taxonID	type	description	language	source
B46FB71AFFFFFF83E18CFD5CFD93FDD5.taxon	description	(Figs 1 – 2)	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFFFF83E18CFD5CFD93FDD5.taxon	diagnosis	Diagnosis (based on Vanin 1976): Integument reddish-brown; middle of elytra with two large, rounded spots of decumbent and yellowish-white scales, and with setae of elytral spot inclined 45 ° in relation to longitudinal axis of body.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFFFF83E18CFD5CFD93FDD5.taxon	discussion	Remarks. In 2006, Mermudes examined a female of Dicordylus serranus from Passa Quatro (elevation 938 m) in Minas Gerais State, which filled an important geographical gap between the southernmost record from Santa Catarina and the northern one in Minas Gerais. The type locality included Serra do Caraça (municipality of Santa Bárbara in Minas Gerais State) and Mafra (municipality in Santa Catarina State), at elevations of 1380 m and 800 m, respectively. Now, I have examined three specimens collected by Zikán in Virginia, municipality of Minas Gerais, at 1500 m elevation. One of these specimens was figured by Lima (1956). I believe these are three females, with only two specimens dissected in this study due to the rarity of the material. The distribution of Dicordylus serranus remains within the Atlantic Forest, ranging from Santa Catarina to Minas Gerais.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFFFF83E18CFD5CFD93FDD5.taxon	materials_examined	Material examined. Minas Gerais. Virginia (Faz [enda] dos Campos, 1500 m), 1 female II. XII. 1920, J. F. Zikán leg., Dicordylus det. C. Lima (ex-coll. Zikán, nº 8669, CEIOC), 2 females 22. X. 1920, with the same data (ex-coll. Zikán nº 8668, nº 8670 CEIOC (dissected).	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFCFF85E18CFDB8FB5AF93F.taxon	description	(Figs 3 – 10)	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFCFF85E18CFDB8FB5AF93F.taxon	materials_examined	Type material. Neotype (Figs 3 – 10) from BRAZIL. Paraná. Ponta Grossa (Vila Velha IAPAR), 1 male, 02. X. 2000 malaise 3 E 2, Ganho & Marinoni leg. (white, printed), (DZUP 085540, white, printed); Dicordylus vanini Mermudes, 2006 (male symbol), Mermudes & Queiroz det. 2011 (white, manuscript). The neotype was hereby designated to guarantee nomenclatural stability and fix the type-locality of this species.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFCFF85E18CFDB8FB5AF93F.taxon	diagnosis	Diagnosis. In 2006, Mermudes distinguished D. vanini from other species in the genus based on the following characteristics: 1) dorsal vestiture with dense, decumbent, shining, reddish-orange scales; 2) elytra with long, decumbent, ferruginous setae near the suture of the basal 1 / 4, these setae being conspicuously denser at the middle of the disk; and 3) absence of a rounded spot on the elytra.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFCFF85E18CFDB8FB5AF93F.taxon	description	Description of terminalia, male. Tergite VIII (Fig. 5) strongly transverse and arched, with slightly sinuous lateral margins, apical margin truncate-rounded with elongated and moderately dense setae. Sternite VIII (Fig. 6) strongly transverse, lateral and apical margins as in the tergite; proximal margin conspicuously emarginate, with the length of the apodeme subequal to the width of the sternite. Spiculum gastrale (Figs 5 – 6) elongate, more than 5 x the width of the proximal process. Tegmen (Fig. 7) elongate, parameres at least 4 x longer than the body of the ring, apodeme short, subequal to the body of the tegmen ring; apex of the paramere rounded and strongly curved in the distal fifth. Aedeagus (Figs 8 – 9), elongate and narrow, with subparallel sides; strongly pointed at the apex, this conspicuously dilated in lateral view (Fig. 9); apodemes very short, less than 1 / 4 the length of the aedeagus.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFCFF85E18CFDB8FB5AF93F.taxon	distribution	Distribution. This species is known only from the type locality of the neotype.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFCFF85E18CFDB8FB5AF93F.taxon	discussion	Remarks. The original holotype specimen of Dicordylus vanini, which was the sole specimen upon which the species description by Mermudes (2006) was based, was tragically destroyed in the fire at the National Museum in Rio de Janeiro on September 2, 2018. This loss leaves the species without its primary name-bearing type, which is essential for ensuring taxonomic stability and accurate identification of the species. As the original description was based on a single, now-destroyed specimen with limited locality data, there is a significant risk of confusion and misidentification of this species in the future. To address this critical issue and maintain nomenclatural stability, I am herein designating a neotype for Dicordylus vanini. This action is taken in strict accordance with The International Code of Zoological Nomenclature (ICZN, 1999). Specifically, I am adhering to Article 74 and 75.3, which outline the conditions and procedures for designating a neotype when the original holotype has been lost or destroyed. Furthermore, I am ensuring that the designated neotype is deposited in a publicly accessible institution (DZUP, Coleção Entomológica Pe Jesus Santiago Moure, Depto de Zoologia, Universidade Federal do Paraná, Curitiba), as required by Article 75.3.7 of the ICZN (1999). The designation of this neotype, a male specimen collected in a malaise trap at Ponta Grossa (Vila Velha IAPAR), Paraná, Brazil, is essential to fix the type locality of Dicordylus vanini and to provide a definitive reference point for future taxonomic studies. This action will prevent ambiguity, ensure correct application of the species name, and facilitate ongoing research on this rare and geographically restricted beetle species.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFAFF84E18CF85EFA95FEA9.taxon	materials_examined	Material examined. BRAZIL. Rio Grande do Sul. S. Salvador, 30. X. 1960, without leg. (MAPA).	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFAFF84E18CF85EFA95FEA9.taxon	discussion	Remarks. Vanin (1976) stated that H. antennalis is the only species characterized by antennomere XI with an elongate length, almost as long as the three preceding antennomeres together. The species was previously known only from the type locality Água Preta (now Uruçuca), State of Bahia. However, Vanin (1976) studied a male and a female from the state of Santa Catarina, municipality of Joinville (10 m elevation). Based on the material from this important collection in Porto Alegre (MAPA), the distribution of the species reaches the state of Rio Grande do Sul and helps to confirm the Atlantic Forest biome for this species at low elevation, unlike Homalocerus lyciformis.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFBFF84E18CFE64FD25FC05.taxon	materials_examined	Material examined. BRASIL. Rio de Janeiro: Itatiaia, 1 male, XII. 1989, Ayr Bello leg. (CEMT), Parque Nacional do Itatiaia (750 m) 1 female, 11 – 14. I. 2008, Monné et al. leg. (DZRJ); Teresópolis (Parnaso, Parque Nacional da Serra dos Órgãos, 1 female, 10. XI. 2010, Silveira, leg. (DZRJ); Estrada Paraty-Cunha (Parque Nacional da Bocaina), 2 males, 23. I. 2010, Mermudes & Mattos leg. (DZRJ).	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
B46FB71AFFFBFF84E18CFE64FD25FC05.taxon	discussion	Remarks. Vanin (1976) had already discussed that Homalocerus lyciformis is one of the best-known species of the genus, noting the variation in its body coloration, but emphasizing that it can be easily identified through the characters presented in the taxonomic key (Vanin 1976: 25). He also described the wide distribution of this species in Brazil, from Bahia to Rio Grande do Sul, and its occurrence in Argentina. In this study, the presence of the species was reported for the first time in two important massifs of the Atlantic Forest biome in Southeastern Brazil: the Mantiqueira massif (based on the records from Itatiaia) and the Serra do Mar massif (based on the records from Teresópolis and Bocaina). Vanin (1976) also mentioned the rarity of Belidae in collections, and this study reiterates that observation, highlighting the scarcity of available material and the still unknown aspect of the group’s biology in Brazil, including the lack of larval descriptions.	en	Mermudes, José Ricardo M. (2025): On the rare species of Dicordylus Lacordaire and Homalocerus Schoenherr (Coleoptera, Belidae) with notes on male terminalia of D. vanini Mermudes. Zootaxa 5646 (1): 144-150, DOI: 10.11646/zootaxa.5646.1.9, URL: https://doi.org/10.11646/zootaxa.5646.1.9
