taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323687/files/figure.png	https://doi.org/10.5281/zenodo.17323687	FIGURES 1–5. Habitus (1) and microsculpture of head (2), pronotum (3), elytra (4), and abdomen (5) of Scopaeus decipiens species group with example of S. kritschi, Cambodia, Angkor.	FIGURES 1–5. Habitus (1) and microsculpture of head (2), pronotum (3), elytra (4), and abdomen (5) of Scopaeus decipiens species group with example of S. kritschi, Cambodia, Angkor.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323689/files/figure.png	https://doi.org/10.5281/zenodo.17323689	FIGURES 6–9. Stridular file of metaventrite in lateral view (6), stridular files of metaventrite (in sublateral position) in ventral view (7), excerpt of metaventrite with left stridular file (8), mesothoracal leg with plectral ridges (9) of Scopaeus kritschi, Cambodia, Angkor (6–8); S. decipiens, Sri Lanka, Galle (9). Abbreviation: mkp—metakatepisternal process. Scale bars: A— Fig. 9 (excerpt of leg with plectral ridges), B—Fig. 8, C—Fig. 7.	FIGURES 6–9. Stridular file of metaventrite in lateral view (6), stridular files of metaventrite (in sublateral position) in ventral view (7), excerpt of metaventrite with left stridular file (8), mesothoracal leg with plectral ridges (9) of Scopaeus kritschi, Cambodia, Angkor (6–8); S. decipiens, Sri Lanka, Galle (9). Abbreviation: mkp—metakatepisternal process. Scale bars: A— Fig. 9 (excerpt of leg with plectral ridges), B—Fig. 8, C—Fig. 7.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400489/files/figure.png	https://doi.org/10.5281/zenodo.17400489	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323693/files/figure.png	https://doi.org/10.5281/zenodo.17323693	FIGURES 27–32. Aedeagus in lateral (27, 30), ventral (28, 31), dorsal view of (29, 32) of Scopaeus decipiens, lectotype, Sri Lanka (27–29); S. complex, lectoype, Japan (30–32). Abbreviations: al—apical lobes, cf—circoforamen, dl—dorsal lobe, f— flagellum, ll—lateral lobes, mf—median foramen, pf—postforamen, vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 27–32. Aedeagus in lateral (27, 30), ventral (28, 31), dorsal view of (29, 32) of Scopaeus decipiens, lectotype, Sri Lanka (27–29); S. complex, lectoype, Japan (30–32). Abbreviations: al—apical lobes, cf—circoforamen, dl—dorsal lobe, f— flagellum, ll—lateral lobes, mf—median foramen, pf—postforamen, vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323695/files/figure.png	https://doi.org/10.5281/zenodo.17323695	FIGURES 33–41. Aedeagus of Scopaeus complex in lateral (33, 36, 39), ventral (34, 37, 40), dorsal view (35, 38, 41) from Taiwan, Kaohsiung (33–35); Vietnam, Cuc Phuong, Ninh Binh (36–38); China, Zhejiang, Siming Mts (39–41). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 33–41. Aedeagus of Scopaeus complex in lateral (33, 36, 39), ventral (34, 37, 40), dorsal view (35, 38, 41) from Taiwan, Kaohsiung (33–35); Vietnam, Cuc Phuong, Ninh Binh (36–38); China, Zhejiang, Siming Mts (39–41). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323699/files/figure.png	https://doi.org/10.5281/zenodo.17323699	FIGURES 42–47. Aedeagus of Scopaeus saaberae in lateral (42, 45), ventral (43, 46), dorsal view (44, 47); holotype, Malaysia, Selangor, Kuala Lumpur (42–44); paratype, Indonesia, Borneo, Pagate (45–47). Abbreviation: vl—ventral lobe. Arrows: a— apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 42–47. Aedeagus of Scopaeus saaberae in lateral (42, 45), ventral (43, 46), dorsal view (44, 47); holotype, Malaysia, Selangor, Kuala Lumpur (42–44); paratype, Indonesia, Borneo, Pagate (45–47). Abbreviation: vl—ventral lobe. Arrows: a— apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323702/files/figure.png	https://doi.org/10.5281/zenodo.17323702	FIGURES 48–53. Aedeagus of Scopaeus kritschi in lateral (48, 51), ventral (49, 52), dorsal view (50, 53); holotype, Cambodia, Angkor (48–50); paratype, Laos, Khammouane, Nakai (51–53). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 48–53. Aedeagus of Scopaeus kritschi in lateral (48, 51), ventral (49, 52), dorsal view (50, 53); holotype, Cambodia, Angkor (48–50); paratype, Laos, Khammouane, Nakai (51–53). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323704/files/figure.png	https://doi.org/10.5281/zenodo.17323704	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD95729FF47F31EE4AF7620.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400496/files/figure.png	https://doi.org/10.5281/zenodo.17400496	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFDE572AFF47F4C5E5C17632.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323689/files/figure.png	https://doi.org/10.5281/zenodo.17323689	FIGURES 6–9. Stridular file of metaventrite in lateral view (6), stridular files of metaventrite (in sublateral position) in ventral view (7), excerpt of metaventrite with left stridular file (8), mesothoracal leg with plectral ridges (9) of Scopaeus kritschi, Cambodia, Angkor (6–8); S. decipiens, Sri Lanka, Galle (9). Abbreviation: mkp—metakatepisternal process. Scale bars: A— Fig. 9 (excerpt of leg with plectral ridges), B—Fig. 8, C—Fig. 7.	FIGURES 6–9. Stridular file of metaventrite in lateral view (6), stridular files of metaventrite (in sublateral position) in ventral view (7), excerpt of metaventrite with left stridular file (8), mesothoracal leg with plectral ridges (9) of Scopaeus kritschi, Cambodia, Angkor (6–8); S. decipiens, Sri Lanka, Galle (9). Abbreviation: mkp—metakatepisternal process. Scale bars: A— Fig. 9 (excerpt of leg with plectral ridges), B—Fig. 8, C—Fig. 7.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFDE572AFF47F4C5E5C17632.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400489/files/figure.png	https://doi.org/10.5281/zenodo.17400489	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFDE572AFF47F4C5E5C17632.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323693/files/figure.png	https://doi.org/10.5281/zenodo.17323693	FIGURES 27–32. Aedeagus in lateral (27, 30), ventral (28, 31), dorsal view of (29, 32) of Scopaeus decipiens, lectotype, Sri Lanka (27–29); S. complex, lectoype, Japan (30–32). Abbreviations: al—apical lobes, cf—circoforamen, dl—dorsal lobe, f— flagellum, ll—lateral lobes, mf—median foramen, pf—postforamen, vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 27–32. Aedeagus in lateral (27, 30), ventral (28, 31), dorsal view of (29, 32) of Scopaeus decipiens, lectotype, Sri Lanka (27–29); S. complex, lectoype, Japan (30–32). Abbreviations: al—apical lobes, cf—circoforamen, dl—dorsal lobe, f— flagellum, ll—lateral lobes, mf—median foramen, pf—postforamen, vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFDE572AFF47F4C5E5C17632.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323704/files/figure.png	https://doi.org/10.5281/zenodo.17323704	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFDE572AFF47F4C5E5C17632.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400496/files/figure.png	https://doi.org/10.5281/zenodo.17400496	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD05726FF47F48DE3267311.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400489/files/figure.png	https://doi.org/10.5281/zenodo.17400489	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD05726FF47F48DE3267311.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323693/files/figure.png	https://doi.org/10.5281/zenodo.17323693	FIGURES 27–32. Aedeagus in lateral (27, 30), ventral (28, 31), dorsal view of (29, 32) of Scopaeus decipiens, lectotype, Sri Lanka (27–29); S. complex, lectoype, Japan (30–32). Abbreviations: al—apical lobes, cf—circoforamen, dl—dorsal lobe, f— flagellum, ll—lateral lobes, mf—median foramen, pf—postforamen, vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 27–32. Aedeagus in lateral (27, 30), ventral (28, 31), dorsal view of (29, 32) of Scopaeus decipiens, lectotype, Sri Lanka (27–29); S. complex, lectoype, Japan (30–32). Abbreviations: al—apical lobes, cf—circoforamen, dl—dorsal lobe, f— flagellum, ll—lateral lobes, mf—median foramen, pf—postforamen, vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD05726FF47F48DE3267311.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323695/files/figure.png	https://doi.org/10.5281/zenodo.17323695	FIGURES 33–41. Aedeagus of Scopaeus complex in lateral (33, 36, 39), ventral (34, 37, 40), dorsal view (35, 38, 41) from Taiwan, Kaohsiung (33–35); Vietnam, Cuc Phuong, Ninh Binh (36–38); China, Zhejiang, Siming Mts (39–41). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 33–41. Aedeagus of Scopaeus complex in lateral (33, 36, 39), ventral (34, 37, 40), dorsal view (35, 38, 41) from Taiwan, Kaohsiung (33–35); Vietnam, Cuc Phuong, Ninh Binh (36–38); China, Zhejiang, Siming Mts (39–41). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD05726FF47F48DE3267311.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323704/files/figure.png	https://doi.org/10.5281/zenodo.17323704	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD05726FF47F48DE3267311.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400496/files/figure.png	https://doi.org/10.5281/zenodo.17400496	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD25720FF47F6B2E2F576F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400489/files/figure.png	https://doi.org/10.5281/zenodo.17400489	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD25720FF47F6B2E2F576F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323699/files/figure.png	https://doi.org/10.5281/zenodo.17323699	FIGURES 42–47. Aedeagus of Scopaeus saaberae in lateral (42, 45), ventral (43, 46), dorsal view (44, 47); holotype, Malaysia, Selangor, Kuala Lumpur (42–44); paratype, Indonesia, Borneo, Pagate (45–47). Abbreviation: vl—ventral lobe. Arrows: a— apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 42–47. Aedeagus of Scopaeus saaberae in lateral (42, 45), ventral (43, 46), dorsal view (44, 47); holotype, Malaysia, Selangor, Kuala Lumpur (42–44); paratype, Indonesia, Borneo, Pagate (45–47). Abbreviation: vl—ventral lobe. Arrows: a— apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD25720FF47F6B2E2F576F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323704/files/figure.png	https://doi.org/10.5281/zenodo.17323704	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD25720FF47F6B2E2F576F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400496/files/figure.png	https://doi.org/10.5281/zenodo.17400496	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD55723FF47F48DE36873F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323687/files/figure.png	https://doi.org/10.5281/zenodo.17323687	FIGURES 1–5. Habitus (1) and microsculpture of head (2), pronotum (3), elytra (4), and abdomen (5) of Scopaeus decipiens species group with example of S. kritschi, Cambodia, Angkor.	FIGURES 1–5. Habitus (1) and microsculpture of head (2), pronotum (3), elytra (4), and abdomen (5) of Scopaeus decipiens species group with example of S. kritschi, Cambodia, Angkor.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD55723FF47F48DE36873F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323689/files/figure.png	https://doi.org/10.5281/zenodo.17323689	FIGURES 6–9. Stridular file of metaventrite in lateral view (6), stridular files of metaventrite (in sublateral position) in ventral view (7), excerpt of metaventrite with left stridular file (8), mesothoracal leg with plectral ridges (9) of Scopaeus kritschi, Cambodia, Angkor (6–8); S. decipiens, Sri Lanka, Galle (9). Abbreviation: mkp—metakatepisternal process. Scale bars: A— Fig. 9 (excerpt of leg with plectral ridges), B—Fig. 8, C—Fig. 7.	FIGURES 6–9. Stridular file of metaventrite in lateral view (6), stridular files of metaventrite (in sublateral position) in ventral view (7), excerpt of metaventrite with left stridular file (8), mesothoracal leg with plectral ridges (9) of Scopaeus kritschi, Cambodia, Angkor (6–8); S. decipiens, Sri Lanka, Galle (9). Abbreviation: mkp—metakatepisternal process. Scale bars: A— Fig. 9 (excerpt of leg with plectral ridges), B—Fig. 8, C—Fig. 7.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD55723FF47F48DE36873F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400489/files/figure.png	https://doi.org/10.5281/zenodo.17400489	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD55723FF47F48DE36873F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323702/files/figure.png	https://doi.org/10.5281/zenodo.17323702	FIGURES 48–53. Aedeagus of Scopaeus kritschi in lateral (48, 51), ventral (49, 52), dorsal view (50, 53); holotype, Cambodia, Angkor (48–50); paratype, Laos, Khammouane, Nakai (51–53). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	FIGURES 48–53. Aedeagus of Scopaeus kritschi in lateral (48, 51), ventral (49, 52), dorsal view (50, 53); holotype, Cambodia, Angkor (48–50); paratype, Laos, Khammouane, Nakai (51–53). Abbreviation: vl—ventral lobe. Arrows: a—apical tooth of apical lobe, b—subapical tooth of apical lobe, c—membranous ventral lobe of apical lobe, d—ventroproximal end of apical lobe, e—distal margin of ventral lobe.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD55723FF47F48DE36873F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323704/files/figure.png	https://doi.org/10.5281/zenodo.17323704	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	FIGURES 54–61. Female abdominal segment IX in ventral (54) and lateral view (55); gonocoxites with bursa, bursal duct and sperm pump in ventral view (56); sperm pump (57); bursa (58–61) of Scopaeus kritschi, paratype, Cambodia,Angkor (54, 55, 57, 61); S. decipiens, Sri Lanka, Kitulgala (58); S. complex, Japan, Okayama (56, 59); S. saaberae, paratype, Indonesia, Kalimantan Tengah, Telang Baru (60). Abbreviations: b—bursa, bd—bursal duct, lgp—lateral gonocoxal plate, lt IX—laterotergite IX, sp—sperm pump. Arrow: subapical, lateral enlargement of laterotergite IX. Scale bars: A—Figs 53–55, B—Figs 56–61.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFD55723FF47F48DE36873F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400496/files/figure.png	https://doi.org/10.5281/zenodo.17400496	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	Figure 62. Distribution of the Scopaeus decipiens species group: °—S. decipiens, ⬡—S. complex, D—S. saaberae,▲—S. kritschi.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFC8573FFF47F7ECE07A74B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17400489/files/figure.png	https://doi.org/10.5281/zenodo.17400489	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	FIGURES 10–26. Male abdominal sternite VII (10, 12, 14, 16, 18, 20, 22, 24) and male abdominal sternite VIII (11, 13, 15, 17, 19, 21, 23, 25, 26) of Scopaeus decipiens, lectotype, Sri Lanka (10, 11); S. complex, lectotype, Japan (12, 13); S. complex, China, Zhejiang, Siming Mts (14, 15); S. complex, Taiwan, Kaohsiung (16, 17); S. complex, Vietnam, Cuc Phuong, Ninh Binh (18, 19); S. saaberae, paratype, Indonesia, Borneo, Pagate (20, 21); S. kritschi, holotype, Cambodia, Angkor (22, 23); S. kritschi, paratype, Laos, Khammouane, Nakai (24, 25); S. velutinus, lectotype, „India Orientales“ (26).	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFC8573FFF47F7ECE07A74B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323706/files/figure.png	https://doi.org/10.5281/zenodo.17323706	FIGURES 63–67. Habitus (63) and microsculpture of head (64), pronotum (65), elytra (66), and abdomen (67) of Scopaeus velutinus, lectotype.	FIGURES 63–67. Habitus (63) and microsculpture of head (64), pronotum (65), elytra (66), and abdomen (67) of Scopaeus velutinus, lectotype.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
BA707837FFC8573FFF47F7ECE07A74B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17323710/files/figure.png	https://doi.org/10.5281/zenodo.17323710	FIGURES 68–70. Aedeagus of Scopaeus velutinus in lateral (68), ventral (69), dorsal view (70); lectotype, „Indes Orientales“. Abbreviations: al—apical lobes, dl—dorsal lobe, ll—lateral lobes, mf—median foramen, p—parameres. Arrow: tip of flagellum.	FIGURES 68–70. Aedeagus of Scopaeus velutinus in lateral (68), ventral (69), dorsal view (70); lectotype, „Indes Orientales“. Abbreviations: al—apical lobes, dl—dorsal lobe, ll—lateral lobes, mf—median foramen, p—parameres. Arrow: tip of flagellum.	2025-09-17	Frisch, Johannes		Zenodo	biologists	Frisch, Johannes			
