identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C31687B2FF8D900DBB46F9BDFD3ADDA5.text	C31687B2FF8D900DBB46F9BDFD3ADDA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triportheus claudiae Lopes & Carvalho 2024	<div><p>Triportheus claudiae,  new species</p><p>urn:lsid:zoobank.org:act: D56A661B-3783-4698-A76E-40E0C911918D</p><p>(Figs. 1–2; Tab. 1)</p><p>Triportheus aff. rotundatus . — Mariguela et al., 2016:132 [phylogenetic relationships and biogeography].</p><p>Holotype. CITL 400, 102.6 mm SL, bay on <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.053886&amp;materialsCitation.latitude=-19.31667" title="Search Plazi for locations around (long -57.053886/lat -19.31667)">Pousada Arara Azul</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.053886&amp;materialsCitation.latitude=-19.31667" title="Search Plazi for locations around (long -57.053886/lat -19.31667)">Pantanal</a>, Corumbá municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.053886&amp;materialsCitation.latitude=-19.31667" title="Search Plazi for locations around (long -57.053886/lat -19.31667)">rio Paraguai</a> basin, Mato Grosso do Sul State, Brazil, 19º19’0.01”S 57º03’14.00”W, 25 Mar 2019, H. Gimenes-Junior, T. T. M. Taveira, F. Severo-Neto &amp; R. Rech.</p><p>Paratypes. Brazil, Mato Grosso do Sul State: CITL 401, 3, 94.3–99.7 mm SL, collected with the holotype .  CITL 1135, 3 c&amp;s, 45.3–52.7 mm.  LBP 3720, 7, 45.2–69.3 mm SL, Aquidauana Municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.021915&amp;materialsCitation.latitude=-19.582863" title="Search Plazi for locations around (long -57.021915/lat -19.582863)">rio Negro</a>, 19º34’54”S 56º15’16.5”W, 3 Aug 2006, C. Oliveira &amp; L. H. G. Pereira.   LBP 3757, 16, 63.4–74.5 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.021915&amp;materialsCitation.latitude=-19.582863" title="Search Plazi for locations around (long -57.021915/lat -19.582863)">rio Negro</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.021915&amp;materialsCitation.latitude=-19.582863" title="Search Plazi for locations around (long -57.021915/lat -19.582863)">rio Paraguai</a> basin, Aquidauana Municipality, 19º34’33.7”S 56º14’49”W, 1 Aug 2006, C. Oliveira &amp; L. H. G. Pereira.   LBP 11917, 5, 37.9–49.4 mm SL, Corumbá municipality, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.021915&amp;materialsCitation.latitude=-19.582863" title="Search Plazi for locations around (long -57.021915/lat -19.582863)">rio Miranda</a>, 19º34’58.3”S 57º01’18.9”W, 25 Nov 2009, C. Oliveira and team  .   Mato Grosso State: MNRJ 21681, 82.5 mm SL, Barão de Melgaço,  rio Cuiabá,  Baía Chacororé, 16 Jan 1978, E. P. Caramaschi &amp; CEPIPAM  .</p><p>Non-types. The following specimens were not included as type-material due to bad preservation of some morphological structures.   Rio Paraguai basin, Brazil, Mato Grosso State: LBP 11916, 3, 55.2–70.7 mm SL.  LBP 18627, 3, 68.0– 96.1 mm. Mato Grosso do Sul State: SL. LBP 3557, 62.2 mm SL. LBP 3795, 4, 59.3–81.7 mm SL. LBP 5802, 4, 32.2–35.0 mm SL.</p><p>Diagnosis.  Triportheus claudiae differs from its congeners by the following combination of characters: two non-elongated scales between the pectoral fin and the ventral keel (vs. one vertically elongated scale between the pectoral fin and the ventral keel in  T. albus,  T. auritus,  T. brachipomus,  T. culter (Cope, 1872), and  T. magdalenae); gill rakers on the lower branch of the first branchial arch, 24–29 (mode = 25, n = 8) (vs. 40–47 in  T. trifurcatus (Castelnau, 1855), 38–48 in  T. nematurus, 52–57 in  T. guentheri (Garman, 1890), 35–44 in  T. signatus); predorsal scales regularly disposed, in series, or with maximum two scales out of series (vs. predorsal scales irregularly disposed, not in series, with more than three scales disorganized in  T. angulatus,  T. rotundatus,  T. nematurus,  T. guentheri,  T. curtus (Garman, 1890),  T. pantanensis,  T. orinocensis Malabarba, 2004, and  T. venezuelensis Malabarba, 2004); premaxilla teeth in three rows (vs. teeth of premaxilla in two rows in  T. curtus and  T. pictus (Garman, 1890); epipleural bones: 20–21 (vs. 16–17 in  T. pantanensis and 22–23  T. signatus); ventral tubules in the laterossensory canal of the preopercule: 7 (vs. 4–5 in  T. nematurus and 5 in  T. signatus).</p><p>Description. Morphometric data in Tab. 1. Deep and short body, laterally compressed, with ventral keel by expansion of coracoids. Greatest body depth at vertical through origin of dorsal fin. Slightly convex dorsal profile. Ventral profile convex and expanded between insertion of head and origin of pelvic fin.</p><p>Two pairs of nostrils, anterior and posterior opening of similar size; anterior opening circular and posterior opening oval. Terminal mouth, lower jaw largest than the upper jaw. Premaxillary teeth in three rows: outer row with 4(2), 5*(26), or 6(6) tricuspid small teeth; medial row with rarely 2(1) or 3*(33) tricuspid, large teeth and inner row with 5(6), 6(24), or 7*(4) pentacuspid teeth, largest from other teeth on premaxilla. Maxillary bone with 0(12), 1(6), 2*(12), or 3(4) tiny tricuspid teeth. Lower jaw teeth in two rows: outer row with 4(2), 5(27), or 6*(5) large pentacuspid teeth, and inner row with one conical symphysial, large tooth. Gill rakers smaller and wider than branchial filaments; gill rakers on lower branch of first branchial arch: 24(4), 25(8), 26(4), 27*(7), 28(6), or 29(5).</p><p>Dorsal fin with ii,9 rays (33); anal fin with iii,26(1), 27*(6), 28(13), 29(9), 30(2) or 31(3) rays, its origin at vertical through last ray of dorsal fin. First branched ray of anal fin larger; pectoral fin with i,10(3), 11*(16) or 12(15) rays, extending more than half length of pelvic fin; pelvic fin with i,6 rays (32); caudal fin with i,9/8,i rays (31), bifurcated margin in juveniles and truncated margin in adults. Middle caudal rays with small extensions in large individuals.</p><p>Scales cycloid, large, with approximately same size of orbit. Lateral line complete, curved, its origin dorsally at flank, on supracleithrum and following to ventral region of body after pectoral fin, in third row of scales in vertical of pelvic fin, finishing at medial rays of caudal fin. Canals of lateral line scales with one to three branches; generally, small individuals (less than 50 mm SL) with one single branch and large individuals (more than 50 mm SL) with two or three branches. Perforated scales on lateral line: 31(7), 32(5), 33(7), or 34*(15); six scales above lateral line and two scales below lateral line; two scales on vertical line between origin of pectoral fin and ventral keel; predorsal scales: 9(1), 10(2), 11(6), 12(7), or 13*(2), in regular series; 10(8) or 11(29) scales around caudal peduncle.</p><p>Total vertebrae: 35(2) or 36(1); precaudal vertebrae: 17(3); caudal vertebrae: 18(2) or 19(1); epineural bones: 35(2) or 37(1), with first three unbranched; epipleural bones: 20(1) or 21(2), with first epipleural unbranched; epipleural bones from second to 12 th branched in dorsal portion; epipleural bones from 13 th to 21 st unbranched. Upper procurrent rays: eight (1) or nine (2); lower procurrent rays: seven (1) or eight (2).</p><p>Coloration in alcohol. Overall ground coloration yellowish silver, darker dorsally. Melanophores on proximal and distal fields of scales, forming five to eight fainted black stripes on flank. Stripes above lateral line generally with grouped melanophores, mainly on third to fifth stripes (from dorsal to ventral region). Remaining stripes with scattered melanophores. One inconspicuous stripe on scales from lateral line, by presence of few scattered melanophores; black stripes on flank converging with diffuse spot at hypural plate and base of caudal fin rays. Melanophores in anterodorsal region of head, from premaxilla to supraoccipital; anterior portion of lower jaw, posterior region of maxilla, antorbital, infraorbitals 4, 5, and 6, supraorbital and all the opercle with scattered melanophores (Fig. 1). Fins mostly hyaline: dorsal fin with distal tip and interradial membrane black; anal fin, occasionally, with black stripe on scales of base and rays generally hyaline, with scattered melanophores on interradial membrane and grouped melanophores on distal tip; adipose fin with scattered melanophores; caudal fin with melanophores grouped on rays, most evident on medial rays and scattered in interradial membrane. Distal tip of caudal fin black; pectoral fin hyaline, with scattered melanophores on interradial membrane and distal tip; pelvic fin with few scattered melanophores.</p><p>Coloration in life. Overall silvery coloration, darker dorsally. Greenish tones on posterior portion of head, in parietal, supraoccipital and opercle. Green metallic chromatophores scattered in scales on flank, between supracleithrum and caudal peduncle most conspicuous on anterior and posterior extremities. Scattered xanthophores on head, mainly in frontal, eyes, infraorbitals 5 and 6 and opercle; scattered xanthophores on scales above the lateral line, most conspicuous in medial portion of body. Scattered melanophores on proximal and distal field of scales, forming five to seven black stripes, inconspicuous, above lateral line. Scattered melanophores in head, conspicuous on lower jaw, maxilla, premaxilla, eyes, antorbital, infraorbitals series, supraorbital, opercle and infraopercle. Fins hyaline, with few scattered melanophores: Dorsal fin hyaline, with rays yellow ground and numerous scattered melanophores. Base of anal fin with diffuse black stripe; anal fin hyaline, with scattered melanophores. Caudal fin hyaline, with yellowish tones on base of interradial membranes and scattered melanophores; medial rays and interradial membrane with scattered melanophores. Adipose fin yellowish, with distal tip hyaline and scattered melanophores. Pectoral fin hyaline, with yellow tones anteriorly and scattered melanophores in all extension. Pelvic fin hyaline, with scattered melanophores (Fig. 2). Small individuals with high concentration of melanophores in dorsal and pectoral fins, and generally, with one black stripe on base of anal fin; in large individuals, fins hyaline, with scattered melanophores and black stripe on base of anal fin inconspicuous or absent.</p><p>Sexual dimorphism and ontogenetic variations. No secondary sexual dimorphism characters were found, including differences in size between females and males.  Triportheus claudiae presents differences in caudal fin form and coloration pattern between adults and juveniles. In small individuals (smaller than 50 mm SL) caudal fin bifurcated, without elongated medial rays; in large individuals, caudal fin truncated, with elongated medial rays (Fig. 3).</p><p>Geographical distribution.  Triportheus claudiae is known to inhabit the floodplain of the upper rio Paraguai basin (Fig. 4).</p><p>Etymology. The specific name, a genitive form for Claudia, is an homage to Maria Claudia de Souza Lima Malabarba, for highly relevant contributions to the knowledge of the  Triportheus species and Neotropical palaeoichthyology. A noun in a genitive case.</p><p>Conservation status.  Triportheus claudiae is a species with wide distribution in the rio Paraguai and tributaries floodplain, with records in Mato Grosso and Mato Grosso do Sul States, Brazil. Despite the Pantanal wetland has suffered a series of impacts of anthropogenic origin in the last years, such as the conversion of native floodplain fields to exotic pasture, large-scale fires, introduction of allochthonous and exotic fish species and mining, we have no evidence that these activities are effectively affecting this species’ population size, distribution, or biology. Additionally, the known distribution of  T. claudiae includes preserved environments, as the Parque Estadual do Rio Negro, Parque Estadual do Encontro das Águas, Parque Nacional do Pantanal Mato-Grossense and several private natural heritage reserves (RPPNs). Using the criteria and categories of International Union for Conservation of Nature (IUCN, 2022),  Triportheus claudiae can be classified as Least Concern (LC).</p><p>Remarks.  Triportheus claudiae was mentioned as  T. aff. rotundatus by Mariguela et al. (2016), based on the examination of specimens from the rio Miranda, an important tributary of left margin of the rio Paraguai. This species is morphologically similar to  T. rotundatus, it is known to inhabit only the rio Amazonas basin, and is genetically closer to  T. nematurus .</p></div>	https://treatment.plazi.org/id/C31687B2FF8D900DBB46F9BDFD3ADDA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lopes, Douglas Alves;Carvalho, Fernando Rogério	Lopes, Douglas Alves, Carvalho, Fernando Rogério (2024): Taxonomy of Triportheus (Ostariophysi: Triportheidae) from the Paraná-Paraguai basin, South America. Neotropical Ichthyology 22 (2): e 230121, DOI: 10.1590/1982-0224-2023-0121, URL: https://doi.org/10.1590/1982-0224-2023-0121
C31687B2FF989012BA7EFCEAFA1BDB76.text	C31687B2FF989012BA7EFCEAFA1BDB76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triportheus Cope 1872	<div><p>Key to identification of  Triportheus from the Paraná-Paraguai basin</p><p>1a. Scales along predorsal line, between tip of supraoccipital and dorsal fin origin, irregularly disposed (not forming a single, continuous line) (Fig. 14A) .................. 2</p><p>1b. Scales along predorsal line, between tip of supraoccipital and dorsal-fin origin, in a single series (a maximum of two scales slightly offset from the main series) (Fig. 14B) ..................................................................................................................... 3</p><p>2a. Perforated lateral-line scales, 33–37 (mode = 34); 38–48 (mode = 43) gill rakers on the lower branch of the first branchial arch........................................  T. nematurus</p><p>2b. Perforated lateral-line scales, 28–32 (mode = 31); 26–32 (mode = 28) gill rakers on the lower branch of the first branchial arch.......................................  T. pantanensis</p><p>3a. Gill rakers on the lower branch of the first branchial arch, 37–44 (mode = 44)....................................................................................................  T. signatus</p><p>3b. Gill rakers on the lower branch of the first branchial arch, 33 or fewer ....  T. claudiae</p></div>	https://treatment.plazi.org/id/C31687B2FF989012BA7EFCEAFA1BDB76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lopes, Douglas Alves;Carvalho, Fernando Rogério	Lopes, Douglas Alves, Carvalho, Fernando Rogério (2024): Taxonomy of Triportheus (Ostariophysi: Triportheidae) from the Paraná-Paraguai basin, South America. Neotropical Ichthyology 22 (2): e 230121, DOI: 10.1590/1982-0224-2023-0121, URL: https://doi.org/10.1590/1982-0224-2023-0121
C31687B2FF87900BBAB9FBCBFA1DDDDB.text	C31687B2FF87900BBAB9FBCBFA1DDDDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triportheus nematurus (Kner 1858)	<div><p>Triportheus nematurus (Kner, 1858)</p><p>(Figs. 5–7; Tab. 2)</p><p>Chalcinus nematurus Kner, 1858:163 [original description]. — Kner, 1860:13–15 [more detailed description and type-locality indicated as: “Cujaba, Suaguragua, Caiçara”]. — Garman, 1890:3 [synonym of Chalcinus angulatus angulatus]. — Eigenmann, Eigenmann, 1891:56 [synonym of Chalcinus angulatus]. — Ulrey, 1895:294 [synonym of Chalcinus angulatus]. — Miranda Ribeiro, 1941:162–69 [morphometric data, synonym of Chalcinus angulatus]. — Fowler, 1950:356 [synonym of Triportheus angulatus angulatus].</p><p>Chalcinus paranensis Günther, 1874:454 [description, type-locality: Rio Paraná]. — Garman, 1890:4 [listed]. — Eigenmann, Eigenmann, 1891:56 [listed: río de La Plata system]. — Travassos, 1940:720 [listed and notes about parasitism]. — Miranda Ribeiro, 1941:171 [diagnosis and comments about morphology].</p><p>Triportheus paranensis. — Fowler, 1950:358 [new combination, listed: rios Paraná-Paraguai basin]. — Géry, 1977:343 [listed and key to identification]. — Portugal, 1990:136 [revision and redescription]. — Fernández, Butí, 1996:252 [listed to Argentina]. — Gómez, Chebez, 1996:48 [listed from Misiones, Argentina]. — Malabarba, 1998:76 [phylogenetic relationships]. — Britski et al., 1999:29 [listing and key to identification]. — Lima et al., 2003:58 [listed]. — López et al., 2003:19 [listed from Argentina]. — Menni, 2004:74. — Malabarba, 2004:182 [synonym of Triportheus nematurus]. — Britski, 2007:44 [listed and key to identification].</p><p>Triportheus nematurus. — Cope, 1878:692 [listed]. — Fowler, 1906:448 [synonym of Chalcinus angulatus]. — Fowler, 1950:356 [synonym of Triportheus angulatus angulatus]. — Portugal, 1990:166–72 [revalidation and redescription, name erroneously attributed to an undescribed species]. — Malabarba, 1998:76 [phylogenetic relationships]. — Britski et al., 1999:29 [listing and key to identification]. — Lima et al., 2003:158 [listed]. — Malabarba, 2004:182 [redescription, lectotype designation; locality of lectotype: “Cujaba”]. — Britski et al., 2007:44 [listed and key to identification]. — Buckup et al., 2007:44 [listed from Brazil]. — Langeani et al., 2007:185 [listed as an allochthonous species in upper rio Paraná basin]. — Graça, Pavanelli, 2007:75 [listed from the upper rio Paraná floodplain]. — Mirande, 2010:485 [phylogenetic relationships]. —Nakagawa, 2011:44 [comments about distribution, identification and study of cephalic musculature]. — Oliveira et al., 2011:13 [phylogenetic relationships and new propose to Triportheidae]. — Litz, Koerber, 2014:10–11 [listed from Uruguay]. — Mariguela et al., 2016:132 [phylogenetic relationships]. — Ota et al., 2018:54 [listed from upper rio Paraná floodplain]. — Reis et al., 2020:463 [listed from Paraná State, Brazil]. — Lopes et al., 2022:247 [illustrated guide and key to identification]. — Brandão et al., 2022:57 [listed from the lower rio Taquari, Paraná State, Brazil]. — Toledo-Piza et al., 2024:436–37 [list]. — Dagosta et al., 2024:69–70, 81 [list and discussion about it is origin in the upper rio Paraná basin].</p><p>Diagnosis.  Triportheus nematurus differs from its congeners by the following combination of characters: two non-elongated scales between the insertion of the pectoral fin and the ventral keel (vs. one elongate scale between the origin of the pectoral fin and the ventral keel in  T. albus,  T. auritus,  T. brachipomus,  T. culter, and  T. magdalenae); 38–48 (mode = 43, n = 23) gill rakers on the lower branch of the first branchial arch (vs. 29–37 in  T. angulatus, 24–32 in  T. rotundatus, 52–57 in  T. guentheri, 22–24 in  T. pictus, 23–28 in  T. curtus, 26–32 in  T. pantanensis, 24–28 in  T. orinocensis, 27–33 in  T. venezuelensis and 24–29 in  T. claudiae). It further differs by the number of perforated scales on the lateral line, 33–37 (mode = 34, n = 62) (vs. 30–33 [mean = 32.1] in  T. guentheri, 30–32 in  T. pictus, 28–32 in  T. pantanensis); irregularly disposed scales on the predorsal line, never in one aligned series (vs. regularly disposed scales of the predorsal line, in one aligned series, in  T. signatus,  T. trifurcatus,  T. pictus, and  T. claudiae); number of scales on the transversal line between the origin of the dorsal fin and the lateral line, six (vs. five in  T. trifurcatus,  T. guentheri, and  T. curtus); 36–37 epineurals (vs. 34–35 in  T. pantanensis), 21 epipleurals (vs. 16–17 in  T. pantanensis, and 22–23 in  T. signatus); number of supraneurals, 9 (vs. 10 in  T. pantanensis and 10–11 in  T. claudiae); and 4–5 ventral tubules in the laterosensory canal of the preoperculum (vs. 7 in  T. pantanensis and  T. claudiae).</p><p>Description. Morphometric and meristic data in Tab. 2. Other characters and color in alcohol in Malabarba (2004). Total vertebrae, 36; precaudal vertebrae, 17. Fifteenth precaudal vertebrae with a hemal arch and canal; caudal vertebrae, 18. Epineurals: 35(2), 36(3) or 37(2), first three unforked; epipleurals 21(5) or 22(2). First epipleural unforked, second to 12 th forked proximally and 13 th to 21 st, unforked. Supraneurals: nine (7). Upper procurrent rays, eight (2) or nine (5); lower procurrent rays, seven (7).</p><p>Coloration in life. Overall coloration silvery, darker dorsally than ventrally. Dorsolateral region of body greenish. Grouped melanophores on proximal field of scales, forming one to six black longitudinal fainted stripes on dorsolateral region of body, above lateral line, converging to one single stripe, which continues on medial caudal fin rays. Fins mostly hyaline, with few scattered melanophores; origin of pectoral fin, base of first five pectoral fin rays and distal tip of pectoral fin with scattered melanophores; grouped melanophores only on base of pectoral fin and first unbranched ray. Distal tip of dorsal and anal fin with scattered melanophores. Last rays of anal fin with scattered melanophores in all extension. Caudal fin with vertical inconspicuous bands of scattered melanophores, barely visible without stereomicroscope; medial caudal fin rays black in all extension, by presence of closely grouped melanophores (Fig. 7). Juveniles with large concentration of grouped melanophores in all fins; in adults, fins hyaline with few scattered melanophores.</p><p>Sexual dimorphism and ontogenetic variations. No characteristics of secondary sexual dimorphism were found, including variations of size. Variations in caudal fin form and coloration between large and small individuals. In small individuals, less than 50 mm SL, caudal fin bifurcated, without elongated medial rays; in large individuals, caudal fin truncated, and medial rays elongated, largest than upper and lower caudal fin rays.</p><p>Geographical distribution.  Triportheus nematurus is widely distributed in lowlands of Paraná-Paraguai basin (Fig. 8). Its natural occurrence known is from the rio Paraguai and lower rio Paraná basins. In the upper rio Paraná basin,  T. nematurus is recognized as a possible introduced species (Dagosta et al., 2024).</p><p>Ecological notes.  Triportheus nematurus is a generalist species and occurs in lentic environments, such as ponds, lakes, bays, and impoundments (Polaz et al., 2014; Severo-Neto et al., 2015), and in lotic environments, such as rivers (Súarez et al., 2013; Polaz et al., 2014). When juveniles, they are associated with beds of aquatic macrophytes (Súarez et al., 2013). The species is omnivorous, its diet based mainly on terrestrial insects, algae, zooplankton, and terrestrial plant matter (including seeds and fruit). The species presents a high feeding plasticity, evidenced mainly by the temporal variations of its diet (Novakowski et al., 2008; Lopes et al., 2017). In the Pantanal wetlands, they can act as seed dispersers, due to their frugivorous interaction during the flood season (Yule et al., 2016). There is no information about the reproductive biology of this species.</p><p>Remarks.  Chalcinus nematurus (=  Triportheus nematurus) Kner, 1858 was described based on three lots (syntypes) with locality not mentioned in the original description. Posteriorly, Kner (1860) mentioned the localities as: “Cujaba, Suaguragua, Caicara”, both in Mato Grosso State, collected by J. Natterer.  Chalcinus paranensis (=  Triportheus nematurus) Günther, 1874 was described based on a unique individual that corresponded to the holotype, collected in a locality defined simply as “from the river Parana ” and probably refers to an exemplar sampled in the lower rio Paraná basin. Malabarba (2004) recognized  T. nematurus as senior synonym of  T. paranensis and designated a lectotype (NMW 69034) and the type-locality was fixed as Cuiabá. In the upper rio Paraná basin, the occurrence of this species was historically attributed to the construction of the Itaipu reservoir in the beginning of the 1980’s, which flooded and consequently eliminated the Sete Quedas waterfalls, the biogeographic divisor between the upper rio Paraná basin and the lower rio Paraná basin (Langeani et al., 2007; Graça, Pavanelli, 2007; Petesse, Petrere, 2012; Ota et al., 2018; Garcia et al., 2018; Pelicice et al., 2018; Jarduli et al., 2019). Dagosta et al. (2024) consider  T. nematurus as non-native species in the upper rio Paraná basin, however, recognize that a molecular approach is necessary to investigate its occurrence within both basins.</p><p>Material examined.   Brazil: Goiás State: upper  rio Paraná basin: MCP 47744, 3, 133.5– 168.7 mm SL.  NUP 8705, 1, 175.8 mm SL.  Mato Grosso State: upper  rio Paraguai basin: MCP 15630, 1, 80.6 mm SL.  MCP 44032, 2, 130.9– 153.5 mm SL. MZUSP 19825, 1, 125.1 mm SL. MZUSP 20440, 1, 125.4 mm SL. MZUSP 28100, 1, 137.7 mm SL. MZUSP 79140, 1, 80.5 mm SL. NUP 1974, 1, 125.2 mm SL. NUP 2935, 1, 220.0 mm SL. NUP 6954, 2, 124.8– 128.7 mm SL. NUP 6994, 1, 147.7 mm SL. NUP 7056, 2, 81.1–84.0 mm SL. NUP 7100, 2, 72.6–113.0 mm SL. NUP 7595, 2, 72.5–96.9 mm SL. NUP 11492, 1, 137.9 mm SL. UFRGS 13453, 2, 56.1–76.8 mm SL. ZUFMS 3274, 7, 39.1–68.9 mm SL. ZUFMS 3438, 1, 41.8 mm SL.  Mato Grosso do Sul State: upper  rio Paraguai basin: CPUEMS not catalogued p.18, 3, 31.9–38.2 mm SL.  CPUEMS not catalogued p.42, 1, 80.1 mm SL. CPUEMS not catalogued p.47, 1, 102.5 mm SL. CPUEMS not catalogued p.89, 3, 61.1–74.6 mm SL. CPUEMS not catalogued p.90, 1, 81.0 mm SL. CPUEMS not catalogued p.92, 4, 75.8–99.6 mm SL. CPUEMS not catalogued p.99, 6, 60.6–70.1 mm SL. CPUEMS not catalogued p.102, 3, 65.5–80.9 mm SL. CPUEMS not catalogued p.110, 1, 98.5 mm SL. DZSJRP 20143, 2, 97.4 mm SL. MZUSP 19740, 1, 134.8 mm SL. NUP 12548, 1, 95.4 mm SL. NUP 12549, 1, 66.5 mm SL. NUP 14220, 2, 132.6– 138.6 mm SL. NUP 14268, 1, 73.5 mm SL. NUP 19898, 1, 140.7 mm SL. NUP 19978, 1, 149.4 mm SL. UFRGS 17443, 2, 56.1–76.8 mm SL. ZUFMS 814, 1, 89.8 mm SL. ZUFMS 856, 3, 74.0– 135.3 mm SL. ZUFMS 3208, 1, 153.3 mm SL. ZUFMS 3209, 1, 126.6 mm SL. ZUFMS 3259, 4, 117.2– 154.2 mm SL. ZUFMS 3268, 5, 108.3– 146.3 mm SL. ZUFMS 3661, 1, 150.8 mm SL. ZUFMS 3704, 11, 74.6–130.2 mm SL. ZUFMS 4857, 3, 119.1– 125.2 mm SL. ZUFMS, 5031, 1, 81.9 mm SL. ZUFMS 5056, 12, 90.0– 194.3 mm SL. ZUFMS 5184, 1, 75.3 mm SL. ZUFMS 5263, 9 (4 c&amp;s), 37.3–59.8 mm SL. ZUFMS 5447, 1, 50.1 mm SL. ZUFMS 5715, 5, 31.8–143.9 mm SL. ZUFMS 5744, 4, 67.9–82.3 mm SL. ZUFMS 5756, 2, 90.3–93.4 mm SL. ZUFMS 5770, 8, 77.0– 102.6 mm SL.  Minas Gerais State: upper  rio Paraná basin: MCP 44034, 1, 153.7 mm SL.  NUP 16876, 2, 48.24–75.4 mm SL. Paraná State: upper and lower rio Paraná basin: MCP 40735, 2, 142.8– 153.6 mm SL. MCP 40765, 2, 189.36– 192.1 mm SL. MZUSP 19854, 1, 171 mm SL. MZUSP 21086, 1, 187.1 mm SL. MZUSP 37181, 1, 134.3 mm SL. NUP 97, 1, 62.5 mm SL. NUP 516, 1, 180.8 mm SL. NUP 1961, 2, 61.2–62.8 mm SL. NUP 7058, 1, 183.6 mm SL. NUP 7146, 2, 185.6– 211.1 mm SL.  São Paulo State: upper  rio Paraná basin: DZSJRP 621, 1, 83.4 mm SL.  DZSJRP 624, 2, 92.7–97.1 mm SL. DZSJRP 4298, 2, 64.9–89.3 mm SL. DZSJRP 10591, 2, 76.9–95.3 mm SL. DZSJRP 13077, 2, 71.9–93.5 mm SL. DZSJRP 13112, 2, 63.9–100.4 mm SL. DZSJRP 13132, 2, 96.0– 102.8 mm SL. ZUFMS 5362, 3, 145.0– 163.6 mm SL.  Paraguay: Alto Paraguay Departament:  Río Paraguay basin: MZUSP 54148, 1, 74.4 mm SL;  MZUSP 54149, 2, 74.3–87.7 mm SL.</p></div>	https://treatment.plazi.org/id/C31687B2FF87900BBAB9FBCBFA1DDDDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lopes, Douglas Alves;Carvalho, Fernando Rogério	Lopes, Douglas Alves, Carvalho, Fernando Rogério (2024): Taxonomy of Triportheus (Ostariophysi: Triportheidae) from the Paraná-Paraguai basin, South America. Neotropical Ichthyology 22 (2): e 230121, DOI: 10.1590/1982-0224-2023-0121, URL: https://doi.org/10.1590/1982-0224-2023-0121
C31687B2FF819017BA92FC1DFC23DCFB.text	C31687B2FF819017BA92FC1DFC23DCFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triportheus pantanensis Malabarba 2004	<div><p>Triportheus pantanensis Malabarba, 2004</p><p>(Figs. 9–10; Tab. 3)</p><p>Triportheus pantanensis Malabarba, 2004:197 [original description; type-locality: Transpantaneira road, pond below bridge, 70 km South of Poconé, Mato Grosso, Brazil]. — Buckup et al., 2007:44 [listed from Brazil]. — Mirande, 2010:485 [phylogenetic relationships]. —Nakagawa, 2011:49 [comments about distribution, taxonomy, and study of cephalic musculature]. — Mariguela et al., 2016:134 [phylogenetic relationships]. — Koerber et al., 2017:27 [list from Paraguay]. — Lopes et al., 2022:247 [illustrated guide and key to identification]. — Toledo-Piza et al., 2024:437 [list].</p><p>Triportheus nematurus (non Kner 1858). — Portugal, 1990:166 [misidentification, redescription]. — Malabarba, 1998:76 [phylogenetic relationships within  Triportheus and  Lignobrycon]. — Britski et al., 1999:29 [listed and key to identification]. — Lima et al., 2003:158 [listed]. — Britski et al., 2007:44 [listed and key to identification].</p><p>Diagnosis.  Triportheus pantanensis differs from its congeners by the following combination of characters: two non-elongated scales between the insertion of the pectoral fin and ventral keel (vs. one elongated scale between the origin of the pectoral fin and ventral keel in  T. albus,  T. auritus,  T. brachipomus,  T. culter, and  T. magdalenae); 26–33 (mode = 28, n = 19) gill rakers on the lower branch of first branchial arch (vs. 40–47 in  T. trifurcatus, 38–48 in  T. nematurus, 52–57 in  T. guentheri, 22–24 in  T. pictus, 35–44 in  T. signatus); perforated scales on the lateral line: 28–32 (mode = 31, n = 29) (vs. 34–37 in  T. angulatus, 33–37 in  T. nematurus, 34–37 in  T. signatus, 34–39 in  T. orinocensis, and 33–36 in  T. venezuelensis); three rows of cuspid teeth on the premaxilla (vs. two rows of cuspid teeth in  T. curtus and  T. pictus); scales of the predorsal series irregularly disposed (vs. scales of the predorsal series regularly disposed in  T. trifurcatus,  T. signatus,  T. pictus, and  T. claudiae); six scale series on the lateral of caudal peduncle (vs. five in  T. angulatus,  T. curtus, and  T. orinocensis, four or five in  T. guentheri,  T. rotundatus,  T. trifurcatus, and  T. venezuelensis); epineural bones, 34–35 (vs. 36–37 in  T. nematurus); epipleural bones: 16–17 (vs. 21 in  T. nematurus, 22–23 in  T. signatus, and 20–21 in  T. claudiae); number of supraneurals: 10 (vs. nine in  T. nematurus and  T. signatus); seven ventral tubules in the laterossensorial canal of the preopercule (vs. 4–5 in  T. nematurus and 5 in  T. signatus).</p><p>Description. Morphometric and meristic data in Tab. 3. Other characters and color in alcohol in Malabarba (2004). Total vertebrae: 36(1) or 37(3); precaudal vertebrae: 17(4). Caudal vertebrae: 18(1) or 19(3); 34(2) or 35(2) epineural bones; first to 25 th or 26 th epineural bones branched in ventral region; 27 th to 34 th or 35 th epineural bone unbranched; epipleural bones 18(3) or 19(1); first epipleural unbranched; second to nineth epipleural branched in dorsal portion; 10 th to 16 th or 17 th epipleural unbranched. Supraneurals: 10(4). Upper procurrent rays: 8(1) or 9(3); lower procurrent rays: six (4).</p><p>Coloration in life. Overall coloration silvery, darker dorsally. Dorsal-middle region of body with greenish or yellowish tones. Scattered or concentrated melanophores on proximal field of scales, forming five or six longitudinal black fainted stripes on dorsolateral portion of body, above lateral line. Fins mostly hyaline, with scattered melanophores. First five pectoral-fin rays darker, with concentrated melanophores. Medial rays of caudal fin, black in all extension. Some individuals with conspicuous melanophores in first dorsal and anal fin rays (Fig. 10). Small individuals with high concentration of melanophores in dorsal and pectoral fins and a black stripe on base of anal fin; in adults, fins hyaline, with scattered chromatophores and black stripe on base of anal-fin inconspicuous or absent.</p><p>Sexual dimorphism and ontogenetic variations. No characteristics of secondary sexual dimorphism were found in  Triportheus pantanensis, including variations in size. This species presents differences in caudal fin form and coloration pattern between large and small individuals. In small individuals (smaller than 50 mm SL) the caudal fin is bifurcated, without elongated medial rays; in adults’ caudal fin is emarginated, with elongated medial rays.</p><p>Geographical distribution.  Triportheus pantanensis presents a wide distribution within the floodplain of the rio Paraguai basin (Fig. 11) and in the lower rio Paraná basin, including rivers from Argentina (Mirande, Koerber, 2015) and Paraguay (Koerber et al., 2017).</p><p>Ecological notes.  Triportheus pantanensis occurs in lentic and lotic environments (Súarez et al., 2013; Polaz et al., 2014; Severo-Neto et al., 2015), including reservoirs (Corrêa et al., 2009). Omnivorous, with tendencies to insectivory or herbivory, depending on the dynamics of the hydrological periods (Corrêa et al., 2009). Despite the issue that its effectiveness as a seed disperser has not been tested, it is a known fact that this species consumes small fruits (Costa-Pereira et al., 2011), therefore we can assume that it possibly plays an interactive biological role in the dispersion of seeds. There is no information on the reproductive biology to this species.</p><p>Material examined.   Brazil: Mato Grosso State:  rio Paraguai basin: MCP 35006, 1, 97.1 mm SL, holotype.  MCP 10733, 3, 88.3–88.7 mm SL, paratypes. MCP 10751, 3, 77.5–78.3 mm SL, paratypes. MCP 15741, 1, 88.5 mm SL, paratype. MCP 35824, 4, 65.9–78.5 mm SL, paratypes. MCP 11119, 1, 99.1 mm SL. MCP 38829, 1, 58.8 mm SL .  MZUSP 38106, 2, 40.4–59.1 mm SL. NUP 888, 2, 112.4– 116.7 mm SL. NUP 3136, 3, 87.8–105.1 mm SL. NUP 6945, 1, 84.7 mm SL. NUP 7053, 1, 76.3 mm SL.  Mato Grosso do Sul State:  rio Paraguai basin: CITL 1136, 4 (c&amp;s) of 17, 52.5–69.1 mm SL.  CPUEMS not catalogued p.24, 1, 41.8 mm SL. CPUEMS not catalogued p.84, 2, 42.5– 44.0 mm SL. CPUEMS not catalogued p.89, 1, 64.1 mm SL. CPUEMS not catalogued p.90, 2, 63.1–70.2 mm SL. MZUEL 12268, 2, 68.3–74.7 mm SL. MZUSP 49968, 1, 50.2 mm SL. MZUSP 59827, 2, 46.7–56.6 mm SL. NUP 12552, 2, 18.9–31.0 mm SL. ZUFMS 857, 1, 51.2 mm SL. ZUFMS 858, 1, 58.9 mm SL. ZUFMS 859, 1, 75.2 mm SL. ZUFMS 880, 2, 45.1–51.0 mm SL. ZUFMS 1055, 8, 68.2–94.3 mm SL. ZUFMS 1525, 3, 65.5–65.8 mm SL. ZUFMS 3723, 3, 84.9–95.5 mm SL. ZUFMS 3886, 7, 67.7– 86.9 mm SL. ZUFMS 5716, 5, 65.6–103.4 mm SL. ZUFMS 5729, 2, 62.3–64.2 mm SL.  Paraguay: Alto Paraguay Department:  Río Paraguay basin: MZUSP 54144, 1, 58.6 mm SL.  MZUSP 54146, 2, 52.9–55.0 mm SL. MZUSP 54147, 2, 60.4–66.9 mm SL. MZUSP 54148, 2, 64.2–70.8 mm SL.</p></div>	https://treatment.plazi.org/id/C31687B2FF819017BA92FC1DFC23DCFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lopes, Douglas Alves;Carvalho, Fernando Rogério	Lopes, Douglas Alves, Carvalho, Fernando Rogério (2024): Taxonomy of Triportheus (Ostariophysi: Triportheidae) from the Paraná-Paraguai basin, South America. Neotropical Ichthyology 22 (2): e 230121, DOI: 10.1590/1982-0224-2023-0121, URL: https://doi.org/10.1590/1982-0224-2023-0121
C31687B2FF9C9012BABBFF6DFB80DD46.text	C31687B2FF9C9012BABBFF6DFB80DD46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triportheus signatus (Garman 1890)	<div><p>Triportheus signatus (Garman, 1890)</p><p>(Fig. 12-13; Tab. 4)</p><p>Chalcinus angulatus signatus Garman, 1890:4 [original description; type-locality: rio Puty, Therezina, Brazil (rio Potí, tributary of the rio Parnaíba, Terezina municipality, Piauí State, Brazil)]. — Eigenmann, Eigenmann, 1891:58 [listed]. — Miranda Ribeiro, 1941:171 [keys to identification and comments regarding subspecies of  C. angulatus].</p><p>Triportheus angulatus signatus . — Fowler, 1941:194 [listed as synonym of  Triportheus angulatus]. — Fowler, 1950:357 [redescription, type-locality: “ Brazil, rio Itapicura”].</p><p>Triportheus signatus . — Portugal, 1990:159–65 [redescription, illustration, and distribution]. — Lima et al., 2003:158 [list]. — Malabarba, 2004:193–94 [redescription and distribution]. — Buckup et al., 2007:44 [listed from Brazil]. —Nakagawa, 2011:57–61 [comments regarding distribution and morphology]. — Ramos et al., 2014:4 [list from rio Parnaíba basin, Brazil]. — Mariguela et al., 2016:132 [phylogenetic relationships]. — Melo et al., 2016:376 [list from lower rio Parnaíba basin, with discussions regarding occurrence of the species in Northeastern Brazil]. — Reis et al., 2020:463 [list from Paraná State, Brazil, invasive within upper rio Paraná basin]. — Silva et al., 2020:9 [list from Bahia State, Brazil]. — Koerber et al., 2022:32 [list from Maranhão State, Brazil]. — Toledo-Piza et al., 2024:437 [list]. — Dagosta et al., 2024:81 [listed as introduced in the upper rio Paraná basin].</p><p>Diagnosis.  Triportheus signatus differs from its congeners by the following combination of characters: two normal scales between the insertion of the pectoral fin and the ventral keel (vs. one elongated scale between the origin of the pectoral fin and the ventral keel in  T. albus,  T. auritus,  T. brachipomus,  T. culter, and  T. magdalenae); 35–44 gill rakers (mode = 39, n = 9) on the lower branch of the first branchial arch (vs. 24–32 gill rakers in  T. rotundatus, 52–57 in  T. guentheri, 22–24 in  T. pictus, 23–28 in  T. curtus, 26–33 in  T. pantanensis, 24–28 in  T. orinocensis, 27–33 in  T. venezuelensis, and 24–29 in  T. claudiae); perforated scales on the lateral line: 34–38 (mode = 36, n = 12) (vs. 30–33 in  T. guentheri, 30–32 in  T. pictus and 28–32 in  T. pantanensis); scales distributed in a regular series on predorsal line (vs. predorsal scales disposed irregularly, not in series, in  T. angulatus,  T. rotundatus,  T. nematurus,  T. guentheri,  T. curtus,  T. pantanensis,  T. orinocensis, and  T. venezuelensis); six scales above the lateral line (vs. 5 scales above the lateral line in  T. trifurcatus); 22–23 epipleural bones (vs. 21 in  T. nematurus, 16–17 in  T. pantanensis, and 20–21 in  T. claudiae), 5 ventral tubules in laterossensorial canal of the preopercule (vs. 7 in  T. pantanensis and  T. claudiae).</p><p>Description. Morphometric and meristic data in Tab. 4. Other characters and color in alcohol in Malabarba (2004). Total vertebrae: 36(3) or 37(1); precaudal vertebrae: 17(4). Hemal arch and canal in the 15 th or 16 th caudal vertebrae. Caudal vertebrae, 19(3) or 20(1); 37(1) or 38(3) epineural bones; first to 22 nd epineural bones branched in ventral region; epipleural bones 22(3) or 23(1); first epipleural unbranched; second to nineth epipleural branched in dorsal portion; 10 th to 16 th, or 17 th epipleural unbranched. Number of supraneurals: nine (4). Upper procurrent rays: 9(1) or 10(3); lower procurrent rays: eight (4).</p><p>Coloration in life. Overall silvery coloration, darker dorsally. Dorsal-middle region of body with greenish or yellowish tones. Scattered melanophores on dorsal portion of head, as in premaxilla, frontal, supraorbital, infraorbitals 4, 5 and 6, parietal and opercle. Melanophores concentrated in proximal and central field of scales, forming four to seven longitudinal fainted black longitudinal stripes on dorsolateral portion of body, above lateral line. Lateral stripes converge on caudal peduncle as a single large stripe, which follow medial rays of caudal fin, black in all extension.</p><p>Sexual dimorphism and ontogenetic variations. No secondary sexual dimorphism characters were found in  Triportheus signatus, including variation in size of females and males. Species present differences in caudal fin form in small and large specimens, similar to identified to all the other  Triportheus species of rios Paraná-Paraguai basin. In small individuals (smaller than 50 mm), caudal fin bifurcated, without elongated medial rays; in large individuals, caudal fin truncated, with elongated medial rays.</p><p>Geographical distribution.  Chalcinus angulatus signatus (=  Triportheus signatus) was described by Garman (1890) with the type-locality in “rio Puty, Therezina”, refering to the rio Potí, a tributary from the rio Parnaíba, in Teresina city, Piauí State, Brazil. The species occurs naturally within the rio Parnaíba basin and other coastal river basins from the Northeast region of Brazil.  Triportheus signatus has also been identified within the upper rio Paraná basin tributaries, mainly in the rio Tietê basin (Fig. 13) since 1990 and it is an invasive species in this river basin.</p><p>Ecological notes. Data about the natural history of  Triportheus signatus are scarce. According to Höfling et al. (2000), the species is generalist, mainly found in environments associated to aquatic macrophytes, with high heterogeneity of habitats. In situations of hypoxia on their natural distribution, they develop lip extensions, called as barbels, which aid surface respiration (Barros-Neto et al., 2019). However, none of the examined specimens in the rio Paraná basin presented barbels. In the upper rio Paraná basin, it occurs mainly in reservoirs with the rios Tietê and Paranapanema basins (Höfling et al., 2000; Nakagawa, 2011; Mariguela et al., 2016). It has an omnivorous or insectivorous diet and feeds on larvae and pupae of aquatic insects, fragments of terrestrial insects, crustaceans, seeds, and fragments of superior terrestrial plants (Höfling et al., 2000; Mendes et al., 2011; Oliveira et al., 2019). According to Höfling et al. (2000), this species is most abundant in the hot and rainy season, between  October and April, when they reproduce.</p><p>Remarks.  Triportheus signatus occurs naturally in the rio Parnaíba basin and other coastal river basins of the Northeast region of Brazil. The occurrence of this species in the upper rio Paraná basin is probably a consequence of an introduction, which is supported by the sampling locations, which are always near to hydroelectric reservoirs, and by the restocking program conducted by energy company of São Paulo State, which begin in 1974 and includes the silver croaker  Plagioscion squamosissimus (Heckel, 1840) as well other species considered as prey for the establishment of this predator (e.g.,  Triportheus signatus and two species of freshwater shrimp) (Queiroz-Sousa et al., 2018). The first citations of this species in the Tietê and Paranapanema basins were made by Portugal (1990) in their unpublished master’s thesis. Published records of  T. signatus in the upper rio Paraná basin date from the early 2000ʼs (Silvano, Begossi, 2001; Buckup et al., 2007). However, specimens deposited at the DZSJRP and MZUSP indicate that the introduction of this species is even older and corroborates that this species was introduced in the region in 1970’s–1980’s (e.g., MZUSP 37221, collected in 1987; DZSJRP 625, collected in 1988, both in rio Tietê). Currently,  T. signatus is known to inhabit the upper rio Paraná basin within the rios Tietê, Paranapanema and the “Canal de Pereira Barreto”, which connects the rios Tietê and São José dos Dourados. The lots of  T. signatus which were collected in the 1980ʼs and 1990ʼs in the upper rio Paraná basin (e.g., DZSJRP 625, DZSJRP 3248, and MZUSP 37221) were mistakenly identified as  T. angulatus,  T. paranensis (=  T. nematurus),  T. nematurus, or just  Triportheus sp.</p><p>Material examined. Brazil: Ceará State:  rio Salgado: MNRJ 44325, 2, 73.7– 90.7 mm SL.  Paraíba State: UFRGS 20345, 2, 138.5–144.0 mm SL.  Paraná State:  rio Paranapanema basin: MZUEL 2485, 2, 144.1– 174.2 mm SL.  MZUEL 2864, 1, 88.0 mm SL.  Piauí State:  rio Parnaíba basin: MNRJ 29130, 2, 58.2–60.8 mm SL.  MNRJ 43204, 2, 61.6–66.4 mm SL.  São Paulo State:  rio Tietê basin: CITL 423, 13 (4 c&amp;s), 49.2–57.8 mm SL.  DZSJRP 625, 1, 122.7 mm SL. DZSJRP 3248, 1, 103.0 mm SL. DZSJRP 17731, 2, 100.8– 117.3 mm SL. MZUEL 5667, 1, 133.1 mm SL. MZUSP 37221, 9, 135.3– 157.8 mm SL. MZUSP 53890, 3, 84.4–148.5 mm SL. NUP 2678, 1, 195.3 mm SL. NUP 16530, 2, 175.0– 186.8 mm SL.</p></div>	https://treatment.plazi.org/id/C31687B2FF9C9012BABBFF6DFB80DD46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lopes, Douglas Alves;Carvalho, Fernando Rogério	Lopes, Douglas Alves, Carvalho, Fernando Rogério (2024): Taxonomy of Triportheus (Ostariophysi: Triportheidae) from the Paraná-Paraguai basin, South America. Neotropical Ichthyology 22 (2): e 230121, DOI: 10.1590/1982-0224-2023-0121, URL: https://doi.org/10.1590/1982-0224-2023-0121
