taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518837	https://doi.org/10.3897/zookeys.1267.170946.figure1	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518838	https://doi.org/10.3897/zookeys.1267.170946.figure2	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518850	https://doi.org/10.3897/zookeys.1267.170946.figure12	Figure 12. A. Boreus borealis female (top) and male (bottom) viewed laterally. B. Collection localities for B. borealis. C. Habitat of Boreus borealis on St. Paul Island. Photo credit: TLK.	Figure 12. A. Boreus borealis female (top) and male (bottom) viewed laterally. B. Collection localities for B. borealis. C. Habitat of Boreus borealis on St. Paul Island. Photo credit: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518860	https://doi.org/10.3897/zookeys.1267.170946.figure19	Figure 19. Boreus spp. pronota, viewed laterally, with anterior to the left. A. Without pronotal bristles as seen in B. borealis; B. With pronotal bristles on the posterior margin only as seen in B. timaryi; C. With pronotal bristles on the posterior and anterior margins as seen in B. tananaensis.	Figure 19. Boreus spp. pronota, viewed laterally, with anterior to the left. A. Without pronotal bristles as seen in B. borealis; B. With pronotal bristles on the posterior margin only as seen in B. timaryi; C. With pronotal bristles on the posterior and anterior margins as seen in B. tananaensis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518864	https://doi.org/10.3897/zookeys.1267.170946.figure23	Figure 23. Tergal apophyses A. Present, as seen on B. timaryi (indicated by arrows); B. Absent, as seen on B. borealis.	Figure 23. Tergal apophyses A. Present, as seen on B. timaryi (indicated by arrows); B. Absent, as seen on B. borealis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518865	https://doi.org/10.3897/zookeys.1267.170946.figure24	Figure 24. Male postabdomen of B. intermedius viewed: A. Laterally; B. Dorsally and of B. borealis viewed: C. Laterally; D. Dorsally. In B and D the left dististyle has been removed to better show the denticular area and tergal pockets.	Figure 24. Male postabdomen of B. intermedius viewed: A. Laterally; B. Dorsally and of B. borealis viewed: C. Laterally; D. Dorsally. In B and D the left dististyle has been removed to better show the denticular area and tergal pockets.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518866	https://doi.org/10.3897/zookeys.1267.170946.figure25	Figure 25. Male left fore- and hind-wings, viewed laterally. A. Boreus intermedius with arrow indicating small hind wing spines on ventral surface, and B. Boreus borealis, without hind wing spines.	Figure 25. Male left fore- and hind-wings, viewed laterally. A. Boreus intermedius with arrow indicating small hind wing spines on ventral surface, and B. Boreus borealis, without hind wing spines.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518868	https://doi.org/10.3897/zookeys.1267.170946.figure27	Figure 27. Distal end of ovipositor viewed laterally. A. B. intermedius; B. B. borealis.	Figure 27. Distal end of ovipositor viewed laterally. A. B. intermedius; B. B. borealis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
CED8F158E1C6517BA98B0585A7CF35F6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518869	https://doi.org/10.3897/zookeys.1267.170946.figure28	Figure 28. Female terminalia viewed laterally. A. Boreus intermedius, with arrow indicating where valvulae narrow; B. Boreus borealis.	Figure 28. Female terminalia viewed laterally. A. Boreus intermedius, with arrow indicating where valvulae narrow; B. Boreus borealis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
6BA4DE3CFB6B50939505E6030441AD84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518837	https://doi.org/10.3897/zookeys.1267.170946.figure1	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
6BA4DE3CFB6B50939505E6030441AD84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518838	https://doi.org/10.3897/zookeys.1267.170946.figure2	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
6BA4DE3CFB6B50939505E6030441AD84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518848	https://doi.org/10.3897/zookeys.1267.170946.figure11	Figure 11. A. Boreus gracilis female (top) and male (bottom) viewed laterally. B. Collection localities for B. gracilis. C. Habitat of B. gracilis. Photo credit: TLK.	Figure 11. A. Boreus gracilis female (top) and male (bottom) viewed laterally. B. Collection localities for B. gracilis. C. Habitat of B. gracilis. Photo credit: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
6BA4DE3CFB6B50939505E6030441AD84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518862	https://doi.org/10.3897/zookeys.1267.170946.figure21	Figure 21. Male postabdomen viewed dorsally. A. B. gracilis; B. B. tananaensis sp. nov.	Figure 21. Male postabdomen viewed dorsally. A. B. gracilis; B. B. tananaensis sp. nov.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
6BA4DE3CFB6B50939505E6030441AD84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518863	https://doi.org/10.3897/zookeys.1267.170946.figure22	Figure 22. Head viewed laterally, arrows indicate lateral end of epistomal suture. A. B. gracilis (UAM: Ento: 457835); B. B. tananaensis sp. nov. (UAM: Ento: 504865).	Figure 22. Head viewed laterally, arrows indicate lateral end of epistomal suture. A. B. gracilis (UAM: Ento: 457835); B. B. tananaensis sp. nov. (UAM: Ento: 504865).	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
6BA4DE3CFB6B50939505E6030441AD84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518867	https://doi.org/10.3897/zookeys.1267.170946.figure26	Figure 26. Female terminalia viewed laterally. A. B. gracilis; B. B. tananaensis sp. nov.	Figure 26. Female terminalia viewed laterally. A. B. gracilis; B. B. tananaensis sp. nov.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518837	https://doi.org/10.3897/zookeys.1267.170946.figure1	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518838	https://doi.org/10.3897/zookeys.1267.170946.figure2	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518852	https://doi.org/10.3897/zookeys.1267.170946.figure13	Figure 13. A. Boreus intermedius female (top) and male (bottom) viewed laterally. B. Collection localities for B. intermedius. C. Habitat of B. intermedius (photo credit: TLK).	Figure 13. A. Boreus intermedius female (top) and male (bottom) viewed laterally. B. Collection localities for B. intermedius. C. Habitat of B. intermedius (photo credit: TLK).	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518853	https://doi.org/10.3897/zookeys.1267.170946.figure14	Figure 14. Size and color difference in B. intermedius specimens collected in the: A. Fall (15 September 2021, UAM: Ento: 451067) and B. Spring (13 May 2021, UAM: Ento: 447163). Photo credits: TLK.	Figure 14. Size and color difference in B. intermedius specimens collected in the: A. Fall (15 September 2021, UAM: Ento: 451067) and B. Spring (13 May 2021, UAM: Ento: 447163). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518865	https://doi.org/10.3897/zookeys.1267.170946.figure24	Figure 24. Male postabdomen of B. intermedius viewed: A. Laterally; B. Dorsally and of B. borealis viewed: C. Laterally; D. Dorsally. In B and D the left dististyle has been removed to better show the denticular area and tergal pockets.	Figure 24. Male postabdomen of B. intermedius viewed: A. Laterally; B. Dorsally and of B. borealis viewed: C. Laterally; D. Dorsally. In B and D the left dististyle has been removed to better show the denticular area and tergal pockets.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518866	https://doi.org/10.3897/zookeys.1267.170946.figure25	Figure 25. Male left fore- and hind-wings, viewed laterally. A. Boreus intermedius with arrow indicating small hind wing spines on ventral surface, and B. Boreus borealis, without hind wing spines.	Figure 25. Male left fore- and hind-wings, viewed laterally. A. Boreus intermedius with arrow indicating small hind wing spines on ventral surface, and B. Boreus borealis, without hind wing spines.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518868	https://doi.org/10.3897/zookeys.1267.170946.figure27	Figure 27. Distal end of ovipositor viewed laterally. A. B. intermedius; B. B. borealis.	Figure 27. Distal end of ovipositor viewed laterally. A. B. intermedius; B. B. borealis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
F65AFAED59895AE297FDEA53ADDA1CA1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518869	https://doi.org/10.3897/zookeys.1267.170946.figure28	Figure 28. Female terminalia viewed laterally. A. Boreus intermedius, with arrow indicating where valvulae narrow; B. Boreus borealis.	Figure 28. Female terminalia viewed laterally. A. Boreus intermedius, with arrow indicating where valvulae narrow; B. Boreus borealis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518837	https://doi.org/10.3897/zookeys.1267.170946.figure1	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	Figure 1. Photos of living Alaskan Boreus spp. A. Female Boreus borealis; B. Male B. borealis; C. Female B. tananaensis sp. nov.; D. Male B. tananaensis sp. nov.; E. Female B. intermedius; F. Female B. gracilis. Photo credits: A: DSS, all others: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518838	https://doi.org/10.3897/zookeys.1267.170946.figure2	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518843	https://doi.org/10.3897/zookeys.1267.170946.figure7	Figure 7. A. Boreus tananaensis sp. nov. female (top) and male (bottom) viewed laterally. B. Collection localities for B. tananaensis. C. Habitat of B. tananaensis. Photo credit: Takeyuki Nakamura.	Figure 7. A. Boreus tananaensis sp. nov. female (top) and male (bottom) viewed laterally. B. Collection localities for B. tananaensis. C. Habitat of B. tananaensis. Photo credit: Takeyuki Nakamura.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518844	https://doi.org/10.3897/zookeys.1267.170946.figure8	Figure 8. Coloration difference in Boreus tananaensis specimens collected in the fall and spring: A. September 9, 2023 (UAM: Ento: 492024); B. September 12, 2024 (UAM: Ento: 504865); C. September 12, 2024 (UAM: Ento: 504856); D. April 27, 2025 (UAM: Ento: 508002). Photo credits: TLK.	Figure 8. Coloration difference in Boreus tananaensis specimens collected in the fall and spring: A. September 9, 2023 (UAM: Ento: 492024); B. September 12, 2024 (UAM: Ento: 504865); C. September 12, 2024 (UAM: Ento: 504856); D. April 27, 2025 (UAM: Ento: 508002). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518845	https://doi.org/10.3897/zookeys.1267.170946.figure9	Figure 9. Boreus tananaensis mating. A. 2 nd phase of copulation (photographed 16 September 2022); B. 3 rd phase of copulation (photographed 27 April 2025) as described by Lestage (1941). Photo credits: TLK.	Figure 9. Boreus tananaensis mating. A. 2 nd phase of copulation (photographed 16 September 2022); B. 3 rd phase of copulation (photographed 27 April 2025) as described by Lestage (1941). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518860	https://doi.org/10.3897/zookeys.1267.170946.figure19	Figure 19. Boreus spp. pronota, viewed laterally, with anterior to the left. A. Without pronotal bristles as seen in B. borealis; B. With pronotal bristles on the posterior margin only as seen in B. timaryi; C. With pronotal bristles on the posterior and anterior margins as seen in B. tananaensis.	Figure 19. Boreus spp. pronota, viewed laterally, with anterior to the left. A. Without pronotal bristles as seen in B. borealis; B. With pronotal bristles on the posterior margin only as seen in B. timaryi; C. With pronotal bristles on the posterior and anterior margins as seen in B. tananaensis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518861	https://doi.org/10.3897/zookeys.1267.170946.figure20	Figure 20. Posterior of male Boreus spp. abdomens viewed laterally. A. 8 th abdominal segments fused, as seen in Boreus timaryi sp. nov. B. 8 th abdominal segments unfused, as seen in B. tananaensis sp. nov.	Figure 20. Posterior of male Boreus spp. abdomens viewed laterally. A. 8 th abdominal segments fused, as seen in Boreus timaryi sp. nov. B. 8 th abdominal segments unfused, as seen in B. tananaensis sp. nov.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518862	https://doi.org/10.3897/zookeys.1267.170946.figure21	Figure 21. Male postabdomen viewed dorsally. A. B. gracilis; B. B. tananaensis sp. nov.	Figure 21. Male postabdomen viewed dorsally. A. B. gracilis; B. B. tananaensis sp. nov.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518863	https://doi.org/10.3897/zookeys.1267.170946.figure22	Figure 22. Head viewed laterally, arrows indicate lateral end of epistomal suture. A. B. gracilis (UAM: Ento: 457835); B. B. tananaensis sp. nov. (UAM: Ento: 504865).	Figure 22. Head viewed laterally, arrows indicate lateral end of epistomal suture. A. B. gracilis (UAM: Ento: 457835); B. B. tananaensis sp. nov. (UAM: Ento: 504865).	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
77B7FFFA98825E1C951F17D3941E12A3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518867	https://doi.org/10.3897/zookeys.1267.170946.figure26	Figure 26. Female terminalia viewed laterally. A. B. gracilis; B. B. tananaensis sp. nov.	Figure 26. Female terminalia viewed laterally. A. B. gracilis; B. B. tananaensis sp. nov.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
9EB93EFDCA7E535D8CB0085CAED2F58A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518838	https://doi.org/10.3897/zookeys.1267.170946.figure2	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	Figure 2. Alaskan Boreus species: Boreus borealis; A. Female (UAM: Ento: 235218); B. Male (UAM: Ento: 235219); Boreus intermedius; C. Female (UAM: Ento: 451067); D. Male (UAM: Ento: 477812); Boreus tananaensis sp. nov.; E. Female (UAM: Ento: 504865); F. Male (UAM: Ento: 504866); Boreus gracilis; G. Female (UAM: Ento: 457836); H. Male (UAM: Ento: 477811); Boreus timaryi sp. nov.; I. Female (UAM: Ento: 406584); J. Male (UAM: Ento: 507961). Photo credits: TLK.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
9EB93EFDCA7E535D8CB0085CAED2F58A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518846	https://doi.org/10.3897/zookeys.1267.170946.figure10	Figure 10. A. Boreus timaryi sp. nov. female (top) and male (bottom) viewed laterally. B. Collection localities for B. timaryi. C. Habitat of B. timaryi holotype. Photo credit: DSS.	Figure 10. A. Boreus timaryi sp. nov. female (top) and male (bottom) viewed laterally. B. Collection localities for B. timaryi. C. Habitat of B. timaryi holotype. Photo credit: DSS.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
9EB93EFDCA7E535D8CB0085CAED2F58A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518860	https://doi.org/10.3897/zookeys.1267.170946.figure19	Figure 19. Boreus spp. pronota, viewed laterally, with anterior to the left. A. Without pronotal bristles as seen in B. borealis; B. With pronotal bristles on the posterior margin only as seen in B. timaryi; C. With pronotal bristles on the posterior and anterior margins as seen in B. tananaensis.	Figure 19. Boreus spp. pronota, viewed laterally, with anterior to the left. A. Without pronotal bristles as seen in B. borealis; B. With pronotal bristles on the posterior margin only as seen in B. timaryi; C. With pronotal bristles on the posterior and anterior margins as seen in B. tananaensis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
9EB93EFDCA7E535D8CB0085CAED2F58A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518861	https://doi.org/10.3897/zookeys.1267.170946.figure20	Figure 20. Posterior of male Boreus spp. abdomens viewed laterally. A. 8 th abdominal segments fused, as seen in Boreus timaryi sp. nov. B. 8 th abdominal segments unfused, as seen in B. tananaensis sp. nov.	Figure 20. Posterior of male Boreus spp. abdomens viewed laterally. A. 8 th abdominal segments fused, as seen in Boreus timaryi sp. nov. B. 8 th abdominal segments unfused, as seen in B. tananaensis sp. nov.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
9EB93EFDCA7E535D8CB0085CAED2F58A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518864	https://doi.org/10.3897/zookeys.1267.170946.figure23	Figure 23. Tergal apophyses A. Present, as seen on B. timaryi (indicated by arrows); B. Absent, as seen on B. borealis.	Figure 23. Tergal apophyses A. Present, as seen on B. timaryi (indicated by arrows); B. Absent, as seen on B. borealis.	2026-01-23	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan		Zenodo	biologists	Kane, Taylor L.;Sikes, Derek S.;Schiff, Nathan			
