: Bryan Espinoza-Prieto.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFBEFFB2891EF9F302E91367.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838956/files/figure.png	https://doi.org/10.5281/zenodo.18838956	FIG. 5. — Distribution of Frullania holostipula S.Hatt. & D.G.Griffin (diamonds), F. pluricarinata Gottsche (stars) and F. winteri Steph. (triangles).	FIG. 5. — Distribution of Frullania holostipula S.Hatt. & D.G.Griffin (diamonds), F. pluricarinata Gottsche (stars) and F. winteri Steph. (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFBEFFB2891EF9F302E91367.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838959/files/figure.png	https://doi.org/10.5281/zenodo.18838959	FIG. 7. — Frullania winteri Steph.: A, habit; B, C, stem leaves; D, E, stem underleaves; F-L, styli; M, N, innermost gynoecial leaves; O, innermost bracteole (upper-half portion). A, F-H, M, N, from Hegewald & Hegewald 7059 (MO); B-E, I-L, M, from lectotype (G00066922). Scale bars: A, 1.25 mm; B-E, M-O, 0.5 mm; F-L, 0.25 mm. Credits: Bryan Espinoza-Prieto.	FIG. 7. — Frullania winteri Steph.: A, habit; B, C, stem leaves; D, E, stem underleaves; F-L, styli; M, N, innermost gynoecial leaves; O, innermost bracteole (upper-half portion). A, F-H, M, N, from Hegewald & Hegewald 7059 (MO); B-E, I-L, M, from lectotype (G00066922). Scale bars: A, 1.25 mm; B-E, M-O, 0.5 mm; F-L, 0.25 mm. Credits: Bryan Espinoza-Prieto.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFBFFFAC8B01FB3207E514E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881526/files/figure.png	https://doi.org/10.5281/zenodo.18881526	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA1FFAC88ACFBB5026A10A2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881526/files/figure.png	https://doi.org/10.5281/zenodo.18881526	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA3FFAE8940FF5607F510A2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881526/files/figure.png	https://doi.org/10.5281/zenodo.18881526	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	FIG. 2. — Distribution of Frullania beauverdii Steph. (diamonds), F. intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. var. intumescens (stars), Frullania lobatohastata Steph. (square) and F. nodulosa (Reinw., Blume & Nees) Nees (triangles).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA3FFAE8B90FE97024C10A2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838952/files/figure.png	https://doi.org/10.5281/zenodo.18838952	FIG. 4. — Distribution of Frullania cuencensis Taylor (diamonds) and F. pyricalycina Steph. (stars).	FIG. 4. — Distribution of Frullania cuencensis Taylor (diamonds) and F. pyricalycina Steph. (stars).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA5FFAA8959FEB602CC10A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881532/files/figure.png	https://doi.org/10.5281/zenodo.18881532	FIG. 8. — Distribution of Frullania tetraptera Nees & Mont. Type (star), other records (diamonds).	FIG. 8. — Distribution of Frullania tetraptera Nees & Mont. Type (star), other records (diamonds).	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA5FFAA8959FEB602CC10A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881534/files/figure.png	https://doi.org/10.5281/zenodo.18881534	FIG. 9. — Frullania tetraptera Nees & Mont: A, C, parts of the same female plant; B, female plant with perianth. All from isolectotype, d’Orbigny s.n. (STR). Corresponding asterisks (*) indicate where the same shoot is continued after a break in the drawing. Scale bar: 1.5 mm. Credits: Yuriy S. Mamontov.	FIG. 9. — Frullania tetraptera Nees & Mont: A, C, parts of the same female plant; B, female plant with perianth. All from isolectotype, d’Orbigny s.n. (STR). Corresponding asterisks (*) indicate where the same shoot is continued after a break in the drawing. Scale bar: 1.5 mm. Credits: Yuriy S. Mamontov.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA5FFAA8959FEB602CC10A5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838965/files/figure.png	https://doi.org/10.5281/zenodo.18838965	FIG. 10. — Frullania tetraptera Nees & Mont: A-C, shoot fragments with androecia, parts of the same male plant. All from isolectotype, d’Orbigny s.n. (STR).Corresponding asterisks (*) indicate where the same shoot is continued after a break in the drawing. Scale bar: 1.5 mm. Credits: Yuriy S. Mamontov.	FIG. 10. — Frullania tetraptera Nees & Mont: A-C, shoot fragments with androecia, parts of the same male plant. All from isolectotype, d’Orbigny s.n. (STR).Corresponding asterisks (*) indicate where the same shoot is continued after a break in the drawing. Scale bar: 1.5 mm. Credits: Yuriy S. Mamontov.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA8FFA58965FF56077A13A7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18838967/files/figure.png	https://doi.org/10.5281/zenodo.18838967	FIG. 11. — Frullania conferta Spruce: A-C, F, J, N-P, V, underleaves; D, G-I, K, Q-U, W, X, parts of shoots showing leaf lobes, lobules and styli; E, fertile shoot with an androecium and two gynoecia;L, median leaf cells; M, fertile shoot with a perianth. All from lectotype (G00047604). Scale bars: A-C, F, J, N-P, V, 0.5 mm; E, 2 mm; D, H, I, K, R, S, U, W, X, 1 mm; G, Q, T, 0.75 mm; L, 0.05 mm; M, 1.5 mm. Credits: Yuriy S. Mamontov.	FIG. 11. — Frullania conferta Spruce: A-C, F, J, N-P, V, underleaves; D, G-I, K, Q-U, W, X, parts of shoots showing leaf lobes, lobules and styli; E, fertile shoot with an androecium and two gynoecia;L, median leaf cells; M, fertile shoot with a perianth. All from lectotype (G00047604). Scale bars: A-C, F, J, N-P, V, 0.5 mm; E, 2 mm; D, H, I, K, R, S, U, W, X, 1 mm; G, Q, T, 0.75 mm; L, 0.05 mm; M, 1.5 mm. Credits: Yuriy S. Mamontov.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
C418D608FFA8FFA58979FAB203E81579.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881536/files/figure.png	https://doi.org/10.5281/zenodo.18881536	FIG. 12. —Frullania mexicana Lindenb: A, B, fertile shoots;C, D, shoot fragments.A,from lectotype of F.semiconnata Lindenb.& Gottsche (G00114899); B-D,from isolectotype of F. mexicana Lindenb. (W-2014-06935). Scale bar: 1.5 mm. Credits: Yuriy S. Mamontov.	FIG. 12. —Frullania mexicana Lindenb: A, B, fertile shoots;C, D, shoot fragments.A,from lectotype of F.semiconnata Lindenb.& Gottsche (G00114899); B-D,from isolectotype of F. mexicana Lindenb. (W-2014-06935). Scale bar: 1.5 mm. Credits: Yuriy S. Mamontov.	2026-01-30	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.		Zenodo	biologists	Espinoza-Prieto, Bryan;Mamontov, Yuriy S.;Opisso, Jasmin;Atwood, John J.			
