taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505700	https://doi.org/10.3897/zse.102.175092.figure5	Figure 5. Holotype of Boulenophrys youran sp. nov. in life (HUNL XB 25061601). A. Lateral view; B. Ventral view; C. Dorsal view; D. Plantar view of right foot; E. Volar view of right hand; F. Dorsal view of left hand. Photos by Leqiang Zhu.	Figure 5. Holotype of Boulenophrys youran sp. nov. in life (HUNL XB 25061601). A. Lateral view; B. Ventral view; C. Dorsal view; D. Plantar view of right foot; E. Volar view of right hand; F. Dorsal view of left hand. Photos by Leqiang Zhu.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505701	https://doi.org/10.3897/zse.102.175092.figure6	Figure 6. Boulenophrys youran sp. nov. in preservation. A. Adult male, holotype HUNL XB 25061601; B. Adult female, paratype HUNL LH 23051202; 1. dorsal view; 2. ventral view. Photos by Shengchao Shi.	Figure 6. Boulenophrys youran sp. nov. in preservation. A. Adult male, holotype HUNL XB 25061601; B. Adult female, paratype HUNL LH 23051202; 1. dorsal view; 2. ventral view. Photos by Shengchao Shi.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505702	https://doi.org/10.3897/zse.102.175092.figure7	Figure 7. Coloration variations of Boulenophrys youran sp. nov. in adult males. Specimens: A. HUNL XB 25061603; B. HUNL XB 24052601; C. HUNL XB 24052602; 1. dorsal view; 2. ventral view. Photos by Hui Li.	Figure 7. Coloration variations of Boulenophrys youran sp. nov. in adult males. Specimens: A. HUNL XB 25061603; B. HUNL XB 24052601; C. HUNL XB 24052602; 1. dorsal view; 2. ventral view. Photos by Hui Li.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505703	https://doi.org/10.3897/zse.102.175092.figure8	Figure 8. Juvenile of Boulenophrys youran sp. nov. (HUNL DB 23092403) in life and preservation. A. Dorsal view; B. Ventral view; C. Lateral view; D. Dorsal view of hand (in preservation); E. Volar view of hand; F. Plantar view of foot. Photos by Dongbo Fei.	Figure 8. Juvenile of Boulenophrys youran sp. nov. (HUNL DB 23092403) in life and preservation. A. Dorsal view; B. Ventral view; C. Lateral view; D. Dorsal view of hand (in preservation); E. Volar view of hand; F. Plantar view of foot. Photos by Dongbo Fei.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505697	https://doi.org/10.3897/zse.102.175092.figure2	Figure 2. ML tree of the genus Boulenophrys reconstructed inferred from 16 S rRNA and COI gene. Bayesian posterior probability (BPP) / ML bootstrap supports (BS) were denoted beside each node is, a dash (–) denotes BS values below 70 % or BPP values below 0.90.	Figure 2. ML tree of the genus Boulenophrys reconstructed inferred from 16 S rRNA and COI gene. Bayesian posterior probability (BPP) / ML bootstrap supports (BS) were denoted beside each node is, a dash (–) denotes BS values below 70 % or BPP values below 0.90.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505704	https://doi.org/10.3897/zse.102.175092.figure9	Figure 9. Violin-plot showing the difference between Boulenophrys youran sp. nov. and its close congeners.	Figure 9. Violin-plot showing the difference between Boulenophrys youran sp. nov. and its close congeners.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505705	https://doi.org/10.3897/zse.102.175092.figure10	Figure 10. Morphological comparisons in adult males of Boulenophrys youran sp. nov. and its relative species. 1. lateral or dorsolateral body, 2. ventral body, 3. ventral foot, 4. ventral hand, 5. nuptial pads and spines on dorsal of inner two fingers (not to scale). A. B. youran sp. nov. (HUNL XB 25061603, paratype from Nanshan National Park, Hunan, China); B. B. dalaolingensis (CIB YL 20240518023, topotype from Dalaoling Nature Reserve, Hubei, China); C. B. fanjingmontis (unvouchered individual from type locality Mt. Fanjing, Tongren, Guizhou, China); D. B. qianbeiensis (CIB WB 2020211, topotype from Tongzhi ,, Guizhou, China); E. B. spinata (CIB LS 20190801002, topotype from Mt. Leigong, Guizhou, China); F. B. sangzhiensis (CIB TPS 20190625004, topotype from Mt. Tianping, Hunan, China); G. B. yezhongensis (CIB SC 20240531005, holotype from Yezhong Nature Reserve, Guizhou, China). Photographer: A by Hui Li; B, C, D, F by Shengchao Shi; E, G by Shize Li.	Figure 10. Morphological comparisons in adult males of Boulenophrys youran sp. nov. and its relative species. 1. lateral or dorsolateral body, 2. ventral body, 3. ventral foot, 4. ventral hand, 5. nuptial pads and spines on dorsal of inner two fingers (not to scale). A. B. youran sp. nov. (HUNL XB 25061603, paratype from Nanshan National Park, Hunan, China); B. B. dalaolingensis (CIB YL 20240518023, topotype from Dalaoling Nature Reserve, Hubei, China); C. B. fanjingmontis (unvouchered individual from type locality Mt. Fanjing, Tongren, Guizhou, China); D. B. qianbeiensis (CIB WB 2020211, topotype from Tongzhi ,, Guizhou, China); E. B. spinata (CIB LS 20190801002, topotype from Mt. Leigong, Guizhou, China); F. B. sangzhiensis (CIB TPS 20190625004, topotype from Mt. Tianping, Hunan, China); G. B. yezhongensis (CIB SC 20240531005, holotype from Yezhong Nature Reserve, Guizhou, China). Photographer: A by Hui Li; B, C, D, F by Shengchao Shi; E, G by Shize Li.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
5F7661D7F33B5F68BBA590F6226590FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1505706	https://doi.org/10.3897/zse.102.175092.figure11	Figure 11. Habitat of Boulenophrys youran sp. nov. in Nanshan National Park, Shaoyang City, Hunan Province, China. A. Microhabitats of the new species, mountain stream in Nanshan National Park; B. Male paratype HUNL XB 24052601 found calling on rocks near stream in situ; C. Montane forests and stream at the type locality. Photographed by Leqiang Zhu.	Figure 11. Habitat of Boulenophrys youran sp. nov. in Nanshan National Park, Shaoyang City, Hunan Province, China. A. Microhabitats of the new species, mountain stream in Nanshan National Park; B. Male paratype HUNL XB 24052601 found calling on rocks near stream in situ; C. Montane forests and stream at the type locality. Photographed by Leqiang Zhu.	2026-01-05	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang		Zenodo	biologists	Xiao, Bei;Zhu, Leqiang;Wang, Lu;Li, Hui;Maimaiti, Ayinuer;Li, Shize;Zhang, Guoai;Liu, Wuzhou;Fei, Dongbo;Liao, Changle;Tang, Xiaoquan;Shi, Shengchao;Mo, Xiaoyang			
