identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7AF80F3DA70F52229EDECD6AFD603523.text	7AF80F3DA70F52229EDECD6AFD603523.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra homage Vanker, Medina-Espinoza, & Evangelista 2026	<div><p>Epilampra homage Vanker, Medina-Espinoza, &amp; Evangelista sp. nov.</p><p>Figs 7, 8, Table 3</p><p>Type material.</p><p>Holotype: • male, pinned, with genitalia in a separate microvial. Original label: “ Peru: MD [Madre de Dios], Estación Biológica Los Amigos: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.10333&amp;materialsCitation.latitude=-12.563056" title="Search Plazi for locations around (long -70.10333/lat -12.563056)">Plataforma Trail</a>. 12°33'47"S, 70°06'12"W [-12.565481, -70.1027]. 265 m. 6. vii. 2021 [6 June 2021]. night hand coll. D. A. Evangelista leg ”. “ AUDE-PE-1-88. ” Paratypes. • 4 adult males. AUDE-PE-3-64, AUDE-PE-3-63, UIRB-PE-24-14, UIRB-PE-26-70 (locality and other data for all specimens are given in Table 4) .</p><p>Determination.</p><p>This specimen is placed in Epilampra (vs Poeciloderrhis) based on the presence of the following characters that differentiate Epilampra from Poeciloderrhis: via not pointed (pointed in Poeciloderrhis), via and lve separated (vs fused in Poeciloderrhis), hooked phallomere (L 3 ’) with a subapical incision (incision absent in Poeciloderrhis), and R’ phallomere cleft not fused (vs fused in Poeciloderrhis). However, this specimen does have a modified T 1 with a triangular medial knob, which was previously only found in Poeciloderrhis . Placement in the burmeisteri species group is supported by autapomorphies in the male genitalia (see below).</p><p>Differential diagnosis.</p><p>This species’ male genitalia are almost identical to E. substrigata . There are some minor differences in via compared to the E. substrigata specimens shown by Roth (1970 d). Without more spatial sampling it is unclear whether the differences in via are distinctive. These species are separated by the presence of a tergal modification on T 1 in E. homage . E. homage also has a transverse carina on T 1, but this may be a plesiomorphy as it is found in Africalolampra, Parapoeciloderrhis, and perhaps other Epilampra species. Epilampra homage is easily separated from Africalolampra, and Parapoeciloderrhis in the morphology of L 2 ’: via separated from lve (vs connected), and the shape of via is with an irregular ridge apically (like a mountain range).</p><p>Description of holotype.</p><p>Male (AUDE-PE-1-88). Head. With large overlapping speckles, densest between the compound eyes, but the ecdysial suture lacks speckles. Dark band connecting ocelli in a U-shape, and underneath it, the speckles are sparser. Clypeus with some brown coloration at the base (dorsal) but otherwise lacking speckles along with the labrum.</p><p>Thorax. Pronotum. Speckled mostly homogenously throughout but densest medially. Other spots form a panther-face pattern. AV margin of forefemur type B 2 with five large spines preceding the row of small spines. On all legs, tarsal claws large, simple, with large arolia and all euplantulae present.</p><p>Abdomen. Ventral abdomen lightly colored with scattered spots of medium density. SG plate with a with a brown patch on the distal third (but brown patch does not meet the posterior margin). SG plate shape and styli typical of Epilampra . T 1 with a transverse carina and a medial modification. Medial modification strongly triangular (almost equilateral), with some hairs originating near the carina pointing towards the anterio-lateral sides of the triangle. Hairs line the depression that houses the triangle, which becomes very narrow at its deepest point. Face of triangle concavely dimpled. T 1 also modified with shallow depressions on either side, anterior to the carina and aligned with the edges of the square projection of the metanotum. Genitalia as shown in Fig. 8.</p><p>Remarks.</p><p>None of the several female Epilampra morphotypes in our collection are obviously attributable to this species. Perhaps some are E. homage females but sexual dimorphism (e. g., female Epilampra are typically larger than males) is making it difficult to associate the sexes.</p><p>Range.</p><p>We have no data for this species outside of Madre De Dios. Within Madre De Dios, this species was only found at Los Amigos Biological Station, and Finca Las Piedras Research station.</p><p>Habitat.</p><p>Low hill forest.</p><p>Etymology.</p><p>The specific epithet homage is meant to imply that this species has a combination of characters present in other taxa ( E. substrigata, Poeciloderrhis, Africalolampra, and Parapoeciloderrhis), as if it were “ paying homage ” to them.</p></div>	https://treatment.plazi.org/id/7AF80F3DA70F52229EDECD6AFD603523	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
8A63D48B9F385490B356AE4435A56311.text	8A63D48B9F385490B356AE4435A56311.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra paititi Vanker, Medina-Espinoza, & Evangelista 2026	<div><p>Epilampra paititi Vanker, Medina-Espinoza, &amp; Evangelista sp. nov.</p><p>Figs 14, 15</p><p>Type material.</p><p>Holotype. • Male, pinned, with genitalia in a separate microvial. Original label: “ Peru: Madre de Dios, Estación Biológica Los Amigos. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.10333&amp;materialsCitation.latitude=-12.563056" title="Search Plazi for locations around (long -70.10333/lat -12.563056)">Plataforma trail</a>, terra firme; 12°33'47"S, 70°06'12"W [-12.563056, -70.103333]; 3. vii. 2021 [3 July 2021], hand coll. D. A. Evangelista leg. ” AUDE-PE-1-83.</p><p>Determination.</p><p>This specimen was placed in Epilampra (vs Poeciloderrhis) based on the presence of the following characters: the absence of medial tergal modifications on T 1 and T 2 (present in Poeciloderrhis), via separated from lve by a membrane (connected in Poeciloderrhis), hooked phallomere (L 3 ’) with a subapical incision (incision absent in Poeciloderrhis), and R’ phallomere cleft not fused (vs fused in Poeciloderrhis). We could not associate any females or juveniles to the adult male based on morphology.</p><p>Differential diagnosis.</p><p>At first, this species seemed to belong in the sodalis group based on the following traits it shares with taxa in that group: L 10 ’ is solidly fused to via and covered in microtrichia (shared with Roth (1970 d) ’ s Epilampra sp. C), and presence of a medial tergal modification on T 3 and T 4 (shared with Roth (1970 d) ’ s Epilampra sp. A). In fact, Roth (1970 d) commented that the latter of these traits was the only known cockroach with such a tergal gland. We do not know if this remains true outside of Epilampra . Yet, Epilampra sp. A and now this new species are unique within Epilampra in having such a dorsal tergal modification. However, this species differs strongly from Epilampra sp. A in the shape of all the genital sclerites (including L 10 ’), although L 3 ’ is more-or-less similar in the two taxa. The new species also differs strongly from Epilampra sp. C in the shape of via and L 1.</p><p>Description of holotype.</p><p>Male (AUDE-PE-1-83). Head. Frons, in dorsal view, predominantly pale brown with some dark brown speckling above the anteclypeus and a dark patch in the interocular space. Ocelli are flat, slightly ovular, and adjacent to the anterior-medial corners of antennal sockets. Labrum and final segment of maxillary palp slightly darker than the pale brown color of most of the face.</p><p>Thorax. Subtrapezoidal anteriorly, with an elongated medial lobe posteriorly. Primarily pale brown with dark speckles and several dark brown patches towards the midline. In the center are two dark parallel lines. Forewings large, extending past the end of abdomen and cerci. Pale brown coloration, speckled across wings with fainter and smaller speckles towards the lateral and posterior margins of the wings. Coloration on all legs similar, primarily pale brown with some sparse speckling that is most dense on the coxa, and a dark brown line on the dorsal margin of all femurs. Forefemur with a dark brown line on anterior margin. The forefemur’s AV margin spination type is B 2 with five large spines proximal to a row of 19 short spines. Forefemur posteroventral margin with four spines, one of which is apical. Fore and hind tarsus with five tarsomeres, and midtarsus with four tarsomeres. Fore- and midtarsus with two parallel rows of short ventral spines on the first tarsomere, short spines on either side of the euplantulae, and a final tarsomere with simple symmetrical claws and an arolium. Hind tarsus damaged without the final clawed tarsomere absent.</p><p>Abdomen. Dorsal abdomen has pale tan coloration at the anterior end of the abdomen, which darkens towards the posterior end, and light speckling on the left and right margins. Tergal modification on T 3. Posterior margin of T 1–5 medially notched. Ventral abdomen is pale brown and speckled. SA plate is semi-elliptical and bilobed with a long medial division. SG plate typical of Epilampra with short straight styli and marginal concavity on the right. Genitalia as in Fig. 15. Morphological measurements are presented in Table 4.</p><p>Range.</p><p>We collected only a single adult male of this species (if we collected an immature, or female, we have not identified it) in Madre De Dios, Peru, and we have no data for this taxon outside the region. We found it only at Los Amigos Biological Station.</p><p>Habitat.</p><p>Low hill forest.</p><p>Etymology.</p><p>The specific epithet paititi draws a parallel between the mythological city of gold lost in Madre De Dios and the now discovered “ lost tergal gland of gold ”.</p></div>	https://treatment.plazi.org/id/8A63D48B9F385490B356AE4435A56311	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
397E99A1B1925D86867CE8D1895BB855.text	397E99A1B1925D86867CE8D1895BB855.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra sofia Evangelista, Medina-Espinoza, & Vanker 2026	<div><p>Epilampra sofia Evangelista, Medina-Espinoza, &amp; Vanker sp. nov.</p><p>Figs 12, 13</p><p>Type material.</p><p>Holotype. • Male, pinned, with genitalia in a separate microvial. Original label: “ within concession, near station at night. -12.5282, -69.0141. 175 M elev. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.0141&amp;materialsCitation.latitude=-12.5282" title="Search Plazi for locations around (long -69.0141/lat -12.5282)">Kawsay Biol. Station</a>, Madre de Dios, Peru. 18 - Jul- 2024 | Coll: Emmy F. Medina E., Jared Martin, Johanna Schwartz, Dominic A. Evangelista [leg]. ” UIRB-PE-26-37 . Paratype. • Female: UIRB-PE-25-77 (locality and other data for all specimens are given in Table 3) .</p><p>Determination.</p><p>The specimen is nearly identical to E. sodalis in the literature (Roth 1970 d; Anisyutkin 2016) except for the L 10 ’, via, and L 2 ’ cleft.</p><p>Differential diagnosis.</p><p>This is a distinct species from E. sodalis and E. carsevennae (Bonfils 1975) based on the following three characters. L 10 ’ large, enveloping most of via and covered with weakly pigmented microtrichia (vs E. sodalis ’ and E. carsevennae ’ s L 10 ’ which are indistinct or entirely reduced). via with a unique shape (lacking the medial knob figured in E. sodalis and not hook-shaped like in E. carsevennae). Finally. The R 1 ’ cleft is fully fused, which Roth showed was the case in some sodalis group representatives, but usually only partially. R 1 ’ in E. carsevennae is not well-illustrated.</p><p>Description of holotype.</p><p>Male (UIRB-PE-26-37). Head. Shape and proportions of head as in Anisyutkin (2016: fig. 6) Frons speckled, almost homogenously, but speckles lacking on the clypeus and labrum. Chestnut colored spots (oval shaped) slightly under ocelli and slightly smaller than the ocelli.</p><p>Thorax. Speckled almost homogenously throughout pronotum, with small areas with larger areas of negative space forming a symmetrical pattern similar to the panther-face pattern on other species (e. g., E. conferta). Speckles larger than E. sodalis in Evangelista et al. (2015) and E. carsevennae . AV margin of foreleg with type B 2 spination (three large spines at base). All four euplantulae very large on all tarsomeres (four on each leg). Tarsomere spination typical of Epilampra, with small spines on either side of euplantulae and two rows of small spines at base. Coxa colored like pronotum and head, but with smaller speckles. Other leg segments similarly colored but lacking speckles. Base color of tegmina a pale, whitish brown, with darker brown highlights on the cell and vein borders. Whitish brown patches lacking other pigmentation throughout the exposed portion of the tegmina, getting larger distally. Portion of right tegmina that overlaps with left tegmina colored distinctly differently, with an orange-brownish base color and large dark-brown spots.</p><p>Abdomen. Abdomen reddish-orange with heterogeneous (in size and distribution) speckles. SG plate typical of Epilamprinae (two simple, subsymmetrical styli with the right one emerging from a concave asymmetry in the margin). SA plate symmetrical and large, the preceding tergite with a large convexity medially, spanning almost the entire width of the SA plate. Genitalia as shown in Fig. 13. Morphological measurements are presented in Table 4.</p><p>Description of female paratype.</p><p>Female (UIRB-PE-25-77). Same as male except larger and with the following departures. Hind legs with only three tarsomeres. Ventral coloration is slightly different, with the thorax having more prominent brown-grey coloration, and the abdomen having a much stronger reddish coloration with an almost purple-pink tinge.</p><p>Remarks.</p><p>We have no data for this species outside of Madre De Dios, Peru. Within the region, we collected only two adult individuals, both from Kawsay Biological Station.</p><p>Habitat.</p><p>Flooded alluvial forest.</p><p>Etymology.</p><p>The specific epithet, sofia, in apposition, is named after the daughter of DAE.</p></div>	https://treatment.plazi.org/id/397E99A1B1925D86867CE8D1895BB855	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
990A0DADB48F57D5B1C550053A4FE72E.text	990A0DADB48F57D5B1C550053A4FE72E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra wandpero Medina-Espinoza, Vanker, & Evangelista 2026	<div><p>Epilampra wandpero Medina-Espinoza, Vanker, &amp; Evangelista sp. nov.</p><p>Fig. 9, Table 3</p><p>Type material.</p><p>Holotype: • male, pinned, with genitalia in a separate microvial. Original label: “ Peru: Madre de Dios, Tambopata, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.17389&amp;materialsCitation.latitude=-12.505555" title="Search Plazi for locations around (long -69.17389/lat -12.505555)">Ecoaventuras Amazonicas</a>, 12°30'20"S, 69°10'26"W [-12.505556, -69.173889], 29. vi. 2021 [29 June 2021], night on forest floor. hand coll. D. A. Evangelista leg ”. AUDE-PE-03-86 . Paratypes. • 16 adult males: AUDE-PE-2-85, AUDE-PE-3-70, AUDE-PE-3-72, AUDE-PE-3-80, AUDE-PE-3-90, UIRB-PE-23-86, UIRB-PE-23-94, UIRB-PE-23-95, UIRB-PE-23-96, UIRB-PE-23-98, UIRB-PE-24-01, UIRB-PE-26-42, UIRB-PE-26-50, UIRB-PE-26-51, UIRB-PE-26-55, UIRB-PE-26-71. 5 adult females: UIRB-PE-26-41, UIRB-PE-26-43, UIRB-PE-26-49, UIRB-PE-26-52, UIRB-PE-26-72 (locality and other data for all specimens are given in Table 4) .</p><p>Determination.</p><p>We identified this taxon as Epilampra based on the mottled coloration, medio-posterior lobe on pronotum, elongated cerci (compared to stout in most other Blaberidae), separated via and lve, and lack of a visible tergal gland in males. We could not associate any juveniles with adults based on morphology. We tentatively associated males and females based on some similarities in color pattern. Epilampra wandpero was determined to be in Roth (1970 d) ’ s subgroup B based on the large via covering most of L 10 ’.</p><p>Differential diagnosis.</p><p>The new taxon can be differentiated from all other Epilampra spp. treated by Roth (1970 d) by the shape of L 3 ’, in which the hook is enlarged, with a very large sub-apical incision, giving the hook a crab-claw-like appearance. In other Epilampra spp. with a robust hook, the smaller lip of the hook (i. e., the inner side of the sub-apical incision) is still slender. It is most closely related to E. campestris (Rocha e Silva and Auguiar 1978) based on the similarity of the SA plate, via, and L 10 ’. Yet, the two species differ strongly in the shape of L 3 ’, pronotal coloration, and head coloration.</p><p>Description of holotype.</p><p>Male (AUDE-PE-03-86). Head. Two ocelli flat and distinctly formed; vertex pale brown with small dark spots; frons pale brown with small dark spots in greater density between the antenna than under the antenna, darker brown band above the epicranial suture fading as it approaches the vertex, with a horizontal thin dark brown band connecting the inferior side of the ocelli; dark brown line in the epistomal and subgenal sulcus not meeting in the middle; labrum yellowish with pale brown areas in the lateral sides and its inferior side; maxillary palps yellowish: first segment with an apical pale-brown ring; second with a dark brown area in ventral side, third yellowish with dark brown ventral side, and fourth dark brown.</p><p>Thorax. Pronotum. Trapezoidal with a rounded anterior edge and a bell-curve shaped posterior edge (typical of Epilamprinae). Pale brown, scattered maculations, with a defined row of dark brown spots on the anterior edge. After the anterior-most quarter of the pronotum, two lateral lines strongly curved posteriorly towards the midline without intersecting and ending shortly after the curve. Two thin parallel dark brown lines in the center of the pronotum, anteriorly curved gently outward; beginning in the posterior quarter, the lines separate, and two parallel lines appear between them, resembling a fork. Mesothorax and metathorax are not visible from dorsal view. Thorax in ventral view pale brown with dark brown spots.</p><p>Legs. Forelegs: forecoxae with small dark brown spots, fore-femoral pale brown, dark brown lines in the edges, five large anteroventral spines towards the proximal side, followed by a row of 14 short spines and two longer spines; four posteroventral spines; one genicular spine. Foretibia pale brown, not uniformly slender, expanding distally, with setae in the anteroventral side, ten strong spines distributed around the tibia, anteroventral side dark brown, distal end with a dark brown ring. Tarsi with five tarsomeres, the first one elongated with a bulbous end, two rows of tiny spines in the anteroventral side, and one apical spine; the next three moniliform with a spine in the distal end, the second one with an apical spine. Mid-legs missing due to damage. Hind-legs: hind coxae with small dark brown spots; hind femoral pale brown with small dark brown spots, with lines along its dorsal and ventral edges; four spines in the anteroventral side and three in the posteroventral side. Foretibia, pale brown with dark brown ventral edge, thirteen large spines on the dorsal side, nine large spines at the ventral side, one large genicular spine, and one short apical spine, distal end with a dark brown ring. Tarsi with five tarsomeres; first and second tarsomeres cylindrical with two parallel rows of tiny spines on the ventral side, the first one the largest; third tarsomere shorter with two tiny spines on the ventral view; a couple of apical spines on each side of the distal border; and fourth tarsomere short without spines; the fifth tarsomere longer without spines. Claws simple, symmetrical, arolium present. Euplantulae present on all tarsomeres. Wings. Longer than the abdomen. Forewing dark-brown, pale brown in the costal area of the wing, a row of black dots roughly following the radial vein until midway towards the tip, marks a sudden transition to the darker tan.</p><p>Abdomen. Pale brown, covered with dark brown dots, increasing in density to the posterior side, a row of larger dark brown dots in the distal margin. No visible external tergal glands, supra-anal plate bilobed. Subgenital plate shape typical for Epilampra, with slender styli.</p><p>Genitalia. L 3 ’ long and slender, with tip strongly curved acutely with a very large sub-apical incision, with the inner portion truncated and bulbous, giving the appearance of a crab-like claw. via large and covering most of L 10 ’, similar to E. opaca, E. substrigata, and E. columbiana in Roth (1970 d). R’ phallomere like E. columbiana but narrower with cleft deeper.</p><p>Remarks.</p><p>Two paratype males with facial coloration pattern like the holotype. Three paratype males with the spots in the center of the frons extend from the inner edge of the eyes, almost meeting in the center. Paratypes show variation in color intensity and spot density on the pronotum.</p><p>Range.</p><p>We have no data for this species outside of Madre de Dios. Within the region, it was most common at Kawsay Biological Station, and present at all other sites except Finca Las Piedras Research Station.</p><p>Habitat.</p><p>Low hill forest and flooded alluvial forest.</p><p>Etymology.</p><p>The specific epithet means cockroach in Harakmbut, the language of the indigenous Harakbut ethnic people living in the Madre De Dios region.</p></div>	https://treatment.plazi.org/id/990A0DADB48F57D5B1C550053A4FE72E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
4D1BE96F215D5931848081E6B9A068AF.text	4D1BE96F215D5931848081E6B9A068AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epilampra yompori Vanker, Medina-Espinoza, & Evangelista 2026	<div><p>Epilampra yompori Vanker, Medina-Espinoza, &amp; Evangelista sp. nov.</p><p>Figs 4, 5, Table 3</p><p>Type material.</p><p>Holotype: • male, pinned, with genitalia in a separate microvial. Original label: “ Trails in forest, near station at night. -12.2285, -69.1143, 265 M. elev. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.1143&amp;materialsCitation.latitude=-12.2285" title="Search Plazi for locations around (long -69.1143/lat -12.2285)">Finca Las Piedras Res. Station</a>, Madre de Dios. 13 - Jul- 2024. Coll. E. Emmy F. Medina E., Jared Martin, Johanna Schwartz, Dominic A. Evangelista ” “ UIRB-PE-23-76 ” . Paratypes. • 1 adult female: UIRB-PE-25-42. 9 juveniles: AUDE-PE-3-76, UIRB-PE-23-93, UIRB-PE-25-35, UIRB-PE-25-36, UIRB-PE-25-78, UIRB-PE-26-23, UIRB-PE-26-25, UIRB-PE-26-38, UIRB-PE-31-51 (locality and other data for all specimens are given in Table 4) .</p><p>Determination.</p><p>This specimen was identified as Epilampra (vs Poeciloderrhis) based on presence of the following characters: the absence of medial tergal modifications on T 1 and T 2 (present in Poeciloderrhis), hooked phallomere (L 3 ’) with a subapical incision (incision absent in Poeciloderrhis), and R’ phallomere cleft not fused (vs fused in Poeciloderrhis).</p><p>Differential diagnosis.</p><p>The species may be differentiated from other Epilampra spp. by a unique combination of the following characters. The pronotal coloration and via + L 10 ’ morphology are most similar to species in the abdomennigrum group. However, the species lacks a setal brush on R’ phallomere, placing it in the burmeisteri group. Roth (1970 d) separated subgroups A and B of the burmeisteri group based on the size of via compared to L 10 ’ (via small in subgroup A, and large in subgroup B). By relative size, this species would fall in subgroup A. However, L 10 ’ is elongated (as in E. grisea – abdomennigrum group), so via may appear small relative to L 10 ’ when in fact it is normally sized when compared to lve. Among the species in the burmeistei group, the L 10 ’ and via shape are most similar to E. opaca or E. columbiana . The species differs strongly from E. opaca in the following characteristics: males 26 mm long (vs 20.5 mm in E. opaca (Evangelista et al. 2015)), central region of pronotum fuscous as in E. taira (speckled with a symmetrical pattern in E. opaca and E. columbiana), and L 10 ’ elongated like in E. grisea (vs mostly rounded in E. opaca and E. columbiana).</p><p>Remarks.</p><p>Roth (1970 d) also noted the similarity in via and L 10 ’ between various species in the abdomennigrum and burmeisteri groups. It is possible that these are close relatives, or that one is nested within the other. Hence, we refrain from placing E. yompori in either group or creating a new one for it.</p><p>Description of holotype.</p><p>Male (UIRB-PE-23-76). Head. Partially visible from dorsal side. Face predominantly pale brown with a dark brown or blackish interocular space ending at the anterior end of the ocelli, except for a circular pale patch in the interocellar space. Ocelli are circular, flat, slightly impressed, and adjacent to the anterior-medial corners of antennal sockets.</p><p>Thorax. Pronotum. Subtrapezoidal anteriorly and with an elongated medial point posteriorly (typical of Epilampra spp.). Coloration pale brown near the lateral borders, but mainly dark brown with speckled coloration toward the lateral, and posterior margins. The middle of the pronotum is more solidly dark brown. The thorax in the ventral view is pale brown. Legs. Primarily pale brown but with medium brown (or slightly reddish brown) stripes on the femora, with darker shaded spines on the femora and tibiae. The forefemur AV margin spination is type B 2 with four large spines proximal to a row of short spines (16 on the right leg). On the posteroventral margin, there are four large spines, one of which is an apical spine. Foretibia has seven dorsal and five ventral spines and is darker than the femur. All tarsi have five tarsomeres and euplantulae, simple symmetrical claws with arolia. Foretarsus has longer first and fifth tarsomeres, with two rows small proximal spines on ventral side of the first tarsomere. Midfemur with four anteroventral spines, one of which is apical, one anterodorsal apical spine, and four posteroventral spines. The mid-tibia has 15 dorsal spines, two of which are apical, and 12 ventral spines, including one apical spine considerably longer than the other spines on the midtibia. First tarsomere of the middle leg considerably longer than the others with a row of small spines on the ventral side, and two short anterior spines and one short posterior spine. Second tarsomere also has two linear rows of short ventral spines. The first, second, and third tarsomeres have small apical spines on either side of the euplantulae. Hind femur with the same coloration and spination as middle leg. Hind tibia with ten dorsal spines, two of which are apical, and eight ventral spines. Hind tarsus with the first tarsomere slightly larger than all other tarsomeres combined and has two parallel rows of short spines along length of ventral margin and a shorter, more spaced-out row of short spines on the anterior margin. Second tarsomere has two parallel rows of short spines on ventral margin and apical spines around the euplantulae. The third tarsomere also has spines on either side of the euplantulae. Macropterous with tegmina terminating much posterior to the end of the cerci.</p><p>Abdomen. Base coloration similar to the legs, but with some scant speckled and larger brown spots laterally and a rusty brown region towards the posterior end on the ventral side. Dorsal side without distinctive markings and lacking any tergal specializations. SA plate semi-circular with a short medial division. SG plate with typical Epilampra shape (slightly asymmetrical, concave on the right side, with pin-like styli). L 3 ’ strongly curved and narrow, with a very noticeable subapical incision.</p><p>Range.</p><p>We have no data from this species anywhere other than Madre De Dios, where it is widespread.</p><p>Habitat.</p><p>Low hill forest and flooded alluvial forest.</p><p>Etymology.</p><p>The specific epithet yompori means “ forest spirit ” in Piro, an indigenous language in southwestern Amazonia, spoken by communities in Madre de Dios and Ucayali. The language is widespread, as is the cockroach.</p></div>	https://treatment.plazi.org/id/4D1BE96F215D5931848081E6B9A068AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
C70D2412B2395E9F8FB3DA228487E4C3.text	C70D2412B2395E9F8FB3DA228487E4C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lanxoblatta patriciae Vanker, Medina-Espinoza, & Evangelista 2026	<div><p>Lanxoblatta patriciae Vanker, Medina-Espinoza, &amp; Evangelista sp. nov.</p><p>Figs 16, 17</p><p>Type material.</p><p>Holotype. • Male, pinned, with genitalia in a separate microvial. Original label: “ Peru: MD [Madre de Dios], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.099724&amp;materialsCitation.latitude=-12.566389" title="Search Plazi for locations around (long -70.099724/lat -12.566389)">Estación Biológica Los Amigos. Trail 10</a>, terra firme forest, 12°33'59"S, 70°05'59"W [-12.566389, -70.099722], 6. vii. 2021 [6 July 2021], night, hand collected. D. A. Evangelista leg ”. AUDE-PE-1-69 . Paratypes: • AUDE-PE-1-89 AUDE-PE-1-90, UIRB-PE-26-61, UI RB-PE-26-62, UIRB-PE-26-74, UIRB-PE-26-75, UIRB-PE-26-76, UIRB-PE-26-77, UIRB-PE-26-81, UIRB-PE-26-82, UIRB-PE-26-83 (locality and other data for all specimens are given in Table 3) .</p><p>Determination.</p><p>This species was placed in Lanxoblatta based on the pronotal morphology (shape, color, and pronotal window shape). We were able to associate juveniles with the adults based on pronotal features.</p><p>Differential diagnosis.</p><p>Lanxoblatta patriciae, which is most similar to L. martinezi (Bolivar, 1881), and can be separated from all known Lanxoblatta based on the body size and pronotal morphology. L. patriciae and L. martinezi share the feature rugae radiating out from the pronotal center into the pronotal window. Lanxoblatta patriciae differs from L. martinezi in males being 20 mm long ( L. martinezi males are 30 mm long; Table 4), and our lone female L. patriciae has semi-circular pronotum without any emargination (notch) in it laterally ( L. martinezi females have an emargination on the pronotum just back of the lateral angles).</p><p>Remarks.</p><p>Lanxoblatta patriciae bears impressive pronotal warts, but we are unclear how they compare to pronotal rugosity in other Lanxoblatta spp. Lanxoblatta frater Hebard, 1933 is described as having “ nodules ”, and L. magnifica Rehn, 1932 bears “ tubercules ” on the pronotum. Lanxoblatta patriciae differs from L. frater Hebard, 1933 in having the warts much more extensive throughout the pronotum. We have not examined the tubercules in L. magnifica Rehn, 1932 but that species is nearly twice the size of L. patriciae . The description of L. martinezi (Bolivar, 1881) mentions granules on the pronotum but perhaps the microscopy available at the time was not sufficient to describe their detail more fully.</p><p>Description of holotype.</p><p>Male (AUDE-PE-1-69). Head. Two flat ocelli. Antennal scape and pedicel dark brown with paler flagellum. Frons and vertex dark brown, palest between the ocelli. The part of the postclypeus directly under the epistomal suture is paler, this pale area shrinks as the dark coloration, continued from the frons into the postclypeus, expands until reaching the anteclypeus where it abruptly stops, being replaced by a pale coloration. The labrum is banded, going from pale posteriorly, to dark, to pale again towards the mandibles. Maxillary and labial palp darker towards center of each segment, growing paler as the joint is approached.</p><p>Thorax. Pronotum. Roughly elliptical, with a greater distance from the vertexes to the top co-vertex than to the bottom co-vertex. Medial-anterior region of pronotum with a kidney shaped semi-transparent and unpigmented region. Pigmented areas of pronotum are mostly chestnut, with some near black-brown areas. Many small dark warts (protrusions that are narrow at the base and bulbous towards the apex) create a rugose texture. Many of the warts are so protruded that the form pillar-like shapes, only visible at high magnification. The greatest concentration are equidistant from the midline to the edge of the pronotum on either side, growing darker as their density increases, towards the edges of the pronotum the warts transition into shallow indentations. At the center of the pronotum, there are two dark sulci. Wings. Tegmina dark brown. Tegmina thickened basally so minor veins and crossveins are not entirely visible. Instead, there is a roughly uniform network of shallow pits. The subcostal and secondary branching of the radial veins support the laterally broad tegmina. Legs. Forecoxae pale brown. The forefemur is pale brown with a row of setae on ventro-anterior margin, with no apical spine or genicular spine. Foretibia with six spines on the dorsal side, one being smaller than the others, and five large ventral spines on the distal side of a dense patch of setae. Foretarsus composed of five segments, the first four having euplantulae and being significantly smaller than the pre-tarsus. Arolium present, claws simple and symmetrical. Mid femur pale brown with a few thin hairs on ventro-anterior margin. Midtibia pale brown with nine large ventral spines and seven dorsal. Midtarsus is broken but has at least four remaining tarsomeres, first tarsomere longer than its equivalent on the foreleg and the other three tarsomeres on mid tarsus with euplantulae present.</p><p>Abdomen. The dorsal abdomen is brown, paler than the pronotum but darker than the ventral abdomen. No visible tergal gland present. SA plate damaged in type. Posterior edge of sub-genital plate convexly arcuate with left stylus in a shallow and moderately narrow invagination. Right stylus missing due to damage (present in other specimens).</p><p>Genitalia. L 3 ’ hooked (see Fig. 17), lve long and slender, via: flat and broad at base with bulbous protrusion curving upwards to lve (see Fig. 17). R phallomere: curving on itself almost in a spiral pattern (see Fig. 17).</p><p>Morphological measurements are presented in Table 4.</p><p>Description of paratype.</p><p>Female. UIRB-PE-26-62. Head identical to description of male. Pronotum wider than in male, and more acutely angled laterally. Texture and color of pronotum indistinguishable from the male. Tegmina similar to male but truncated, only reaching to the posterior edge of T 5. Forelegs like male except only three spines in the dense patch of setae on the ventral tibia, and six spines on the dorsal side. Abdominal flanges more pronounced than in males, with some mottled coloration near the edges. SA plate also more expanded than male, extending past the ends of the cerci, which are also shorter in the female.</p><p>Range.</p><p>This is the first published record of Lanxoblatta in Peru. L. patriciae is widespread in Madre de Dios, but we did not find it at the two suburban sites near Puerto Maldonado. We hypothesize that it may be part of an evolutionary grade (parapatry) with L. martinezi because of the similarity with that species. This could suggest that its range to extends northward throughout the Peruvian Amazon.</p><p>Habitat.</p><p>Low hill forest and flooded alluvial forest.</p><p>Etymology.</p><p>The specific epithet is in honor of the grandmother of KV.</p></div>	https://treatment.plazi.org/id/C70D2412B2395E9F8FB3DA228487E4C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
B154E197A18B5C9BB7AAB3948D8471CE.text	B154E197A18B5C9BB7AAB3948D8471CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phoetalia amigo Vanker, Medina-Espinoza, & Evangelista 2026	<div><p>Phoetalia amigo Vanker, Medina-Espinoza, &amp; Evangelista sp. nov.</p><p>Figs 18, 19</p><p>Type material.</p><p>Holotype. • Male, pinned, with genitalia in a separate microvial. Original label: “ Peru: MD [Madre de Dios], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.10036&amp;materialsCitation.latitude=-12.569139" title="Search Plazi for locations around (long -70.10036/lat -12.569139)">Estación Biológica Los Amigos. In / around buildings</a>, terra firme, 12°34'08.9"S, 70°06'01.3"W [-12.569139, -70.100361]; 5–8. vii. 2021 [5–8 July 2021], night. D. A. Evangelista leg ”. AUDE-PE-1-95 . Paratype. • Female. UIRB-PE-30-77 (locality and other data for both specimens are given in Table 3) .</p><p>Determination.</p><p>This species was placed in Blaberinae based on the presence of sclerite L 4 ’ (Klass 1997) and by the large crown of spines on L 10 ’. We tentatively placed it in Phoetalia after comparisons of genitalia in Phoetalia (L 10 ’ spines present; (Roth 1970 c), Parasphaeria (L 10 ’ spines absent), and Schultesia (L 10 ’ spines absent and has a differently shaped via; (Roth 1973). Grandcolas and Pellens (2002) mentioned the left paraproct forming a blunt, curved hook as a synapomorphy of Parasphaeria but it is also present in P. amigo .</p><p>Differential diagnosis.</p><p>Parasphaeria amigo differs from P. circumvagans (Burmeister, 1838) and P. pallida (Brunner von Wattenwyl, 1865) in the shape of via, which is highly elongated in P. amigo . P. amigo and P. circumvagans are similar in pronotal coloration, which both have a pale boarder with a uniform width on the anterior-lateral margin (vs P. pallida in which the interface between the pale and dark regions is sinusoidal).</p><p>Remarks.</p><p>Grandcolas and Pellens (2002) mentioned the left paraproct forming a blunt, curved hook as a synapomorphy of Parasphaeria but it is also present in P. amigo .</p><p>Description of holotype.</p><p>Male (AUDE-PE-1-95). Pronotum. Sub-semicircular, with no pigmentation on anterior-lateral margin. Central mahogany-color region reaches posterior margin. Tegmina extends past end of abdomen, and mostly lacks pigmentation except for brown highlights on the veins, which is more prominent basally. Head with frons uniformly black smooth, with slight texture and sparse dimples throughout. A slight concavity connecting the ocelli create a slight W shape (Fig. 18 C). Ocelli white without a defined border, placed next to the superior inner part of the antenna. Clypeus and labrum paler than remainder of head. Ventral body. Almost uniformly dark chestnut colored, abdomen bordered by buff all around. Foreleg femur with no genicular spines, and AV margin spination type B 1 with one large basal spine and 20–24 small hairs / spinules. Legs with euplantulae on all legs and arolia on middle and hind legs (foreleg tarsus broken). Genitalia. SA plate with a patch of hairs, densest medially. Paraprocts asymmetrical, left paraproct forming a stout, slightly curved hook. Right paraproct amorphous. SG plate slightly asymmetrical, with slightly more concave margin on right. Base of subgenital plate long and without pigmentation. Right stylus short and projecting from the nadir of the concavity. Left style projects from left corner of subgenital plate. Styli otherwise symmetrical. Morphological measurements are presented in Table 4.</p><p>Description of paratype.</p><p>Female (UIRB-PE-30-77). Same as male except larger, and with the following departures. Subtle concavity between ocelli is U-shaped. Tegmina only reaching to slightly before SA plate. Foreleg AV margin type B 1 with two basal spines. Arolia on fore and mid-legs (hindlegs broken) and euplantulae on all tarsomeres (likely same as male).</p><p>Range.</p><p>We do not have any data on this species outside of Madre De Dios, Peru. We collected only a single adult male of this species at Los Amigos Biological Station. The female described above is from Kawsay Biological station. We have numerous immatures (all from Finca Las Piedras) tentatively identified to Phoetalia, and they appear more like P. amigo than P. pallida .</p><p>Habitat.</p><p>Low hill forest.</p><p>Etymology.</p><p>The specific epithet amigo, Spanish for friend (masculine) is a reference to the collection area of the male type, Los Amigos Biological Station.</p></div>	https://treatment.plazi.org/id/B154E197A18B5C9BB7AAB3948D8471CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Vanker, Katharine;Medina-Espinoza, Emmy Fiorella;Schwartz, Johanna;Martin, Jared;Evangelista, Dominic A.	Vanker, Katharine, Medina-Espinoza, Emmy Fiorella, Schwartz, Johanna, Martin, Jared, Evangelista, Dominic A. (2026): Blaberidae (Blattodea) of Madre de Dios, Peru and a key to Epilamprinae s. s. ZooKeys 1281: 171-232, DOI: 10.3897/zookeys.1281.178573
