identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BE25B46BFF98FF846ADCFACB02C6ACDC.text	BE25B46BFF98FF846ADCFACB02C6ACDC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaneropterinae	<div><p>Subfamily Phaneropterinae</p><p>Comments. This subfamily currently comprises 49 tribes, 384 genera, and nearly 3000 valid species worldwide, with the greatest diversity occurring in tropical regions (Cigliano et al. 2025). Members of the Phaneropterinae, commonly known as bush katydids, range from tiny species, such as those in the tribes Phaneropterini or Pycnopalpini, to large-bodied forms found in tribes like Steirodontini or Vossiini (Bey-Bienko 1954, Nickle 1992, Cohn et al. 2014, Cadena-Castañeda 2014a, 2015a, d, 2016).</p><p>They are characterized by a generally globose head, although some tribes, such as Dysoniini and Aegimiini, exhibit a pronounced vertex (Nickle 1992, Cadena-Castañeda 2011, Dias et al. 2012). The lateral lobes of the pronotum almost entirely cover the auditory spiracle, which is elliptical. The first tarsomere is cylindrical and not laterally sulcate; the prosternum is unarmed, and the tympanum is generally exposed on both sides of the fore tibia, although in some taxa it may be covered by cuticle, leaving only a slit open through which sound can reach the tympanic organ. The female ovipositor varies greatly, adapting to the oviposition requirements of each species, so no single ovipositor morphology characterizes the subfamily (Brunner von Wattenwyl 1878, Bey-Bienko 1954, Ragge 1980, Nickle 1992). Phaneropterine eggs are typically flattened and ovoid (although in some rare cases they may be asymmetrical), which serves as a diagnostic trait for the subfamily (Heller &amp; Hemp 2024). Most species are green, but yellowish, brown, pink (usually due to erythrism), white, and black individuals may also occur. It is common for species to blend into vegetation, while others resemble lichens and bryophytes, or in some cases, mimic ants in their nymphal stages and wasps in adulthood (Brunner von Wattenwyl 1878, Leroy 1985, Nickle 1992, Cadena-Castañeda et al. 2022).</p><p>Phaneropterines are unique among Tettigoniidae in that females produce response signals to the conspecific male calling song. To produce these sounds, females possess a structure analogous to the male stridulatory file, consisting of a complex of rigid, recurved spines on the anal area of the tegmina (Nickle &amp; Carlysle 1975). Males respond to the female reply by actively moving toward the sound source, rather than calling from a fixed location (Heller &amp; Helversen 1986, Spooner 1995). Male calls are diverse and involve remarkable acoustic adaptations in the stridulatory apparatus (Heller &amp; Hemp 2014). Some species produce among the most complex songs known in insects (Nickle 1984, Walker 2004). In many cases, species are morphologically very similar, making acoustic signals an essential tool for species identification (Leroy 1970, Walker et al. 2003, Heller et al. 2015).</p><p>As evidenced above, the Phaneropterinae are highly diverse in multiple aspects and are clearly distinguishable from other subfamilies of Tettigoniidae . However, due to the high number of taxa, the systematics and delimitation of this group remain in progress. This contribution focuses on the classification and delineation of short-winged tribes and other related groups, to resolve longstanding classification challenges and clarify the identity of traditional and derivative taxa.</p><p>Key for the identification of short-winged Phaneropterinae tribes derived from Odonturini</p><p>1. Male terminalia generally without modifications in the last abdominal segments, epiproct, cerci, and subgenital plate. Male genitalia entirely membranous, titillator (ti) membranous or poorly defined; apodemes (AP) absent. Female subgenital plate generally triangular, without any medial division......................................................... 2</p><p>– Male terminalia generally modified, with the last abdominal segments bearing projections or divisions; epiproct usually with a denticulated posterior margin or elongated posterior prolongation; cerci variable, simple or sharply serrated at apex, or divided in various ways (Figs. 12F–H, 13F–H, 15C–E, 15J–L, 15P–R). Male genitalia membranous but with a developed, sclerotized titillators’ sclerite (TS), sometimes with marginal denticulations; apodemes’ sclerite (AP) present and well developed (Figs. 12I–J, 13I–J, 15G–H, 15N–O, 15T–U). Female subgenital plate divided into two sclerotized lobes separated by a medial longitudinal membrane (Figs. 14G, M, 15F, M, S). Distributed in North America, from Mexico to the United States ............................................................. Dichopetalini trib. nov.</p><p>2. Body generally slender. Head rounded or subrectangular and mostly narrow; fastigium of the vertex narrower than the antennal scape (with some exceptions such as Anisophya s.s. and Anisophya (Brevianisophya) subgen. nov.). Pronotum with variable length and proportions, posterior margin not conspicuously elevated or swollen, and not markedly covering the base of the tegmina. Female tegmina variable in length but never completely covered by the pronotum. Legs elongated, usually extending significantly beyond the tip of the abdomen. Male cerci generally simple and unmodified, bearing an apical spine, without serrations at the apex ( Caimanellus with bifurcated cerci). Male genitalia with the ventral lobe (vl) of similar length to the dorsal lobe (dl), not projecting conspicuously........................................... 3</p><p>– Body generally robust (Figs. 3, 4), though some species may be slender (Fig. 5). Head rounded and globose in frontal view; fastigium of the vertex almost as wide as or even wider than the antennal scape (Figs. 6A, D, G). Pronotum usually longer than high or wide, with the posterior margin moderately elevated or swollen in males (Figs. 6B, E, H, K); posterior part of the male pronotum often covering the basal portion or half of the tegmina (Figs. 6C, F, I). Female tegmina much shorter than the pronotum or completely hidden beneath it. Legs not elongated and do not project significantly beyond the tip of the abdomen. Male cerci cylindrical and armed with an apical spine or serrations (Figs. 7B, F, J, N, R); in some species, terminalia may be specialized. Male genitalia with the ventral lobe (vl) projecting from the phallic complex, subrectangular, with undulated or slightly divided posterior margin (Figs. 8). Widely distributed across the Palearctic region, Eurasia............................................................................................ Barbitistini</p><p>3. Eyes conspicuous, with a maximum diameter exceeding the length of the antennal scape. Mesosternum quadrangular, almost as wide as high, with rounded lateral margins; metasternum 1.5 to 2 times wider than the mesosternum; sternal grooves short, with apophyseal pits positioned close together. Ovipositor variable in length, with finely serrated margins, except Lorieriella gen. nov., Cohnia, and Caimanellus .................................................... 4</p><p>– Eyes inconspicuous, with a maximum diameter not exceeding the length of the antennal scape (Figs. 1B, 2B). Mesosternum and metasternum rectangular, 4 to 5 times wider than high, with anterior and posterior margins nearly straight; meso- and metasternal grooves narrow and almost entirely covered, apophyseal pits well separated (Figs. 1E, 2E). Ovipositor slender, moderately widened in lateral view, slightly curved, with toothed apex on all margins (Fig. 2F). Western Mediterranean distribution, extending through North Africa, the Iberian Peninsula, Balearic Islands, Sicily, and Sardinia .... Odonturini</p><p>4. Head subrectangular or ovoid in frontal view; fastigium narrower than one-third the width of the scape (Fig. 9B). Pronotum mostly rugose, with well-defined sulci (Fig. 9D). Lateral lobes rectangular, nearly as tall as wide, with anterior, ventral, and posterior margins almost straight (Figs. 9C, 10A). Subgenital plate quadrangular, almost as wide as long, usually with a median notch on the posterior margin (Figs. 9H, M). Male genitalia with upper folds of the ventral lobe (up.vl.) rounded, surrounding but not covering the lower folds (lw.vl.); lw.vl. short and poorly differentiated (Figs. 9I–J, 9N–O). Ovipositor short and lamellate, with slightly serrated or nearly smooth margins. Distributed in East Africa.. Monticolariini trib. nov.</p><p>– Head rounded or ovoid in frontal view; fastigium of the vertex generally narrower than the scape (Figs. 16C, F, K, 18 G, O), except Anisophya and Anisophya (Brevianisophya) subgen. nov. (Fig. 18A). Pronotum smooth and sulci not defined or only faintly outlined (Figs. 16G, 18D, F, L). Lateral lobes variable in height and width, ranging from nearly as tall as wide to much wider than tall, with ventral and posterior margins gradually curved and rounded, lacking a distinct angular junction (Figs. 18C, E, K, M). Subgenital plate rectangular, longer than wide, posterior margin usually with a median notch (Figs. 19B, E, H, O, S). Male genitalia with upper folds of the ventral lobe (up.vl.) rounded, surrounding but not covering the lower folds (lw.vl.); lw.vl. projects medially between up.vl., reaching mid-length of the dorsal lobe (Figs. 19U–X). Ovipositor variable in length, equal to, slightly longer than (Figs. 20D, F), or more than twice the length of the pronotum (Figs. 20B, H); margins finely serrated, except in Lorieriella gen. nov., Cohnia, and Caimanellus (Figs. 20J). Distributed in South America................................................................. Anisophyini trib. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFF98FF846ADCFACB02C6ACDC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFF96FF876ADCFD9D02C6A962.text	BE25B46BFF96FF876ADCFD9D02C6A962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odonturini Brunner von Wattenwyl 1878	<div><p>Odonturini Brunner von Wattenwyl, 1878 n. sensu.</p><p>Redescription. Males slender (Fig. 1A) and females generally robust and stout (Fig. 2A). Head rounded and slender in frontal view; fastigium sulcate, narrower than the width of the scape or pedicel; the fastigium does not exceed the average width of the antennal scape. Eyes rounded and slightly protrusive (Figs. 1B, 2B). Thorax. Pronotum without modifications on any of its margins; lacking marked lateral carinae, though sometimes indicated by different coloration or a slight suggestion of relief, especially posteriorly; lateral lobes longer than high; humeral sinus not developed (Figs. 1C, 2C). Mesosternum and metasternum rectangular, four to five times wider than high, with anterior and posterior margins almost straight; meso- and metasternal grooves narrow and almost entirely covered; margins of the lateral lobes not raised (Figs. 1E, 2E). Wings. Tegmina short in both sexes, not covering the first or second abdominal segment, always fully exposed (Figs. 1D, 2D). Legs long and slender, especially in males; hind femur greatly exceeding the apex of the abdomen; fore coxae unarmed; fore tibiae with open tympana. Abdomen. Tenth tergite without modifications or projections, and posterior margin straight or slightly undulated (Fig. 1F). Males: Epiproct triangular; cerci simple, unmodified, each bearing a small apical spine (Fig. 1G). Subgenital plate rectangular, longer than wide, without modifications or styles, posterior margin usually with a moderately deep notch (Fig. 1H). Genitalia. Entirely membranous; dorsal lobe (dl) simple, rounded, and not exceeding the length of the lateral folds of dorsal lobe (ldl). Titillator’s sclerite (TS) and sclerite of apodemes (AP) absent. Upper folds of ventral lobe (up.vl.) rounded, surrounding the lower folds of ventral lobe (lw.vl.) without covering them; lw.vl. short and poorly differentiated; ejaculatory vesicles (ejv) conspicuous and rounded (Figs. 1I, J). Females: Epiproct triangular; cerci conical and short; ovipositor slender, moderately broadened in lateral view, slightly curved, with the apex toothed on all margins (Fig. 2F). Subgenital plate triangular, without modifications or divisions (Fig. 2G).</p><p>Taxa included. Two genera and 16 species: Odontura Rambur, 1838 (type genus) and Odonturella Bolívar, 1900 stat. rev.</p><p>Distribution. Western Mediterranean. It extends across North Africa, the Iberian Peninsula, the Balearic Islands, Sicily, and Sardinia (Map 1).</p><p>Comments. This tribe had not yet been morphologically differentiated from other short-winged Phaneropterinae taxa, which were traditionally included in Odonturini or Barbitistini . Although studies such as those by Grzywacz et al. (2014, 2018) provide evidence that Odonturini is not monophyletic and define the tribe as monogeneric based on molecular analyses, it remains important to establish a clear morphological distinction for Odonturini . This will help prevent the future inclusion of superficially similar but unrelated taxa.</p><p>It is proposed to elevate Odonturella Bolívar, 1900 stat. rev. to genus status as already done by Kirby 1906, as it presents distinguishing characteristics that fully separate it from Odontura, following the suggestion by Grzywacz et al. (2014, 2018) and Cassar &amp; Massa (2022). The two Odonturella stat. rev. species are found in the central and western regions of the Iberian Peninsula, in contrast to Odontura species, which are distributed across northwestern Africa, the Balearic Islands, Sicily, and Sardinia. Thus, the Strait of Gibraltar and the Mediterranean as a whole have likely acted as a diversification barrier between the two genera.</p><p>The only exception is Odontura glabricauda (Charpentier, 1825), a species recorded in Morocco as well as in Spain and Portugal (Llorente &amp; Pinedo 1990). The specimens used for its description, particularly the only known male, are not in good condition, which led to doubts regarding the species’ validity for a long time (Pinedo 1987). However, Llorente &amp; Pinedo (1990) confirmed its presence in both Africa and Europe, raising an important question: If the African and European specimens indeed belong to a single species, Could O. glabricauda have originated in Africa, like its congeners, and later been introduced to the Iberian Peninsula? If this hypothesis is correct, it would support the proposed differentiation of both genera on opposite sides of the Mediterranean, as previously mentioned.</p><p>Below is a key for distinguishing the genera, based on the characteristics described by Bolívar (1900), Llorente &amp; Pinedo (1990), and Cassar &amp; Massa (2022):</p><p>Key to Odonturini genera</p><p>1. Pronotum short, with a slight trace of lateral carinae on the dorsum; lateral lobes broad and almost always uniformly colored. Male tegmina with weakly developed veins. Ovipositor curved ventrally from about the second third, female cerci three times as long as wide................................................................... Odontura</p><p>– Pronotum long, laterally extended in a roof-like shape; dorsum without traces of lateral carinae; lateral lobes very narrow, with a pale band along the lower margin. Male tegmina with strongly marked veins. Ovipositor curved ventrally throughout its length, female cerci no more than twice as long as wide................................ Odonturella stat. rev.</p></div>	https://treatment.plazi.org/id/BE25B46BFF96FF876ADCFD9D02C6A962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFF95FF986ADCFBCF02C6AE26.text	BE25B46BFF95FF986ADCFBCF02C6AE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Barbististini Jacobson 1905	<div><p>Barbististini Jacobson, 1905</p><p>Redescription. Males and females generally robust (Figs. 3, 4), though some species are slender (Fig. 5). Head rounded and globose in frontal view; fastigium almost as wide (Figs. 6G, J, M) as or even wider than the antennal scape (Figs. 6A, D). Thorax. Pronotum generally longer than it is high or wide, with the posterior margin moderately elevated (Figs. 6B, E, H) or swollen in males (Figs. 6K, N); lateral carinae are faintly marked in some genera, sometimes highlighted by a band of different coloration. Lateral lobes longer than high, and the humeral sinus is undeveloped. Mesosternum rectangular, nearly twice as wide as it is high, with rounded lateral margins; metasternum broader than the mesosternum, with divergent lateral margins; sternal grooves usually broad, visible, and with paired pits at each end, appearing as widely separated apophyses; lower margins of the meso- and metasternal lobes unarmed and elevated (Figs. 7A, E, I, M, Q). Wings. Tegmina short in both sexes, not surpassing half of the second abdominal segment; in males, the posterior section of the pronotum often covers the basal portion or half of tegmina (Figs. 6C, F, I, L, O). Female tegmina are much shorter than the pronotum or even completely hidden by it (Figs. 5B, C). Legs. Slender, not very elongated, and not extending significantly beyond the tip of the abdomen. Fore coxa unarmed; fore tibiae with open tympana. Abdomen. Tenth tergite is generally unmodified, with few exceptions. Males: tenth tergite usually straight, though in some species it may be prolonged (Figs. 7B, F, J, N, R); epiproct rectangular (Figs. 7C, G, K, S); cerci cylindrical and armed with a spine (Figs. 7B, F, J, R) or serrated at the apex (Fig. 7N); some species showing specialized terminalia. Subgenital plate rectangular, wider than long, and lacks styles; it usually curves outward, projecting between the cerci (Figs. 7D, H, L, P, T). Genitalia. Completely membranous; dorsal lobe (dl) simple, rounded, with the apex slightly divided, and shorter than the lateral folds of dorsal lobe (ldl). Titillator’s sclerite (TS) and sclerite of apodemes (AP) absent. ldl generally quadrangular or rectangular shaped. Ventral lobe (vl) projects from the phallic complex, subrectangular, posterior margin undulated or slightly divided in the middle. Upper folds of ventral lobe (up.vl.) rounded or subrectangular and cover the lower folds of ventral lobe (lw.vl.); ejaculatory vesicles (ejv) conspicuous and rounded (Fig. 8). Females: epiproct triangular; cerci short and conical; ovipositor either elongate, with strongly serrate margins (Fig. 4C), or short and lamelliform (Figs. 5B, C), with slightly serrated or almost smooth margins.</p><p>Taxa included. 14 genera, 293 species: Barbitistes Charpentier, 1825 (type genus), Ancistrura Uvarov, 1921, Andreiniimon Capra, 1937, Dasycercodes Bey-Bienko, 1951, Euconocercus Bey-Bienko, 1950, Isoimon Bey-Bienko, 1954, Isophya Brunner von Wattenwyl, 1878, Kurdia Uvarov, 1916, Leptophyes Fieber, 1853, Metaplastes Ramme, 1939, Orthocercodes Bey-Bienko, 1951, Phonochorion Uvarov, 1916, Poecilimon Fischer, 1853 and Polysarcus Fieber, 1853 .</p><p>Distribution. Widely distributed in the Palearctic, Europe, and Asia, extending to eastern China with notable diversification in the Mediterranean (Map 3 and 4).</p><p>Comments. Several genera currently grouped within this tribe were originally included in Odonturini (= Odonturae), along with other short-winged (macropterous and brachypterous) genera from various parts of the world (Brunner von Wattenwyl 1878, 1891). Later, Jacobson (1905) proposed the name Barbitistini, grouping the taxa known at that time. However, he did not provide a formal description of the tribe nor any clear differentiation between Barbitistini and Odonturini . The first formal description of the tribe was provided by Bey-Bienko (1954), who also included a taxonomic key for identifying the Phaneropterinae of Russia and neighbouring regions. However, the distinction between Odonturini and Barbitistini remained diffuse due to their superficial morphological similarities and overlapping distributions, particularly in the Iberian Peninsula and some Mediterranean islands.</p><p>Bey-Bienko (1954) mentioned that the tribe could be divided into two groups connected by transitional forms, although he did not propose any names or diagnoses to distinguish them. One of the groups suggested by Bey-Bienko (1954), included Isophya, Barbitistes, Poecilimon, and related genera, represent the more robust and diversified barbitistines. The other is treated as the Leptophyes group and related genera; it also includes Orthocercodes, which he considered to occupy an intermediate position. Bey-Bienko noted that last group is characterized by an ovipositor that is “lamelliform” with very fine marginal serration, similar to most winged Phaneropterinae, in contrast to the other group with an elongated and distinctly serrate ovipositor. That distinction may be useful for differentiating some genera; however, phylogenetic analyses such as those of Grzywacz et al. (2018) do not support those groups as constituting monophyletic units.</p><p>The taxonomy of Barbitistini has remained relatively stable since its establishment, despite comprising highly speciose genera that have been the subject of extensive revisions. Within these revisions, bioacoustics has proven to be a widely used tool, as well as cytogenetic analyses, both of which have contributed to the differentiation and delimitation of taxa (Heller 1990, Warchalowska-Sliwa et al. 2013, Zhantiev &amp; Korsunovskaya 2005, 2015). Nonetheless, the high diversity, complex evolutionary histories, and overlapping distributions of these groups make species delimitation a challenging task.</p><p>Below a key for identifying the genera, based on Bey-Bienko (1954), is also provided:</p><p>Key to Barbitistini genera</p><p>1. Fore tibiae not more than 1.5–2 times as long as pronotum (Figs. 3, 4). Ovipositor with strong pointed marginal denticles on apical part (Fig. 4C). Body medium-sized and robust................................................... 2</p><p>– Fore tibiae 2.5–3 times as long as pronotum (Fig. 5); females in which the fore tibiae are only twice or less than twice as long as the pronotum have an ovipositor with very finely serrate or almost smooth margins (Figs. 5B, C). Body small and slender.......................................................................................... 9</p><p>2. Fore femora shorter than the pronotum (Fig. 3A). Fore tibiae with strong spines, not widened basally, with middle part at least as wide as ocular diameter. Fastigium much wider than antennal scape (Fig. 6A).................... Polysarcus</p><p>– Fore femora not shorter than pronotum, often longer. Fore tibiae thinner, with small spines, widened basally; middle part of fore tibiae narrower than ocular diameter. Fastigium narrower or wider than antennal scape....................... 3</p><p>3. Pronotum normal in males without vertically elevated posterior parts and females with slight or shallow transverse grooves. First abdominal tergite of the male concealed by tegmina or partially exposed, always without a thick transverse elevated fold............................................................................................. 4</p><p>– Pronotum of males with vertically elevated posterior part, of females with curved upper side and deep transverse groove. First abdominal tergite of males partially exposed, with large wide transverse fold protruding upward.... Phonochorion</p><p>4. Transverse groove behind or along the middle of the pronotum. Tegmina of female almost entirely protruding from under the pronotum, in male, quite free, with exposed stridulatory organ........................................... 5</p><p>– Transverse groove in front of middle pronotum. Tegmina of female fully concealed or at least half hidden by the pronotum, in male with partially or entirely concealed stridulatory organs.............................................. 8</p><p>5. Lower margin of ovipositor straight up to the serrate part. Subgenital plate of male with sharp lamelliform median carina in lower part. Stridulatory organ on tegmina of male with obliterated veins, not defined by sharp convex vein along outer margin.......................................................................................... 6</p><p>– Lower margin of ovipositor arched. Subgenital plate of the male flat in apical part, with no median longitudinal carina or with very slight one in lower part. Stridulatory organ on tegmina of male with well-developed venation and clearly defined by convex vein along the outer margin........................................................... Isophya</p><p>6. Subgenital plate of male long, narrow, with median apical protuberance. Cerci of male not undulate, curved near the middle. Ovipositor with deep basal pit near upper margin of lower valve...................................... 7</p><p>– Subgenital plate of male short, transverse, swollen. Cerci of male sharply bent into shape of an S. Ovipositor with impressed longitudinal basal slit under the upper margin of the lower valve (Fig. 4C)............................. Barbitistes</p><p>7. Cerci of the male slightly curved, with obtuse apex. Epiproct of male without basal protuberance on each side. Ovipositor nearly twice as long as pronotum............................................................... Ancistrura</p><p>– Cerci of male strongly curved, with pointed or bifurcate apex. Epiproct of male with think protuberance on each side. Ovipositor not more than 1.5 times as long as pronotum.......................................... Metaplastes</p><p>8. Pronotum and exposed part of tegmina identical in both sexes; pronotum with strongly curved upper side even in female; tegmina in female swollen and protruding in male with fully concealed stridulatory organ. Mesonotal plate with posterior denticle on each side......................................................................... Isoimon</p><p>– Pronotum and tegmina different in the two sexes; pronotum of female simpler, almost cylindrical; tegmina in female fully concealed or barely protruding, not swollen. Mesonotal plate without posterior lateral denticle............. Poecilimon</p><p>9. Anterior margin of lateral lobes of pronotum quite vertical, not protruding obliquely forward. Pronotum without swelling between bases of forelegs; meso- and postnotal plates flat. Subgenital plate of male moderate length or short, not markedly curved upward and not protruding above the level of the posterior abdominal segment.......................... 10</p><p>– Anterior margin of lateral lobes of pronotum not vertical, protruding slightly and obliquely forward. Pronotum, mesonotal, and postnotal plates strongly convex. Subgenital plate of the male produced backward in a very long, narrow process bent upward and almost emerging above the upper side of the abdomen................................. Dasycercodes</p><p>10. Ovipositor with very finely serrate, sometimes almost smooth margins in apical part.Tegmina partially covered by pronotum or quite free; in the latter case, the subgenital plate of male has a slight longitudinal median carina on the lower side or is deeply notched posteriorly.......................................................................... 11</p><p>– Ovipositor with strongly serrate apex. Tegmina quite free. Subgenital plate of male with strong longitudinal carina in lower part............................................................................................ 13</p><p>11. Last abdominal tergite of male normal, without wide median lobes in posterior part. Subgenital plate of female triangular, not transverse, even elongate........................................................................ 12</p><p>– Last abdominal tergite of male with large median lobe in posterior part; apex of lobe transversely notched, bent sharply downward. Subgenital plate of female strongly transverse, with moderately produced acute apex......... Andreiniimon</p><p>12. Cerci of male without basal tubercle on upper side. Subgenital plate of female triangular or moderately pointed posteriorly but not produced apically.................................................................... Leptophyes</p><p>– Cerci of male with basal tubercle on upper side. Subgenital plate of female elongate, pointed, produced posteriorly into a narrow process......................................................................... Euconocercus</p><p>13. Epiproct small, elongatedly triangular, in male quite normal size and much narrower than distance between bases of cerci. Ovipositor without longitudinal serrate carinae on lateral surfaces of apical part...................... Orthocercodes</p><p>– Epiproct large, with arcuate round posterior margin, in male occupying the whole area between the cerci and situated in part above the posterior abdominal tergite. Ovipositor with two serrate longitudinal carinae on lateral surfaces of apical part.............................................................................................. Kurdia</p></div>	https://treatment.plazi.org/id/BE25B46BFF95FF986ADCFBCF02C6AE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFF8AFF9B6ADCFC3702C6AE5B.text	BE25B46BFF8AFF9B6ADCFC3702C6AE5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Monticolariini Cadena-Castañeda 2026	<div><p>Monticolariini trib. nov.</p><p>Description. Body small and slender (Figs. 9A, 10A, 11A). Head subrectangular or ovoid and slender in frontal view; fastigium sulcate, narrower than a third of the width of scape (Fig. 9B). Eyes rounded and slightly protrusive (except Odonturoides) (Figs. 9C, 11B). Thorax. Pronotum without modifications on any of its margins; lacking marked lateral carinae, though sometimes indicated by different coloration or a slight suggestion of relief, especially posteriorly; lateral lobes longer than high; humeral sinus not developed (Figs. 9C, 11B). Mesosternum quadrangular, almost as wide as high, with rounded lateral margins; metasternum 1.5 times broader than mesosternum; sternal grooves short with apophyses pits close together; lateral margins of meso- and metasternum lateral lobes elevated (Fig. 9E). Wings. Tegmina short in both sexes, covering from the first abdominal segment or two-thirds of the abdomen, always fully exposed (Figs. 9A, 10A, 11A). Legs long and slender, hind femur greatly exceeding the apex of the abdomen; fore coxae unarmed or with a small spinule; fore tibiae with open tympana. Mid-tibia of some males with one or two modified inner apical spurs. Abdomen. Tenth tergite without modifications or projections (except Peropyrrhicia with elongated and modified terminalia) (Figs. 9F, K). Males: Epiproct subtriangular and posterior margin rounded; cerci simple, unmodified, bearing a small apical spine (Figs. 9K, L, 10C). Subgenital plate quadrangular, almost as wide as long; without stylli, posterior margin generally with a median notch (Figs. 9H, M) ( Peropyrrhicia with deeply emarginate, with pronounced distal lateral edges curved upward). Genitalia. Entirely membranous; dorsal lobe (dl) simple, rounded, not exceeding the length of the lateral folds of dorsal lobe (ldl), and usually covered by several microstructures like small bristles. Titillator’s sclerite (TS) and sclerite of apodemes (AP) absent. Upper folds of ventral lobe (up.vl.) are rounded, surrounding the lower folds of ventral lobe (lw.vl.) without covering them. short and poorly differentiated; ejaculatory vesicles (ejv) conspicuous and rounded (Figs. 9I, J, N, O). Females: Epiproct triangular. Cerci conical and thin; ovipositor short and lamelliform, with slightly serrated or almost smooth margins. Subgenital plate subtriangular or quadrangular, rarely strongly divided.</p><p>Taxa included. Seven genera and 20 species (Check list 1): Monticolaria Sjöstedt, 1910 (type genus), Arostratum Massa, 2015, Atlasacris Rehn, 1914, Hempia gen. nov. Massaium gen. nov., Odonturoides Ragge, 1980, and Peropyrrhicia Brunner von Wattenwyl, 1891 .</p><p>Distribution. East Africa, between Tanzania, Kenya, Ethiopia, and Somalia. Also, Peropyrrhicia parva Ragge, 1980, from Yemen (South Arabian Peninsula) (Map 4).</p><p>Comments. The relationships among several genera now grouped within this new tribe were hypothesized in the past. Ragge (1980) was the first to suggest affinities among Atlasacris, Ivensia Bolívar, 1890, Monticolaria, and Odonturoides, highlighting the modification or specialization of one of the apical spurs of the mid tibiae.</p><p>Ragge also proposed a relationship between Monticolaria and Meruterrana Sjöstedt, 1912, again emphasizing the structure of the apical spurs of the mid tibiae. However, the development of modified spurs on the mid-tibiae may have occurred convergently. For example, both Atlasacris and Hempia gen. nov. exhibit developed spurs, but with different configurations, and the two genera are not entirely similar, as can be observed in the key to the genera of Monticolariini trib. nov.</p><p>Although Ivensia and Meruterrana show modifications to their spurs, they do not meet the other diagnostic characters required for inclusion in this new tribe. Ivensia may be more closely related to Ducetiini, despite its short subgenital plate. Meruterrana, on the other hand, is morphologically closer to Pronomapyga Rehn, 1914, than to the other short-winged phaneropterines of eastern Africa.</p><p>In the morphological and molecular study of Monticolaria by Hemp et al. (2009), the genus was recognized as well-differentiated, showing morphological affinities with Atlasacris, Meruterrana, and Odonturoides, which partially agrees with the perspective presented here. Recently, Massa (2015) included the genera Monticolaria, Atlasacris, Austrodontura, Odonturoides, Arostratum, and Odontura within Odonturini, providing a preliminary framework for understanding this group of katydids.</p><p>Here, we define and formally establish the new tribe Monticolariini trib. nov., which includes several African genera previously placed in Odonturini, and provides diagnostic characters to distinguish them from other short-winged phaneropterines. This tribe is well differentiated from other Odonturini s.l. taxa based on both external and genital morphology. Additional taxa likely remain undescribed in eastern Africa, as many are restricted to isolated mountain ranges (Hemp 2009, Hemp et al. 2009), particularly in primary forests (Felix &amp; Massa 2016). Moreover, given their peculiar morphology, the reproductive behaviour of species within this tribe is likely to be complex and fascinating. Modified spurs in some genera, glandular hairs on the exposed hind wings of Massaium gen. nov., and the highly modified male terminalia in Peropyrrhicia are examples of the morphological and behavioural diversity expected in this group.</p><p>Below, a key to the identification of the genera included in this new tribe is provided:</p><p>Key to Monticolariini, trib. nov. genera</p><p>1. Posterior part of the lateral lobes of the pronotum inflated (Fig. 11B)......................................... 2</p><p>– Posterior part of the lateral lobes of the pronotum not inflated (Figs. 9C, 10A).................................. 4</p><p>2. Mid tibiae without an enlarged apical spur. Posterior part of the lateral lobes of the pronotum moderately inflated; posterior margin of the last male tergite with lateral edges not prolonged.............................................. 3</p><p>– Mid tibiae with a markedly enlarged internal apical or subapical spur. Posterior part of the lateral lobes of the pronotum conspicuously inflated; posterior margin of the last male tergite with laterally angled prolongations that do not exceed the median length of the cerci................................................................... Atlasacris</p><p>3. Posterior margin of the last male tergite prolonged and slightly curved downward, not exceeding the median length of the cerci, covering most of the epiproct. Posterior margin of the lateral lobe of the pronotum obliquely curved forward. Last tooth on the posterior edge of the stridulatory file of the left tegmen hook-shaped, larger, and longer than the preceding ones. Hind wings reduced to minute scales...................................................... Arostratum</p><p>– Posterior margin of the last male tergite almost straight, not prolonged, and does not cover the epiproct (Figs. 11D, E). Posterior margin of the lateral lobe of the pronotum curved downward (Figs. 11A, B). Last tooth on the posterior edge of the stridulatory file of the left tegmen normally developed and unmodified. Hind wings extending beyond the tegmina (Figs. 11B, 11C)........................................................................ Massaium gen. nov.</p><p>4. Males with one or two modified internal apical spurs on the mid tibiae........................................ 5</p><p>– Males with unmodified internal apical spurs on the mid tibiae............................................... 6</p><p>5. Tegmina in both sexes extending beyond the first abdominal segment, covering half to two-thirds of the abdomen (Fig. 9A); laterally without whitish stripes (Fig. 9C), but with a black spot at the base of the tegmina (Fig. 9D). Mid tibiae with two internal apical spurs, the ventral one markedly enlarged and curved upward.......................... Monticolaria</p><p>– Male tegmina not extending beyond the first abdominal segment (Fig. 10A), inflated and forming an acoustic chamber (Figs. 10A, B), laterally with whitish stripes between the costal and subcostal veins, and lacking black spots at the base. Mid tibiae with three internal apical spurs, the middle one markedly enlarged and curved upward..... Hempia gen. nov.</p><p>6. Eyes circular and pronotum rugose. Male terminalia strongly modified; last abdominal segment prolonged; subgenital plate deeply emarginate with pronounced distal lateral edges curved upward............................. Peropyrrhicia</p><p>– Eyes ovoid and pronotum smooth. Male terminalia unmodified; last abdominal segment with a straight posterior margin; subgenital plate rectangular, posterior margin nearly straight...................................... Odonturoides</p></div>	https://treatment.plazi.org/id/BE25B46BFF8AFF9B6ADCFC3702C6AE5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFF89FF946ADCFC1003E9AFC0.text	BE25B46BFF89FF946ADCFC1003E9AFC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hempia Cadena-Castañeda 2026	<div><p>Hempia gen. nov.</p><p>Type species. Hempia jagoi (Ragge, 1980) comb. nov., here designated.</p><p>Description. Small to medium-sized (12–17 mm), body slender (Fig. 10A). General coloration green; base of the legs whitish-green; area between the costal and subcostal margins of the tegmina white; dorsal surface of the abdomen whitish, usually with distinctive spots along the mid-dorsal stripe. Antennae reddish-brown, especially the scape; flagellum with dark and reddish to reddish-brown patches (Fig. 10B). Eyes and femoro-tibial joints reddish to reddish-brown. Head ovoid; fastigium of the vertex compressed, grooved dorsally; eyes circular and prominent.</p><p>Thorax. Pronotum without lateral carinae (Fig. 10B); lateral lobes of the pronotum almost as high as its wide; posterior part of the lateral lobes of the pronotum not inflated (Fig. 10A). Wings. Tegmina not extending beyond the first abdominal segment, inflated and forming an acoustic chamber; hind wings absent (Figs. 10A, B). Legs. Fore coxa with a small spine; femora unarmed; fore and mid tibiae with three to five external ventral spurs. Mid tibiae with three inner apical spurs, the middle one markedly enlarged and curved upward. Hind tibiae with 13 to 18 outer dorsal spines. Abdomen. Last abdominal segment moderately emarginate at the center. Epiproct broader than long, with a rounded posterior margin. Cerci cylindrical, bearing a small internal apical spine (Figs. 10C). Subgenital plate quadrangular, nearly as long as wide, posterior margin with a notch and two subtriangular projections on each side; stylli absent.</p><p>Female. Unknown.</p><p>Species included. Hempia jagoi (Ragge, 1980) comb. nov., and Hempia hanangensis (Hemp, 2009) comb. nov.</p><p>Distribution. Tanzania.</p><p>Etymology. The genus is dedicated to Claudia Hemp as recognition for her significant contribution to Orthoptera from East Africa. Also, as an acknowledgment of her help and support with African specimens. The gender of this name is being established as feminine.</p><p>Comparison. Hempia gen. nov. differs from Peropyrrhicia and Odonturoides by lacking modified apical spurs on the mid tibiae. It differs from Peropyrrhicia by the absence of a rugose pronotum and by not having a markedly modified terminalia. It also differs from Odonturoides in having rounded, rather than ovoid eyes, and in possessing inflated tegmina, which do not occur in Odonturoides .</p><p>The genus most closely related to Hempia gen. nov. is Monticolaria . They differ in tegmina length: in Monticolaria, the tegmina are not inflated, reach at least half the length of the abdomen or up to two-thirds of its length, and bear a black spot at the base without white stripes on costal margin. In contrast, Hempia gen. nov. with short tegmina that do not extend beyond the first abdominal segment, and the costal margin is whitish. Both genera share a modified median spur on the mid tibiae, but differ in the number and configuration of spines on the mid tibiae. Monticolaria has two internal apical spurs, the ventral one being markedly enlarged and falciform. In contrast, the new genus has three internal apical spurs, with the middle one markedly enlarged and hook-shaped.</p><p>Comments. Ragge (1980), when describing Odonturoides, suspected that some of the species included in the genus might actually belong to other genera. Following the study by Hemp (2009) and the description of Odonturoides hanangensis Hemp, 2009, as well as the broader examination of short-winged phaneropterine genera worldwide, Ragge’s suspicions are confirmed. Therefore, upon analyzing the morphological differences among the species previously assigned to Odonturoides, it became necessary to divide them into two new genera: Hempia gen. nov., which is more closely related to Monticolaria and characterized by modified mid-tibial spurs; and Massaium gen. nov., another derived genus with the posterior part of the lateral lobes of the pronotum inflated, showing closer affinities to Atlasacris and Arostratum .</p><p>As a result of these taxonomic revisions, Odonturoides becomes monotypic and is the only genus within the tribe characterized by ovoid eyes and a male subgenital plate with an almost straight posterior margin. The two species included in the new genus are described originally in Odonturoides, here, the new combinations are effective: Hempia jagoi (Ragge, 1980) comb. nov., and Hempia hanangensis (Hemp, 2009) comb. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFF89FF946ADCFC1003E9AFC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFF84FF966ADCFF0206BEABF0.text	BE25B46BFF84FF966ADCFF0206BEABF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Massaium Cadena-Castañeda 2026	<div><p>Massaium gen. nov.</p><p>Type species. Massaium insolitum (Ragge, 1980) comb. nov., here designated.</p><p>Description. Small to medium-sized (13–18 mm), body slender (Fig. 11A). General coloration green; base of the legs, coxae, thoracic pleura, and sternites whitish; tegmina apex white; dorsal surface of abdomen with a white stripe, extending to epiproct. Eyes, knees, tarsi, and hind tibiae brown. Scape and pedicel segments of antennae red or pale orange-brown; flagellomeres black. Head ovoid; fastigium of the vertex compressed, grooved dorsally; eyes circular and prominent. Thorax. Pronotum without lateral carinae; (Figs. 11B, C) lateral lobes of the pronotum wider than high; posterior part of the lateral lobes of the pronotum inflated (Fig. 11B). Wings. Tegmina not extending beyond the second abdominal segment; hind wings present, extending beyond tegminae, reaching hind margin of second abdominal tergite, apparently with glandular hairs towards tip (Figs. 11B, C). Legs. Fore coxa unarmed (Fig. 11B), femora with ventral spinules; fore tibiae with four external ventral spurs; mid tibiae with five or six external ventral spurs. Mid tibiae with three internal apical and unmodified spurs. Hind tibiae with 22 to 27 external dorsal spines. Abdomen. Last abdominal segment almost straight (Figs. 11D, E). Epiproct broader than long, with a rounded posterior margin (Figs. 11D). Cerci cylindrical, bearing a small internal apical spine (Figs. 11D, E). Subgenital plate quadrangular, nearly as long as wide, posterior margin with a notch and two conical projections on each side; stylli absent (Figs. 11F).</p><p>Female. As the male. Tegmina reduced to small overlapping flaps not reaching hind margin of first abdominal tergite. Cerci slender at tip but not pointed. Ovipositor upcurved, finely denticulated on margins.</p><p>Species included. Only the type species.</p><p>Distribution. Tanzania.</p><p>Etymology. The genus is dedicated to Bruno Massa an esteemed orthopterist. Also, this is an acknowledgment for his help and support with European specimens. The gender of this name is neuter.</p><p>Comparison. Massaium gen. nov. is closely related to Atlasacris and Arostratum; all three genera share the characteristic of having the posterior part of the lateral lobes of the pronotum inflated. Massaium gen. nov. differs from Atlasacris in lacking a long apical spur on the mid tibiae, a conspicuous feature in males of Atlasacris .Although both genera possess short tegmina, they can be distinguished by the coloration of the area between the costal and subcostal margins: in Atlasacris it is white, while in the new genus, only the apex of the tegmina is whitish. In addition, the hind wings in Massaium gen. nov. protrude beyond the middle of the tegmina, a condition not observed in other genera of the tribe. Another distinguishing feature is the male terminalia: in Atlasacris, the posterior margin of the last male tergite bears laterally angled prolongations, which are absent in Massaium gen. nov.</p><p>Massaium gen. nov. and Arostratum both possess short tegmina and lack a long apical spur on the mid tibiae. However, they differ in the shape of the pronotal lateral lobes: in Arostratum, these lobes are less inflated, and the posterior margin of the lateral lobe is obliquely curved forward. In contrast, Massaium gen. nov. exhibits more inflated lateral lobes, with the posterior margin curved downward. Regarding wing characters, the two genera differ in the stridulatory file and hind wings. In Arostratum, the last tooth on the posterior edge of the stridulatory file of the left tegmen is hook-shaped, and the hind wings are reduced to minute scales. In contrast, in Massaium gen. nov., the last tooth of the stridulatory file is unmodified, and the hind wings protrude at the middle of the tegmina.</p><p>Comments. As mentioned in the remarks on Hempia gen. nov., Massaium gen. nov. is established as new and distinct from Odonturoides based on the aforementioned characters. Males of the only known species in this genus, Massaium insolitum comb. nov., exhibit hind wings that protrude at the middle of the tegmina, and this exposed section bears hairs which may well be glandular, possibly producing an attractant used during courtship and copulation (Ragge 1980).</p></div>	https://treatment.plazi.org/id/BE25B46BFF84FF966ADCFF0206BEABF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFF84FFAC6ADCF97E02C7A983.text	BE25B46BFF84FFAC6ADCF97E02C7A983.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dichopetalini Cadena-Castañeda 2026	<div><p>Dichopetalini trib. nov.</p><p>Description. Body small or medium-sized, slender, and moderately robust (Figs. 12A, 13A, 14A). Head subrectangular or ovoid and slender in frontal view; fastigium sulcate, narrower than half of width of scape (Figs. 12B, 13B, 14B, 14H). Eyes rounded and slightly protrusive. Thorax. Pronotum without modifications on any of its margins; lacking marked lateral carinae (except Pterodichopetala), though sometimes indicated by different coloration or a slight suggestion of relief, especially posteriorly; lateral lobes longer than high; humeral sinus not developed (Figs. 12C, 12D, 13C, 13D, 14C, 14D, 14I, 14J). Mesosternum quadrangular, wider than high, lateral margins rounded or divergent from ventral to upper margins; metasternum 1.5 times broader than mesosternum, lateral margins divergent from upper to ventral margins; sternal grooves short with pits apophyses close together; lateral margins of meso- and metasternum lateral lobes not elevated (Figs. 12E, 13E, 14E, 14K). Wings. Tegmina short in both sexes, always fully exposed, mainly covering the first or second segments (Figs. 12C, 12D, 13C, 13D, 14C, 14D, 14I, 14J). Some exceptions exist, such as Arachnitus being very reduced or absent, or Tedcohnia gen. nov. and Pterodichopetala covering half or the abdomen or completely (Figs. 15A). Legs long and slender, hind femur exceeding the apex of the abdomen; fore coxae unarmed, fore tibiae with open tympana; and mid tibia spurs unmodified. Abdomen. Tenth tergite variable, usually modified, partially divided, or prolonged (Figs. 14F, 14G, 13F, 15C, 15J, 15P). Males: epiproct variable, from unmodified, subtriangular or rectangular, or modified, with posterior margin denticulated or with elongated posterior prolongation. Cerci simple and unmodified, may sharply from the mesial to apex (Figs. 12F, 12G, 12P, 12Q); most genera with modified cerci, apex serrate (Figs. 13F, G), or divided in different ways (Figs. 15C, E, K, L). Some genera with a basal appendage or projection, conspicuous to Rhabdocerca, present but poorly developed in Dichopetala . Subgenital plate without stylli, mostly rectangular, with different apex shape, with distal lobes divergent and conspicuous (Fig. 12H) or normally developed (Figs. 13H, 15D, 15L, 15R). Genitalia. Membranous but with some sclerotized structures. Dorsal lobe (dl) simple, rounded, short of the lateral folds of dorsal lobe (ldl), usually with a medium-sized notch at apex. Titillator’s sclerite (TS) and sclerite of apodemes (AP) present and well developed ( Planipollex and Arachnitus with TS not strongly sclerotized, submembranose, but differentiated). Upper folds of ventral lobe (up.vl.) rounded, surrounding the lower folds of ventral lobe (lw.vl.) and mostly covering them, only partially exposed; lw.vl. short and rounded; ejaculatory vesicles (ejv) conspicuous and rounded (Figs. 12I, 12J, 12I, 12J). Females: Epiproct triangular. Cerci conical and robust; ovipositor either elongated, longer than pronotum length, apex with strongly serrate margins (Figs. 14F, L). Subgenital plate variable in shape, but divided into two separated sclerotized lobes by a medial-longitudinal membrane (Figs. 14G, 14M, 15F, 15M, 15S).</p><p>Taxa included. 10 genera and 47 species (Check list 2): Dichopetala Brunner von Wattenwyl, 1878 (type genus), Acanthorintes Cohn, Swanson &amp; Fontana, 2014, Arachnitus Hebard, 1932, Gymnocerca Cohn, Swanson &amp; Fontana, 2014, Mactruchus Cohn, Swanson &amp; Fontana, 2014, Maxium gen. nov., Obolopteryx Cohn, Swanson &amp; Fontana, 2014, Planipollex Cohn, Swanson &amp; Fontana, 2014, Pterodichopetala Buzzetti, Barrientos-Lozano &amp; Rocha-Sánchez, 2010, Rhabdocerca Cohn, Swanson &amp; Fontana, 2014 and Tedcohnia gen. nov.</p><p>Distribution. North America, ranging from Mexico to the southern United States (Maps 5 and 6).</p><p>Comments. A preliminary mention of this tribe was made under the informal name “dichopetalines” by Cohn et al. (2014), emphasizing that it was an unofficial term referring to taxa derived from Dichopetala, and that the use of this term did not constitute the formal establishment of a tribal taxon. Similarly, that informal group name is unavailable under the rules of nomenclature, as it was published without a formal description or diagnosis. Here, based on additional morphological evidence and in comparison, with other short-winged taxa worldwide, the group is recognized as a distinct tribe, separate from Odonturini, which also includes Arachnitus a genus not covered in the comprehensive monograph by Cohn et al. (2014).</p><p>An early revisionary study was carried out by Rehn &amp; Hebard (1914), which provided a broad and significant contribution, including the grouping of species within Dichopetala s.l. that partially corresponds to several currently recognized genera. Many of the species known today were described by those authors, who also provided a key to the distribution of the species and abundant distributional data. For nearly a century, that work served as the primary reference for the identification of these insects.</p><p>Recently, the group was extensively revised by Cohn et al. (2014), who clarified the delimitation of currently recognized genera. Prior to their contribution, all known species were primarily placed within the genus Dichopetala . Their study was comprehensive, offering detailed morphological and distributional characteristics and addressing multiple issues of taxonomic definition and differentiation. As a result, it currently stands as the most complete and authoritative reference on the group.</p><p>Additional contributions have focused on the description of new species within two genera, Obolopteryx (Barrientos-Lozano et al. 2015, 2016) and Pterodichopetala (Buzzetti et al. 2010b; Barrientos-Lozano &amp; Rocha-Sánchez 2013; Rocha-Sánchez et al. 2015). Molecular systematics studies have also been conducted by Kensinger et al. (2017) and Rocha-Sánchez et al. (2019), which will be discussed further in this contribution.</p><p>Below, two new genera are described, and a key for their identification is also provided.</p><p>Key to Dichopetalini trib. nov. genera</p><p>1. Tegmina completely covering the abdomen or at least half of it.............................................. 2</p><p>– Tegmina covering up to the second abdominal segment at most, or it is absent.................................. 3</p><p>2. Lateral margins of the pronotal disc straight and well-defined; pronotal disc flat. Tegmina fully covering the abdomen, narrowing toward the apex, and almost pointed (Fig. 15I). Male’s last abdominal segment prolonged, reaching or surpassing half the length of the cerci (Fig. 15J). Male cerci with two distinct branches (Figs. 15J–L)........... Pterodichopetala</p><p>– Lateral margins of the pronotal disc curved and rounded; pronotal disc slightly arched. Tegmina not covering the entire abdomen, usually reaching half to two-thirds (Fig. 15A) (except T. alfredoi comb. nov., where the tegmina reach close to the abdominal apex); tegmina rounded, not tapering apically (Fig. 15B). Male’s last abdominal segment not conspicuously prolonged (Fig. 15C). Male cerci with three well-defined branches (Figs. 15C–E)............... Tedcohnia gen. nov.</p><p>3. Body slender to moderately robust, medium-sized. Tegmina covering the first to second abdominal segment. Legs long, hind femur 1 to 1.5 times the body length. Male cerci modified (e.g., branched, thickened, flattened, or distally denticulate). Ovipositor as long as half to two-thirds of the hind femur.................................................. 4</p><p>– Body very small and slender. Tegmina lobiform, reduced or absent. Legs extremely long and slender; hind femur two times or more the body length. Male cerci simple and unmodified. Ovipositor reduced, not exceeding one-third of the hind femur length................................................................................... Arachnitus</p><p>4. Male cerci simple, without branching or additional appendages (except in some Matrachus species with a small dorsal appendage)....................................................................................... 5</p><p>– Male cerci mostly robust, with branching or a well-defined dorsal appendage.................................. 8</p><p>5. Head and body slender; distal half of male cerci thin and pointed............................................ 6</p><p>– Head and body moderately robust; distal half of male cerci not pointed, simple, denticulate, or with a tiny preapical dorsal projection........................................................................................ 7</p><p>6. Male’s last abdominal segment with triangular, moderately prolonged lateral posterior edges; medial posterior margin concave and without projections (Figs. 12F–G). Male subgenital plate with a V-shaped notch, lateral lobes divergent and pointed (Fig. 12H). Titillator’s sclerite (TS) composed of two fin-shaped plates, non-denticulate (Fig. 12I). Female terminalia with rectangular lateral lobes, poorly separated (Figs. 14L–M)............................. Dichopetala</p><p>– Male’s last abdominal segment without lateral projections (Fig. 15P); medial posterior margin with two small central projections (Fig. 15Q). Male subgenital plate with a U-shaped notch, lateral lobes sub-rectangular, not strongly divergent (Fig. 15R). Titillator’s sclerite (TS) fused, with two anterior and two posterior denticulate projections (Figs. 15T–U). Female terminalia with triangular lateral lobes, widely separated (Fig. 15S)...................... Maxium gen. nov.</p><p>7. Male cerci simple, tapering, without appendages; apex acute, subapically notched, or blunt with a minute subapical scale. Apex of male subgenital plate entire or truncate, or apical region narrowed with rounded or angulate lobes.. Gymnocerca</p><p>– Male cerci variable, with or without dorsal appendage; acuminate or broad with blunt teeth. Apex of male subgenital plate with small lateral projections (Fig. 13I)......................................................... Matrachus</p><p>8. Rod-shaped projection at the base of the male cerci absent. Male titillator’s sclerite (TS) variable, denticulate or absent.. 9</p><p>– Rod-shaped projection arising proximally from the basal constriction of the male cerci present. Male titillator’s sclerite (TS) consists of paired, weakly sclerotized, roundly quadrate plates with slightly upturned cephalic margins................................................................................................. Rhabdocerca</p><p>9. Male epiproct always simple and unarmed. Male cerci bifurcate, both branches approximately equal in length, usually with a concave dorsal branch; titillator’s sclerite (TS) non-denticulate and weakly sclerotized. Female tegmina short, rounded, either separated or barely touching................................................................... 10</p><p>– Male epiproct elongate, reflexed, ending in paired short projections. Male cerci cheliform, robust, without concave regions; titillator’s sclerite (TS) with paired distal and erect proximal fingers, both spiny and with lateral flanges. Female tegmina short to moderate, sinuate or squarish, and slightly overlapping................................... Acanthorintes</p><p>10. Dorsal body surface in males with a black, brown, or reddish stripe extending from the head to the last abdominal segment, laterally bordered by a white stripe. Female dorsal body surface with a similar or distinct pattern as male, or entirely green, but always with a white lateral stripe on each side. Male cerci with a short to long dorsolateral, ventrally concave “thumb” near the base. Titillator’s sclerite (TS) W-shaped and sclerotized.................................... Obolopteryx</p><p>– Dorsal body surface green in both sexes; males with a narrow white stripe on each side, females lacking the stripe or with a faint one. Male cerci with a long lateral, ventrally concave, flattened “thumb” near the base. Titillator’s sclerite (TS) not sclerotized.............................................................................. Planipollex</p></div>	https://treatment.plazi.org/id/BE25B46BFF84FFAC6ADCF97E02C7A983	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFBEFFAF6ADCFAA80398AC0C.text	BE25B46BFFBEFFAF6ADCFAA80398AC0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tedcohnia Cadena-Castañeda 2026	<div><p>Tedcohnia gen. nov.</p><p>Type species. Tedcohnia cultricerca (Strohecker, 1945) comb. nov., here designated.</p><p>Description. Medium-sized species (14–23 mm), and body robust (Fig. 15A). General coloration green. Most species exhibit a contrasting dorsal stripe along the body, ranging in color from reddish-brown to black. In some species, this stripe may be faint or poorly colored on pronotum or abdomen; in such cases, the dorsal margin is outlined by lateral white stripes. Tegmina with a white costal stripe, which may be broad, located between the costal and subcostal veins, or narrow and accompanied by black spots. Antennae reddish-brown, especially the scape, and pedicel; the flagellum shows dark and reddish to reddish-brown patches. Femoro-tibial joints reddish to reddish-brown, rarely black. Head rounded; fastigium of the vertex compressed; eyes rounded and moderately prominent.</p><p>Thorax. Pronotum without lateral carinae or humeral notch; pronotal disc somewhat constricted mesally in dorsal view and slightly arched in transverse section; lateral lobes of the pronotum wider than high. Wings. Tegmina do not cover the entire abdomen, usually extending to half or two-thirds of its length (except in T. alfredoi comb. nov., where the tegmina reach near the apex of the abdomen); tegmina are rounded apically, not tapering (Fig. 15B). Hind wings vestigial. Legs. Fore coxae unarmed; tympana open on both sides of the fore tibia; apical spurs of all tibiae unmodified. Hind femora noticeably surpass the abdominal apex. Abdomen. Last abdominal segment is not conspicuously prolonged; it may be truncated or bear small lateral lobes on the posterior margin (Figs. 15C, D). Epiproct rectangular, wider than long, with the posterior margin usually bearing short teeth. Male cerci with three well-defined branches (Fig. 15E): the basal projection is longer than the distal ones; the dorsal projection is wider than the basal one and divided into two lobes. Subgenital plate is subtriangular, with a small median notch on the posterior margin. Stylli absent, represented only by minute styliform projections (Fig. 15E). Titillator sclerite (TS) present as a pair of fused median plates, bearing dorsoproximal, appressed, spined fingers, in addition to robust, paired distal projections with spines (Figs. 15G–H).</p><p>Female. Similar in general appearance to the male, differing mainly in sexual traits and tegmina. Female tegmina shorter than males, but not shorter than the median abdominal length; they cover at least the first three abdominal tergites. Cerci are conical, slender, and pointed. Ovipositor is as long as or nearly as long as the median length of the hind femur; its distal third is denticulate. Subgenital plate is ovoid or subtriangular; lateral lobes widely separated, rectangular shaped, and positioned on either side of the plate (Fig. 15F).</p><p>Species included. Tedcohnia cultricerca (Strohecker, 1945) comb. nov., T. alfredoi (Barrientos-Lozano &amp; Rocha-Sánchez, 2013) comb. nov., T. geovanyi (Barrientos-Lozano &amp; Rocha-Sánchez, 2015) comb. nov., T. hypsibates (Cohn, Swanson &amp; Fontana, 2014) comb. nov., T. padrisima (Cohn, Swanson &amp; Fontana, 2014) comb. nov., T. pityophila (Cohn, Swanson &amp; Fontana, 2014) comb. nov. and T. robertoi (Barrientos-Lozano &amp; Rocha-Sánchez, 2015) comb. nov.</p><p>Distribution. Northeastern Mexico, between San Luis Potosí, Tamaulipas, Nuevo León, and Coahuila states (Map 6).</p><p>Etymology. The genus is dedicated to the memory of the great and legendary orthopterist Ted Cohn (1930– 2012), who, together with his collaborators, conducted an impeccable revision of this group of katydids. Beyond his scientific contributions, he was a generous mentor, supporter, and source of inspiration for many orthopterists. His kindness, approachability, and unwavering passion for Orthoptera made him an esteemed colleague and an honorable man in every sense. The gender of this name is being established as feminine.</p><p>Comparison. Tedcohnia gen. nov. is similar to Pterodichopetala mainly due to its elongated tegmina, which are longer than the head and pronotum combined. However, the new genus differs from Pterodichopetala in several morphological aspects. The Pterodichopetala pronotal disc is flat with straight, well-defined lateral margins. In contrast, Tedcohnia gen. nov. exhibits a pronotum more typical of most genera within the tribe, with the pronotal disc slightly constricted medially in dorsal view and slightly arched in transverse section.</p><p>Regarding the tegmina, Pterodichopetala has elongate tegmina that cover the abdomen; the costal margin is curved towards the apex, the anal margin is nearly straight, and both margins converge to form an almost pointed apex (Fig. 15I). In Tedcohnia gen. nov., the tegmina are ovoid, and the apex is not tapered. Both the costal and anal margins are curved (Fig. 15B), and the A1 vein or stridulatory file tends to be longer and extends closer to the anal margin compared to Pterodichopetala .</p><p>There are also marked differences in the male terminalia between the two genera. In Pterodichopetala, the last abdominal segment is prolonged, reaching or exceeding half the length of the cerci (Fig. 15J). In contrast, Tedcohnia gen. nov., this segment is not significantly extended and may appear truncated or bear small lateral lobes on the posterior margin (Fig. 15C). The male cerci in Pterodichopetala bear two distinct branches, with the basal or inner branch shorter than the outer one (Figs. 15K–L). Tedcohnia gen. nov., cerci exhibit three well-defined projections: the basal projection is longer than the distal ones, and the dorsal projection is wider than the basal one and divided into two lobes (Figs. 15D–E).</p><p>The male genitalia also present key differences between the two genera. Pterodichopetala titillators sclerite (TS) consist of two plates strongly fused medially, with short basal fingers and narrow posterior edges, each bearing rows of small teeth (Figs. 15N–O). In contrast, Tedcohnia gen. nov., the TS are either gently joined or separated medially, with long basal fingers and posterior margins forming two elevated, denticulate folds (Figs. 15G–H).</p><p>Tedcohnia gen. nov. is also morphologically similar to Acanthorintes in terms of internal genitalia, as both genera have a denticulate posterior margin and basal elevations resembling fingers. However, Acanthorintes differs by having notably elongated fingers and only a small denticulate section on the posterior margin, whereas Tedcohnia gen. nov. has the entire posterior margin denticulate. Regarding external morphology, the two genera are easily distinguished: Acanthorintes has short wings that do not fully cover the second abdominal segment, cheliform cerci, and a conspicuously modified epiproct.</p><p>Comments. Tedcohnia gen. nov. is herein proposed as a new genus, and new combinations are established for the species transferred to it. The new genus is entirely derived from Pterodichopetala . Of the 11 species currently recognized in the original genus, only four remain in Pterodichopetala: P. cieloi Buzzetti, Barrientos-Lozano &amp; Rocha-Sánchez, 2010; P. monternach Barrientos-Lozano &amp; Zaldívar-Riverón, 2015; P. tuliensis Barrientos-Lozano &amp; Rocha-Sánchez, 2015; and P. strepsidactyla Cohn, Swanson &amp; Fontana, 2014 .</p><p>As a result of this taxonomic revision, the distribution of Pterodichopetala is now restricted primarily to the southern portion of northeastern Mexico, particularly between the states of San Luis Potosí and southern Tamaulipas. In contrast, Tedcohnia gen. nov. displays a broader specific diversity and distribution, ranging from San Luis Potosí to Coahuila, with both genera inhabiting the mountainous regions of the Sierra Madre Oriental.</p></div>	https://treatment.plazi.org/id/BE25B46BFFBEFFAF6ADCFAA80398AC0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFBDFFA86ADCFE220408A95C.text	BE25B46BFFBDFFA86ADCFE220408A95C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maxium Cadena-Castañeda 2026	<div><p>Maxium gen. nov.</p><p>Type species. Maxium neomexicanum (Barrientos-Lozano &amp; Ramírez-Núñez, 2013) comb. nov.</p><p>Description. Body slender and medium-sized (11–13 mm). General coloration green, with a contrasting dorsal stripe along the body, lateral margins of the band are outlined in white, followed by a broader black outline. The strip on pronotal disc is brown-reddish, narrowing medially; the abdominal stripe keeps the same pattern of lateral edging, but the dorso-central section is yellowish-green with numerous black punctuations. Tegmina with a white stripe between costal and subcostal veins; dorsal field of tegmina reddish-brown.Antennal scape pale orange, pedicel reddish, and flagellomeres with long black sections alternating with short whitish segments. Fore and mid femora orange, hind femur only with the distal half orange, the basal one green; femoro-tibial joints dark brown; all tibiae and tarsomeres black dorsally. Head rounded; fastigium of the vertex compressed; eyes rounded and moderately prominent. Thorax. Pronotum without lateral carinae or humeral notch; pronotal disc somewhat constricted mesally in dorsal view and slightly arched in transverse section; lateral lobes of the pronotum almost as wide as high.</p><p>Wings. Tegmina not extending beyond the second abdominal segment, and hind wings vestigial. Legs. Fore coxae unarmed; tympana open on both sides of the fore tibia; apical spurs of all tibiae unmodified. Hind femora noticeably surpass the abdominal apex. Abdomen. Last abdominal tergite divided into three plates: two lateral plates, dorsally separated and narrowing toward the middle, and a posterior median plate that separates the lateral ones and extends along the entire posterior half. Posterior margin with two small, conical, and distinctive projections at the middle (Fig. 15Q). Cerci basally robust, curving outward, then inward, and gradually tapering to a sharp apex (Figs. 15P, Q). Subgenital plate rectangular, longer than wide, curving upward between the cerci; posterior section with a Ushaped notch, lateral lobes sub-rectangular, not strongly divergent (Fig. 15R). Titillator sclerite (TS) fused, with two anterior and two posterior denticulate projections (Figs. 15T, U).</p><p>Female. Similar in general appearance to the male, differing mainly in sexual traits and tegmina. Female tegmina shorter than males, rectangular shaped, slightly separated; costal margin with a conspicuous white band, followed by a reddish-brown band between veins Sc and M; the remaining half dark green with a broad black basal border along the anal margin. Cerci conical, moderately robust, and pointed. Ovipositor nearly as long as the median length of the hind femur; its distal section is denticulate. Subgenital plate divided, forming two ovoid projections; lateral lobes widely separated and triangular shaped (Fig. 15S).</p><p>Species included. Only the type species.</p><p>Distribution. Northeastern Mexico, Tamaulipas state, only recorded to Sierra de Tamaulipas (Map 6).</p><p>Etymology. The genus is dedicated to Max Gonzalez, a dear friend and loyal companion of the first author, in recognition and appreciation for his support during the first author’s stay in Mexico. The gender of this name is being established as neuter.</p><p>Comparison. Maxium gen. nov. is externally similar to Dichopetala, Obolopteryx, and Planipollex, which are among the slenderest genera occurring in the same region as the new genus. Maxium gen. nov. differs from Obolopteryx and Planipollex in that its cerci are not divided into two branches. In Obolopteryx, the titillator sclerite (TS) is W-shaped and lacks denticulations, while in Planipollex, the TS is not sclerotized. In contrast, Maxium gen. nov. has a well-sclerotized and denticulate TS.</p><p>The new genus is more similar to Dichopetala in the shape of the male cerci, which are unbranched and gradually acuminate. However, the two genera differ in other aspects: males of Dichopetala have the last abdominal segment with triangular, moderately prolonged posterolateral margins and lack median projections. Maxium gen. nov., the posterolateral margins are not prolonged, and the posterior edge of the tergite bears two small, conical median projections. The male subgenital plate also differs: Dichopetala has a V-shaped notch and divergent, pointed lateral lobes, features not present in the new genus. Female terminalia further aid in distinguishing the two genera: Dichopetala has rectangular, poorly separated lateral lobes, while in the new genus the lateral lobes are triangular and widely spaced. Another key distinguishing feature is the structure of the titillator sclerite. To Dichopetala, it consists of two fin-shaped, non-denticulate plates, whereas in Maxium gen. nov., it is a single, fused, denticulate structure.</p><p>It is also necessary to compare Maxium gen. nov. with Acanthorintes, as both differ markedly in the shape of the male cerci and the internal genitalia. Acanthorintes possesses divided, cheliform cerci, and its TS consists of two separated plates with paired distal and erect proximal fingers, both spinose and bearing lateral flanges. The new genus does not share this combination of features. The cerci are not cheliform or branched, but rather slender and gradually tapered in their distal half. The TS is not divided into paired plates but is strongly fused. Although the two projections resemble those of Acanthorintes, in Maxium gen. nov. they are longer, with fewer distal denticulations; the anterior projections are divergent, broader, and with denticulations on the distal dorsal margin, a condition not observed in Acanthorintes . Overall, the structure of the TS in Maxium gen. nov. is more similar to that of Tedcohnia gen. nov. than to Acanthorintes .</p><p>Comments. Maxium gen. nov. is described here as a new, monotypic genus, currently known only from the Sierra de Tamaulipas, a small isolated mountain range separated from the Sierra Madre Oriental. This geographic isolation likely facilitated the differentiation of this taxon.</p><p>With the description of this new genus, the distribution of Acanthorintes is once again restricted to the areas originally defined by Cohn et al. (2014), confined to the southeastern portion of the Central Plateau desert. Its range extends only slightly into the western limits of the Sierra Madre Oriental, north of Ciudad del Maíz (San Luis Potosí), in southwestern Tamaulipas, and into the Trans-Mexican Volcanic Belt near Carapán (Michoacán), within pine-oak forests, between 997 and 2450 meters in elevation.</p><p>This revision excludes Acanthorintes from the coastal plains of Tamaulipas, where it might otherwise be expected to occur as part of a continuous distribution that would connect with Maxium neomexicanum comb. nov. This species was originally described as Dichopetala by Barrientos-Lozano et al. (2013), later transferred to Acanthorintes by Cohn et al. (2014), and is now placed in its own genus based on a unique set of diagnostic characters that do not align with any known genus in the tribe.</p></div>	https://treatment.plazi.org/id/BE25B46BFFBDFFA86ADCFE220408A95C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFBAFFAB6ADCFB1302C6AC40.text	BE25B46BFFBAFFAB6ADCFB1302C6AC40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisophyini Cadena-Castañeda 2026	<div><p>Anisophyini trib. nov.</p><p>Description. Body small to medium-sized, mostly slender (Figs. 16A, 16E, 16I, 16J, 17). Head rounded or ovoid; fastigium of the vertex generally narrower than the width of the scape (Figs. 16C, 16F, 16K, 18O), except in Anisophya s.s. and Anisophya (Brevianisophya) subgen. nov. (Fig. 18A). Eyes rounded (Figs. 16C, 18K) or ovoid (Fig. 18M), slightly protruding. Thorax. Pronotum without modifications on any of its margins. In Angara and Parangara, the anterior and posterior margins of the pronotal disc are curved and slightly raised, giving the pronotum a saddle-like appearance (Fig. 16A). Lateral carinae of the pronotal disc are not strongly defined, generally rounded, and usually bordered by whitish stripes. Lateral lobes longer than high; humeral sinus not developed (Figs. 18C, M E, I, K, M, Q). Mesosternum quadrangular, nearly as wide as high, with rounded lateral margins; metasternum rectangular, twice as wide as the mesosternum; sternal grooves short with apophyses pits placed close together; lateral margins of the meso- and metasternum lobes not elevated (Figs. 18B, H, P). Wings. Tegmina short in both sexes, covering between the first and fifth abdominal segments, always fully exposed (Fig. 17) ( Nanoleptopoda males, tegmina covering abdomen (Fig. 16E)). Legs. Long and slender; hind femora exceed the apex of the abdomen; fore coxae unarmed; fore tibiae with open tympana. Abdomen. Tenth tergite unmodified and without projections (except in Hellerium gen. nov. (Fig. 19L)). Males: Epiproct subtriangular with rounded posterior margin; cerci generally simple and unmodified, bearing an apical spine (Figs. 19A–F), although exceptions occur, such as Anisophya (Brevianisophya) subgen. nov. with a preapical notch (Figs. 19G–K), Anisophya s.s. with an apically dilated cercus (Fig. 19C), and Caimanellus with bifurcated cerci (Figs. 19N–O). Subgenital plate rectangular, longer than wide, without styli; posterior margin usually with a median notch (Figs. 19B, E, H, S). Genitalia. Entirely membranous; dorsal lobe (dl) simple, rounded, not exceeding the lateral folds of the dorsal lobe (ldl), and generally covered by microstructures such as small bristles. Titillator sclerite (TS) and apodeme sclerite (AP) absent. Titillator (ti) rarely differentiated; when present, submembranous. Upper folds of the ventral lobe (up.vl.) rounded, surrounding but not covering the lower folds of the ventral lobe (lw.vl.); lw.vl. protruding medially between up.vl., reaching the mid-length of the dl. Ejaculatory vesicles (ejv) conspicuous and rounded (Figs. 19U–X). Females. Epiproct triangular; cerci conical and slender. Ovipositor variable in length, equal to, slightly longer than (Figs. 20D, F), or more than twice the length of the pronotum (Figs. 20B, H); margins finely serrated, except in Lorieriella gen. nov., Cohnia, and Caimanellus (Fig. 10J). Subgenital plate subtriangular or quadrangular, not deeply divided at the apex.</p><p>Taxa included. 8 genera, 21 species, and two subtribes (Check list 3): Anisophyina subtr. nov. and Angarina subtr. nov.</p><p>Distribution. South America: Andean region from Chile to Ecuador, and lowlands across Bolivia, Paraguay, Argentina, and southern Brazil (Map 7).</p><p>Comments. This new tribe includes most South American taxa traditionally grouped within Odonturini . Based on additional morphological evidence and comparisons with short-winged taxa from other regions of the world, the group is herein recognized as a distinct tribe, separate from Odonturini .</p><p>=Studies focused on resolving the taxonomy of South American short-winged phaneropterines have been scarce. Early descriptions, mostly by Brunner von Wattenwyl (1878, 1891), often included these species within Old World genera of the tribes Barbitistini or Odonturini, followed by sporadic descriptions of species and genera (e.g., Hebard 1924, Rehn 1945, 1955). The work of Karabag (1960) marked the beginning of an approach that distinguished South American taxa from Old World ones, through his clarification of Isophya and the description of Anisophya, which included a few species. However, the lack of clearly defined taxonomic boundaries, the inclusion of South American species in superficially similar foreign genera, poor original descriptions, and the absence of illustrations made the study of this group nearly impossible.</p><p>This highlights the significant impact of the Orthoptera Species File (Cigliano et al. 2025 and ongoing), which has provided open access to images of type specimens, references, and taxonomic tools, democratizing essential knowledge that was once accessible only to those able to visit European and North American collections.</p><p>Recently, Braun (2010, 2011) transferred the remaining South American Isophya species to Anisophya, reallocated species from Dichopetala to Cohnia, and revitalized the description of new taxa after decades of inactivity, ultimately completing the distinction between South American short-winged katydids and those from the Old World and North America. The contributions by Braun (2010, 2011), along with those of Buzzetti et al. (2010a), marked a turning point in the study of the so-called “South American Odonturini ”, paving the way for further taxonomic work, often accompanied by acoustic data (Nickle 2011; Deloya &amp; Braun 2023; Fianco 2023, Fianco &amp; Melo 2025, Fianco et al. 2019). Most recently, Battiston &amp; Fontana (2024) described a new genus in this group, marking a rare event not seen in at least six decades.</p><p>Below, two new subtribes and two genera are described, along with a key for their identification.</p><p>Key to Anisophyini trib. nov. subtribes and genera</p><p>1. Genicular lobes of the femora, especially the hind ones, armed with sharp spines. Hind tibiae strongly armed with dorsal sublamellate spines. Cryptic coloration with black and brown tones, accompanied by green patches in various shades that aid in camouflage among bryophytes (Figs. 16E, I, J) Angarina subtr. nov. .................................... 2</p><p>– Genicular lobes of the femora unarmed or with small, acuminate spines. Hind tibiae armed with short dorsal spines. Color predominantly green; some species with white, brown, or greenish stripes. Not camouflaged with bryophytes (Fig. 17) Anisophyina subtr. nov. ............................................................................ 4</p><p>2. Tegmina short, not surpassing the first or second abdominal segment (Fig. 16B). Pronotum with anterior and posterior margins elevated, more prominently so on the posterior margin, with a saddle-like appearance (Fig. 16A). Pronotal disc gradually narrowing in the middle, with a distinct transverse sulcus. Ovipositor not abruptly upcurved, apex acute or angled, and denticulated............................................................................. 3</p><p>– Tegmina covering abdomen in males (Figs. 16E, G); in females, covering from the third segment to slightly beyond midabdomen (Figs. 16I, J, L). Pronotal anterior and posterior margins not elevated. Pronotal disc with lateral margins nearly parallel, not noticeably narrowing, without a transverse sulcus (Figs. 16G, L). Ovipositor broadly and abruptly upcurved at the base, its dorsal margin and apex finely serrated, apex rounded (Figs. 16I, J)..................... Nanoleptopoda</p><p>3. Abdominal tergites with a paired series of small pointed dorsal projections. Male epiproct lanceolate, longer than wide, with sigmoid lateral margins, narrowing near mid-length. Ovipositor uniformly upcurved................. Parangara</p><p>– Abdominal tergites without paired dorsal projections (Figs. 16A, B). Male epiproct rounded, nearly as long as wide, with uniformly curved lateral margins. Ovipositor slightly upcurved, almost straight............................ Angara</p><p>4. Fastigium of vertex narrow, as wide as or narrower than the antennal scape (Figs. 18G, O)........................ 5</p><p>– Fastigium of vertex broad, 1.5 to three times as wide as the antennal scape (Fig. 18A)............................ 9</p><p>5. Last abdominal segment rarely constricted. Epiproct subtriangular and medium-sized. Male cerci variable. Male genitalia membranous, with TS absent. Ovipositor variable in length, margins not coarsely serrated (except Caimanellus and Lorieriella gen. nov.)............................................................................... 6</p><p>– Last abdominal tergite medially constricted at the posterior margin. Epiproct conspicuous, ovoid, with rounded posterior margin. Male cerci simple and armed with a distal spine. Male genitalia with a submembranous TS, collar-shaped, fused anteriorly and posteriorly, with a dorsal median groove. Ovipositor thin, regularly curved upward in its middle portion, distal half coarsely serrated..................................................................... Cohnia</p><p>6. Color predominantly green, rarely brown, or with conspicuous black stripes. Male cerci simple and cylindrical. Ovipositor as long as or longer than half of the hind femur.......................................................... 7</p><p>– Color grayish-brown, in varying shades (Figs. 17I, 18M). Dorsal surface of abdominal tergites greyish-cream, with triangular and angled lateral edges (Figs. 18N). Male cerci moderately flattened, internally excavated, with two conspicuous triangular branches (Figs. 19N, O). Ovipositor straight, not exceeding half the length of the hind femur, distal third strongly serrated................................................................................ Caimanellus</p><p>7. Auditory spiracle medium size, almost completely covered by the lateral lobe of the pronotum (Figs. 18E, K). Ovipositor exceeding the length of the pronotum by 1.5 times or more, finely to strongly serrated (Figs. 20H, J). Male cerci gradually tapering toward the apex; subgenital plate with a broad V- or U-shaped notch.................................. 8</p><p>– Auditory spiracle conspicuous, protruding below the lateral lobe of the pronotum (Fig. 18I). Ovipositor short, almost as long as the pronotum, finely serrated (Fig. 20D). Male cerci moderately expanded at the apex (Figs. 19D, F); subgenital plate with a shallow, rectangular notch (Fig. 19E)........................... Anisophya (Castellium) subgen. nov.</p><p>8. Pale green or yellowish coloration, without conspicuous lateral stripes; if present, thin and diffused. Tenth abdominal tergite strongly inflated and ventrally deflexed, almost completely concealing the cerci (Fig. 19L). Male cerci with an elongated, medially directed preapical spine (Fig. 19M). Ovipositor with finely denticulated margins distally (Figs. 20H)......................................................................................... Hellerium gen. nov.</p><p>– Green coloration with two conspicuous white lateral stripes extending from the postocular stripe or pronotum to the last abdominal segment (Fig. 17H). Tenth tergite simple, not concealing the cerci (Fig. 19R). Male cerci with short preapical spine (Fig. 19T). Ovipositor strongly serrated (Fig. 20J)................................... Lorieriella gen. nov.</p><p>9. Color variable, mostly brown or green tones, but with conspicuous black stripes. Male cerci with a preapical depression or notch (Figs. 19G, I), sometimes occupying a large portion of the cerci (as in A. una) (Fig. 19K). Ovipositor distinctly shorter, 1.5 times longer than pronotum (Fig. 20F)....................... Anisophya (Brevianisophya) subgen. nov.</p><p>– Color predominantly green. Male cerci without preapical depressions, usually slightly swollen or distally modified (Figs. 19A, C). Ovipositor two to three times longer than the pronotum (Fig. 20B).......... Anisophya (Anisophya) stat. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFFBAFFAB6ADCFB1302C6AC40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFB9FFA46ADCFEE90675AECC.text	BE25B46BFFB9FFA46ADCFEE90675AECC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Angarina	<div><p>Subtribe Angarina subtr. nov.</p><p>Description. Body small and slender. Coloration cryptic, combining black and brown tones with green patches of various shades that facilitate camouflage among bryophytes (Figs. 16A, B, E, I, J). Head rounded; fastigium of the vertex very narrow, about one-third the width of the antennal scape in frontal view. Face black or whitish, varying among species (Figs. 16C, F, K). Thorax. Pronotum longer than high; anterior and posterior margins elevated, especially the posterior one, giving the disc a saddle-like appearance (Fig. 16A) (except in Nanoleptopoda (Figs. 16E, I)). Mesosternum rectangular, almost as wide as high; metasternum as wide as the mesosternum, with rounded and non-divergent lateral margins; sternal grooves short. Wings. Tegmina short, covering between the first and fifth abdominal segments (Figs. 16A, B, I). However, in the known males of Nanoleptopoda, the tegmina fully cover the abdomen (Figs. 16E). Legs. Genicular lobes of the femora, especially the hind femora, armed with sharp spines. Hind tibiae strongly armed, bearing sublamellate dorsal spines (Figs. 16E, I, J). Abdomen. Male terminalia unspecialized; cerci cylindrical, bearing a small inner apical spine. Male subgenital plate flat and unspecialized, with conical styliform processes on the posterior margin (Figs. 16D, H). Ovipositor short, slightly longer than the pronotum, progressively curved from base to apex (Figs. 16I, J). Although, the ovipositor in Angara is nearly straight and more than twice the length of the pronotum.</p><p>Taxa included. Angara Brunner von Wattenwyl, 1891 (type genus), Parangara Rehn, 1945 and Nanoleptopoda Braun, 2011 .</p><p>Distribution. Andes of Ecuador and Peru, and southeastern Brazil (Map 7).</p><p>Comments. The taxa grouped within this subtribe are distinguishable by the aforementioned morphological characters, but most notably, they share a common ecological adaptation: camouflage among bryophytes. Their distribution is primarily Andean, with Parangara and Nanoleptopoda, and two Angara species occurring in southeastern Brazil. The disjunct distribution suggests a formerly broader ancestral range, potentially connecting the central Andes with southeastern Brazil.</p><p>A similar pattern is observed in Machimoides species ( Tettigoniidae: Phaneropterinae: Dysoniini), another genus of phaneropterines adapted for camouflage, this time with lichens (Cadena-Castañeda 2013a, b; Cadena-Castañeda et al. 2022), whose distribution is likewise interrupted in the same geographic region as the taxa in this new subtribe.</p></div>	https://treatment.plazi.org/id/BE25B46BFFB9FFA46ADCFEE90675AECC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFB6FFA66ADCFC630792AC74.text	BE25B46BFFB6FFA66ADCFC630792AC74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisophyina	<div><p>Subtribe Anisophyina subtr. nov.</p><p>Description. Body mostly mid-sized and robust. Coloration predominantly green, although some species may exhibit black stripes or body coloration in varying shades of brown, black, or gray (Fig. 17). Head rounded; face without distinctive color markings. Fastigium of the vertex variable, generally narrower than the scape but not less than half its width in frontal view (Figs. 18G, O). In genera such as Anisophya s.s. and Anisophya (Brevianisophya) subgen. nov., fastigium is broader than the scape (Fig. 18A). Thorax. Pronotum longer than high (Figs. 18C, E, I, K, M), although Lorieriella gen. nov. it is distinctly longer than high (Figs. 18Q, R). Mesosternum rectangular, almost as wide as high; metasternum broader than the mesosternum, with rounded and non-elevated lateral margins; sternal grooves short (Figs. 18B, H, P). Wings. Tegmina short, covering between the first and third abdominal segments (Figs. 18D, F, J, L, N). Legs. Genicular lobes of the femora unarmed or bearing small, acuminate spines. Hind tibiae armed with short dorsal spines. Abdomen. Last abdominal segment usually unmodified (Figs. 19A, D, G), though in some species it is medially constricted or inflated, covering the cerci, as observed in Hellerium gen. nov. (Fig. 19L). Cerci predominantly cylindrical and generally unmodified, although in some Anisophya s.s. species the cerci have distal expansions (Figs. 19A–C); in Anisophya (Brevianisophya) subgen. nov. with a preapical notch (Figs. 19I, K), and Caimanellus, conspicuously bifurcated (Figs. 19N, O). Subgenital plate unmodified and usually bearing styliform projections (Figs. 19B, E, H, O, S). Ovipositor length and shape variable, ranging from as long as the pronotum (Figs. 20D, F) to more than twice its length; margins generally finely serrated (Figs. 20B, H), except in females of Cohnia, Caimanellus, and Lorieriella gen. nov., with conspicuous denticulations (Figs. 20J).</p><p>Taxa included. Anisophya Karabag, 1960 (type genus with three subgenera), Caimanellus Battiston &amp; Fontana, 2024, Cohnia Buzzetti, Fontana &amp; Carotti, 2010, Hellerium gen. nov., and Lorieriella gen. nov.</p><p>Distribution. Mainly in the southern half of South America, ranging from southern Bolivia and southern Brazil to Argentine Patagonia, with some representatives occurring in the Andes between Ecuador and Chile (Map 7).</p><p>Comments. Most of the relevant information concerning this subtribe was discussed in the comments on the tribe Anisophyini trib. nov. As indicated in the keys to subtribes and genera, the main differences between Anisophyina subtr. nov. and Angarina subtr. nov. are outlined, justifying the establishment of these two subtribes to distinguish the taxa and facilitate genus-level identification.</p><p>Additionally, Anisophyina subtr. nov. exhibits a distinct distribution pattern compared to Angarina subtr. nov., being primarily concentrated in the lowlands of southern South America, particularly in the Pampean, Chacoan, and southern Atlantic Forest regions. Few taxa occur in montane environments, such as Cohnia and Hellerium gen. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFFB6FFA66ADCFC630792AC74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFB5FFA76ADCFF0202C6AB45.text	BE25B46BFFB5FFA76ADCFF0202C6AB45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisophya Karabag 1960	<div><p>Anisophya Karabag, 1960 s.s.</p><p>Taxa included. Anisophya (Anisophya) hamata (Giglio-Tos, 1894) (type species), A. (A.) brasiliensis (Brunner von Wattenwyl, 1878), A. (A.) bolivia Gorochov, 2016, A. (A.) borellii (Giglio-Tos, 1894), A. (A.) hemanuelae Fianco, 2023, A. (A.) pulchella (Giglio-Tos, 1894) .</p><p>Comments. This genus was originally described to distinguish some New World species from the predominantly European genus Isophya (Karabag, 1960; Braun, 2010). Later, many short-winged South American species were included in Anisophya (Braun 2010, Nickle 2011, Gorochov 2016, Fianco et al. 2019).</p><p>Upon reviewing the short-winged taxa and comparing the characters useful for genus-level differentiation, Anisophya and other genera are herein delimited. Anisophya is distinguished from different South American genera by its broad fastigium of the vertex, in contrast to the narrower fastigium observed in other taxa. The only comparable taxon in this regard is Anisophya (Brevianisophya) subgen. nov. Males of Anisophya s.s. lack a preapical notch or depression on the cerci, although some species exhibit a distal modification where the cerci is expanded (Figs. 19A– C). This contrasts with other genera derived from Anisophya s.s. and is only comparable to Nickleium gen. nov., although in the latter the cerci are longer, and the subgenital plate projects between them, a character not observed in Anisophya s.s. species. The ovipositor is elongate, two to three times longer than the pronotum (Figs. 20B), and differs from species now placed in other subgenera but formerly included in Anisophya . Only Hellerium gen. nov. also has a long ovipositor, but it is characterized by a basal modification on the ventral valve in the form of a plate that projects downward; this is not the case in Anisophya s.s.</p><p>The combination of these characters allows for a clear delimitation and diagnosis of Anisophya s.s., thus justifying the establishment of additional genera and subgenera, which are described in the present contribution. A key to distinguish the Anisophya s.s. species is provided below. A. pulchella is excluded, as its type specimens are subadult and come from various localities in Argentina and Paraguay, making its taxonomic status doubtful; it is possibly an immature form of another species.</p><p>Key to Known Anisophya s.s. species</p><p>1. Male cerci with the distal portion moderately flattened, expanded, and obliquely truncated at the apex............... 2</p><p>– Male cerci not flattened or distally expanded; apex rounded or obtuse......................................... 3</p><p>2. Lateral lobes of the pronotum wider than high; ventral and posterior margins rounded. Apex of the male cerci fish-tail shaped, with a small spine on the inner projection. Ovipositor gradually curved upward........... A. (A.) hemanuelae</p><p>– Lateral lobes of the pronotum nearly as high as wide; ventral and posterior margins nearly straight and obliquely curved. Apex of the male cerci hook-shaped, subtriangular, with a reduced spine on the inner triangular projection. Ovipositor almost straight......................................................................... A. (A.) hamata</p><p>3. Epiproct smooth, without keel or projections. Male cerci gradually tapering toward the apex. Female subgenital plate triangular, almost as long as wide..................................................................... 4</p><p>– Epiproct with a rather long and high median keel in the middle part (Fig. 19A). Male cerci with an inflated, almost globular apical part, slightly curved medially (Fig. 19C). Female subgenital plate trilobate.................... A. (A.) bolivia</p><p>4. Males with a faint white stripe along the dorsal margins of the body, without scattered spots. Vein pattern and surface of the head and pronotum smooth. Apex of cerci rounded......................................... A. (A.) brasiliensis</p><p>– Males with a white and a black stripe on each side of the body, and white spots scattered across the body. Venation and surface texture of the head and pronotum rugose. Apex of the cercus obtuse......................... A. (A.) borellii</p></div>	https://treatment.plazi.org/id/BE25B46BFFB5FFA76ADCFF0202C6AB45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFB5FFA16ADCF9E202C6A814.text	BE25B46BFFB5FFA16ADCF9E202C6A814.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisophya (Brevianisophya) Cadena-Castañeda 2026	<div><p>Anisophya (Brevianisophya) subgen. nov.</p><p>Type species. Anisophya (Brevianisophya) arreguii (Delaloye &amp; Braun, 2023) stat. nov. here designated.</p><p>Description. Body robust,small to medium-sized(12–18mm)(Figs.17A–B).Coloration variable,predominantly green with brown and black tones. Some species are entirely green and lack dorsal stripes, although faint whitish lateral stripes may be present; portions of the femora often show black or brown punctuations, less frequently on the tibiae (mainly in A. (B.) arreguii stat. nov.). Conspicuous black stripes are also observable, especially in A. (B.) melanochloris stat. nov., or the body may be predominantly brown and black, as in A. (B.) una stat. nov. Head rounded; fastigium of the vertex broad, 1.5 to 2 times wider than the scapus in frontal view; eyes subcircular and moderately prominent (Fig. 18A). Thorax. Pronotum lacking lateral carinae or humeral notch (Fig. 18C); lateral margins of the pronotal disc parallel, without medial constrictions; anterior and posterior margins nearly straight (Fig. 18D). Lateral lobes almost as tall as wide, with posterior and ventral margins uniformly rounded (Fig. 18C). Wings. Tegmina not extending beyond the second abdominal segment; hind wings absent (Figs. 17A–B). Legs. Fore coxae unarmed; tympana open on both sides of the fore tibia; apical spurs on all tibiae unmodified. Hind femora noticeably surpass the apex of the abdomen. Abdomen. Last abdominal tergite unmodified, posterior margin straight or with a slight medial undulation (Fig. 19G, J). Cerci cylindrical, with a preapical notch or depression of varying degrees, ranging from narrow (Fig. 19I) to broad and occupying most of the cercus, comma-shaped (Fig. 19K). Subgenital plate rectangular, longer than wide, with a shallow notch on the posterior margin, mostly rectangular or U-shaped; stylli absent, only a stilliform process present (Fig. 19H). Genitalia membranous and symmetrical (Figs. 19W, X).</p><p>Female. Similar in general appearance and size to the male, differing mainly in sexual characters (Fig. 17D). Tegmina of similar length to those of males, partially overlapping one another (Fig. 17E). Cerci conical and pubescent; epiproct triangular, almost as wide as long, with a rounded apex. Ovipositor distinctly shorter, 1.5 times the length of the pronotum or one-third the length of the hind femur, curved upward, with finely serrated margins and acute apex from which the upper valve protrudes (Fig. 17F). Subgenital plate triangular, as wide as it is long.</p><p>Species included. Anisophya (Brevianisophya) arreguii (Delaloye &amp; Braun, 2023) stat. nov., A. (B.) melanochloris (Rehn, 1911) stat. nov., and A. (B.) una (Fianco, Faria &amp; Braun, 2019) stat. nov.</p><p>Distribution. Southeastern Brazil, Paraguay, Uruguay, and the northeastern and Pampas regions of Argentina (Map 7).</p><p>Etymology. The name of the subgenus refers to the group of Anisophya species with short ovipositors. The gender of this name is being established as femenine.</p><p>Comparison. Anisophya (Brevianisophya) subgen. nov. is more similar to Anisophya s.s. than to the other genera or subgenera of Anisophyina subtr. nov. The new subgenus differs from the other genera and subgenera in the subtribe for its wide fastigium and a distinctive configuration of the male and female terminalia. Anisophya (Brevianisophya) subgen. nov. resembles Anisophya s.s. in the structure of the terminalia. Still, males differ in that the new genus has a preapical depression or notch, sometimes occupying a large portion of the cerci (Figs. 19I, K). This character is absent in Anisophya s.s. males, whose cerci lack preapical depressions and are usually only slightly swollen or modified distally (Fig. 19C). The cerci of A. (A.) hammata and A. (A.) hemanuelae may be somewhat flattened distally, but they do not possess a preapical notch. Females of Anisophya (Brevianisophya) subgen. nov. have a short ovipositor, only slightly longer than the pronotum (Fig. 20F), in contrast to the females of Anisophya s.s., whose ovipositor is two to three times longer than the pronotum (Fig. 20B). Other genera with which Anisophya (Brevianisophya) subgen. nov. could be compared is Nickelium gen. nov. and Anisophya (Castellium) subgen. nov., but its differentiation is discussed in the description of each new taxa.</p><p>Key to Known Anisophya (Brevianisophya) subgen. nov. Species</p><p>1. Color predominantly green, with or without stripes of different colors. Male cerci with a small preapical notch, occupying only the preapical section............................................................................ 2</p><p>– Color brown and black; males with a yellowish dorsal band across the abdominal tergites. Male cerci with a broad notch occupying most of the inner margin, giving the cercus a comma-shaped appearance in dorsal view.. A. (B.) una stat. nov.</p><p>2. Body with a black stripe on the dorsal surface of the abdomen, as wide as the pronotal disc; this stripe may also cover the pronotal disc and vertex in some specimens, although coloration is variable. Males with a gland-like structure on the first abdominal tergite. Male cerci with obliquely truncated apex, bearing two processes, and a central notch..................................................................................... A. (B.) melanochloris stat. nov.</p><p>– Body lacking black dorsal stripes. Males without gland-like structures on the first abdominal tergites. Male cerci with a single inner distal spine; apex of cerci rounded....................................... A. (B.) arreguii stat. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFFB5FFA16ADCF9E202C6A814	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFB3FFA36ADCFA5A0262AEE9.text	BE25B46BFFB3FFA36ADCFA5A0262AEE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisophya (Castellium) Cadena-Castañeda 2026	<div><p>Anisophya (Castellium) subgen. nov.</p><p>Type species. Anisophya (Castellium) punctinervis (Stål, 1861) stat. nov.</p><p>Description. Body slender and small (9–11 mm) (Fig. 17E). Coloration green with white stripes along the sides of the body, extending from the postocular stripe to the last abdominal segments, including the costal margin of the tegmina. Another white stripe outlines the boundary between the abdominal tergites and sternites ventrally. Scape, pedicel, and at least the first segments of the antennal flagellum are greenish-white, becoming pale greenish-brown distally. The rest of the body, including legs, is covered with small white spots or punctuations. Head rounded; fastigium of the vertex slightly narrower than the first antennal segment in both frontal and dorsal views (Fig. 18G); eyes circular and prominent. Thorax. Pronotum lacking lateral carinae or humeral notch; lateral margins of the pronotal disc parallel, without medial constrictions; anterior and posterior margins nearly straight (Figs. 18I, J). Lateral lobes wider than tall, with posterior and ventral margins uniformly rounded (Fig. 18I). Auditory spiracle conspicuous, largely exposed at the posterior edge of the lateral lobe of the pronotum (Fig. 18I). Wings. Tegmina not extending beyond the second abdominal segment; hind wings absent (Fig. 18J). Legs. Fore coxae unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora noticeably surpass the abdominal apex. Abdomen. Last abdominal tergite unmodified, posterior margin nearly straight (Fig. 19D). Male cerci cylindrical, with a moderately expanded apex bearing a small inner spine (Figs. 19D, F). Subgenital plate rectangular, longer than wide, with a shallow rectangular notch on the posterior margin; stylli absent, only a conical stilliform process present (Fig. 19E). Genitalia membranous and symmetrical (Figs. 19U, V).</p><p>Female. Similar in general appearance and size to the male (Fig. 17F). Stripe along the dorsal margins of the body thinner than in males, present only from the postocular stripe through the lateral margins of the pronotum and along the costal margin of the tegmina (Fig. 20C). Cerci conical, slender, and elongate; epiproct triangular, almost as wide as long, apex rounded. Ovipositor short, nearly as long as the pronotum, curved upward, and finely serrated (Fig. 20D). Subgenital plate triangular, as wide as long.</p><p>Species included. Only the type species.</p><p>Distribution. Uruguay, southeastern Brazil, and northeastern Argentina (Map 7).</p><p>Etymology. Named in honor of my dear friend and Argentine orthopterist Lucas Emanuel Castelli. The gender of this name is being established as neuter.</p><p>Comparison. Anisophya (Castellium) subgen. nov. It is morphologically similar to genera derived from Anisophya s.s., characterised by a narrow fastigium and the absence of a preapical notch on the male cerci. It differs from Nickleium gen. nov. by the absence of a humeral sinus on the pronotum and by lacking a preapical notch on the male cercus.Additionally, the male subgenital plate of Nickleium gen. nov. is apically produced and curved upward, differing from that of Anisophya (Castellium) subgen. nov., which has a subgenital plate that is mid-sized and straight in lateral and frontal views. Although both genera share a similarly short ovipositor, Nickleium gen. nov. has nearly straight margins, especially the lower valve, and the apex ends in a sharp point. In contrast, Anisophya (Castellium) subgen. nov. has upward-curved ovipositor margins, a nearly rounded apex, and a more prominent upper valve.</p><p>Anisophya (Castellium) subgen. nov. also differs from Anisophya s.s. in the width of the fastigium and by having a short, curved ovipositor rather than a long, nearly straight one. Anisophya (Castellium) subgen. nov. and Anisophya (Brevianisophya) subgen. nov. are similar in the shape and size of the female ovipositor. However, Anisophya (Brevianisophya) subgen. nov. has a fastigium 1.5 to over 2 times as wide as the antennal scape, while in Anisophya (Castellium) subgen. nov. the fastigium is narrower than the scape. Furthermore, males of Anisophya (Castellium) subgen. nov. lack the preapical notch and additional projections present in Anisophya (Brevianisophya) subgen. nov. Both sexes of Anisophya (Castellium) subgen. nov. are slenderer and more gracile, while those of Anisophya (Brevianisophya) subgen. nov. are more robust.</p><p>Comments. Superficially, Anisophya (Castellium) subgen. nov. could be confused with Anisophya (Brevianisophya) subgen. nov., especially because species in both subgenera were formerly treated as shortovipositor Anisophya . However, as demonstrated above, comparison of terminalia characters and the shape and width of the fastigium reveals differences that justify separating these species into two distinct subgenera.</p></div>	https://treatment.plazi.org/id/BE25B46BFFB3FFA36ADCFA5A0262AEE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFB1FFBC6ADCFC470785AF54.text	BE25B46BFFB1FFBC6ADCFC470785AF54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hellerium Cadena-Castañeda 2026	<div><p>Hellerium gen. nov.</p><p>Taxa included. Hellerium schoenemanni (Karsch, 1889) comb. nov.</p><p>Description. Body slender and small (10–11 mm) (Fig. 17G). Coloration pale green or yellowish; pronotal disc bordered laterally by a distinct white stripe; thorax and abdomen covered with numerous small reddish punctuations. Head rounded; fastigium of the vertex slightly narrower than the first antennal segment in both frontal and dorsal views; eyes circular and prominent. Thorax. Pronotum lacking lateral carinae and humeral notch (Fig. 18K); lateral margins of the pronotal disc parallel, without median constrictions; anterior and posterior margins nearly straight (Fig. 18L). Lateral lobes wider than tall, with uniformly rounded posterior and ventral margins (Fig. 18K). Auditory spiracle of moderate size, partially exposed at the ventral edge of the pronotal lateral lobe (Fig. 18K). Wings. Tegmina short, not extending beyond the third or fourth abdominal segment (Fig. 18L); hind wings absent. Legs. Fore coxae unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora distinctly surpass the apex of the abdomen. Abdomen. Last abdominal tergite strongly inflated and ventrally deflexed, almost entirely concealing the cerci (Fig. 19L). Cerci with a long, medially directed preapical spine (Fig. 19M). Subgenital plate rectangular and spatulate, with a broad V-shaped notch at the apex; styli absent and without conical, styliform projections.</p><p>Female. Similar in general appearance and size to the male (Fig. 19H). Tegmina ovoid, overlapping dorsally, not surpassing the second abdominal segment. Cerci conical, robust, and short; epiproct triangular, nearly as wide as long, apex rounded. Ovipositor more than twice the length of the pronotal disc, nearly straight, slightly curved upward at the distal half, with finely denticulated margins. The basal ridge of the ventral valve of the ovipositor is triangular and modified (Fig. 19H). Subgenital plate triangular, as wide as long, with a truncate apex.</p><p>Species included. Only the type species.</p><p>Distribution. Central Chile (Map 7).</p><p>Etymology. The genus is dedicated to the orthopterist Klaus-Gerhard Heller, in recognition of his valuable contributions, particularly to the study of phaneropterines in Europe and Africa, as well as his significant work in orthopteran bioacoustics. The gender of this name is being established as neuter.</p><p>Comparison. Hellerium gen. nov. differs from Anisophya (Castellium) subgen. nov. in the females’ long ovipositor, a character that aligns it more closely with Anisophya s.s. species. However, Hellerium gen. nov. is distinguished from Anisophya by its fastigium, which is narrower than the antennal scape. Males of Hellerium gen. nov. are easily separated from those of Anisophya s.s. and Anisophya (Castellium) subgen. nov. by their terminalia: in the latter two subgenera, the cerci lack an elongated preapical spine, and the tenth tergite does not cover most of the cerci.</p><p>Females of Hellerium gen. nov. also differ in a triangular and modified basal ridge on the ventral valve of the ovipositor, this condition is not observed in the other compared genera, where the basal ridge is represented only by a narrow, normally developed fold.</p><p>Comments. The new genus includes H. schoenemanni comb. nov., one of the three Chilean species previously placed in Anisophya, thereby excluding the presence of that genus in Chile. As previously discussed, Hellerium gen. nov. clearly differs from Anisophya . This taxon appears to be the only member of the tribe that has managed to cross to the western side of the Andes, a region where species of the tribes Cosmophyllini and Aniarellini trib. nov., considered by some authors to be the most “primitive” in South America, are more commonly found (Nickle 2011).</p></div>	https://treatment.plazi.org/id/BE25B46BFFB1FFBC6ADCFC470785AF54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFAEFFBC6ADCFD1B0423ABF0.text	BE25B46BFFAEFFBC6ADCFD1B0423ABF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caimanellus Battiston & Fontana 2024	<div><p>Caimanellus Battiston &amp; Fontana, 2024</p><p>Type species. Caimanellus australis Battiston &amp; Fontana, 2024 .</p><p>Redescription. Body moderately robust and small-sized (15 mm) (Fig. 17I). Coloration grayish-brown, with varying shades. Dorsal surface of abdominal tergites grayish-cream, with triangular and angled lateral edges. The body is covered with numerous black punctuations. Head rounded; fastigium of the vertex narrow, about half the width of the scape; eyes subcircular and prominent. Thorax. Pronotum without humeral notch (Fig. 18M); lateral margins of the pronotal disc parallel, well-defined, with a median constriction; anterior and posterior margins nearly straight (Fig. 18N). Lateral lobes wider than tall, with uniformly rounded posterior and ventral margins. Auditory spiracle of moderate size, mostly covered by the ventral edge of the pronotal lateral lobe (Fig. 18M). Wings. Tegmina not extending beyond the third abdominal segment; hind wings highly reduced (Fig. 18N). Legs. Fore coxa unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora surpass the abdominal apex. Abdomen. Abdominal tergites with a small medial pointed projection on the posterior margin. Last abdominal tergite constricted and narrowed medially (Fig. 19N). Epiproct triangular, as wide as long, with a rounded and narrow apex. Male cerci moderately flattened, internally excavated, with two conspicuous triangular branches (Fig. 19N): the inner branch is thinner, longer, nearly straight, and diverges inward at mid-length of the cerci (Fig. 19Q); the outer branch has a small conical projection at the apex on the dorsal margin and a medium-sized, hook-shaped sclerotized spine on the ventral distal margin (Fig. 19P). Subgenital plate rectangular, longer than wide, with a broad and shallow U-shaped notch on the posterior margin; styli absent (Fig. 19O).</p><p>Female. Similar in general appearance and size to the male. Tegmina ovoid, not surpassing the first abdominal segment, partially overlapping. Cerci conical, robust, and short; epiproct triangular, wider than long, with a rounded apex. Ovipositor slightly longer than the pronotum, nearly straight, slightly curved upward at the distal portion, with strongly serrated margins. Subgenital plate rectangular, wider than long, with a truncate apex.</p><p>Species included. Only the type species.</p><p>Distribution. Argentina, the Patagonian provinces of Chubut and Santa Cruz (Map 7).</p><p>Comments. This monotypic genus was recently described and represents the southernmost known Phaneropterinae of the world. The original description included only the female (Battiston &amp; Fontana 2024). Here, the conspecific male is described for the first time, providing additional diagnostic features that reinforce its distinction from other genera of the tribe distributed farther north.</p></div>	https://treatment.plazi.org/id/BE25B46BFFAEFFBC6ADCFD1B0423ABF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFAEFFBD6ADCF97F047DAB38.text	BE25B46BFFAEFFBD6ADCF97F047DAB38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lorieriella Cadena-Castañeda 2026	<div><p>Lorieriella gen. nov.</p><p>Type species. Lorieriella transfuga (Brunner von Wattenwyl, 1878) comb. nov.</p><p>Description Body robust and medium-sized (12–15 mm). Coloration green, with two conspicuous white lateral stripes extending from the postocular stripe or pronotum to the last abdominal segment; a thinner reddish line usually borders these stripes. Vertex and median line of the pronotal disc with a thin white stripe, which may continue dorsally on the abdominal tergites, becoming broader; lower margins of the pronotal lateral lobes also outlined in white. Head rounded; fastigium of the vertex narrow, about half the width of the scape (Fig. 18O); eyes circular and prominent. Thorax. Pronotum subcylindrical, lacking lateral carinae or humeral notch (Fig. 18Q); lateral lobes twice as wide as tall, with uniformly rounded posterior and ventral margins (Fig. 18Q). Auditory spiracle mid-sized and narrow, almost entirely covered by the lateral lobes of the pronotum. Wings. Tegmina not extending beyond the second abdominal segment; hind wings absent (Fig. 18R). Legs. Fore coxae unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora clearly surpass the abdominal apex. Abdomen. Last abdominal tergite unmodified, posterior margin nearly straight (Fig. 19R). Male cerci cylindrical, bearing a short preapical spine (Figs. 19R, T). Subgenital plate rectangular, longer than wide, gradually tapering toward the apex, with a V-shaped notch on the posterior margin; styli absent, with only a conical styliform process present (Fig. 19S). Genitalia membranous and symmetrical.</p><p>Female. Similar in general appearance and size to the male (Fig. 17J). Tegmina rectangular, overlapping dorsally, not surpassing the second abdominal segment (Fig. 20I). Cerci conical, robust, and short; epiproct triangular, nearly as wide as long, apex rounded. Ovipositor longer than the pronotum, nearly straight, slightly curved upward in the distal half, with strongly serrated margins (Fig. 20J). Subgenital plate rectangular, wider than long, with a truncate apex.</p><p>Species included. Only the type species.</p><p>Distribution. Uruguay and Buenos Aires Province, Argentina (Map 7).</p><p>Etymology. The genus is dedicated to my dear friend Estrellita Lorier, in recognition of her kindness and her contributions to the study of Uruguayan grasshoppers. The gender of the name is hereby established as feminine.</p><p>Comparison. Lorieriella gen. nov. is a morphologically well-differentiated genus, easily distinguishable from other genera in the tribe. It differs primarily by its elongated pronotum, twice as long as wide, whereas in most other taxa the pronotum is slightly longer than wide. Additionally, the whitish stripes are conspicuous and more prominent in the new genus; in other genera, these stripes may be absent, moderately developed, or restricted to the lateral margins of the pronotal disc and tergites.</p><p>Another distinguishing character is the ovipositor, with strongly serrated margins, a condition only comparable to Caimanellus and Cohnia, but differing from both by its greater length in Lorieriella gen. nov. The new genus also differs from Caimanellus in its green coloration, in contrast with the grayish-brown body and conspicuous grayish-cream dorsal abdominal stripe of Caimanellus . Furthermore, male cerci in Caimanellus are modified, flattened, and branched, unlike the cylindrical and unbranched cerci of Lorieriella gen. nov.</p><p>Lorieriella gen. nov. also differs from Cohnia in that males of the latter have a conspicuous, ovoid epiproct with a rounded posterior margin, and submembranous tenth tergite in the genitalia, which are absent in the new genus. Pronotal structure, ovipositor morphology, and coloration patterns further differentiate Lorieriella gen. nov. from Cohnia, as previously described.</p><p>Comments. Lorieriella transfuga comb. nov. was originally described under the genus Odontura, being at the time the only species of that genus in South America (Brunner von Wattenwyl 1878). Later, Kirby (1906) transferred it to Odonturella, a placement followed by Pirán (1942). Subsequently, Fontana &amp; Buzzetti (2004) placed the species in Dichopetala, based on pronotal morphology. More recently, Cohn et al. (2014) transferred it to Cohnia, where it has remained until now.</p><p>However, as indicated in the description and comparison, Lorieriella gen. nov. is a well-defined and morphologically distinct genus, justifying its formal recognition. Accordingly, Cohnia now comprises only three species and is restricted to the Andes of Ecuador and Peru.</p></div>	https://treatment.plazi.org/id/BE25B46BFFAEFFBD6ADCF97F047DAB38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFACFFB86ADCFF0202C6AF97.text	BE25B46BFFACFFB86ADCFF0202C6AF97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cosmophyllini Brunner von Wattenwyl 1878	<div><p>Cosmophyllini Brunner von Wattenwyl, 1878</p><p>Description. Body small to medium-sized, mostly slender (Figs. 21A, B, G, K). Head rounded; fastigium of the vertex narrower than the width of the scape and sulcate. Eyes subcircular (Figs. 21C, L) or ovoid (Fig. 21H), not protruding. Thorax. Pronotum with smooth margins, posterior margin rounded; pronotal disc slightly flattened, with lateral carinae weakly defined (Figs. 21D, M) (more distinct in Marenestha (Figs. 21I)). Lateral lobes longer than high; humeral sinus developed (Figs. 21C, H, L). Mesosternum rectangular, four to five times wider than high, narrowing medially at the level of the upper lobes; lateral lobes ovoid with rounded margins. Metasternum rectangular, almost twice as wide as the mesosternum; upper margin not narrowed at the level of the superior lobes; lateral lobes subrectangular, progressively elongate and diagonally arranged, with slightly elevated rounded margins. Sternal grooves short; apophyseal pits close together. Wings. Tegmina usually as long as or slightly longer than the abdomen; only a few species of Nickleium gen. nov. with shortened wings that do not reach half the length of abdomen. In long-winged species, tegmina are either broad and ovate (Figs. 21A, B, G) or narrow and subrectangular (Fig. 21K). Legs medium-sized and slender; hind femora surpass the abdominal apex. Ventral margins unarmed, though a few species may bear small spines on the inner distal third. Fore coxae unarmed; fore tibiae with open tympana. Abdomen. Tenth tergite unmodified, without projections. Males: Epiproct subtriangular with rounded posterior margin. Cerci simple and slender, with an apical spine (in Nickleium gen. nov., a conspicuous preapical depression or notch is present on the inner margin). Subgenital plate longer than wide, progressively narrowing and curving upward at the apex, without styli; posterior margin with a small median notch (Fig. 21E). Females: Epiproct triangular; cerci conical, elongate, and slender. Ovipositor short and broad, slightly longer than the pronotum, curving upward, with finely serrated margins and either rounded or pointed apices (Figs. 21F, J). Subgenital plate usually rectangular, longer than wide, with a median notch on the posterior margin; lateral lobes moderately divergent and rounded.</p><p>Distribution. Chile (Map 8).</p><p>Taxa included. Cosmophyllum Blanchard, 1851 (type genus), Marenestha Brunner von Wattenwyl, 1878, Stenophyllia Brunner von Wattenwyl, 1878, and Nickleium gen. nov.</p><p>Comments. This tribe currently exhibits a disjunct distribution across South America, Madagascar, and eastern Australia. However, in this contribution, its distribution is partially restricted, following in part the original concept proposed by Brunner von Wattenwyl (1878). As a result, the following genera are excluded from Cosmophyllini and reassigned as justified below:</p><p>Engonia Brunner von Wattenwyl, 1878 . Originally included in the Cosmophylla group, this genus is here transferred to the tribe Aniarellini trib. nov. The justification for this reassignment is provided in the comments section of that tribe.</p><p>Paracosmophyllum Brunner von Wattenwyl, 1891, this monotypic genus was the first taxon described outside South America within the Cosmophylla group. Its inclusion was based on similarities with Cosmophyllum, particularly in tegmina and pronotum shape. However, several morphological characters justify its exclusion from Cosmophyllini . For example, the meso- and metasternum are narrow and quadrangular (in contrast with the rectangular and moderately broadened condition in Cosmophyllini), and the hind femur bears small ventral spines along its entire length, absent in the genera retained in Cosmophyllini . These characters suggest a closer affinity with Madagascarantia Massa, 2017, also sharing pronotal shape and tympanal structure, although Paracosmophyllum lacks the anterior coxal spine present in Madagascarantia . Based on these differences, Paracosmophyllum is herein considered unplaced at the tribal level, pending further comparisons with additional taxa to determine its position.</p><p>Ozphyllum Rentz, Su &amp; Ueshima, 2007, that the Australian genus was originally proposed as related to Cosmophyllum, as part of a pattern of biogeographic affinity between Chilean and Australian faunas, similar to that observed in the tribe Coniungopterini ( Conocephalinae) (Rentz et al. 2007). However, after examining specimens of Cosmophyllini and comparing them with Ozphyllum, it is clear that these taxa are not closely related. They differ in the morphology of the sternum and wing venation. Additionally, Ozphyllum exhibits a reduced ovipositor and a spine on the anterior coxa, characters absent in Cosmophyllini . Further distinguishing traits such as the globose head, the fastigium of the vertex, eye shape, and the reduced mirror on the tegmina support the alternative view proposed by Rentz et al. (2007), aligning Ozphyllum more closely with Acripeza Guérin-Méneville, 1832 . Consequently, it suggests transferring Ozphyllum to the tribe Acripezini .</p><p>With the taxonomic changes introduced here, the distribution of Cosmophyllini is now restricted to Chile.A key to the genera included is provided below.</p><p>Key to Cosmophyllini genera</p><p>1. Coloration predominantly in shades of green and yellow; tegmina broad and ovate (Figs. 21A, B, G); ovipositor with rounded apex (Figs. 21F, J).......................................................................... 2</p><p>– Coloration predominantly black, gray, or rarely green; tegmina narrow and subrectangular, with some brachypterous forms (Fig. 21K); ovipositor with pointed apex................................................................ 3</p><p>2. Pronotal humeral sinus well-developed (Fig. 21C); tegmina with a rounded apex, not narrowing; hind wings well-developed, reaching near but not beyond the tegminal apex (Figs. 21A, B); male subgenital plate elongated, projecting upward between the cerci (Fig. 21E)...................................................................... Cosmophyllum</p><p>– Pronotal humeral sinus poorly defined (Fig. 21H); tegmina narrowing apically; hind wings short, not exceeding half the length of the tegmina (Fig. 21G); male subgenital plate short, not curving upward between the cerci....... Marenestha</p><p>3. Body predominantly shiny black, tegmina greenish, hind wings reddish; wings always well-developed, covering all or most of the abdomen (Fig. 21 K); male cerci simple, without modifications............................... Stenophyllia</p><p>– Coloration variable but not shiny (black, yellowish-brown, or green) (Figs. 22A–D); hind wings hyaline; wing development variable, with both brachypterous (Figs. 22C–D) and long-winged forms (Figs. 22A–B); male cerci with a conspicuous preapical depression or notch on the inner margin (Figs. 22E, H)............................ Nickleium gen. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFFACFFB86ADCFF0202C6AF97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFABFFB96ADCFF0202C6AA84.text	BE25B46BFFABFFB96ADCFF0202C6AA84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nickleium Cadena-Castañeda 2026	<div><p>Nickleium gen. nov.</p><p>Type species. Nickleium biforma (Nickle, 2011) comb. nov. here designated.</p><p>Description. Body slender, small to medium-sized (10–15 mm). Coloration variable, with olive-green, yellowish-brown, or black forms occurring within the same species (Figs. 22A, C). The two known species share a conspicuous white stripe along the lower margin of the pronotal lateral lobes. Head ovoid; fastigium of the vertex about as wide as the antennal scape, in contact with the fastigium of the frons; eyes subcircular and moderately prominent. Thorax. Pronotum with weakly defined lateral carinae and a median constriction that can be prominent or slight; anterior margin of the pronotal disc nearly straight, posterior margin convex; humeral notch present. Lateral lobes of the pronotum wider than tall, with posterior and ventral margins uniformly rounded. Wings. Tegmina usually short, not surpassing the fifth abdominal segment (Fig. 22C). However, one species ( N. biforma comb. nov.) has fully developed wings and tegmina that exceed the abdominal apex (Fig. 22A). Legs. Fore coxae armed with a small spine; tympana open on both sides of the fore tibiae; apical spurs on all tibiae unmodified. Hind femora clearly surpass the apex of the abdomen. Abdomen. Last abdominal tergite unmodified, with a nearly straight posterior margin. Cerci cylindrical, with a conspicuous preapical depression or notch on the inner margin (Figs. 22E, H). Subgenital plate rectangular, longer than wide, without styli, apically produced and curved upward (Figs. 22F, I).</p><p>Female. Similar in general appearance and size to the male, differing mainly in sexual characters (Figs. 22B, D). Tegmina similar in length to those of the male, partially overlapping. Cerci conical and pubescent; epiproct triangular, almost as wide as long, with a rounded apex. Ovipositor robust, at most only slightly longer than the pronotal disc; basal half inflated, distal half distinctly flattened, gradually curving upward, and apically pointed; margins smooth to weakly serrated. Modified ridge at base of ventral valve of ovipositor forming a medium-sized fold, gently expanding laterally. Subgenital plate triangular, as wide as long (Figs. 22G, J).</p><p>Species included. Nickleium biforma (Nickle, 2011) comb. nov. and Nickleium careomacula (Nickle, 2011) comb. nov.</p><p>Distribution. Central Chile.</p><p>Etymology. The genus is dedicated to David A. Nickle, an esteemed entomologist who made important contributions to the study of Orthoptera and Thysanoptera. He also described the two species now grouped in this new genus. The gender of the name is hereby established as neuter.</p><p>Comparison. Nickleium gen. nov. differs from Cosmophyllum and Marenestha by its slenderer body, shorter wings (typically), narrow tegmina, and pointed ovipositor apex. In contrast, Cosmophyllum and Marenestha are more robust, with fully developed, broad, ovoid tegmina, and ovipositors with rounded apices. Additionally, male cerci in those two genera are slender and lack a preapical notch.</p><p>Nickleium gen. nov. is more similar to Stenophyllia, as both genera display chromatic variation among individuals, have a slender appearance, and, when tegmina are long, they are narrow. The male subgenital plate in both is similarly shaped and curved. However, Stenophyllia males lack the preapical notch on the cerci, which is a distinctive feature of Nickleium gen. nov.</p><p>In females, the ovipositor is likewise similar in shape, being pointed apically, but in Stenophyllia, it is longer and wider, with margins gradually curving upward, unlike in Nickleium gen. nov. The female subgenital plate in Stenophyllia is rectangular with a notched posterior margin, contrasting with the triangular plate in Nickleium gen. nov.</p><p>Comments. The two species included in this new genus were originally described under Anisophya (Originally placed under Odonturini). Upon reviewing their morphology, it became evident that they differ from the defining characters of Anisophya and related genera, and instead align morphologically with Cosmophyllini, being most closely related to Stenophyllia .</p><p>Key to known species of Nickleium gen. nov. (adapted from Nickle 2011)</p><p>1. Wing development variable, with both brachypterous and fully winged forms (Figs. 22A–D). Pronotal disc constricted at midpoint, then gradually expanding posteriorly; male subgenital plate with a broad U-shaped notch on the posterior margin (Fig. 22F). Basal lobe of ovipositor trapezoidal, its distal ventral corner pointed (Fig. 22G)...... N. biforma comb. nov.</p><p>– Brachypterous; tegmina extending only to the sixth abdominal tergite. Pronotal disc not notably constricted, with lateral margins straight along their entire length; male subgenital plate with a nearly rectangular posterior notch (Fig. 22I). Basal lobe of ovipositor oval, with straight ventral margin and a rounded, non-projecting distal ventral corner (Fig. 22J)................................................................................ N. careomacula comb. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFFABFFB96ADCFF0202C6AA84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFA9FFB56ADCFF0202C6A809.text	BE25B46BFFA9FFB56ADCFF0202C6A809.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aniarellini Cadena-Castañeda 2026	<div><p>Aniarellini trib. nov.</p><p>Redescription. Body small to medium-sized and slender, with some taxa exhibiting an elongated form (Figs. 23A, 23B, 24A, 24D, 24F, 25C, 25D, 25H, 25J). Head rounded, projecting forward at an angle of approximately 30°, orthognathous in type (Figs. 23A, 25K). The lower portion of the head covering the mouthparts progressively narrows. Fastigium of the vertex narrower than the width of the scape and sulcate (Fig. 23C). Eyes rounded or ovoid, ranging from moderately sized to conspicuous. Thorax. Pronotum flat (Figs. 23B, 25B, 26B); lateral lobes distinctly longer than high, closely appressed to the body and not projecting laterally (Figs. 23A, 24A, 24D, 25G). The humeral sinus may be well or poorly defined. Meso- and metasternum narrow, quadrangular, almost as tall as wide, and of similar width; lateral lobes rounded, though in some taxa the lateral lobes of the metasternum taper posteriorly, giving the metasternum a subtriangular appearance. Sternal grooves short; apophyseal pits close together. Wings. Tegmina generally narrow and distinctly surpass the apex of the abdomen (Figs. 23A, 24A, 24D, 24F, 25G, 25H). In some taxa (e.g., Coryphoda, Engonia, Buzzettium gen. nov.), the tegmina are ovoid and extend beyond the abdomen, though not as prominently as in most genera (Figs. 24B, 25J, 25K). In contrast, Xenicola has short wings that do not extend beyond the second or third abdominal tergite (Figs. 25C, D). Legs long, slender, and gracile; hind femora extending beyond the abdominal apex. Fore coxae unarmed; fore tibiae with open tympana.</p><p>Abdomen. Tenth tergite unmodified, lacking projections (Figs. 23D, 25E) (except in Coryphoda and Hyperophora (Figs. 24C, H)). Males: Epiproct subtriangular with a rounded posterior margin. Cerci simple and slender, usually with an apical spine (Figs. 23E, 25E) (in Hyperophora, cerci exhibit distal expansions and branches (Fig. 24H); Buzzettium gen. nov. has a preapical notch and a slender inner apical branch forming a pincer-like structure (Fig. 26C)). Subgenital plate rectangular and nearly straight (not curved or projecting between the cerci), without styli; posterior margin flat or with a shallow median notch (Figs. 23F, 25I). Females: Epiproct triangular; cerci conical, elongated, and slender. The ovipositor occurs in two morphotypes: narrow, slightly compressed and curved, with a rough surface and sharp denticles along the margins (Fig. 24E); or broad, strongly compressed, curved upward at the base, with a smooth surface and finely crenulated edges (Fig. 24G).</p><p>Distribution. Southern half of South America, including Bolivia, Brazil, Paraguay, Uruguay, Argentina, and Chile (Map 9).</p><p>Taxa included. Three subtribes Aniarellina subtrib. nov., Hyperophorina subtr. nov. and Tetanina subtr. nov., ten genera and 39 species.</p><p>Comments. This tribe was originally described by Brunner von Wattenwyl (1888) as the group Aniarae, comprising primarily South American taxa, with Corymeta Brunner von Wattenwyl, 1878 as the only African representative. Only Brunner von Wattenwyl (1878, 1891) conducted comprehensive studies on this group. However, species descriptions for some genera were later provided by Rehn (1907, 1913, 1917) with additional Burgilis and Hyperophora species, and Piza (1950, 1971), describing Brazilian Hyperophora species. The general habitus of several taxa in this group resembles members of the Elimaeini, which, despite not being closely related, may indicate convergent ecological adaptations in different regions of the world.</p><p>Based on a morphological review and redefinition of its taxa, the group is here recognized at tribal level as Aniarellini trib. nov., comprising three subtribes. However, the type genus was changed from Aniara Brunner von Wattenwyl, 1878 to Aniarella Bolívar, 1906 due to homonymy with a coleopteran genus (Bolívar 1906). The tribal name retains the new type genus name, since it is a preoccupied name, so it is treated as a new tribal name, which in essence preserves part of the classification of Brunner von Wattenwyl (1878, 1891), with some adjustments noted below.</p><p>It proposes the exclusion of Corymeta Brunner von Wattenwyl, 1878, from Aniarellini trib. nov., as it does not conform to the diagnostic characters of the tribe. The pronotum of the only known Corymeta species lacks a flat disc and sharply defined lateral margins; instead, the margins are rounded, and a sub-elevated ridge runs from the anterior to posterior edge of the disc. The male terminalia are elongated and modified, a trait uncommon in other Aniarellini trib. nov. genera. Furthermore, the female ovipositor does not conform to either of the two ovipositor morphotypes defined for the tribe, and the subgenital plate is broadly emarginate posteriorly, with two pointed lateral projections, also not observed in female Aniarellini trib. nov.</p><p>The genus Engonia is included in Aniarellini trib. nov. based on its morphological affinity with this tribe, rather than Cosmophyllini, where it was initially placed.The head morphology and orientation are consistent with Aniarellini trib. nov., and the venation, particularly between the costal and subcostal areas, resembles that of Aniarella . The description of Buzzettium gen. nov., which shares venational structure and meso-metasternal morphology with Engonia, further supports this placement and differentiates both genera from those of Cosmophyllini .</p><p>Euxenica Bruner, 1915 is a monotypic genus that has not been recorded since its original description. Based on its morphological characteristics, it should be included within Aniarellini trib. nov. However, according to the original description, it appears to be more closely related to Engonia, particularly due to the shape of the tegmina. Although Euxenica lacks fully developed hind wings (Bruner 1915). Euxenica is include within Aniarellina subtrib. nov. and incorporate it into the following key for the identification of subtribes and genera, based on the characters provided in its generic description.</p><p>Key to Aniarellini trib. nov. subtribes and genera</p><p>1. Predominantly green coloration. Pronotum not subcylindrical; lateral lobes slightly longer than high; tympanal spiracle medium-sized, usually covered by the lateral lobes of the pronotum or only slightly visible under it. Fore tibiae unarmed dorsally. Femora unarmed ventrally, except in Pseudoburgilis, which has small spines along the ventral margin of the hind femur........................................................................................... 2</p><p>– Coloration in dark brown and greyish tones (Figs. 23A, B). Pronotum subcylindrical; lateral lobes much longer than high (Fig. 23A); tympanal spiracle conspicuous and projecting at the posterior corner of the pronotum (Fig. 23A). Fore tibiae densely armed with robust dorsal spines (Fig. 23B). Femora ventrally armed with spines (Fig. 23A) Tetanina subtr. nov. ............................................................................................ Tetana</p><p>2. Head robust; eyes generally ovoid and medium-sized (Figs. 24A, D, F). Antennae without distinctive coloration. Lateral margins of the pronotal disc outlined by a whitish cream line, usually accompanied by a reddish line Hyperophorina subtr. nov. ............................................................................................ 3</p><p>– Head slender; eyes rounded and conspicuous (Figs. 25B, K). Scape and pedicel entirely black, or at least with the ventral half black; first flagellomeres usually black (except in Pseudoburgilis, where only the flagellomeres are black). Lateral margins of the pronotal disc outlined in yellow or white, accompanied by a thinner black stripe, sometimes with two black stripes bordering the lighter band (Figs. 25J, K). Aniarellina subtrib. nov. .................................... 5</p><p>3. Lateral lobes of pronotum at most twice as long as high, commonly 1.5 times as longer than high. Tegmina with parallel anal and costal margins, lacking conspicuous or abrupt dilations. Hind wings projecting beyond the apex of the tegmina. Stridulatory mirror of moderate size................................................................... 4</p><p>– Lateral lobes of pronotum three times as longer than high (Fig. 24A). Tegmina strongly expanded basally; costal margin narrowing rapidly towards the apex; anal margin almost straight; in repose, tegmina appear obliquely curved upward at the distal third (Fig. 24B), and hind wings project near the point where curvature begins. Stridulatory mirror conspicuous and expanded................................................................................ Coryphoda</p><p>4. Hind wings clearly projecting beyond the apex of the resting tegmina (Fig. 24D). Lateral lobes of pronotum quadrangular, nearly as long as high (Fig. 24D). Male cerci unmodified, without distal branches. Ovipositor slender and almost straight, with strongly serrated margins (Fig. 24E)......................................................... Burgilis</p><p>– Hind wings moderately projecting beyond the apex of the tegmina (Fig. 24F). Lateral lobes of pronotum rectangular, wider than high (Fig. 24G). Male cerci with distal modifications, usually with a conspicuous inner notch and apical or lateral projections or expansions (Fig. 24H). Ovipositor moderately thickened, upward-curved, and margins typically finely serrated (Fig. 24G)....................................................................... Hyperophora</p><p>5. Scape and pedicel black or at least with the ventral half black; flagellomeres entirely black or at least the first few segments. Lateral margins of the pronotum outlined by a yellowish or whitish band, accompanied by one or two black stripes. Hind femur unarmed ventrally. Hind wings extending beyond the apex of the tegmina. Tegmina consistently shaped, not noticeably narrowing at the apex; radial veins usually bifurcated; spurious vein absent. Hind wings translucent........ 6</p><p>– Scape and pedicel greenish, lacking black markings; flagellomeres black. Lateral margins of the pronotum outlined by a whitish band, without additional black stripes, but with a dark green stripe instead (Fig. 25B). Hind femur ventrally armed with medium-sized spines. Tegmina covering the hind wings, apex narrowing and pointed; radial veins disjoint, not bifurcated; spurious vein present between radial and cubital veins (Fig. 25A). Hind wings opaque and pinkish (Fig. 25A)...................................................................................... Pseudoburgilis</p><p>6. Tegmina extending beyond the apex of the abdomen. Lateral margins of the pronotal disc well defined, never denticulate. Ovipositor upward-curved, not exceeding half the length of the hind femur, usually one-third its length.............. 7</p><p>– Tegmina shorter than the pronotum, scale-like, not reaching halfway along the second abdominal tergite (Figs. 25C, D). Lateral margins of the pronotal disc angular and well defined, denticulate in some species. Ovipositor rectangular in lateral view, nearly straight, about half or more the length of the hind femur (Fig. 25F).......................... Xenicola</p><p>7. Tegmina rectangular, extending beyond the apex of the hind femur (Figs. 25G, H). Posterior margin of the male subgenital plate straight or with a shallow notch (Fig. 25I)................................................... Aniarella</p><p>– Tegmina ovoid or lanceolate, not extending beyond the apex of the hind femur (Figs. 25J, K). Posterior margin of the male subgenital plate with a moderately deep and narrow notch, bearing well-defined styliform projections............... 8</p><p>8. Hind wings present and fully developed, slightly or even greatly longer than the tegmina......................... 9</p><p>– Hind wings absent or rudimentary, tegmina as long as the abdomen and acuminate....................... Euxenica</p><p>9. Pronotum with a well-defined humeral notch (Fig. 26A). Male cerci cylindrical, with a preapical notch and a slender inner branch at the apex, forming a pincer-like structure (Fig. 26C)............................... Buzzettium gen. nov.</p><p>– Pronotum lacking a distinct humeral notch (Figs. 25J, K). Male cerci tapering distally and curved at the tip, without modifications or accessory branches............................................................. Engonia</p></div>	https://treatment.plazi.org/id/BE25B46BFFA9FFB56ADCFF0202C6A809	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFA7FFB56ADCFA3D07BAAA93.text	BE25B46BFFA7FFB56ADCFA3D07BAAA93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetanina	<div><p>Subtribe Tetanina subtr. nov.</p><p>Description. Body medium-sized and slender. Coloration in dark brown and grayish tones (Figs. 23A, B). Head elongated and moderately projected forward; fastigium of the vertex very narrow, acuminate but not sulcate. Eyes globose and conspicuous (Fig. 23C). Thorax. Pronotum subcylindrical; lateral lobes much longer than high (Figs. 23A, B). Tympanal spiracle conspicuous and projects at the posterior corner of the pronotum (Fig. 23A). Wings. Tegmina narrow with nearly parallel margins, slightly narrowing toward the apex and terminating in a pointed tip (Fig. 23A). Legs. Fore tibiae densely armed with robust dorsal spines (Fig. 23B); femora ventrally armed with spines (Fig. 23A). Abdomen. Male terminalia without specializations (Figs. 23D–F). Female ovipositor elongated, longer than the pronotum, and nearly straight.</p><p>Included taxa. Tetana Brunner von Wattenwyl, 1878 (type genus).</p><p>Distribution. Originally recorded from Chile, with more recent records from Argentina, particularly in the Northeast and Cuyo regions (Map 9).</p><p>Comments. This group is among the few known phaneropterines bearing rows of spines along the dorsal margin of the fore tibiae. Similarly, the presence of spines on the ventral margin of the femora is uncommon within the subfamily, a trait otherwise only observed in the tribe Dysoniini in the Neotropics (Cadena-Castañeda et al. 2022).</p></div>	https://treatment.plazi.org/id/BE25B46BFFA7FFB56ADCFA3D07BAAA93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFA4FFB76ADCFAEF06E9A8E3.text	BE25B46BFFA4FFB76ADCFAEF06E9A8E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperophorina	<div><p>Subtribe Hyperophorina subtr. nov .</p><p>Description. Body elongated, medium-sized, and slender (Figs. 24A, D, F). Predominantly green coloration, though some individuals may exhibit ochre tones; antennae without distinctive coloration. Lateral margins of the pronotal disc outlined by a whitish cream line, usually accompanied by a reddish line. Head elongated and moderately projected forward; fastigium of the vertex narrow and acuminate. Eyes generally ovoid and medium-sized (Figs. 24A, F). Thorax. Pronotum flattened; lateral lobes almost twice as long as high (in Coryphoda, they may be even longer). Tympanal spiracle medium-sized, generally covered by the lateral lobes of the pronotum or only slightly visible beneath them. Wings. Tegmina narrow (Figs. 24D, F) (in Coryphoda, they are usually broadened at the base (Fig. 24B)); hind wings extend beyond the apex of the tegmina. Legs. Femora unarmed ventrally, and tibiae lack conspicuous dorsal spines. Abdomen. Male terminalia without specializations, although some Hyperophora males exhibit dorsal projections arising from the last two tergites, and the cerci show distal modifications (Fig. 24H). The female ovipositor is variable: in Hyperophora and Coryphoda, it is curved and finely denticulate (Fig. 24G); in Burgilis, it is narrow, nearly straight, and strongly serrated along the margins (Fig. 24E).</p><p>Taxa included. Hyperophora Brunner von Wattenwyl, 1878 (type genus), Burgilis Stål, 1873, and Coryphoda Brunner von Wattenwyl, 1878 .</p><p>Distribution. Southern half of South America (Map 9).</p><p>Comments. Hyperophora is the most speciose genus, with 15 described species and the broadest distribution. Coryphoda, with only two species, is restricted to central Chile, while Burgilis, with four species, has only been recorded from Argentina.</p></div>	https://treatment.plazi.org/id/BE25B46BFFA4FFB76ADCFAEF06E9A8E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFA5FFB16ADCFA490451AF70.text	BE25B46BFFA5FFB16ADCFA490451AF70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aniarellina Cadena-Castañeda 2026	<div><p>Subtribe Aniarellina subtrib. nov.</p><p>Description. Body medium-sized and slender. Coloration predominantly green. Scape and pedicel entirely black or, at minimum, with the ventral half black; the first flagellomeres are typically black (except in Pseudoburgilis, where only the flagellomeres are black). Lateral margins of the pronotal disc are bordered by a yellow or white band, usually accompanied by a thinner black stripe, and sometimes by two black stripes bordering the lighter band. Head slender and moderately projected forward; fastigium of the vertex narrow and acuminate. Eyes rounded and conspicuous. Thorax. Pronotal disc flat; lateral lobes quadrangular, nearly as tall as wide (Figs. 25G, H, J) ( Xenicola has lateral lobes that are wider than tall, and some species exhibit denticulations along the margins of the pronotal disc (Figs. 25C, D)). Tympanal spiracle medium-sized, generally covered by the lateral lobes of the pronotum or only slightly visible beneath them. Wings. Tegmina follow two main patterns: rectangular in Aniarella (Fig. 25G, H), and ovoid or lanceolate (Figs. 25J, K) in Engonia, Euxenica, and Buzzettium gen. nov. In Xenicola, the wings are short and do not extend beyond the second abdominal segment (Figs. 25C, D). The hind wings are developed in most species, although rudimentary in Euxenica, and pinkish in color in Pseudoburgilis (Figs. 25A). Legs. Femora unarmed ventrally; tibiae lack conspicuous dorsal spines. Only Pseudoburgilis presents small spines along the ventral margin of the hind femur. Abdomen. Male terminalia without specializations (Figs. 25E, I) ( Buzzettium gen. nov. has cerci with a preapical notch and a slender inner branch at the apex, forming a pincer-like structure (Fig. 26C)). Female ovipositor narrow, slightly compressed and curved, with small denticles along its margins (Figs. 25L). In Xenicola, the ovipositor is elongated and nearly straight (Figs. 25D, F).</p><p>Included taxa. Aniarella Bolívar, 1906 (type genus), Engonia Brunner von Wattenwyl, 1878, Euxenica Bruner, 1915, Pseudoburgilis Brunner von Wattenwyl, 1878, Xenicola Uvarov, 1940, and Buzzettium gen. nov.</p><p>Distribution. Southern half of South America, including Bolivia, Brazil, Uruguay, Paraguay, and Argentina (Map 9).</p><p>Comments. Most taxa are concentrated in the Atlantic Forest and Pampas regions of Brazil, Uruguay, and Argentina. Records from outside these regions are scarce and include Buzzettium borelli comb. nov., Euxenica aptera Bruner, 1915, and Aniarella minor Bruner, 1915 .</p></div>	https://treatment.plazi.org/id/BE25B46BFFA5FFB16ADCFA490451AF70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFA3FFB26ADCFDFE06A8AE50.text	BE25B46BFFA3FFB26ADCFDFE06A8AE50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Buzzettium Cadena-Castañeda 2026	<div><p>Buzzettium gen. nov.</p><p>Type species. Buzzettium borellii (Giglio-Tos, 1897) comb. nov. here designated.</p><p>Description. Body slender and small-sized (13 mm) (Figs. 26A, B). Predominantly green coloration, with antennal and pronotal color patterns typical of the subtribe Aniarellina subtrib. nov.: scape and pedicel mostly black with a narrow dorsal greenish stripe, similarly colored first flagellomeres. A postocular band consists of a thin whitish line above a broader black band, both extending along the lateral margins of the pronotal disc. Ventral margin of the femoral apex, upper half of the tympanum, and ventral surface of all tarsomeres black. Head ovoid; fastigium of the vertex narrow, in contact with the fastigium of the frons; eyes circular and moderately prominent. Thorax. Pronotum with well-defined lateral carinae (Fig. 26A); anterior margin of the pronotal disc nearly straight, posterior margin convex (Figs. 26B); humeral notch present. Lateral lobes of the pronotum wider than high, ventral margin straight, posterior margin slightly rounded, almost straight (Figs. 26A). Wings. Tegmina lanceolate, extending beyond the abdomen but not surpassing the apex of the hind femur; costal margin curved from the mid-length onward, narrowing toward the apex; anal margin nearly straight (Figs. 26A). Hind wings well developed and projecting beyond the tegmina apex. Legs. Fore coxae unarmed; tympana open on both sides of the fore tibiae; apical spurs of all tibiae unmodified. Hind femora clearly surpass the abdominal apex. Abdomen. Last abdominal tergite unmodified, with a nearly straight posterior margin. Cerci cylindrical, with a conspicuous preapical notch or depression on the inner margin; after the notch, a short cylindrical extension is present just before the apex (Fig. 26C). Subgenital plate rectangular, with medium-sized, styliform, non-articulated projections, and a shallow median notch between them.</p><p>Female. Unknown.</p><p>Species included. Only the type species.</p><p>Distribution. Andean foothills in Jujuy Province, northern Argentina, near the border with Bolivia (Map 9).</p><p>Etymology. This genus is dedicated to the Italian entomologist Filippo Maria Buzzetti, who contributed to Orthoptera and Heteroptera. He also made several contributions to different groups of short-winged phaneropterines. The gender of the name is hereby established as neuter.</p><p>Comparison. Buzzettium gen. nov. resembles Engonia and Euxenica primarily in the shape and length of the tegmina, differing from Aniarella, which has rectangular tegmina in males (broader in females) that extend well beyond both the abdominal apex and the hind femur. In contrast, the other three genera ( Engonia, Euxenica, and Buzzettium gen. nov.) exhibit lanceolate to ovoid tegmina that moderately surpass the abdomen but not the apex of the hind femur. The new genus differs from Euxenica in having fully developed hind wings, whereas Euxenica lacks them. Compared to Engonia, although both have developed hind wings, those of Engonia do not extend beyond the tegmina apex, unlike those in Buzzettium gen. nov.</p><p>A further distinctive character separating Buzzettium gen. nov. from other Aniarellina subtrib. nov. members is the morphology of the male cerci. Other genera in the subtribe lack cercal modifications or branching; their cerci are cylindrical, tapering toward the apex, and usually curving inward at the tip. In contrast, the male cerci of Buzzettium gen. nov. exhibit a preapical inner notch preceded by a thickened section, followed by a short cylindrical projection just before the apex. This cercal structure is only comparable to taxa such as Anisophya (Brevianisophya) subgen. nov. ( Anisophyini trib. nov.) and Nickleium gen. nov. ( Cosmophyllini), which also possess a preapical notch, although narrower and less pronounced. Furthermore, those two genera lack the preapical projection present in Buzzettium gen. nov.; instead, they bear a small, sclerotized apical spine.</p><p>Buzzettium gen. nov. differs from Hyperophora in characters that distinguish the subtribes Hyperophorina subtr. nov. and Aniarellina subtrib. nov., such as eye shape, characteristic antennal coloration, and pronotal disc margins. Species of Hyperophora are typically slender and elongated, with ovoid eyes, and very long, narrow legs and wings. Although both genera have modified cerci, their structure differs: Hyperophora lacks a preapical notch and exhibits two general patterns: either a medially thickened, inward-curved apex forming an ovoid mass, or an internal branch near mid-length that extends and tapers toward the apex.</p><p>Comments. The only known species of Buzzettium gen. nov. was originally described under the genus Hyperophora . In the description of B. borellii comb. nov., it was compared with Hyperophora minor Brunner von Wattenwyl, 1891, based on similarities in cerci shape (Giglio-Tos 1897). However, both species differ in generic and subtribal characters, as highlighted above. While superficially similar, the cerci differ in shape and length: the basal thickening in the new genus is unlike that in H. minor, where it is represented as a basal branch. In H. minor, there is no true notch, but rather a continuous tapering and inward-curving structure, without the accessory preapical projection seen in B. borellii comb. nov. Additionally, the spiniform extension of the last abdominal segments, common in several Hyperophora species, is absent in B. borellii comb. nov.</p><p>These comparisons and remarks support the generic distinctiveness of Buzzettium gen. nov., which shows closer affinity to genera of the subtribe Aniarellina subtrib. nov. Therefore, we establish the new combination: Buzzettium borellii comb. nov.</p></div>	https://treatment.plazi.org/id/BE25B46BFFA3FFB26ADCFDFE06A8AE50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
BE25B46BFFA1FFCD6ADCFF02055EAE85.text	BE25B46BFFA1FFCD6ADCFF02055EAE85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenicola Uvarov 1940	<div><p>Xenicola Uvarov, 1940 stat. rev.</p><p>Comments. Xenicola was originally described as Xenica Brunner von Wattenwyl, 1891, within the Odonturae group (Brunner von Wattenwyl 1891). The change in the generic name occurred because the name had already been used for a lepidopteran (Uvarov, 1940). Its initial inclusion in Odonturae was based on the presence of short wings in its species (Brunner von Wattenwyl, 1891). However, having short wings does not necessarily imply a close relationship within a particular group of phaneropterines, as demonstrated throughout this contribution. Wing reduction is a condition that has independently arisen multiple times in unrelated taxonomic groups.</p><p>A recent and noteworthy study, by Fianco &amp; Melo (2025), offered a molecular systematics analysis of short-winged neotropical katydids. Their phylogenetic reconstruction yielded interesting insights, the most striking of which, from my perspective, was the synonymization of Xenicola under Angara . From a morphological standpoint, although both genera share short wings, they are distinct enough to warrant placement in different tribes, as its propose here.</p><p>Specifically, the synonymy of Xenicola under Angara is not supported by morphological compatibility. In the resulting tree from Fianco &amp; Melo (2025), the genus appears composed of two main clades. In clade B, the type species Angara albofasciata Brunner von Wattenwyl, 1891, and the morphologically similar Angara nigrocercata Fianco, 2025, cluster together, and among them are other species that are more consistent with the morphology of Xenicola .</p><p>Therefore, the two species we consider to belong to Angara are nested within a group of species that we attribute to Xenicola . This leads me to suspect that, in the absence of clear morphological correlation, there may have been a labelling error with the samples of A. albofasciata and A. nigrocercata, and that these sequences may in fact correspond to one or more species of Xenicola rather than to those two specific species.</p><p>Based on the arguments above, its propose the revalidation of the genus Xenicola stat. rev., with the following species included:</p><p>Xenicola superba (Brunner von Wattenwyl, 1891) comb. reinst. (type species).</p><p>Xenicola brauni Fianco, Engelking &amp; Faria, 2020 comb. reinst.</p><p>Xenicola dohrni (Brunner von Wattenwyl, 1891) comb. reinst.</p><p>Xenicola nunoi Fianco, 2023 comb. reinst.</p><p>Xenicola taroba Fianco, Faria &amp; Braun, 2019 comb. reinst.</p><p>Xenicola xukrixi Fianco, Faria &amp; Braun, 2019 comb. reinst.</p><p>Xenicola caayari (Fianco, 2025) comb. nov.</p><p>Xenicola flavolineata (Fianco, 2025) comb. nov.</p><p>Xenicola rubra (Fianco, 2025) comb. nov.</p><p>It is important to emphasize that the contribution by Fianco &amp; Melo (2025) is both valuable and informative. Any potential error should not undermine the value of their work, which was exhaustive and comprehensive, providing critical insights into the phylogenetic placement of short-winged South American katydids in relation to both short- and long-winged phaneropterines worldwide. I agree with their conclusion regarding the relationship between Xenicola (treated as Angara in their analysis) and the genera Aniarella and Engonia, which supports our decision to include Xenicola in the tribe Aniarellini trib. nov.</p><p>Short-winged taxa not included in the traditional or derived tribes</p><p>Comments. After reviewing and classifying most of the genera included in some of the short-winged phaneropterine tribes such as Odonturini, Barbitistini, and related groups, three genera still require comment or clarification regarding their tribal placement: Brinckiella Chopard, 1955, Austrodontura Fontana &amp; Buzzetti, 2004, and Paraperopyrrhicia Ebner, 1915 .</p><p>1. Brinckiella is a South African katydid genus originally described as monotypic and placed in the subfamily Meconematinae (Chopard 1955) . Nearly four decades later, Naskrecki (1996) transferred it to Phaneropterinae under the tribe Barbitistini . Subsequently, Naskrecki &amp; Bazelet (2009) redescribed the genus and described seven additional species, reinforcing its inclusion in Phaneropterinae . Recently, Heller &amp; Hemp (2024) proposed a provisional reassignment back to Meconematinae, based on the unusual morphology of the eggs. It is suggested here to retain Brinckiella within Meconematinae until more robust evidence justifies a different placement.</p><p>2. Austrodontura, another short-winged phaneropterine genus from South Africa, was established for the type species originally described as Odontura capensis Walker, 1869, based on a female with a prominently serrated ovipositor (Walker 1869). More than a century later, Fontana &amp; Buzzetti (2004) erected the genus Austrodontura, transferring O. capensis as Austrodontura capensis and describing an additional species, Austroduntura raggei Fontana &amp; Buzzetti, 2004. Later, Naskrecki &amp; Bazelet (2011) redescribed the genus, synonymized the two previously known species under A. capensis, and described an additional species, A. castletoni, including the song record.</p><p>Currently, Austrodontura is unassigned to any tribe (Cigliano et al. 2025). However, its morphological traits do not align with those of traditional tribes such as Odonturini or Barbitistini, nor with the derived African tribe Monticolariini trib. nov. Instead, it shares several characters with members of Phaneropterini and Ducetiini . The shape of the head, eyes, and ovipositor structure are particularly reminiscent of Ducetiini, especially Ducetia Stål, 1874 . Nevertheless, it differs in having short wings and a subgenital plate without a deep emargination or curved cerci, which are typical of Ducetiini . For the moment, keep as incertae sedis.</p><p>3. Paraperopyrrhicia is another short-winged genus currently placed in Odonturini . It includes a single species, Paraperopyrrhicia papua Ebner, 1915 (Fig. 27), described from New Britain Island, Papua New Guinea. This genus is peculiar in that it is the only short-winged katydid in the region with an elongated and strongly serrated ovipositor. Most phaneropterines in these islands have fully developed wings and reduced ovipositors. Since its original description, P. papua has not been collected again, and the male remains unknown, limiting the availability of diagnostic characters for proper tribal placement. The single known female displays very short tegmina with thick, widely spaced veins, unlike other short-winged phaneropterines, which typically exhibit thin and reticulate venation. Based on current evidence, it is clear that Paraperopyrrhicia does not belong to Odonturini . However, due to the lack of additional specimens and information, it is best regarded as incertae sedis within Phaneropterinae until further data become available.</p><p>The inclusion and analysis of these last three genera complete the treatment of all taxa historically assigned to either Odonturini or Barbitistini .</p></div>	https://treatment.plazi.org/id/BE25B46BFFA1FFCD6ADCFF02055EAE85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cadena-Castañeda, Oscar J.	Cadena-Castañeda, Oscar J. (2026): Studies on Phaneropterinae: The short-winged bush katydids, Odonturini sensu lato, Barbististini and their allies (Orthoptera: Tettigoniidae). Twelfth contribution to the suprageneric organization of phaneropterines. Zootaxa 5762 (1): 1-82, DOI: 10.11646/zootaxa.5762.1.1, URL: https://doi.org/10.11646/zootaxa.5762.1.1
