identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B9168023FFAFFF9DD760FD5B7C769854.text	B9168023FFAFFF9DD760FD5B7C769854.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aeduella Westoll 1937	<div><p>Genus Aeduella Westoll, 1937 sensu Gonçalves &amp; Luccisano (2025b)</p><p>TYPE SPECIES. — Palaeoniscum blainvillei Agassiz, 1833 by monotypy.</p></div>	https://treatment.plazi.org/id/B9168023FFAFFF9DD760FD5B7C769854	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gonçalves, Daniel	Gonçalves, Daniel (2026): Paleontological content of the Stephano-Permian basins from the Armorican Massif (France): review, actinopterygian taxonomy, paleobiogeography and correlations. Geodiversitas 48 (19): 463-497, DOI: 10.5252/geodiversitas2026v48a19
B9168023FFAFFF81D742FC147D5E9CF1.text	B9168023FFAFFF81D742FC147D5E9CF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aeduella blainvillei (Agassiz 1833)	<div><p>Aeduella blainvillei (Agassiz, 1833) sensu Gonçalves &amp; Luccisano (2025b)</p><p>(Figs 4H; 7; 8)</p><p>Palaeoniscum blainvillei Agassiz, 1833: 4, 48-55, pl. V, figs 1-7.</p><p>Paramblypterus cf. decorus – Racheboeuf &amp; Crasquin 2012: 230, fig. 158.</p><p>Aeduella blainvillei – Gonçalves &amp; Luccisano 2025b: 17-26 (last diagnosis).</p><p>TYPE MATERIAL. — Lectotype. France • 1 specimen (almost complete skeleton); Saône-et-Loire, Dracy-Saint-Loup, Muse; Muse Formation, Grès et Schistes de Muse Member, Muse OSB; Early Asselian, 298.57 ± 0.16 Ma; UNISTRA [UNISTRA.2013.0.133] . Paralectotypes. France • 6 specimens (complete to subcomplete skeletons); Saône-et-Loire, Dracy-Saint-Loup, Muse; Muse Formation, Grès et Schistes de Muse Member, Muse OSB; Early Asselian, 298.57 ± 0.16 Ma; MNHN [MNHN.F.AUT25, AUT676, AUT704], SU [SU.PAL.2017.0.1.16], UNISTRA [UNISTRA.2013.0.135, UNISTRA.2013.0.136] .</p><p>LECTOTYPE. — An almost complete skeleton (UNISTRA.2013.0.133).</p><p>PARALECTOTYPES. — Six complete to subcomplete skeletons (MNHN.F.AUT25, AUT676, AUT704, SU.PAL.2017.0.1.16, UNISTRA.2013.0.135, UNISTRA.2013.0.136), followingBrignon (2019).</p><p>TYPE LOCALITY AND HORIZON. — Muse, Dracy-Saint-Loup, Saône-et-Loire, France (Muse Formation, Grès et Schistes de Muse Member, Muse OSB). Early Asselian, 298.57 ± 0.16 Ma (Pellenard et al. 2017; Mercuzot et al. 2023).</p><p>NEWLY REFERRED SPECIMENS. — Two complete skeletons in a single slab (ML-PAL-0451; Figs 4H; 7; 8).</p><p>NEW LOCALITY AND HORIZON. — Les Effretais (precise layer unknown, Saint-Pierre-la-Cour Formation, Saint-Pierre-la-Cour Basin). Stephanian C regional substage (Vetter 1957), early to middle Gzhelian (Aretz et al. 2020).</p><p>REMARKS</p><p>This slab was donated to the Musée de Laval by Louis Saminn (1816-1887), former director of the coal mines at Saint-Pierre-la-Cour. It may have been discovered during the mining of Les Effretais mine, which was in operation during the second half of the 19 th century (e.g., Étienne &amp; Limasset 1979).</p><p>COMPARATIVE DESCRIPTION</p><p>General body shape</p><p>The ML-PAL-0451 slab contains two specimens; ML-PAL-0451a of 13.2 cm in body length and its posterior-ventral skull covered by a coprolite; and ML-PAL-0451b (5.7 cm preserved body length) with almost all its caudal fin covered by ML-PAL-0451a (Fig. 4H, 7A, B). Complete ossification of the skull bones, short lepidotrichia articles, and deep body indicate that both specimens are skeletally mature (Growth Stage 3, following Gonçalves &amp; Luccisano 2025b). The size and body length/depth ratio (3.2) of ML-PAL-0451a are typical of derived aeduellids, such as Aeduella blainvillei, Bourbonnella fourrieri Poplin, 2001, B. guilloti Heyler, 1967, B. hirsuta Štamberg, 2007, Decazella vetteri Heyler, 1967, Neslovicella rzehaki Štamberg, 2007, and Westollia crassa (Pohlig, 1892) (Heyler 1969; Poplin 2001; Štamberg 2007, 2024; Gonçalves &amp; Luccisano 2025b).</p><p>Rostral region</p><p>The rostral region of ML-PAL-0451a is poorly preserved and absent in ML-PAL-0451b. A possible fragment of the ventral part of the rostral and a preorbital are present in ML-PAL-0451a.</p><p>The V-shape of the ventral edge of the rostral of ML-PAL-0451a is found in other aeduellids (Fig. 8A, B) (e.g. Heyler 1969; Štamberg 2007, 2010, 2018, 2024; Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2025, 2026c).</p><p>The preorbital of ML-PAL-0451a is ornamented with long dorsoventral ridges, a characteristic unique to Aeduellidae but invariable within the family (Fig. 8A, B; e.g., Heyler 1969; Heyler &amp; Poplin 1983; Štamberg 2007, 2010, 2018, 2024; Gonçalves &amp; Luccisano 2025b).</p><p>Skull roof</p><p>The skull roof of ML-PAL-0451a is well preserved in its posterior region, and the bones sutures are not preserved in ML-PAL-0451b.</p><p>The parietal is elongated with straight lateral and medial margins and sigmoid posterior margins (Fig. 8A, B). This morphology is the most common among aeduellids (e.g., Heyler 1969; Štamberg 2007; Gonçalves &amp; Luccisano 2025b), but differs from the broad parietal of Westollia crassa and Puertollanichthys ritchiei Forey &amp; Young, 1985 and from the parietal with lobed interparietal suture in Otovice aeduellid (Forey &amp; Young 1985; Štamberg 2021, 2024).</p><p>The postparietal of ML-PAL-0451a and ML-PAL-0451b is rectangular (length/width ratio of 2.1) with straight lateral, medial and posterior margins (Fig. 8). Its anterior margin is oblique, like that of some specimens of Aeduella blainvillei, Bourbonnella guilloti and B. hirsuta (Fig. 8A, B; Heyler 1969; Štamberg 2007, 2018). It differs from the postparietal with a V-shaped ventral margin of Amelangia ornata Štamberg &amp; Werneburg, 2023 and Platysella lallyi Heyler &amp; Poplin, 1983 (Heyler &amp; Poplin 1983; Štamberg &amp; Werneburg 2023). The postparietals of ML-PAL-0451a and ML-PAL-0451b have only two and one pit lines, respectively. The presence of two pit lines, the medial and posterior ones, is a diagnostic character of Aeduellidae (e.g., Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2026b). The absence of the posterior pit line in ML-PAL-0451b is also observed in certain specimens of aeduellids and constitutes an intraspecific variation (Fig. 8C, D; e.g., Heyler 1969).</p><p>The triangular intertemporo-dermosphenotic of ML-PAL-0451a is similar to that of other aeduellids (Fig. 8A, B; e.g., Heyler 1969; Štamberg 2007, 2010, 2024; Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2026b, c), except that of ‘ Neslovicella ’ elongata Štamberg, 2010, which is more crescent-shaped (Štamberg 2010). The anterior part of the intertemporo-dermosphenotic is not preserved in ML-PAL-0451a, and this bone is not distinct in ML-PAL-0451b.</p><p>The supratemporal is complete in ML-PAL-0451a but its margins are not distinct in ML-PAL-0451b (Fig. 8). It is rectangular (length/width ratio of 2.0), with straight medial, lateral and posterior margins, and a V-shaped anterior margin. The supratemporal pit line is present in ML-PAL-0451a, and common among aeduellids (Fig. 8A, B; e.g., Heyler 1969; Gonçalves &amp; Luccisano 2025b). The supratemporal of ML-PAL-0451a is distinct from the supratemporal with M-shaped anterior margin of Decazella vetteri (Gonçalves &amp; Luccisano 2025b), and similar to the supratemporal of other aeduellids (e.g., Heyler 1969; Forey &amp; Young 1985; Štamberg 2007, 2010, 2024; Štamberg &amp; Werneburg 2023; Gonçalves et al. 2026b, c).</p><p>The extrascapulars are preserved in ML-PAL-0451a and indistinct in ML-PAL-0451b. They are arranged in a mediolateral band, with at least three extrascapulars in each side: one medial and two lateral extrascapulars (Fig. 8A, B). Aeduella blainvillei is the only aeduellid to possess six or more extrascapulars (Heyler 1969). The most common arrangement in aeduellids is the presence of four extrascapulars (Heyler 1969; Štamberg 2007, 2010, 2024; Gonçalves et al. 2026c).</p><p>The course of the sensory canals is not visible in ML-PAL-0451a and poorly defined in ML-PAL-0451b: its supraorbital canal runs along the lateral margin of the parietal in (Fig. 8C, D). The presence of a pit line in the supratemporal of ML-PAL-0451a indicates a supraorbital canal course from the parietal to the supratemporal (Fig. 8A, B). This is characteristic of aeduellids (Heyler 1969; Štamberg 2007; Gonçalves &amp; Luccisano 2025b).</p><p>Circumorbital series</p><p>The circumorbital bones of ML-PAL-0451a-b are poorly preserved: fragments of ventral infraorbitals are present in ML-PAL-0451b while the dorsalmost posterior infraorbital and the jugal of ML-PAL-0451a are present. Therefore, the number of ventral infraorbitals and posterior infraorbitals cannot be counted and compared with aeduellids.</p><p>The ventral infraorbitals are narrow, rectangular elongated antero-posteriorly,and without ornamentation (Fig.8C, D). The infraorbital canal runs along the ventral infraorbitals margins.</p><p>The jugal is L-shaped and unornamented, but its margins are not preserved in ML-PAL-0451a,making unclear its exact shape and relationships with the posteriormost ventral infraorbital and ventralmost posterior infraorbital (Fig. 8A, B). The infraorbital canal runs along the middle of the jugal, following the bone curvature.Numerous branches of this canal are regularly positioned, starting from the main infraorbital canal branch to the posterior and ventral margins of the jugal. While the ambiguous shape of the jugal of ML-PAL-0451a does not allow comparisons, its sensory canal disposition is characteristic: Aeduella blainvillei is the only aeduellid with numerous branches of the infraorbital canal regularly positioned, starting from the main infraorbital canal branch to the posterior and the ventral margin of the jugal (Heyler 1969).</p><p>Cheek bones</p><p>The cheek bones of ML-PAL-0451b are indistinct, and the preoperculum is the only preserved cheek bone in ML-PAL-0451a. The preoperculum of ML-PAL-0451a reach around the fifth of the operculum depth, as in Aeduella blainvillei, Amelangia ornata, B.hirsuta, Franchessella pohlcremeri Gonçalves, Lemaître, Štamberg,Logghe,Rebillard, Pouillon, Fernandez&amp; Steyer,2026 and Neslovicella spp. (Fig. 8A, B; Heyler 1969; Štamberg 2007, 2010, 2018; Štamberg &amp; Werneburg 2023; Gonçalves &amp; Luccisano 2025b).The preopercular canal runs dorso-ventrally along the posterior margin of the preoperculum.The preoperculum of ML-PAL-0451a has a bending angle of 156° and an inclination of 52°. This inclination is low because aeduellids usually have an inclination of around 80°, but not uncommon (e.g., Štamberg 2010; Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2026b).</p><p>Jaws</p><p>The jaw bones are the best preserved skull elements in ML-PAL-0451b and poorly preserved in ML-PAL-0451a. Their maxilla is triangular, with a curved ventral margin and tubular teeth (Fig. 8). This morphology is a synapomorphy of Aeduellidae (Heyler 1969; Štamberg 2007; Gonçalves et al. 2026b). The complete maxilla of ML-PAL-0451b has a length/height ratio of 3.0 (Fig. 8C, D). Most of the specimens of A. blainvillei have maxillae with length/height ratio of 3.0-3.5, as in Amelangia ornata, Platysella spp. and Stephaniella melanocephala Gonçalves, Pouzadoux &amp; Luccisano, 2026 (Heyler 1969; Heyler &amp; Poplin 1983; Štamberg 2018; Štamberg&amp;Werneburg 2023; Gonçalves et al. 2026c).</p><p>A dentalosplenial is preserved in ML-PAL-0451b, but its posterior part is overlapped by the maxilla (Fig. 8C, D). It is elongated antero-posteriorly, with the ventral margin curved, as in other aeduellids (e.g., Heyler 1969; Štamberg 2007, 2010, 2024; Gonçalves &amp; Luccisano 2025b).</p><p>Opercular series</p><p>The opercular bones of ML-PAL-0451b are poorly defined. In ML-PAL-0451a, they are incompletely preserved – the branchiostegal rays and the medial gular are not preserved – and bear ornamentation made of flat tubercles. This ornamentation is identical compared to other aeduellids, excepted with Amelangia ornata which bears pointed tubercles (Štamberg &amp; Werneburg 2023).</p><p>The operculum of ML-PAL-0451a is slightly elevated (depth/ length of 1.6) with a rounded dorsal margin, an oblique ventral margin, and a bending angle of 126° (Fig. 8A, B).This morphology is common among aeduellids (e.g., Heyler 1969; Heyler &amp; Poplin 1983; Štamberg 1986, 2007, 2010, 2018, 2024; Gonçalves et al. 2026c).</p><p>The suboperculum of ML-PAL-0451a is trapezoidal, with a larger posterior margin: the posterior margin/anterior margin depth ratio is 1.8 (Fig. 8A, B). The suboperculum is lower than the operculum, with an operculum/suboperculum depth ratio of 1.5. Among aeduellids, Aeduella blainvillei, Otovice taxon, Puertollanichthys ritchiei, Stephaniella melanocephala, Spinarichthys dispersus (Fritsch, 1895) and Westollia crassa also possess trapezoidal subopercula with conspicuously sloping dorsal margin (e.g., Heyler 1969; Štamberg 2007, 2018, 2021, 2024; Gonçalves &amp; Luccisano2025a, 2025b; Gonçalves et al. 2026c). The posterior margin/anterior margin depth ratio of 1.8 of ML-PAL-0451a falls within the range of that of A. blainvillei, Otovice taxon, P. ritchiei and W. crassa (Heyler 1969; Štamberg 2018, 2021, 2024; Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2026c).</p><p>The lateral gular of ML-PAL-0451a is ovoid and slightly rhombic, narrowing anteriorly at the middle of the anterior margin (Fig. 8A, B). This morphology is unique to Aeduella blainvillei (Heyler 1969; Štamberg 2018; Gonçalves et al. 2025; Gonçalves &amp; Luccisano 2025b). Gular elements of ML-PAL-0451b are preserved but indistinct (Fig. 8C, D).</p><p>Pectoral girdle</p><p>The pectoral girdle of ML-PAL-0451a is not preserved, and poorly defined in ML-PAL-0451b, the posttemporal being very incomplete and the cleithrum being the only bone with well-defined outlines</p><p>This cleithrum is ovoid and has a large ridge that runs dorso-ventrally and separates into the lateral and ventral branches (Fig. 8C, D). This morphology is shared by all aeduellids (e.g., Heyler 1969; Štamberg 2007, 2010, 2024; Štamberg &amp; Werneburg 2023; Gonçalves &amp; Luccisano 2025b).</p><p>Fins</p><p>Like almost all aeduellids, the fins of ML-PAL-0451a and ML-PAL-0451b are composed of lepidotrichia with sigmoidal sutures, which divide dichotomously (Fig. 7; e.g., Heyler 1969; Heyler &amp; Poplin 1983; Forey &amp; Young 1985; Štamberg 1986, 2007, 2010, 2024; Poplin &amp; Dutheil, 2005). The pectoral fins are not preserved in ML-PAL-0451a and ML-PAL-0451b,and a large portion of the caudal fin of ML-PAL-0451b is overlapped by the body of ML-PAL-0451a.Dorsal and anal fin bases of ML-PAL-0451a are located on a field of minute scales (Fig. 7C, D). These minute scales are not observable in ML-PAL-0451b, but this absence may be linked to the poor preservation.</p><p>The pelvic fins are triangular, with a leading edge length/base ratio of 1.6-1.8 and around 22 lepidotrichia (Fig. 7A, B). It is located on 4 scale rows in the middle of the body, between the pectoral and anal fins.</p><p>The anal fin is triangular with a leading edge length/base ratio of 1.0-1.3 (Fig. 7A, B, D). The anal fin has 20 lepidotrichia and its base sits on 9 scale rows. The base of the anal fin is located just after the posterior end of the base of the dorsal fin. The dorsal fin is also triangular, with a length/base ratio of 1.0-1.5 and is composed of 26 to 28 lepidotrichia (Fig. 7 A-C). Its base sits on 10-11 scale rows and is located in the middle of the body length.</p><p>The paired and unpaired fins shape and size of ML-PAL-0451a and ML-PAL-0451b are similar to those of the other aeduellids (e.g., Heyler 1969; Heyler &amp; Poplin 1983; Poplin 2001; Štamberg 2007, 2024; Štamberg &amp; Werneburg 2023; Gonçalves &amp; Luccisano 2025b).</p><p>The caudal fin of ML-PAL-0451a is large (caudal fin/body length ratio of 3.1) and heterocercal, with a dorsal lobe/ventral lobe length ratio of 1.5 (Fig. 7A). It differs from the caudal fin of Decazella vetteri, Bourbonnella hirsuta and ‘ Neslovicella’ elongata in which the ventral lobe is almost as elongated as the dorsal lobe (Štamberg 2007, 2010; Gonçalves &amp; Luccisano 2025b). The ventral lobe of the caudal fin of ML-PAL-0451a has 12 lepidotrichia and its dorsal lobe 44 preserved lepidotrichia, the dorsalmost lepidotrichia being missing. Its caudal inversion is gradual as in Aeduella blainvillei, Franchessella pohlcremeri, Platysella lallyi and P. descusi Heyler &amp; Poplin, 1983 (e.g., Heyler 1969, 2002; Heyler &amp; Poplin 1983; Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2026b).</p><p>Squamation</p><p>The body of ML-PAL-0451a and ML-PAL-0451b is covered with large, rhombic smooth scales (Fig. 7A, B). The lateral sensory canal crosses the body through its middle and extends into the caudal inversion. The posterior edge of the posterior flank scales is straight. The posterior margin of the scales is always broken in the anterior flank scales. Presence or absence of serration on these scales cannot be therefore assessed. The squamation of ML-PAL-0451a and ML-PAL-0451b is identical to that of most of the aeduellids, being different from the ornamented scales of Franchessella pohlcremeri (Gonçalves et al. 2026b). The scale count of ML-PAL-0451a is:</p><p>23 ––––––––––– 38 8 23 34</p><p>The scale count of ML-PAL-0451b is:</p><p>21 –––––––––––? 8 21 34</p><p>These scale counts are within the range of Aeduella blainvillei, Decazella vetteri, Neslovicella rzehaki, and Stephaniella melanocephala (Heyler 1969; Štamberg 2007; Gonçalves &amp; Luccisano 2025b; Gonçalves et al. 2026c). The number of scales rows at the anal fin base is higher than in Puertollanichthys ritchiei, and that at the dorsal fin base is lower than in Platysella lallyi (Heyler &amp; Poplin 1983; Forey &amp; Young 1985). The scale counts of ML-PAL-0451a and ML-PAL-0451b are higher than in Platysella descusi (Heyler &amp; Poplin 1983), and lower than in Amelangia ornata, Bourbonnella spp., Franchessella pohlcremeri, Platysella poplinae and Westollia crassa (Heyler 1969, 2002; Poplin 2001; Štamberg 2007, 2024; Štamberg &amp; Werneburg 2023; Gonçalves et al. 2026b).</p><p>Ridge scales are poorly preserved in ML-PAL-0451a and ML-PAL-0451b. Few are present in front of the dorsal fin base, and two large scutes are located before the ventral lobe of the caudal fin (Fig. 7A, B). The dorsal lobe of the caudal fin is protected by numerous fulcral scales. These specialized scales of ML-PAL-0451a and ML-PAL-0451b are typical of aeduellids (e.g., Heyler 1969; Poplin 2001; Štamberg 2007, 2024; Gonçalves &amp; Luccisano 2025b)</p></div>	https://treatment.plazi.org/id/B9168023FFAFFF81D742FC147D5E9CF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gonçalves, Daniel	Gonçalves, Daniel (2026): Paleontological content of the Stephano-Permian basins from the Armorican Massif (France): review, actinopterygian taxonomy, paleobiogeography and correlations. Geodiversitas 48 (19): 463-497, DOI: 10.5252/geodiversitas2026v48a19
