identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
164443B41626568BAF8F147971421480.text	164443B41626568BAF8F147971421480.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexapinus clivus Prema & Cubelio & Ng 2026	<div><p>Hexapinus clivus sp. nov.</p><p>Figs 6 C, 6 F, 6 J, 9, 10, 11 A, 12 A, 13 A – C</p><p>Material examined.</p><p>Holotype. • 1 female (8.8 × 5.6 mm) (CMLRE -BC-BRY-8), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.15381&amp;materialsCitation.latitude=11.9605" title="Search Plazi for locations around (long 80.15381/lat 11.9605)">station 11, cruise number 392</a>, dredge, 433 m, sandy bottom, off Tamil Nadu, Bay of Bengal 11°57'37.8"N, 80°09'13.7"E, coll. S. S. Cubelio, 29 November 2019 . Paratype. • 1 juv. female (6.7 × 4.6 mm) (ZRC 2019.1840), from trawl bycatch, Shankarpur coast, West Bengal, India, coll. M. Dev Roy, March 2018 .</p><p>Comparative material.</p><p>Hexapinus simplex Rahayu &amp; Ng, 2014 . Indonesia: Paratypes. • 2 males (8.6 × 5.1 mm, 5.5 × 3.5 mm) (CMLRE -BC-BRY-9); • 2 females (10.1 × 5.8 mm, 6.9 × 3.8 mm), (CMLRE - BC-BRY- 10, ex. ZRC 2010.0297), Ekas, Eastern Lombok, 8 ° 52 ' 09 " 5 N, 116°27'26.1"E; • 1 male (9.3 × 5.8 mm) (ZRC 2014.0203), Sulawesi, Lembeh Straits. coll. H. H. Tan, 17 July 2023; Singapore: • 1 female (10.2 × 5.7 mm) (ZRC 2013.1686), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.85132&amp;materialsCitation.latitude=1.2186" title="Search Plazi for locations around (long 103.85132/lat 1.2186)">St. John Island</a>, north lagoon, 1°13.116'N, 103°51.079'E, coll. P. K. L Ng and A. Anker, 31 May 2013; • 1 ovigerous female (8.0 × 4.5 mm) (ZRC 2013.1687), DR 2, near Raffles Lighthouse, R. V. Galaxea, 20 May 2013; • 1 male (9.1 × 5.7 mm) (ZRC 2014.0227), stn. SEN – AA 04, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.73631&amp;materialsCitation.latitude=1.1750667" title="Search Plazi for locations around (long 103.73631/lat 1.1750667)">Pulau Senang</a>, north side near jetty, intertidal sand-mud flat with abundant rubble and rocks, from burrow together with chaetopterid worm, 0.5 m, yabby pump, 01°10.504'N, 103°44.1790'E, A. Anker, 30 March 2014; • 1 male (4.5 × 3.3 mm) (ZRC 2014.0224), station. 0523 DR 3, between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.09&amp;materialsCitation.latitude=1.40235" title="Search Plazi for locations around (long 104.09/lat 1.40235)">Pulau Tekong</a> east and Pengerang (Malaysia), 10.6–10.7 m, trawl, sandy with broken shell substrate, 01°23.914'N – 01°24.141'N, 104°05.353'E – 104°05.400'E, coll. S. C. Lim et al. 25 March 2014; China: • 1 male (10.3 × 6.8 mm), 1 female (10.3 × 9.0 mm) (ZRC 2012.1016), Taiwan Strait, Fujian Province, Dongshan, pearl shell culture ground, station 75 C – 2368, 1 Jun. 1975; • 1 female (11.4 × 9.8 mm) (ZRC 2012.1017), southeast Hainan <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.5&amp;materialsCitation.latitude=18.5" title="Search Plazi for locations around (long 110.5/lat 18.5)">Island</a>, station 6160, 18°30'N, 110°30'E, South China Sea, coll. D. L. Rahayu; 1 September 2013; Hong Kong: • 1 male (9.1 × 5.6 mm), 1 female (10.3 × 6.7 mm) (ZRC 2019.1102), from Chaetopterus tubes, J. W Qiu, 20 April 2015; Japan: • 2 females (11.3 × 8.8 mm, 10.3 × 8.6 mm) (ZRC 2012.1018), Takebu Amami Islands, 25 April. 2012; • 1 male (9.1 × 8.7 mm) (ZRC 2013.1688), Iriomote Island, Funaura Bay, coll. T. Naruse, 1 July 2011. Hexapinus okinawa Sato, Reimer &amp; Naruse, 2025 . Japan: Holotype. • female (12.0 × 17.6 mm) (RUMF-ZC- 8244), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.911705&amp;materialsCitation.latitude=26.446043" title="Search Plazi for locations around (long 127.911705/lat 26.446043)">Red Beach</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.911705&amp;materialsCitation.latitude=26.446043" title="Search Plazi for locations around (long 127.911705/lat 26.446043)">Kin Bay</a>, Okinawa <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.911705&amp;materialsCitation.latitude=26.446043" title="Search Plazi for locations around (long 127.911705/lat 26.446043)">Island</a>, Ryukyu Islands, Japan, 26.446043°N, 127.911707°E, ca. 10 m, SCUBA and yabbie pump from Chaetopterus tube, coll. T. Sato and H. Nakajima, 16 April 2022 [photographs examined].</p><p>Diagnosis.</p><p>Carapace subquadrate, region indistinct, surface smooth; antero-posterolateral margin protuberance bilobed, densely packed with granules; pterygostomial region with a row of 4 oblique striae, entire; thoracic sternum wide, sternites 1 and 2 fused, subtriangular, laterally forming gentle slope, separated from sternite 3 by distinct curved lateral margin; pleon subovate, all somites free; vulva semilunate, oblique with elevated protrusion. P 2 – P 4 short, stout, P 3 longest; P 4 merus about 2.5 times as long as wide.</p><p>Description.</p><p>Carapace subquadrate, 1.6 times as wide as long, dorsal surface smooth, region indistinct, median H-shaped depression visible; larger granules present on anterolateral margins, only pitted surface on posterior margin (Figs 9 A, 10 A); anterolateral margin entire, arcuate, posterolateral margin subtrapezoidal protuberance with bilobed lateral margin, densely packed with granules; posterior margin weakly convex ventrally, lateral ends concave, accommodating small condyle of thoracic sternite 7 (Figs 9 A, 10 A, 11 A); frontal margin deflexed, establishing triangular lobe in frontal view, forming part of interantennular septum, medially concave, surface smooth; eyes slender, small, slightly movable, pigmented cornea, supra- and infraorbital margins smooth, entire, mesial end of infraorbital margin upwardly produced as subtriangular inner orbital tooth; suborbital and subhepatic regions with granules; pterygostomial region with a row of 4 oblique striae (Fig. 10 B); branchial region covered with very short, dense setae (Fig. 9 C). Third maxillipeds relatively wide, densely granulated, slightly setose, ischium 1.5 times as wide as long, slightly longer than merus, strongly expanded distomesially, with rounded mesial margin; combined length of dactylus, propodus and carpus shorter than that of merus and ischium; combined length of dactylus, propodus and carpus shorter than that of merus and ischium; flagellum well developed (Figs 9 B, 10 B, 10 E). Chelipeds stout, unequal (Fig. 9), dactylus with tufts of setae on upper and lower margins proximally; major chela; slightly gaping when fingers closed; palm slightly wider than long, outer surface densely covered with tubercles; minor chela with wider gap between closed fingers (Fig. 10 C, D). P 2 – P 4 (Fig. 9 A, C) short, stout, P 3 longest; P 4 merus about 2.5 times as long as wide; dactylus of P 2 – P 4 straight (Fig. 10 F).</p><p>Thoracic sternum wide, sparsely pitted medially, and tuberculated marginally (Figs 9 C, 10 G – I, 12 A, 13 A – C), sternites 1 and 2 fused, subtriangular, pointed towards buccal cavity, slightly concave medially, separated from sternite 3 by distinct curved lateral margin; lateral margin of fused sternites 1 and 2 gradually joining that of sternite 3, forming gentle curve; sternite 3 separated from sternite 4 laterally, medially shallow groove, extends small transverse ridges from sternoabdominal cavity to suture of sternites 3 and 4 (Fig. 13 A – C); sternites 4–7 very wide, separated from other somites by distinct granulated sutures, medially appears triangular projection; sternite 8 invisible; sternopleonal cavity subovate, smooth, anterior end spatulate, reaches base of sternite 2 (Figs 10 B, 10 E, 10 H, 13 A – C); sternopleonal locking spine well developed, hooked structure on suture 4 / 5, appears close to vulva; vulva located on suture 5 / 6, below to locking spine, medially closure, elevated protrusion, semilunate, oblique (Fig. 13 A – C).</p><p>Pleon subovate, all somites free, lateral margins covered with long setae; somites 3, 4 widest; laterally somite 3 slightly concave, somite 4 straight; somite 6 longest, subtrapezoidal, convergent distally, lateral margin weakly concave; telson subovate, shorter than somite 6 (Fig. 9 C, G).</p><p>Colour.</p><p>Unknown.</p><p>Variation.</p><p>The small paratype female (ZRC 2019.1840) is similar to the holotype female (CMLRE -BC-BRY-8) except that its pleon is less expanded, the pleopods are poorly developed, and the vulvae do not appear mature.</p><p>Etymology.</p><p>The species name is derived from the Latin word clivus, sloping side of a mountain; alluding to the sides of the triangular structure of thoracic sternites 1 and 2, which is a diagnostic character of the species. The name is used as a noun in apposition.</p><p>Type locality.</p><p>Off, Kochi, Kerala, Arabian Sea.</p><p>Distribution.</p><p>Presently known only from the type locality.</p><p>Remarks.</p><p>The recognition of the present new species is connected to the identity of Hexapus edwardsi, which was described from a single small subadult male 4.8 × 3.0 mm from the Andaman Sea. Rahayu and Ng (2014: 409) transferred the species to Hexapinus based on the structure of the third maxilliped as well as the form of the male pleon. The male pleon as figured by Serène and Soh (1976: 25, fig. 27 B) is unusual because somite 5 seems separate from fused somites 3 and 4 (somite 1 apparently not figured). In typical Hexapinus, male pleonal somites 3–5 are fused with somite 6 elongate (e. g., see Rahayu and Ng 2014: fig. 21 E – G). Even in small subadult males, this is apparent (Rahayu and Ng 2014: fig. 21 E). Females, on the other hand, have all the somites free, so it seems unlikely Serène and Soh’s (1976) type specimen is a female because at least somites 3 and 4 are fused. Whether the figure in Serène and Soh (1976) is accurate will require re-examination of the specimen. Serène and Soh (1976: 25) described the anterolateral margin of H. edwardsi as “ like an open continuous arch; the lateral borders converge regularly forward and are not parallel ” and “ the antero-lateral region and the external surface of the palm of chelipeds are ornamented with a covering of acute granules, disappearing under a short tomentum ”. This description generally matches that of Hexapinus clivus sp. nov., except that its carapace appears somewhat more rectangular with the anterolateral margins more angular (Figs 9 A, 10 A) whereas in H. edwardsi, the carapace is more semicircular with the anterolateral margin clearly more rounded (cf. Serène and Soh 1976: pl. 7 D). In addition, the merus of the third maxilliped is distinctly more quadrate and longer (Figs 6 F, 10 B, 10 E) than that of H. edwardsi (cf. Serène and Soh 1976: fig. 27 A). Hexapinus clivus sp. nov. is clearly close to H. edwardsi but the differences noted here indicate that we should treat them as different species for the time being.</p><p>The characters of the present new species agree best with two West Pacific species: H. simplex Rahayu &amp; Ng, 2014 (from East and Southeast Asia) and H. okinawa Sato, Reimer &amp; Naruse, 2025 (from the Ryukyu Islands), but it can be clearly separated by a suite of distinct characters. Hexapinus clivus sp. nov. is distinguished with its subquadrate carapace, as wide as long, with lateral margins moderately curved and densely granulated (Figs 9 A, 10 A, 11 A) (versus carapace elongated, lateral margins slightly curved and granulated in H. simplex; Fig. 11 B; with carapace wider, lateral margin deeply curved and moderately granulated in H. okinawa; Fig. 11 C); the third maxilliped ischium as wide as long and is strongly expanded distomesially with a rounded mesial margin (Figs 6 F, 10 B, 10 E) (versus third maxilliped ischium 1.1 times as wide as long, slightly shorter in H. simplex; Fig. 12 B; and ischium 0.7 times as long as ischium and much shorter in H. okinawa; Fig. 12 D); the lateral margin of fused female thoracic sternites 1 and 2 gradually joining lateral margin of sternite 3 forming a gentle curve (Fig. 12 A) (versus lateral margin of fused sternites 1 and 2 joining that of sternite 3 at a distinct obtuse angle in H. simplex; Fig. 12 B, C; and H. okinawa; Fig. 12 D); P 3 much longer than P 4, lower margin with serrated granules, and P 4 about as long as wide (Figs 9 A, 9 C, 10 F, 11 A) (versus P 3 slightly longer than P 4, lower margin smooth, P 4 shorter in H. simplex; Fig. 11 B; and P 3 slightly longer than P 4, lower margin with serrated granules, with P 4 short and wide in H. okinawa; Fig. 11 C); and the vulva semilunate with elevated protrusion, medially closed, with a well-developed, hook-like locking spine (Fig. 13 A – C) (versus vulva subovate without protrusion, medially wider, and locking spine not hook-like in adult H. simplex; Fig. 13 D – F; with the condition in juveniles subovate, not fully developed with the locking spine weakly hooked; Fig. 13 G, H; and vulva elliptical, with posterolateral margin elevated to form hood-like protrusion in H. okinawa; Sato et al. 2025: fig. 1 D).</p></div>	https://treatment.plazi.org/id/164443B41626568BAF8F147971421480	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Cubelio, Sherine Sonia;Ng, Peter K. L.	Prema, Mani, Cubelio, Sherine Sonia, Ng, Peter K. L. (2026): First records of the genera Mariaplax Rahayu & Ng, 2014 and Hexapinus Manning & Holthuis, 1981 from India, with descriptions of three new species (Crustacea, Decapoda, Brachyura, Hexapodidae). ZooKeys 1292: 349-372, DOI: 10.3897/zookeys.1292.204005
4EABB2017A10567A8B69C964236A298F.text	4EABB2017A10567A8B69C964236A298F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexapinus Manning & Holthuis 1981	<div><p>Genus Hexapinus Manning &amp; Holthuis, 1981</p><p>Type species.</p><p>Hexapinus latipes (De Haan, 1835) (type species Hexapus latipes De Haan, 1835, by original designation; gender masculine).</p><p>Remarks.</p><p>The genus Hexapinus Manning &amp; Holthuis, 1981 is represented by seven known species, only three species, H. edwardsi (Serène &amp; Soh, 1976), H. simplex Rahayu &amp; Ng, 2014, and H. okinawa Sato, Reimer &amp; Naruse, 2025, possess a proportionately wider carapace and a granulated anterolateral margin with a bilobed angle on the posterolateral margin (see Rahayu and Ng 2014; Rahayu and Widyastuti 2018; Sato et al. 2025). We here recognise a fourth species from this group.</p></div>	https://treatment.plazi.org/id/4EABB2017A10567A8B69C964236A298F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Cubelio, Sherine Sonia;Ng, Peter K. L.	Prema, Mani, Cubelio, Sherine Sonia, Ng, Peter K. L. (2026): First records of the genera Mariaplax Rahayu & Ng, 2014 and Hexapinus Manning & Holthuis, 1981 from India, with descriptions of three new species (Crustacea, Decapoda, Brachyura, Hexapodidae). ZooKeys 1292: 349-372, DOI: 10.3897/zookeys.1292.204005
F4BEE1FB1AF05682B49FB24D2A177FA3.text	F4BEE1FB1AF05682B49FB24D2A177FA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mariaplax albucus Prema & Cubelio & Ng 2026	<div><p>Mariaplax albucus sp. nov.</p><p>Figs 4, 5, 6 B, E, J, 7 D – F, 8 D – F</p><p>Material examined.</p><p>Holotype. • 1 female (6.2 × 3.5 mm) (CMLRE -BC-BRY-6) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=75.592&amp;materialsCitation.latitude=9.965" title="Search Plazi for locations around (long 75.592/lat 9.965)">station 24, cruise number 343</a>, grab, 30 m, sandy bottom, off Kochi, Arabian Sea, 9°57'54.0"N, 75°35'31.2"E, 16 August 2015 . Paratypes. • 2 females (5.8 × 3.2 mm, 6.3 × 3.5 mm, both fragile) (CMLRE -BC-BRY-7), same data as holotype .</p><p>Comparative material.</p><p>Mariaplax sundaica Rahayu &amp; Widyastuti, 2018: • 1 female (4.4 × 2.5 mm) (ZRC 2018.0988) station DW 06, on mud and clay, 266–294 m, 5°43.413'S, 104°49.712'E – 5°43.779'S, 104°50.060'E, Sunda Strait (between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.834335&amp;materialsCitation.latitude=-5.72965" title="Search Plazi for locations around (long 104.834335/lat -5.72965)">Tabuan Island</a> and Sumatra), Java, coll. SJADES 2018 cruise, Baruna Jaya VIII, 25 March 2018; • 1 female (3.8 × 2.3 mm), (ZRC 2018.0989), on mud, station DW 01, 100–104 m, 6°20.489'S, 105°26.890'E – 6°20.431'S, 105°26.214'E, Sunda Strait (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.4369&amp;materialsCitation.latitude=-6.3405166" title="Search Plazi for locations around (long 105.4369/lat -6.3405166)">South of Krakatau Island</a>), Java, coll. SJADES 2018 cruise, Baruna Jaya VIII, 25 March 2018; • 1 female (3.8 × 2.4 mm) (ZRC 2020.0398), station DW 06, on mud and clay, Sunda Strait, between Tabuan Island and Sumatra, Java, Indonesia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.834335&amp;materialsCitation.latitude=-5.72965" title="Search Plazi for locations around (long 104.834335/lat -5.72965)">Indian Ocean</a>, 5°43.413'S, 104°49.712'E – 5°43.779'S, 104°50.060'E, 266–294 m, coll. SJADES 2018 cruise, Baruna Jaya VIII, 25 March 2018.</p><p>Diagnosis.</p><p>Carapace as wide as long, dorsal surface strongly granulated, pterygostomial with 6 oblique striae, P 4 merus short and stout, vulvae large with operculum cover, located distally on bulged, sloped margin between sternites 6–8.</p><p>Description.</p><p>Carapace subquadrate, ca. 1.8 times as wide as long, dorsal surface strongly granulated; regions indistinct, with shallow grooves on branchial region (Figs 4 A – C, 5 A, 6 B, 8 D). Eyes small, slightly movable, cornea black, equal in size to granular peduncle (Fig. 5 B). Pterygostomial regions (Figs 4 B, 5 B, 8 E) relatively smooth, covered by granules, with 6 oblique striae. Epistome longer, pointed distally, with bilobed lateral margin (Figs 5 B, 8 E). Third maxillipeds wide (Figs 5 B, 6 E, 8 E); merus slightly rectangular; ischium longer than merus, more pronounced subproximally, with oblique mesial margin, combined length of carpus, propodus and dactylus, shorter than merus and ischium; exopod relatively longer, stouter, granulated margins, flagellum well developed. Chelipeds asymmetrical, covered by dense tubercles with fringe of setae (Figs 4 B, 5 C, 5 D); major chela (Fig. 5 C) with gaping finger when closed, cutting edge at third proximal with 2 larger teeth; both dactyli with longitudinal groove medially, almost reaching proximal part but not to tip, tubercles on proximal third; palm as long as wide; outer surface of palm covered with closely-spaced tubercles; carpus outer surface with closely-spaced granules with denticulate mesial margin; merus short, covered with granules; minor chela (Fig. 5 D) with relatively narrower gap when fingers closed; both dactyli with longitudinal ridge medially, not reaching proximal part, cutting edges of dactylus and fixed finger with row of small teeth, outer surface of palm, carpus and merus covered by closely-spaced tubercles; fringe of setae on each upper inner margin of merus. P 2 – P 4 (Figs 5 E, 8 F) relatively long, slender, covered by dense tubercles, dactylus gently upcurved; P 3 longest; dactylus of P 2, P 3 longer than propodus and carpus; merus with distinct longitudinal median groove on dorsal view in P 2, merus with distinct median groove on lateral and dorsal face in P 3 (Fig. 5 E); P 4 dactylus same in length as propodus and carpus, margins denticulate, merus relatively stout, about 4.0 times longer than wide, with distinct longitudinal median groove on latero-dorsal surface (Figs 4 E, 8 F).</p><p>Thoracic sternum wide (Figs 5 F, 7 D), surface densely packed with tubercles, sternites 1 and 2 fused, separated from sternite 3 by shallow groove; sternite 3 separated from sternite 4; sternites 4–7 well developed, separated from each other by well-developed suture; sternite 8 not exposed; sternopleonal cavity smooth, reaches base of sternite 3 (Figs 5 H, 7 D – E). Vulvae large, operculum cover present, located distally on bulged, sloping region of inner margin between sternites 6–8, on sternite 5; pleonal locking spine on sternite 4 (Figs 5 H, 7 D – F). Pleon wide, granulated, with 6 free somites, somite 3 widest; telson longer than wide, semicircular (Fig. 6 J). Male unknown.</p><p>Colour.</p><p>Unknown.</p><p>Variation.</p><p>Males are unknown for this species, and the type females do not vary substantially among the known specimens.</p><p>Etymology.</p><p>The name of the species is derived from the Latin word albucus, bulb of the lily; alluding to the structure of the bulging vulva which is diagnostic of the species. The name is used as a noun in apposition.</p><p>Type locality.</p><p>Off Kochi, Kerala, Arabian Sea.</p><p>Distribution.</p><p>Presently known only from the type locality.</p><p>Remarks.</p><p>Mariaplax albucus sp. nov. differs from M. sundaica and M. kochazhi sp. nov. in that the subquadrate carapace, about 1.8 times as wide as long (Figs 4 A, 5 A, 6 B, 8 D) (versus the carapace about 1.7 times as long as wide in M. sundaica; Fig. 8 G; see Rahayu and Widyastuti 2018: figs 9 A, B, 10 A; and 1.5–1.7 times as wide as long in M. kochazhi; Fig. 8 A); the pterygostomial region is granulated with a row of six oblique striae M. albucus sp. nov. (Figs 4 B, 5 B, 8 E) (versus with relatively reduced granulation and a row of four oblique striae in M. sundaica; Fig. 8 H; Rahayu and Widyastuti 2018: figs 9 B, F; and with 5–7 oblique striae with the upper margin weakly granulated in M. kochazhi sp. nov.; Figs 1 C, 8 B); the epistome is relatively longer, distally pointed, with the lateral margins distinctly bilobed (Figs 4 B, 5 B, 8 E) (versus epistome relatively shorter, distally pointed and the lateral margins entire in M. sundaica; Fig. 8 H; see Rahayu and Widyastuti 2018: fig. 9 F; and being distally more pointed with weakly bilobed lateral margins in M. kochazhi sp. nov.; Figs 1 C, 8 B); the ischium of the third maxilliped as long as wide, more rectangular, with a more pronounced subproximal expansion and forming a triangular projection (Figs 5 B, 6 E, 8 E) (versus the ischium of the third maxilliped slightly narrower, with the merus as long as wide in M. sundaica; Fig. 8 H, see Rahayu and Widyastuti 2018: figs 9 B; and the ischium slightly longer (0.7 times width) and more rectangular in shape, with more densely packed larger granules on the margins in M. kochazhi sp. nov.; Figs 1 D, 6 D, 8 B); the P 4 merus is comparatively longer, 4.0 times width with the median groove distinct on the lateral surface and visible in the dorsal surface, with densely packed granules, dactylus slightly wide and upcurved (Figs 4 A, 5 C, 8 F) (versus P 4 merus short, 3.4 times as long as wide, the longitudinal groove being absent on the lateral surface, dactylus wide, short, upcurved, spatuliform in M. sundaica; Fig. 8 I, see Rahayu and Widyastuti 2018: figs 9 A, 10 H; and the P 4 merus is short and stout, with deep grooves on lateral surface of the merus, densely packed granules, the dactylus being gently upcurved and narrow in M. kochazhi sp. nov.; Fig. 8 C); and the vulva is comparatively large, with a well-developed operculum cover and positioned distally on the surface of the bulged, sloped margin of sternite 5 (Figs 5 H, 7 D – F) (versus vulva large, subovate in shape, operculum absent and positioned on the surface of slightly bulged, sloped distal margin of sternite 5 in M. sundaica; Rahayu and Widyastuti 2018: fig. 9 E; and vulva is simple, rounded, completely exposed, with the operculum and the bulged sloping area absent in M. kochazhi; Figs 2 C, 7 A – C).</p></div>	https://treatment.plazi.org/id/F4BEE1FB1AF05682B49FB24D2A177FA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Cubelio, Sherine Sonia;Ng, Peter K. L.	Prema, Mani, Cubelio, Sherine Sonia, Ng, Peter K. L. (2026): First records of the genera Mariaplax Rahayu & Ng, 2014 and Hexapinus Manning & Holthuis, 1981 from India, with descriptions of three new species (Crustacea, Decapoda, Brachyura, Hexapodidae). ZooKeys 1292: 349-372, DOI: 10.3897/zookeys.1292.204005
3AA92146A08D55E5A4DB3C26B62FC675.text	3AA92146A08D55E5A4DB3C26B62FC675.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mariaplax kochazhi Prema & Cubelio & Ng 2026	<div><p>Mariaplax kochazhi sp. nov.</p><p>Figs 1, 2, 3, 4, 6 A, D, G, H, K – N, 7 A – C, 8 A – C</p><p>Material examined.</p><p>Holotype. • 1 male (5.5 × 3.5 mm) (CMLRE -BC-BRY-1), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=75.592&amp;materialsCitation.latitude=9.965" title="Search Plazi for locations around (long 75.592/lat 9.965)">station 24, cruise number 343</a>, grab, 30 m, sandy bottom, off Kochi, Arabian Sea, 9°57'54.0"N, 75°35'31.2"E, 16 August 2015 . Paratypes. • 1 female (5.3 × 2.8 mm) (CMLRE -BC-BRY-2), 1 male (6.7 × 4.0 mm) (CMLRE -BC-BRY-3), same data as holotype . Non-types. • 6 females (10.1 × 5.7 mm, 6.6 × 4.1 mm, 5.9 × 3.0 mm, 4.9 × 2.8 mm, 4.5 × 2.6 mm, 4.0 × 2.1 mm), (CMLRE -BC-BRY-4) 1 male (5.0 × 3.1 mm) (CMLRE -BC-BRY-5) same data as holotype (all very fragile so not suitable to be type material).</p><p>Diagnosis.</p><p>Carapace subtrapezoidal, 1.5–1.7 times as wide as long, dorsal surface granular; pterygostomial region smooth with 5–7 striae; third maxilliped ischium rectangular, merus granulated; telson of male pleon subpentagonal; female pleon telson rounded; G 1 strongly curved, S-shaped, distal tip rounded; vulvae simple, rounded, fully exposed, operculum absent.</p><p>Description.</p><p>Carapace subtrapezoidal, about 1.5–1.7 times as wide as long, dorsal surface densely granulated, anterolateral margin lined with distinct rounded granules; regions indistinct, median H-shaped elongated furrow towards anterolateral margin visible (Figs 1 A, 1 B, 2 A, 6 A, 8 A). Each lobe of frontal margin with gently convex margin; antero-posterolateral margins arcuate, armed with larger tubercles; posterolateral margin with corner prominently protruding, forming distinct angle before strongly curved towards posterior margin (Fig. 1 B); eye small, slightly movable, globular cornea, peduncle granular (Fig. 1 C). Pterygostomial region generally smooth, with 5–7 oblique, long striae, upper margin with scattered granules; epistome longer, more pointed medially, shallowly bilobed lateral margins (Figs 1 C, 8 B). Third maxillipeds as long as wide, almost covering buccal cavity (Figs 1 C, 1 D, 6 D, 8 B); ischium of third maxillipeds distinctly wide, longer than merus, mesial margin straight; combined length of dactylus, propodus, and carpus relatively shorter than that of merus and ischium; exopod relatively narrow, flagellum well developed. Chelipeds stout, unequal (Fig. 1 F, G); major chela (Fig. 1 F) forming gap when fingers closed; dactylus and fixed finger with longitudinal groove on outer surface; palm slightly longer than wide, covered by sparse granules; minor chela (Fig. 1 G) with small gap between closed fingers, dactylus and fixed finger with shallow, wider, longitudinal groove, palm more or less equally wide to its length, covered by densely packed, prominent, larger granules. P 2 – P 4 (Figs 1 A, 2 A, 8 C) relatively short, slender; deep longitudinal groove on merus with granular outer surface; P 3 merus outer surface distinctly separated by granules, carpus and propodus smooth; P 4 merus, carpus, propodus densely packed granules, with dactylus gently upcurved.</p><p>Male thoracic sternum wide (Fig. 1 E), covered by dense tubercles; sternites 1 and 2 fused, separated from sternite 3 by distinct ridge, sternal spine for locking pleon present; thoracic sternal groove (anterolateral extension of sterno-pleonal cavity to accommodate G 1) wide, deep, extends from sternopleonal cavity to base of sternite 3 (Fig. 1 D, E); sternite 4 wide, sternopleonal cavity reaching distal margin of sternites 3. Male pleon relatively wide (Figs 1 E, 6 G), somites 1 and 2 free, somites 3–5 completely fused, wider than somite 6, lateral margin slightly convex to sinuous, somite 6 pentagonal, shorter than length of somites 3–5, lateral margin subproximally convex to blunt triangular projection; telson lateral margin slightly sinuous, lined with larger granules, distolateral margin gently concave, forming subpentagonal structure, setose densely.</p><p>G 1 (Figs 3 A – D, 6 K – M), curved outwards medially at 60–70 °, basal part stout, gently curved outwards, outer margin smooth, inner margin slightly serrated, curved; distal half slender, margin covered by straight, long and short plumose setae; distal tip broad, rounded, directed upwards, apex not concealed under closed pleon, exposed on thoracic sternal groove, obscured by setae of telson. G 2 (Figs 3 E – G, 6 N) small, slightly curved, ridge separates distal region and tip.</p><p>Female thoracic sternum (Figs 2 B, 2 C, 7 A – C), wide, densely covered by tubercles or rounded granules, sternites 1 and 2 fused, separated from sternite 3 by distinct ridge; sternopleonal cavity wide, smooth, reached to sternite 3; thoracic sutures 4 / 5 and 5 / 6 parallel and directed slightly posteromesially outside sternopleonal cavity, then bent anteromesially in sternopleonal cavity. Vulvae simple, rounded and completely exposed, operculum absent, located adjacent to mesial end of sutures 4 / 5 and 5 / 6, which is level with middle of sternite 5 outside cavity, pleonal locking spine present, on sternite 5 (Figs 2 C, 7 A – C); female pleon with 6 free somites and telson with rounded distal margin (Figs 2 B, 6 H).</p><p>Variation.</p><p>No obvious non-sexual differences exist between the sexes.</p><p>Colour.</p><p>Unknown.</p><p>Etymology.</p><p>The species is named after its type locality, off the coast of Kochi, using the original Malayalam form. The name Kochi is derived from the Malayalam word ക ൊ ച ് ച ു ഴ ി or “ Koch-azhi, ” meaning “ small sea ” (kochu = small; azhi = sea). Kochi (formerly Cochin), a significant port city historically known as the “ Queen of the Arabian Sea, ” lies along the southwestern coast of India. The name is treated as a noun in apposition.</p><p>Type locality.</p><p>Off Kochi, Kerala, Arabian Sea.</p><p>Distribution.</p><p>Presently known only from the type locality.</p><p>Remarks.</p><p>The general appearance of Mariaplax kochazhi sp. nov., with its carapace shape, distinctly granulated surface and structure of the G 1, allies it most closely with M. granulifera Campbell &amp; Stephenson, 1970, originally described from Queensland, Australia (Campbell and Stephenson 1970; Rahayu and Ng 2014). Mariaplax kochazhi sp. nov. however, differs in that the carapace is slightly wider (1.5–1.7 times), with relatively larger granules on the dorsal surface and the median H-shaped depression deep with a more elongated groove (Figs 1 A, 1 B, 2 A, 6 A) (versus carapace relatively narrower, 1.3 times as wide as long, more quadrate, with relatively smaller granules and the H-shaped median depression less prominent without an elongated groove in M. granulifera; see Rahayu and Ng 2014: figs 34 A, 35 A); the ischium of the third maxilliped is slightly longer (0.7 times width) and more rectangular in shape, with more densely packed larger granules on the margins (Figs 1 D, 6 D) (versus ischium relatively shorter (0.5 times width), more quadrate in shape with only sparsely distributed granules on the margins in M. granulifera; see Rahayu and Ng 2014: figs 35 C, 36 E); the distolateral margin of the male telson is gently concave with the structure appearing subpentagonal (Fig. 6 G) (versus clearly rounded in M. granulifera; see Rahayu and Ng 2014: fig. 36 D); and the distal tip of the G 1 is clearly more rounded with the basal part proportionately stouter and the outer margin more strongly curved (Figs 3 A – D, 6 K – M) (versus part more pointed, with the basal part of the G 1 more slender and outer margin less curved in M. granulifera; see Rahayu and Ng 2014: fig. 36 I, J).</p><p>In the proportions of the carapace, M. kochazhi sp. nov. is also similar to M. hispida, M. propinqua, and M. chenae, but they are very different in other characters. In M. hispida, the carapace is subquadrate in shape, the third maxilliped ischium is proportionately longer, and the vulva is semilunate in shape (see Rahayu and Widyastuti 2018: fig. 6 A – E) (carapace subtrapezoidal, third maxilliped ischium shorter and vulva rounded in M. kochazhi sp. nov.; Figs 1 A – D, 2 C, 6 A, 6 D, 7 A – C). In M. propinqua, the carapace is distinctly more tuberculate, the pterygostomial region has four oblique short striae and the upper margin has scattered granules, and the G 1 is S-shaped with a tapered distal part and medially, there are strong three large teeth present (see Rahayu and Ng 2014: figs 43, 45 I, J) (carapace relatively less tuberculate, the pterygostomial region has 5–7 long oblique striae with the upper margin covered with scattered granules, (Fig. 1 C); and the G 1 is curved outwards medially with the distal tip broad and rounded, without teeth on the medial margin in M. kochazhi sp. nov.; Figs 3 A – D, 6 K – M). In M. chenae, the carapace is subquadrate in shape, the pterygostomial region has four oblique short striae with fewer granules on the upper margins, the G 1 is S-shaped distally, with a tapered distal part and there is a row spines medially (see Rahayu and Ng 2014: figs 25 A, B, I, J) (versus carapace subtrapezoidal, the pterygostomial region has 5–7 long oblique striae with the upper margin covered with scattered granules, and the G 1 is curved outwards medially with the distal tip broad and rounded, without teeth on the medial margin in M. kochazhi sp. nov.; Figs 1, 3, 6 K – M).</p><p>The vulva of M. kochazhi sp. nov. (Figs 2 C, 7 A – C), being slightly bulging, gently sloping mesially, somewhat subtriangular in form and lacking an operculum, with the two vulvae well separated and positioned adjacent to the mesial end of sutures 4 / 5 and 5 / 6 and level to the middle part of sternite 5, is similar in structure to that of M. chenae (see Rahayu and Widyastuti 2018: fig. 4 D – F). In this aspect, its vulva is also similar to M. pitrai Rahayu &amp; Widyastuti, 2018, although that of the latter is relatively larger and more subcircular vulvae in shape (see Rahayu and Widyastuti 2018: fig. 4 C). The vulvae of M. hispida, M. propinqua, M. chenae and M. pitrai, however, are not bulging, with the surfaces not sloping mesially between sternites 6–8 in the sternopleonal cavity.</p></div>	https://treatment.plazi.org/id/3AA92146A08D55E5A4DB3C26B62FC675	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Cubelio, Sherine Sonia;Ng, Peter K. L.	Prema, Mani, Cubelio, Sherine Sonia, Ng, Peter K. L. (2026): First records of the genera Mariaplax Rahayu & Ng, 2014 and Hexapinus Manning & Holthuis, 1981 from India, with descriptions of three new species (Crustacea, Decapoda, Brachyura, Hexapodidae). ZooKeys 1292: 349-372, DOI: 10.3897/zookeys.1292.204005
1C14583FD80F5FB49A5F1EAD3A9A2839.text	1C14583FD80F5FB49A5F1EAD3A9A2839.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mariaplax Rahayu & Ng 2014	<div><p>Genus Mariaplax Rahayu &amp; Ng, 2014</p><p>Type species.</p><p>Lambdophallus anfractus Rathbun, 1909; type by original designation.</p><p>Remarks.</p><p>The genus Mariaplax is currently represented by 19 species, all from the Indo-West Pacific (cf. Rahayu and Ng 2014; Ng and Rahayu 2015; Rahayu and Widyastuti 2018; Ng and Wong 2019). The relatively wide and granular carapaces of the two new species, M. kochazhi sp. nov. and M. albucus sp. nov. (1.5–1.8 times longer than wide) ally them with M. chenae Rahayu &amp; Ng, 2014, M. narusei Rahayu &amp; Ng, 2014, M. propinqua Rahayu &amp; Ng, 2014, M. secus Rahayu &amp; Ng, 2014, M. hispida Rahayu &amp; Widyastuti, 2018, and M. sundaica Rahayu &amp; Widyastuti, 2018, all of which have carapaces that are 1.5 times or more longer than wide (see Rahayu and Ng 2014; Rahayu and Widyastuti 2018). In general, the appearance of M. albucus sp. nov., with its wider rectangular carapace with granulation pattern, third maxilliped, and sternopleonal cavity, resembles M. sundaica, which was originally described from Selat Sunda, Indonesia (Rahayu and Widyastuti 2018); while M. albucus sp. nov. is closer to M. kochazhi sp. nov. by its wider, densely tuberculated carapace and wider sternopleonal cavity. Detailed comparisons of the two new species with congeners are done under the Remarks section for each species.</p><p>Rahayu and Ng (2014: 442) tentatively assigned the record of “ Hexapus? granuliferus ” from the Andaman Sea by Serène and Soh (1976) to it because of the shapes of the third maxilliped and female pleon, but it needs to be verified because this record is some distance from the type locality, Queensland, Australia. It is possible that this record is actually M. anfracta (Rathbun, 1909) as there is a record of this species from nearby Penang (Rahayu and Ng 2014: 424), conspecific with one of the two new species of Mariaplax described here, or even an undescribed taxon.</p></div>	https://treatment.plazi.org/id/1C14583FD80F5FB49A5F1EAD3A9A2839	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Prema, Mani;Cubelio, Sherine Sonia;Ng, Peter K. L.	Prema, Mani, Cubelio, Sherine Sonia, Ng, Peter K. L. (2026): First records of the genera Mariaplax Rahayu & Ng, 2014 and Hexapinus Manning & Holthuis, 1981 from India, with descriptions of three new species (Crustacea, Decapoda, Brachyura, Hexapodidae). ZooKeys 1292: 349-372, DOI: 10.3897/zookeys.1292.204005
