taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
D595BEB991E15B2EA03F48844D6B95B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666611	https://doi.org/10.3897/zookeys.1281.175455.figure5	Figure 5. Micro- CT images of skull, occipital bone, and pectoral region of male Calotes thailandensis sp. nov. (A, C. CUMZ-R - 2738 and E, G. CUMZ-R - 2769) and male C. maehongsonensis sp. nov. (B, D. CUMZ-R - 2825 and F, H. CUMZ-R - 2820). Abbreviations: Cl, Clavicle; F, Frontal; In, Interclavicle, Mx, Maxilla; Na, Nasal; P, Parietal, Px, Premaxilla; So, Supraoccipital bones. Scale bars: 5 mm.	Figure 5. Micro- CT images of skull, occipital bone, and pectoral region of male Calotes thailandensis sp. nov. (A, C. CUMZ-R - 2738 and E, G. CUMZ-R - 2769) and male C. maehongsonensis sp. nov. (B, D. CUMZ-R - 2825 and F, H. CUMZ-R - 2820). Abbreviations: Cl, Clavicle; F, Frontal; In, Interclavicle, Mx, Maxilla; Na, Nasal; P, Parietal, Px, Premaxilla; So, Supraoccipital bones. Scale bars: 5 mm.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
D595BEB991E15B2EA03F48844D6B95B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666613	https://doi.org/10.3897/zookeys.1281.175455.figure7	Figure 7. Ventral head and lateral views (A – F) of paratypes of Calotes thailandensis sp. nov. (CUMZ-R - 2769 [SA 149], 2872 [SA 221], 2767 [SA 147]) (A – C) and Calotes maehongsonensis sp. nov. (CUMZ-R - 2820 [SA 169], 2827 [SA 176], 2839 [SA 188]) (D – F) are showed consisting of adult males (A, B, D, E) and adult females (C, F). Scale bars: 10 mm.	Figure 7. Ventral head and lateral views (A – F) of paratypes of Calotes thailandensis sp. nov. (CUMZ-R - 2769 [SA 149], 2872 [SA 221], 2767 [SA 147]) (A – C) and Calotes maehongsonensis sp. nov. (CUMZ-R - 2820 [SA 169], 2827 [SA 176], 2839 [SA 188]) (D – F) are showed consisting of adult males (A, B, D, E) and adult females (C, F). Scale bars: 10 mm.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
D595BEB991E15B2EA03F48844D6B95B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666615	https://doi.org/10.3897/zookeys.1281.175455.figure9	Figure 9. Holotype of adult male Calotes maehongsonensis sp. nov. (CUMZ-R - 2825 [SA 174]). A. Head dorsal; B. Head ventral; C. Head lateral; D. Full body dorsal; E. Body ventral, and F. Lateral region. Scale bars: 10 mm.	Figure 9. Holotype of adult male Calotes maehongsonensis sp. nov. (CUMZ-R - 2825 [SA 174]). A. Head dorsal; B. Head ventral; C. Head lateral; D. Full body dorsal; E. Body ventral, and F. Lateral region. Scale bars: 10 mm.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
D595BEB991E15B2EA03F48844D6B95B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666616	https://doi.org/10.3897/zookeys.1281.175455.figure10	Figure 10. Habitat (A) of the type locality and male (B) (CUMZ-R - 2820) and female (C) (CUMZ-R - 2839) of Calotes maehongsonensis sp. nov. at Khun Yuam District, Mae Hong Son Province, Thailand.	Figure 10. Habitat (A) of the type locality and male (B) (CUMZ-R - 2820) and female (C) (CUMZ-R - 2839) of Calotes maehongsonensis sp. nov. at Khun Yuam District, Mae Hong Son Province, Thailand.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
D595BEB991E15B2EA03F48844D6B95B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666610	https://doi.org/10.3897/zookeys.1281.175455.figure4	Figure 4. Females: PCA (A, C) and DAPC (B, D) of female Calotes species based on mensural (A, B) and meristic (C, D) characters, respectively, containing C. irawadi from central Myanmar (n = 14), C. thailandensis sp. nov. from central and southern Thailand (n = 27) and C. maehongsonensis sp. nov. from Mae Hong Son, northwestern Thailand (n = 10). PERMANOVA support the significantly morphological difference with p < 0.001.	Figure 4. Females: PCA (A, C) and DAPC (B, D) of female Calotes species based on mensural (A, B) and meristic (C, D) characters, respectively, containing C. irawadi from central Myanmar (n = 14), C. thailandensis sp. nov. from central and southern Thailand (n = 27) and C. maehongsonensis sp. nov. from Mae Hong Son, northwestern Thailand (n = 10). PERMANOVA support the significantly morphological difference with p < 0.001.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
D595BEB991E15B2EA03F48844D6B95B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666607	https://doi.org/10.3897/zookeys.1281.175455.figure1	Figure 1. Sample locality map of Calotes thailandensis sp. nov. (blue) (1–9), C. maehongsonensis sp. nov. (brown) (10–15), C. irawadi (white) (16–20), C. wangi wangi (21–23) (red), and C. wangi hainanensis (24–26) (yellow) from Thailand and nearby countries. Symbols correspond to molecular (triangle for ND 2; circle for COI genes) and morphological (star) analyses. Localities include Thailand: Songkhla (type locality of C. thailandensis sp. nov.) (1), Krabi (2), Ranong (3), Chon Buri (4), Saraburi (5), Kamphaeng Phet (6), Sakon Nakhon (7), Mukdahan (8), Roi Et (9), Mae Hong Son (type locality of C. maehongsonensis sp. nov.) (10), and Kanchanaburi (11); Myanmar: Mon, Mawlamyine (12), Mon, Kyaik (13), Bago (14), Yangon (15), Mandalay (16), Sagaing (type locality of C. irawadi) (17), Sagaing (18), and Magway (19); China: Western Yunnan (20); Vietnam: Lang Son (21); China: Nanning, Guangxi (22), Hong Kong (23); China: Wanling, Hainan (24), Jiachai, Hainan (25), Tunchang, Hainan (26). Details of all samples used in this study are given in Table 1.	Figure 1. Sample locality map of Calotes thailandensis sp. nov. (blue) (1–9), C. maehongsonensis sp. nov. (brown) (10–15), C. irawadi (white) (16–20), C. wangi wangi (21–23) (red), and C. wangi hainanensis (24–26) (yellow) from Thailand and nearby countries. Symbols correspond to molecular (triangle for ND 2; circle for COI genes) and morphological (star) analyses. Localities include Thailand: Songkhla (type locality of C. thailandensis sp. nov.) (1), Krabi (2), Ranong (3), Chon Buri (4), Saraburi (5), Kamphaeng Phet (6), Sakon Nakhon (7), Mukdahan (8), Roi Et (9), Mae Hong Son (type locality of C. maehongsonensis sp. nov.) (10), and Kanchanaburi (11); Myanmar: Mon, Mawlamyine (12), Mon, Kyaik (13), Bago (14), Yangon (15), Mandalay (16), Sagaing (type locality of C. irawadi) (17), Sagaing (18), and Magway (19); China: Western Yunnan (20); Vietnam: Lang Son (21); China: Nanning, Guangxi (22), Hong Kong (23); China: Wanling, Hainan (24), Jiachai, Hainan (25), Tunchang, Hainan (26). Details of all samples used in this study are given in Table 1.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
AE0E76922E9D55798DE17EBE3EA9E729.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666611	https://doi.org/10.3897/zookeys.1281.175455.figure5	Figure 5. Micro- CT images of skull, occipital bone, and pectoral region of male Calotes thailandensis sp. nov. (A, C. CUMZ-R - 2738 and E, G. CUMZ-R - 2769) and male C. maehongsonensis sp. nov. (B, D. CUMZ-R - 2825 and F, H. CUMZ-R - 2820). Abbreviations: Cl, Clavicle; F, Frontal; In, Interclavicle, Mx, Maxilla; Na, Nasal; P, Parietal, Px, Premaxilla; So, Supraoccipital bones. Scale bars: 5 mm.	Figure 5. Micro- CT images of skull, occipital bone, and pectoral region of male Calotes thailandensis sp. nov. (A, C. CUMZ-R - 2738 and E, G. CUMZ-R - 2769) and male C. maehongsonensis sp. nov. (B, D. CUMZ-R - 2825 and F, H. CUMZ-R - 2820). Abbreviations: Cl, Clavicle; F, Frontal; In, Interclavicle, Mx, Maxilla; Na, Nasal; P, Parietal, Px, Premaxilla; So, Supraoccipital bones. Scale bars: 5 mm.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
AE0E76922E9D55798DE17EBE3EA9E729.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666612	https://doi.org/10.3897/zookeys.1281.175455.figure6	Figure 6. Holotype of adult male Calotes thailandensis sp. nov. (CUMZ-R - 2738 [SA 118]). A. Head dorsal; B. Head ventral; C. Head lateral; D. Full body dorsal; E. Body ventral, and F. Lateral region. Scale bars 10 mm.	Figure 6. Holotype of adult male Calotes thailandensis sp. nov. (CUMZ-R - 2738 [SA 118]). A. Head dorsal; B. Head ventral; C. Head lateral; D. Full body dorsal; E. Body ventral, and F. Lateral region. Scale bars 10 mm.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
AE0E76922E9D55798DE17EBE3EA9E729.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666613	https://doi.org/10.3897/zookeys.1281.175455.figure7	Figure 7. Ventral head and lateral views (A – F) of paratypes of Calotes thailandensis sp. nov. (CUMZ-R - 2769 [SA 149], 2872 [SA 221], 2767 [SA 147]) (A – C) and Calotes maehongsonensis sp. nov. (CUMZ-R - 2820 [SA 169], 2827 [SA 176], 2839 [SA 188]) (D – F) are showed consisting of adult males (A, B, D, E) and adult females (C, F). Scale bars: 10 mm.	Figure 7. Ventral head and lateral views (A – F) of paratypes of Calotes thailandensis sp. nov. (CUMZ-R - 2769 [SA 149], 2872 [SA 221], 2767 [SA 147]) (A – C) and Calotes maehongsonensis sp. nov. (CUMZ-R - 2820 [SA 169], 2827 [SA 176], 2839 [SA 188]) (D – F) are showed consisting of adult males (A, B, D, E) and adult females (C, F). Scale bars: 10 mm.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
AE0E76922E9D55798DE17EBE3EA9E729.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666614	https://doi.org/10.3897/zookeys.1281.175455.figure8	Figure 8. An uncollected male (A, B) on breeding season (March, 2025) and female (D) (CUMZ-R - 2767) of Calotes thailandensis sp. nov. in rubber plantation (C) at Khlong Hoi Kong District, Songkhla Province, Thailand.	Figure 8. An uncollected male (A, B) on breeding season (March, 2025) and female (D) (CUMZ-R - 2767) of Calotes thailandensis sp. nov. in rubber plantation (C) at Khlong Hoi Kong District, Songkhla Province, Thailand.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
AE0E76922E9D55798DE17EBE3EA9E729.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1666607	https://doi.org/10.3897/zookeys.1281.175455.figure1	Figure 1. Sample locality map of Calotes thailandensis sp. nov. (blue) (1–9), C. maehongsonensis sp. nov. (brown) (10–15), C. irawadi (white) (16–20), C. wangi wangi (21–23) (red), and C. wangi hainanensis (24–26) (yellow) from Thailand and nearby countries. Symbols correspond to molecular (triangle for ND 2; circle for COI genes) and morphological (star) analyses. Localities include Thailand: Songkhla (type locality of C. thailandensis sp. nov.) (1), Krabi (2), Ranong (3), Chon Buri (4), Saraburi (5), Kamphaeng Phet (6), Sakon Nakhon (7), Mukdahan (8), Roi Et (9), Mae Hong Son (type locality of C. maehongsonensis sp. nov.) (10), and Kanchanaburi (11); Myanmar: Mon, Mawlamyine (12), Mon, Kyaik (13), Bago (14), Yangon (15), Mandalay (16), Sagaing (type locality of C. irawadi) (17), Sagaing (18), and Magway (19); China: Western Yunnan (20); Vietnam: Lang Son (21); China: Nanning, Guangxi (22), Hong Kong (23); China: Wanling, Hainan (24), Jiachai, Hainan (25), Tunchang, Hainan (26). Details of all samples used in this study are given in Table 1.	Figure 1. Sample locality map of Calotes thailandensis sp. nov. (blue) (1–9), C. maehongsonensis sp. nov. (brown) (10–15), C. irawadi (white) (16–20), C. wangi wangi (21–23) (red), and C. wangi hainanensis (24–26) (yellow) from Thailand and nearby countries. Symbols correspond to molecular (triangle for ND 2; circle for COI genes) and morphological (star) analyses. Localities include Thailand: Songkhla (type locality of C. thailandensis sp. nov.) (1), Krabi (2), Ranong (3), Chon Buri (4), Saraburi (5), Kamphaeng Phet (6), Sakon Nakhon (7), Mukdahan (8), Roi Et (9), Mae Hong Son (type locality of C. maehongsonensis sp. nov.) (10), and Kanchanaburi (11); Myanmar: Mon, Mawlamyine (12), Mon, Kyaik (13), Bago (14), Yangon (15), Mandalay (16), Sagaing (type locality of C. irawadi) (17), Sagaing (18), and Magway (19); China: Western Yunnan (20); Vietnam: Lang Son (21); China: Nanning, Guangxi (22), Hong Kong (23); China: Wanling, Hainan (24), Jiachai, Hainan (25), Tunchang, Hainan (26). Details of all samples used in this study are given in Table 1.	2026-06-03	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai		Zenodo	biologists	Prakobkarn, Arpapan;Zug, George R.;Tandavanitj, Nontivich;Ngamprasertwong, Thongchai			
