identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7EF056A323FB5E0AA9DE86402065A669.text	7EF056A323FB5E0AA9DE86402065A669.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaepsila gropensis Varga 2026	<div><p>Chamaepsila gropensis Varga sp. nov.</p><p>Figs 2 A – C, 3 A – D</p><p>Material examined.</p><p>Holotype. Albania • ♂; Tirane-Shengjergj, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.027166&amp;materialsCitation.latitude=41.359665" title="Search Plazi for locations around (long 20.027166/lat 41.359665)">Mali me Gropa</a>; ca. 1400 m; 41°21.58'N, 20°01.63'E (coordinate uncertainty ca. 10 m), from ferns; 11 Oct. 2025; Varga N., Balogh B., Horváth Á. leg.</p><p>Diagnosis.</p><p>Chamaepsila gropensis sp. nov. can be immediately distinguished from all other Palearctic Chamaepsila species by its strongly and uniformly darkened, blackish wings (Fig. 2 A – C). This pigmentation is caused by dense black microtrichia covering nearly the entire wing surface (Fig. 2 C). By contrast, similar microtrichia in species such as C. rosae are light, yellowish to brownish, sparsely distributed, resulting in fully transparent or slightly darkened wings (Fig. 2 D).</p><p>Based on overall habitus, male genitalia, coloration of head, thorax, setae (except number of dc), legs and even phenology, C. gropensis sp. nov. belongs to the C. hebraica group. It shows the closest resemblance to Chamaepsila homochroa . However, C. homochroa has transparent wings and 3–4 dorsocentral setae (exclusively 4 in Albanian specimens, mainly 3 in Israeli specimens), whereas C. gropensis sp. nov. bears only two. Additionally, the phallus of C. gropensis sp. nov. tapers toward apex (from middle), and not forked (i. e., terminal parts not crossing each other).</p><p>Using the European key provided by Beuk (2023), C. gropensis sp. nov. runs either to couplet 44, if the two large vertical setae are considered and the minute middle seta is regarded as a hair, or to couplet 55, if the middle small seta is counted. In both cases, additional differences beyond wing darkening allow the new species to be distinguished from those resulted by the key.</p><p>In couplet 44, it differs from Chamaepsila nigrotaeniata (Strobl, 1899) by having predominantly black femora, whereas the latter species bears only a narrow ventral black stripe. C. gropensis sp. nov. has a large, black frontal triangle, while only the ocellar spot is black in C. nigrotaeniata (Strobl 1899) . It also differs from C. sardoa, which has generally lighter legs, with femora and tibiae mostly reddish-yellow, and different male genitalia (see figures by Shatalkin 1986).</p><p>In couplet 55, in addition to the blackish wings, C. gropensis sp. nov. differs from C. rosae and C. nigricornis by leg coloration: femora and tibiae mostly dark in C. gropensis sp. nov., whereas completely yellow in the other two species. The structure of male genital parts is also different (see figures by Shatalkin 1986).</p><p>Given that Psilidae species may have large distributional ranges (e. g., Chamaepsila tenebrica Shatalkin, 1986, was originally described from the Russian Far East and later recorded over 7,000 km away in the Czech Republic by Roháček et al. (2016)), the new species was also run with the comprehensive Palearctic and Oriental key ( Psila Meigen, 1803 sensu lato) by Shatalkin (2008). According to this key, if two vertical setae are considered, the species runs to C. sardoa, and if three, to Chamaepsila mixta Soós, 1974 . Besides darkened wings, C. gropensis sp. nov. differs from C. mixta by having entirely black antennae, whereas the basal two segments are yellow in C. mixta .</p><p>Description.</p><p>Body length 4.2 mm. Wing length 3.8 mm. Head. Dark yellowish; frontal triangle large, black. Lunula bright yellow. Gena yellow, ca. 0.8 × eye height (widest point). Ocellar tubercle black; 2 black ocellars. Two black fronto-orbitals. Palpi yellow basally, apically blackish; small setae yellowish, 1 larger blackish preapically and 1 apically. Antennal segments 1–3 entirely black; first flagellomere ca. 1.5 × as long as wide. Basal thickened part of arista blackish, remainder yellowish; aristal setulae blackish, as on first flagellomere. Postocellars blackish, converging; 2 pairs of verticals with a small setae between them (only in one side of the specimen). Small setae on facial carina yellowish. Occiput and postcranium entirely blackish, with blackish setae.</p><p>Thorax. Completely glossy black. Scutum and lateral pleura with yellow setae, few blackish ones near the two strong black postsutural dorsocentrals. Notopleural, supraalar, postalar setae 1, black. One pair of strong black scutellars; scutellum itself glossy black. Postscutellar area and anatergite strongly pruinose, except lowest postscutellar part shiny black. Halteres relatively large, whitish – yellowish.</p><p>Legs. Fore coxae brownish anteriorly, with small yellowish setulae and some weaker blackish ones near apex. Lateral side of coxa with black transverse stripe on anterior margin, remainder yellowish.</p><p>Mid and hind coxae mostly shiny black, setae mostly yellowish, few long blackish weaker ones near apices. Trochanters yellowish, setae mostly yellow. Femora blackish, apices yellow; yellow continues ventrally as a narrow line reaching to ca. mid-point of femur. Small setulae mostly black dorsally, yellow ventrally. Tibiae yellow at basal 1 / 3, black at apical 2 / 3; small setae on them mostly blackish; as well as spurs. Tarsi dark brownish; setae blackish.</p><p>Wings. Uniformly dark grayish, almost black; veins dark brownish. Microtrichia blackish, dense, covering practically the entire wing surface in both dorsal and ventral sides. Calypter fringe composed of long black setae. Section of R 4 + 5 between r-m and wing apex&gt; 4 × length between r – m and junction with R 2 + 3. Penultimate section of M 1 + 2 slightly longer than ultimate.</p><p>Abdomen. Abdomen completely glossy black; with blackish setae of moderate length uniformly distributed, no denser patches.</p><p>Male genitalia (Fig. 3 A – D). Epandrium rounded, with black setae, without thickened or strongly projecting parts (Fig. 3 A). Parameres in dorsal view elongated, with basal protruding part, apically pointed. Parameres connected to epandrium via a broad parameral arm. Derivates of gonopods long, narrow. Phallapodeme not strongly elongated, ca. as long as epandrium. Hypandrium rather short, rounded. Phallus weakly sclerotised, in lateral view, widest at ca. midpoint, narrowing towards apex, pointed (Fig. 3 B). In dorsal view (Fig. 3 C), apices do not cross each other (i. e. phallus not forked). Cerci and subepandrial sclerite with black setae, shape as in Fig. 3 D.</p><p>Female. Unknown.</p><p>Etymology.</p><p>The species-group name gropensis is a Latin adjective derived from the name of the locus typicus (Mali me Gropa massif, meaning “ hill full of pits ” in Albanian), specifically using the suffix – ensis.</p><p>Ecology.</p><p>The specimen was collected by sweeping a bracken field ( Pteridium aquilinum L.) growing on a steep west-facing karstic slope in a recently burnt area of Mali me Gropa (Fig. 4 A), at an altitude of approximately 1400 m. The flight period is likely similar to that of other members of the C. hebraica group.</p></div>	https://treatment.plazi.org/id/7EF056A323FB5E0AA9DE86402065A669	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Varga, Nimród;Horváth, Áron;Balogh, Botond	Varga, Nimród, Horváth, Áron, Balogh, Botond (2026): The occurrence of the family Psilidae (Diptera) in Albania, with description of a new Chamaepsila Hendel, 1917 species. Evolutionary Systematics 10 (1): 93-99, DOI: 10.3897/evolsyst.10.188012
07F7953BFA7A59569D0E02D161BEBEEA.text	07F7953BFA7A59569D0E02D161BEBEEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaepsila Hendel 1917	<div><p>Genus Chamaepsila Hendel, 1917</p><p>Chamaepsila Hendel, 1917: 37 . Type species: Musca rosae (Fabricius, 1794) (by original designation).</p></div>	https://treatment.plazi.org/id/07F7953BFA7A59569D0E02D161BEBEEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Varga, Nimród;Horváth, Áron;Balogh, Botond	Varga, Nimród, Horváth, Áron, Balogh, Botond (2026): The occurrence of the family Psilidae (Diptera) in Albania, with description of a new Chamaepsila Hendel, 1917 species. Evolutionary Systematics 10 (1): 93-99, DOI: 10.3897/evolsyst.10.188012
41F0FEB919C55DB1B62FC850C814E092.text	41F0FEB919C55DB1B62FC850C814E092.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaepsila homochroa Freidberg & Shatalkin 2008	<div><p>Chamaepsila homochroa Freidberg &amp; Shatalkin, 2008</p><p>Figs 1 A – E, 4 B – D</p><p>Material examined.</p><p>Albania • 10 ♂♂ 1 ♀; Shkallë, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.071667&amp;materialsCitation.latitude=41.323334" title="Search Plazi for locations around (long 20.071667/lat 41.323334)">near Liqeni i Shkallës</a>; 41°19.65'N, 19°57.72'E (coordinate uncertainty ca. 10 m); 9. Oct. 2025; Varga N., Balogh B., Horváth Á. leg. 1 ♂ 1 ♀; Tirane-Shengjergj, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.071667&amp;materialsCitation.latitude=41.323334" title="Search Plazi for locations around (long 20.071667/lat 41.323334)">Mali me Gropa</a>; ca. 1400 m; 41°21.58'N, 20°01.63'E (coordinate uncertainty ca. 10 m), from ferns; 11 Oct. 2025; Varga N., Balogh B., Horváth Á. leg. 1 ♂; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.071667&amp;materialsCitation.latitude=41.323334" title="Search Plazi for locations around (long 20.071667/lat 41.323334)">Shengjin</a>; 41°19.40'N, 20°04.30'E; 11 Oct. 2025; Varga N., Balogh B., Horváth Á. leg.</p><p>Diagnosis.</p><p>Chamaepsila homochroa is named after its characteristic, uniformly dark body coloration (Fig. 1 A, B). The species is further characterised by three dorsocentral setae (rarely four), black setae on the head, elongated frontal triangle (Fig. 1 C) and late flight period (Freidberg and Shatalkin 2008). However, the most diagnostic character is the apically forked phallus (Fig. 1 D, E). According to Freidberg and Shatalkin (2008), this character is unique within the genus Chamaepsila . The Albanian specimens possess four dorsocentral setae; all other external characters and male genitalia fully correspond to the original description, figures, and photographs provided by Freidberg and Shatalkin (2008) and Çerçi (2017: as “ Friedberg ” [misspelling of the first author’s name]).</p><p>Remarks.</p><p>The record from Turkey reported by Çerçi (2017) represents the first occurrence of the species in Europe. The present findings from Albania constitute the currently known westernmost occurrence of C. homochroa and the second record from Europe. The species is expected to occur in neighbouring countries; however, it has likely remained undetected due to limited research on Psilidae in the region and the late flight period characteristic of the Chamaepsila hebraica Hennig, 1941 species group (sensu Freidberg and Shatalkin (2008)). The type specimens were collected from November to January, the earliest Turkish record dated 22 October, whereas the Albanian specimens were collected on 10–11 October, representing the earliest known records of the species. This further confirms that the flight period may shift earlier, farther from the equator (Çerçi 2017).</p><p>Distribution.</p><p>Israel, Turkey (Freidberg and Shatalkin 2008; Çerçi 2017); newly recorded from Albania.</p></div>	https://treatment.plazi.org/id/41F0FEB919C55DB1B62FC850C814E092	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Varga, Nimród;Horváth, Áron;Balogh, Botond	Varga, Nimród, Horváth, Áron, Balogh, Botond (2026): The occurrence of the family Psilidae (Diptera) in Albania, with description of a new Chamaepsila Hendel, 1917 species. Evolutionary Systematics 10 (1): 93-99, DOI: 10.3897/evolsyst.10.188012
