taxonID	type	description	language	source
BA2F8796FF83FF8B9E8FFCA12D8EB879.taxon	description	Redescription of Tsivoka Villiers, 1892 (Revised from Villiers et al. 2011): Head with an elongate and subparallel rostrum. Genae extended beyond the lower lobe of the eyes, which are finely faceted, narrow and weakly indented. Frons almost flat, divided by a dorsal furrow that extends from the inter-antennary area up to the level of the posterior edge of the eyes, short and subparallel temples. Mandibles large, falcate, inner margin sinuate. Maxillary and labial palps densely covered with fine setae. Antennae inserted in front of the eyes, closer to them than to the base of the mandibles; base of antennae prominent and close together; scape long, curved at apex; antennomere III slightly shorter than scape, subequal to IV; V and the following ones longer than IV, and projected at the external apical angle from VI to X. Strongly convex pronotum with barely marked disc tubercles, the two medians close together, often obsolete, the two posterior ones further apart and a more evident; anterior margins and posterior margins strongly depressed, finely rimmed; almost straight posterior half of sides, strongly convergent forward, this line break forming an obtuse angle. Scutellum as long as it is wide. Elytra strongly narrowed from anterior to posterior, humeri slightly prominent in males, more strongly so in females; apex truncate. Fusiform femora; straight tibiae, metatibiae widened at apex and, in males, with a small process at the end of the inner edge; protarsomeres strongly dilated in males; metatarsomeres I as long or longer than the metatarsomeres II-V together; onychium strongly dilated and toothed in males. Tergite VII of males subrectangular, longer than wide; sternite VII with a wide but shallow indent; sternite VIII short and deeply indented. Apically lobed tegmen very long, arched, and pubescent. Male genitalia of the same type as in Mastododera.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF83FF8B9E8FFCA12D8EB879.taxon	discussion	Remarks: Tsivoka has been observed on the flowers of Mimosa latespinosa Lamarck, 1806 (Fabaceae) in bright sunlight (Villiers et al. 2011).	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFEB5296CBA26.taxon	description	Redescription (translated from Villiers et al. 2011): Length 15 – 22 mm. Same proportions and structure as in T. simplicollis (Fig. 7) but differs in having less dense elytral pubescence and in coloring: head, pronotum, and scutellum orange, with short pubescence; yellow antennae, covered with fine golden pubescence; yellow ocher elytra, with golden pubescence, lateral slope and a medial suture with black band: yellow forelegs; midlegs and hindlegs with black femora and yellow base, yellow tibiae, and tarsi; ventral side dark yellow ocher.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFEB5296CBA26.taxon	materials_examined	Material examined: Madagascar Est Belalona, 30 km S. O. Sambava, 11.1969, Vadon et Peyrieras, male (holotype, MNHN) (Fig. 1 b)	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFEB5296CBA26.taxon	distribution	Distribution: Northeastern Madagascar lowland forests (Fig. 8). Ten specimens from MNHN all have the same locality information.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFC492D85BE2F.taxon	description	Redescription (translated from Villiers et al. 2011): Length 19 – 25 mm. Black head with red labrum and palps. Red antennae. Black pronotum with prostrate pubescence. Black scutellum. Elytra black with a narrow yellow band, clearly defined, extending from the shoulder almost to the apex; the yellow band with golden pubescence, and the black area with silvery pubescence. Red forelegs. The mid and hind legs black, tarsi red. Ventral surface black with fine silvery pubescence. Head densely punctate; genae one and a half times longer than the lower eye lobe. Frons approximately one and a half times (male) to two times (female) wider than an eye seen from a dorsal view. Antennae extend beyond the apex of the elytra (male) or only reach the level of their apical quarter (female). Pronotum approximately as long as wide at the base (male) or transverse (50 / 60) (female), with slightly convex sides on their posterior half, very strongly converging forward on their anterior half; very convex disc, with very weak tubercles; collar and basal margins strongly depressed, clearly delimited. Elytra narrow, 2.8 times as long as they are broad when held together (closed) at the base, with strongly protruding shoulders; male tibia with a small dentiform process at the apex of the inner surface.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFC492D85BE2F.taxon	materials_examined	Material examined: Madagascar, Forêt Didy, Antsevabe, I. 2000, E. Vives, male (CEV) (Fig. 7 a); Madagascar, Antananarivo Province, Manankazo, 26. XI. 2002, I. Jenis, female (CEV) (Fig. 7 b); Madagascar, Toamasina, Tamatave, 9. XI. 2012, P. Senft, male (CEV); Madagascar, Moramanga, Alaotra Mangoro, IX. 1991, local collector, female (CEV).	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFC492D85BE2F.taxon	distribution	Distribution: Eastern, northern, and southern Madagascar (Fig. 8). Tsikova simplicicollis occurs in subhumid lowland forests and spiny thickets.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF82FF8B9E8FFC492D85BE2F.taxon	discussion	Remarks: The morphology of adult, larva, and pupa was examined by Švácha and Lawrence (2014), (Fig. 6).	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF85FF8D9E8FFF752A75BD00.taxon	description	Redescription (revised from Villiers et al. 2011): Length 13 – 22 mm. Light brown to black, red mouthparts, concolorous or elytra with the periscutellar and suture area dark, or dark with a light stripe starting from the humeral depression towards the apex, this band sometimes reduced to a line towards the rear of the elytron, red fore-, mid- and hindlegs, with brown or black femurs, tibias, and red tarsi; black ventral side. Head very finely and densely pitted. Gena is one and a half times longer than the lower eye lobe. Frons twice (male) to just over twice (female) wider than an eye seen from dorsal view. Temple with less setae in females. Antenna orange reaches the level of the apical sixth of the elytra (male) or only the apical third (female). Scape longer than flagellomere 1; flagellomere 3 through 8 sharp outer edges with 2 small pits; Mandible and maxillary palpi reddish brown with dense yellow setae. Pronotum densely punctate, with short pubescence, only partially masking the integument, as long or a little longer than broad at base in both sexes; protrusions obsolete, contiguous discs; oval, oblique basal protuberances, highlighted by a depression along their anterior edge; sides of the pronotum straight and subparallel behind, strongly concave and convergent forward; pronotum lateral tubercles protruded with obtuse angle. Procoxae about each other. Elytra with prominent shoulders, two and a half times (male) to 2.2 times (female) longer than wide together at the base, covered with a lying, silky pubescence which hides the integument, except in the region of the humeral depression where relatively strong and loose punctation is distinguished. End of the inner edge of the male hind tibia with a tiny triangular projection.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF85FF8D9E8FFF752A75BD00.taxon	materials_examined	Material examined: Madagascar, Fianarantsoa, Fitovavy Fitovinany Region, District of Ifanadiana Belle Vue area 1200 m S of Ranomafana National Park entrance, 1018 m, (- 21.2665, 47.42017), VI. 3.2002 – VI. 13.2002, Rin’ha, Mike, female (CAS); Madagascar, Fianarantsoa, Fitovavy Fitovinany Region, District of Ifanadiana, 12 km W of Ranomafana, 1127 m, (- 21.25083, 47.40717), X. 7.2004 – X. 20.2004, Rin’ha, Mike, female (CAS); Madagascar, Toamasina, Analanjirofo Region, District of Toamasina, MOBOT Site, Analalava humid dense forest low altitude on the sand 7 km SW of Foulpointe, 75 m, (- 17.69333, 49.46028), Rin’ha, Mike, female (CAS); Madagascar, Toamasina, Réserve Naturelle Betampona, 34.08 km 332 ° Toamasina, 525 m, (- 17.91977, 49.20039), I. 13.2008 – I. 20.2008, B. L. Fisher, male (CAS); ditto II. 17.2008 – II. 24.2008, B. L. Fisher, female (CAS); Réserve Naturelle Betampona, 34.1 km 332 ° Toamasina, 550 m (- 17.916135, 49.20185), I. 18.2009 – I. 25.2009, B. L. Fisher, female (CAS); ditto, female (CAS).	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF85FF8D9E8FFF752A75BD00.taxon	distribution	Distribution: Eastern and southwestern Madagascar (Fig. 8). T. testaceipes can be found in subhumid, lowland, and dry deciduous forests in Madagascar.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF84FF8F9E8FFB2C2C82BE95.taxon	materials_examined	Type series: HOLOTYPE: Madagascar, Toliara, Atsimo Andrefana Region, District of Betioky, 30 km E Betioky, Beza Mahafaly Special Reserve (Around Research Station), 165 m, (- 23.6865, 44.591), VIII. 16 – VIII. 28.2002, Rin’ha, Mike, male (CAS) (Fig. 3). PARATYPES: Madagascar, Toliara, Atsimo Andrefana Region, District of Betioky, 30 km E Betioky, Beza Mahafaly Special Reserve (Around Research Station), 165 m, (- 23.6865, 44.591), X. 15. – XI. 10.2001, Rin’ha, Mike, male (CAS); ditto, VI. 8 – VI. 18.2002, female (Fig. 4) (CAS); ditto, IX. 9 – IX. 20.2002, male (CAS); Madagascar Ouest Reserve spéciale du Zombitsy Est de Sakaraha Matsabory, 640 m, VII. 10 – 11.1974, P. Viette et A. Peyrieras, female (MNHN) (Fig. 1 a); Madagascar Zombites Vohibasia National Park, 792 m, (- 22.8884, 44.6945), XI. 29.2024, Jeong, male, (DDM)	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF84FF8F9E8FFB2C2C82BE95.taxon	description	Description: 18 – 20 mm. Dark brown to black, orange antennae, elytra with bright orange marks on the humeral area, legs black, red fore femur. Head very finely and densely punctuated. Genea one and a half times longer than the lower eye lobes; covered with brown pubescence. Frons twice (male) to just over twice (female) wider than an eye seen from above. Mandible and maxillary palpi black to dark brown with dense yellow setae. Antennae reaching the apex of elytra in males, and the posterior one third in females. Scape as long as flagellomere 1; flagellomere 3 through 8 sharp outer edges with two small pits; flagellomere 9 longer than flagellomere 7 in males and shorter in females. Pronotum densely punctate, with short pubescence, only partially obliterating the integument, as long or a little longer than broad at base in both sexes; tubercles obsolete and contiguous disc protrusion with medial furrow; oval, oblique basal protuberances; sides of the pronotum straight and subparallel posteriorly, strongly concave and convergent anteriorly; lateral tubercles obsolete but slightly protruded with obtuse angle. Sternum black with yellowish pubescence; mesosternum 1.5 x as broader as prosternum. Procoxae adjacent to each other with a narrow intercoxal area. Elytra with prominent humeri, two and a half times (male) to 2.2 times (female) longer than wide together at the base, covered with a prostrate silky pubescence obliterating the integument, except in the region of the humeral depression; black with bright orange marks on each humeral area. Wings dark brown. Legs dark brown to black except for the reddish-brown front femur. The hind femur is long and slender; as long as the abdomen. Tibiae long and slender; end of the inner edge of the male hind tibiae with a tiny triangular projection with two black spines. Male onychium dilated and toothed. Abdomen black; male tergite 8 orange, laterally narrowed in medial area and round apex; abdominal sternites 3 and 4 similar lengths and sternites 5 and 6 slightly shorter. Male sternite 7 about 1.2 times longer than sternite 3 and abdomen flat in lateral view. Female sternite 7 similar lengths with sternite 3 and abdomen slightly curved between sternite 3 and 4 in lateral view.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF84FF8F9E8FFB2C2C82BE95.taxon	etymology	Etymology: Having a cape in Latin. The name describes the distinctive orange mark on the humeri.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF84FF8F9E8FFB2C2C82BE95.taxon	discussion	Remarks: The species was first recognized as a new species and named by late Dr. Villiers in 1898 based on a single specimen housed at the NMHN. However, it has never been formally described or published (Fig. 1 a). The species is named Tsivoka humeralis sp. nov. in honor of Dr. Villiers with the specimen serving as a paratype. The type specimens from CAS were collected using Malaise traps in southern dry deciduous gallery forests. A male specimen was collected during the daytime (11: 54 pm) following the first two days of rain in the year. The beetle was observed flying across a trail and landed on a branch near fallen wood.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF84FF8F9E8FFB2C2C82BE95.taxon	distribution	Distribution: Southwest Madagascar (Fig. 8). T. humeralis sp. nov. occurs in succulent woodlands and dry deciduous forest in Southwestern Madagascar.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
BA2F8796FF84FF8F9E8FFB2C2C82BE95.taxon	discussion	Discussion: The species is easily distinguished by the unique pattern on the elytra which sets it apart from others in the same genus. Its distribution pattern is also distinctive within the genus. While T. simplicicollis and T. testacipes are widely distributed from the humid Northeastern regions to the drier Southern areas, T. humeralis is exclusively found in the Southwestern succulent woodlands and dry deciduous forests. This differs from T. peyrierasi, which is only observed in the humid lowland forests of the northeastern region.	en	Jeong, Soo-Hyun, Vives, Eduard, Mckenna, Duane D. (2025): A taxonomic revision of the Madagascar-endemic longhorn beetle genus Tsivoka Villiers (Coleoptera: Cerambycidae) with the description of a new species. Zootaxa 5632 (1): 159-170, DOI: 10.11646/zootaxa.5632.1.9, URL: https://doi.org/10.11646/zootaxa.5632.1.9
