identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B368F1650769FFE1FCDFFAF93E5BF8A5.text	B368F1650769FFE1FCDFFAF93E5BF8A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterovianaida Montemayor & Carpintero 2007	<div><p>Clade  Pterovianaida +  Thaumamannia</p><p>The clade  Pterovianaida +  Thaumamannia is supported by one synapomorphy ‘head position declined’ (char 1, state 1 or 1-1 – Fig. 2B) and one homoplastic synapomorphy, ‘the large scales on the anterior region of the hemelytral margins’ (25-2). Internally to this clade, the genus  Pterovianaida is recovered monophyletic, supported by two synapomorphies and one homoplastic synapomorphy: abundant head setae (0-2 – Fig. 3C), presence of a paranota constriction (18-1 – Fig. 4B) and frons at same height as clypeus (8-0), respectively.  Thaumamannia, including its two new species, is also found to be monophyletic, supported by one synapomorphy and two homoplastic synapomorphies (presence of large scales on paranota borders, 17-2 – Fig. 5C; scutellum narrower than half of the maximal width of the head, 15-0, and scent gland peritreme with a shorter posterior branch than the upper part of the anterior branch, 23-1, as the two homoplastic synapomorphies).  Thaumamannia insolita is found to be the sister-group to all remaining congeners, supported by one synapomorphy: pronotum more than two times wider than long (14-1 – Fig. 6B).  Thaumamannia urucuana and  T. manni are recovered as one clade, the sister-group to  T. vanderdrifti based on two synapomorphies: posterior region of the bucculae round, 6-0 – Fig. 7A, and posterior region of the bucculae wider than anterior half (7-1).</p></div>	https://treatment.plazi.org/id/B368F1650769FFE1FCDFFAF93E5BF8A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650768FFECFCCDFA573A41F951.text	B368F1650768FFECFCCDFA573A41F951.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Henryvianaida Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Clade  Henryvianaida +  Anommatocoris</p><p>The clade composed by the  Henryvianaida +  Anommatocoris is supported by two synapomorphies and one homoplastic synapomorphy (pronotum finely punctate, 13-0 – Fig. 8A, and anterior region of hemelytra slightly constricted, 26-1 – Fig. 11B; collar not projected towards the head, 10-0, respectively). The new genus  Henryvianaida is supported as monophyletic by two synapomorphies: mandibular plates laterally compressed (2-1 – Fig. 9B) and a straight margin on the posterior region of bucculae (6-1 – Fig. 7B).  Anommatocoris is recovered with the most synapomorphies, seven in total: a flat posterior region of pronotum (11-1 – Fig. 10B), anterior branch of the scent gland peritreme almost perpendicular to sagittal body plane (22-0), the presence of a constriction on the anterior region of hemelytra (26-1 – Fig. 11B), hypocosta narrow (27-0 – Fig. 12A) and smooth (28-1 – Fig. 13B), costal area thickened and narrow (30-1 – Fig. 14B) and subcostal area subvertical (31-1 – Fig. 15B). Additionally, one homoplastic synapomorphy also supports the monophyly of this genus: frons at same height as clypeus (8-0).  Anommatocoris bolivianus is found to be sister to the rest of the genus, which then has two different clades: [ A. schuhi + [ A. zeteki +  A. coibensis] supported by one homoplastic synapomorphy (posterior branch of scent gland peritreme shorter than the upper part of anterior branch, 23-1) whereas the internal [ A. zeteki +  A. coibensis] clade is supported by a different homoplastic synapomorphy (posterior region of paranota comprising a small acute humeral angle, 19-1) and  A. sucreanus + the polytomy [ A. coleopteratus,  A. minutissimus,  A. serratus,  A. knudsoni,  A. araguanus], supported by one synapomorphy (paranota borders bearing small scales, 17-1 – Fig. 5B).</p><p>TAXONOMY</p><p>VIANAIDINAE KORMILEV</p><p>Diagnosis: The subfamily is mainly characterized by its unique scent gland peritreme (Figs 16, 17) composed by an anterior and a posterior branch, the latter transversally connected to the first, forming a sulcus and varying in inclination, curvature and swelling, giving a T- or Y-shaped aspect. The pronounced clypeus, punctate pronotum and hemelytra and visible scutellum are also diagnostic characters for the subfamily. It presents both coleopteroid and macropterous forms, which hampers the comparison of hemelytral characters. However, in coleopteroid forms, a carina-like vein extending from its anterior border is always present, and in macropterous forms, a well-defined clavus and vein-less membrane can also be indicated as a diagnostic feature.</p><p>Redescription: Head: Pubescent, clypeus convex, strongly delimited from vertex, extending beyond mandibular plates; antenniferous process facing downwards; antennae four-segmented; bucculae strongly developed, extending the entire length of the head ventrally, open anteriorly, divergent posteriorly, punctate; rostrum four-segmented always reaching abdominal segments. Thorax: Pronotum punctate; pronotal carinae absent; scutellum visible, smooth; sternal laminae present, punctate, sinuous, widening posteriorly. Hemelytra: Coleopteroid or macropterous; in coleopteroid forms, clavus and membrane absent, carina-like vein extending variably from the anterior border; in macropterous forms, clavus and veinless membrane well-developed in addition to discoidal, subcostal and costal areas, punctate in coriaceous parts. Scent gland: Peritreme distinct, composed by an anterior and a posterior branch, the latter transversally connected to the first, these bearing a sulcus and varying in shape, curvature, inclination and swollenness; evaporatorium covering the entire metapleuron and hind mesopleuron, also advancing ventrally; scent gland ostiole conspicuously big. Legs: Long, slender, pubescent and unarmed; coxae widely separated; trochanter not fused; femora usually swollen to some degree; tarsi two segmented, second segment many times longer, claws long and slender. Abdomen: In coleopteroid forms, completely enclosed in the hemelytra and roundly ovate; in macropterous forms, long, rectangular; spiracles located ventrally near lateral margins of abdominal sternites; usually pubescent.</p><p>Type genus:  Vianaida Kormilev, a junior synonym of  Anommatocoris China.</p><p>Distribution: Central and South America (Fig. 18).</p></div>	https://treatment.plazi.org/id/B368F1650768FFECFCCDFA573A41F951	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650765FFEDFC5EF9703AACF88D.text	B368F1650765FFEDFC5EF9703AACF88D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris China 1945	<div><p>Anommatocoris China, 1945</p><p>Diagnosis: This genus can be characterized by the straight position of the head, the flat pronotum, the lack of explanate paranota, the constriction on the anterior part of hemelytra, the mostly carinate costal area, the nearly vertical subcostal area, the narrow and smooth hypocosta, in addition to the almost perpendicular (to the sagittal body plane) anterior branch of the scent gland which is only slightly laterally projected.</p><p>Redescription: Head: Triangular in dorsal view, pubescent; clypeus usually in different colour than head; pedicel subequal to basiflagellomere, both slightly smaller than distiflagellomere and usually twice as big as scape; eyes reduced with none or few scarcely distributed ommatidia or fully developed, compound; bucculae usually with one row of punctures; rostrum reaching at least the first abdominal segment. Thorax: Flat, widening posteriorly, sometimes also slightly widened laterally; punctate, mostly on posterior region of pronotum; collar indistinct, punctate; anterior border usually straight, posterior usually convex; paranota carinate. Hemelytra: Either</p></div>	https://treatment.plazi.org/id/B368F1650765FFEDFC5EF9703AACF88D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650761FFE8FDD0FF733AB4FEC2.text	B368F1650761FFE8FDD0FF733AB4FEC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris China 1945	<div><p>KEY TO THE SPECIES OF  ANOMMATOCORIS</p><p>1. Hemelytra strongly modified, coriaceous, coleopteroid, membrane not entirely developed......................... 2</p></div>	https://treatment.plazi.org/id/B368F1650761FFE8FDD0FF733AB4FEC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650762FFF6FC78F9543DE6F97F.text	B368F1650762FFF6FC78F9543DE6F97F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris coleopteratus (Kormilev 1955)	<div><p>Anommatocoris coleopteratus</p><p>(Kormilev, 1955) (Fig. 22)</p><p>Diagnosis: This species is the only  Anommatocoris species with a visible scent gland in dorsal view. It also presents an unusually large tip of the anterior branch of the peritreme for the genus and a long carina-like vein on the hemelytra, reaching its posterior region.</p><p>Redescription: Body: Light brown; head whitish; antennae, rostrum and legs yellowish (Fig. 22A). Head: Pubescent, with small and well-spaced hairs; clypeus darker than head, same colour as body; antenniferous process length more than half of pedicel; pedicel subequal to basiflagellomere, both smaller than distiflagellomere and at least twice as big as scape; eyes with few scarcely distributed ommatidia; bucculae rounded, with scarce hairs on its border and deeply impressed punctures near head insertion, concave posteriorly (Fig. 22E); rostrum reaching at least second abdominal segment (Fig. 22B, C). Thorax: Punctate, with small punctures, one row at collar and then only on the posterior lobe of pronotum; anterior border straight, posterior sinuous; paranota carinate with hairs and few irregularly distributed small scale-like projections on its border (Fig. 22F); scutellum large, a little less than one-fifth the maximum pronotum width (Fig. 22D). Hemelytra: Coriaceous and coleopteroid, evenly pubescent; laterally constricted anteriorly; carina-like vein extending from the anterior border to the posterior third of hemelytra, fading abruptly; finely punctate entirely, punctures small, larger anteriorly, up to the hemelytra constriction. Scent gland: Anterior branch tip tilted horizontally, large, same size as the rest of the anterior branch, presenting a cracked texture; posterior branch slightly curved, with considerably enlarged bottom edge on its tip; sulcus prominent on both branches; evaporatorium with curved border anteriorly, advancing to most of the upper part of the mesopleuron (Figs 16C – 22C).</p><p>Measurements: BL, 1.78 (1.87; 1.55–1.87 xM; 1.74– 1.96 xF); BW, 0.74 (0.84; 0.51–0.84 xM; 0.74–0.89 xF); HL, 0.25 (0.25; 0.21–0.27 xM; 0.23–0.29 xF); HW, 0.32 (0.37; 0.27–0.37 xM; 0.32–0.36 xF); ID, 0.23 (0.29; 0.17–0.29 xM; 0.23–0.25 xF); PL, 0.34 (0.36; 0.29–0.36 xM; 0.32–0.40 xF); PW, 0.55 (0.57; 0.46–0.68 xM; 0.53– 0.61 xF); AS, 0.11 (0.10; 0.10–0.11 xM; 0.10–0.11 xF); AP, 0.23 (0.23; 0.22–0.24 xM; 0.22–0.26 xF); AB, 0.24 (0.24; 0.21–0.24 xM; 0.23–0.27 xF) and AD, 0.34 (0.32; 0.32–0.34 xM; 0.34–0.38 xF).</p><p>Distribution: Described from Buenos Aires province, Argentina (Kormilev, 1955) and reported from Rocha province, Uruguay (San Martin, 1966).</p><p>Material examined: Holotype xM: ARGENTINA: Buenos Aires: Tigre, Rio Luján, 25 March 1955, M. J. Vianna col. (USNM). Allotype xF: ARGENTINA: Buenos Aires: Tigre, Rio Luján, 25 March 1955, M. J. Vianna col. (USNM). Paratypes: ARGENTINA: Buenos Aires: Tigre, Rio Luján, June 1955, M. J. Vianna col. [Guidoti PhD –  Vianaidinae 039–046 and 048, 9 glued to card boards and thus, gender undefined] (039- 046 USNM; 048 MNRJ); ARGENTINA: Buenos Aires: Tigre, Rio Luján, June 1955, M. J. Vianna col. [Guidoti PhD –  Vianaidinae 047, 1 glued to card board and thus, gender undefined] (USNM). Other Specimens: ARGENTINA: Buenos Aires: Res. Punta Lara, III-2001, Carpintero col. [Guidoti PhD – 6xF,  Vianaidinae 023 and 026-030; 7xM,  Vianaidinae 024-025 and 031- 035] (MNHN).</p><p>Anommatocoris knudsoni Guidoti et al.,</p></div>	https://treatment.plazi.org/id/B368F1650762FFF6FC78F9543DE6F97F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F165077EFFF0FC78F9173A4DF8BA.text	B368F165077EFFF0FC78F9173A4DF8BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris minutissimus China 1945	<div><p>Anommatocoris minutissimus</p><p>China, 1945 (Fig. 24)</p><p>Diagnosis: This species is distinguishable from the other  Anommatocoris species by the lack of acute humeral angle, the lightly pronounced punctations on pronotum, the narrow scutellum and the scent gland peritreme, which is similar to the one presented by  A. coibensis due to the almost straight upper part of the anterior branch, but it is equally divided by the posterior branch and lacks a sinuous margin.</p><p>Redescription: Body: Light brown; antennae, rostrum and legs even lighter; head whitish (Fig. 24A). Head: Pubescent, with small and well-spaced hairs; clypeus darker than head, same colour as body; antenniferous process half of pedicel length; pedicel twice the size of scape; eyes with just a few scarcely distributed ommatidia; bucculae rounded, widest anteriorly, with scarce hairs on its border and fine marks of punctures organized in a single complete row in the middle, concave posteriorly (Fig. 24D); rostrum reaching at least second abdominal segment (Fig. 24B). Thorax: Finely punctate, with tiny punctures on collar and on the posterior lobe of pronotum; anterior border concave, posterior only slightly sinuous; paranota carinate with hairs on its border (Fig. 24E); scutellum large, a little less than one-sixth the maximum pronotum width (Fig. 24C). Hemelytra: Coriaceous and coleopteroid, pubescent mostly on subcostal and costal areas; laterally constricted anteriorly; carina-like vein extending from the anterior border to the middle of hemelytra, fading abruptly; finely impressed by punctures anteriorly up to the hemelytra constriction, smooth posteriorly to the hemelytra constriction, bearing punctures marks but not punctures. Scent gland: Anterior branch tip tilted horizontally, large, slightly smaller than the rest of the anterior branch, presenting a lightly cracked texture; posterior branch straight, with evenly enlarged edges through its whole extant; sulcus much more prominent on posterior branch than in anterior branch; evaporatorium with curved border anteriorly, advancing to up to one-third of the mesopleuron on its upper part (Figs 16E – 24B).</p><p>Measurements: BL, 1.64 (1.58 xM; 1.71 xF); BW, 0.70 (0.67 xM; 0.74 xF); HL, 0.19 (0.19 xM; 0.19 xF); HW, 0.30 (0.30 xM; 0.30 xF); ID, 0.21 (0.21 xM; 0.21 xF); PL, 0.32 (0.32 xM; 0.32 xF); PW, 0.50 (0.49 xM; 0.51 xF); AS, 0.10 (0.10 xM; 0.11 xF) and AP, 0.20 (0.19 xM; 0.21 xF).</p><p>Distribution: Known only from Trinidad.</p><p>Material examined: Syntypes: TRINIDAD: B. W. I., St. Augustine, in litter soil, Cacao Plantation, 11.1943 – 2.1944, A. H. Strickland. [Guidoti PhD – 1xF,  Vianaidinae 037; 1xM,  Vianaidinae 038] (USNM).</p></div>	https://treatment.plazi.org/id/B368F165077EFFF0FC78F9173A4DF8BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650778FFF2FED4FF573D64FC32.text	B368F1650778FFF2FED4FF573D64FC32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris schuhi Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Anommatocoris schuhi</p><p>Guidoti et al., sp. nov. (Fig. 25)</p><p>lsid: urn:lsid:zoobank.org:act: 47B40CAA-0B7C-4623-A578-D3D1D86E6BF2</p><p>Diagnosis: This species resembles the most  A. coibensis by its darker coloration, larger size, increased number of ommatidia on the eyes and general shape of the scent gland peritreme. It differs from this species by the even darker aspect of its habitus, and by the short hemelytra carina-like vein, which seems to be unique among its congeners. The unequally divided peritreme also differs from  A. coibensis and  A. zeteki, the other species bearing such unequally divided peritreme. The sinuosity of the anterior branch is more pronounced, and the tip of the anterior branch is rounded and not acutely defined as in  A. coibensis, and the posterior region of the posterior branch is not enlarged like in  A. coibensis and  A. zeteki .</p><p>Description: Body: Dark brown; head same as body; antennae, rostrum and legs brown (Fig. 25A). Head: Pubescent, with well-spaced hairs; clypeus slightly lighter than head; antenniferous process less than one-fourth of the pedicel length; pedicel subequal to basiflagellomere, both smaller than distiflagellomere and more than twice as big as scape; eyes with few scarcely distributed ommatidia, more than what is usually observed among its congeners; bucculae rounded, widest anteriorly, slightly concave posteriorly; with scarce hairs on its border, at least three rows of punctures not so deeply impressed, concave posteriorly (Fig. 25E); rostrum reaching at least the third abdominal segment (Fig. 25B, C). Thorax: Punctate, one row at collar and then only on the posterior lobe of pronotum; anterior border usually straight, posterior sinuous; paranota carinate, with hairs on its border (Fig. 25F); scutellum conspicuously large, about one-third the maximum pronotum width (Fig. 25D). Hemelytra: Coriaceous and coleopteroid, pubescent almost entirely, glabrous only on the middle of the hemelytra; laterally constricted anteriorly; carina-like vein extending from the anterior border to only the first third of hemelytra, smoothly fading; coarsely punctate anteriorly, up to the hemelytra constriction, completely smooth thereafter. Scent gland: Anterior branch edges sinuous and considerably enlarged if compared to posterior branch; posterior straight, edges conspicuously narrow, even narrower than sulcus; sulcus prominent on posterior branch, weakly impressed on anterior branch, fading entirely way before the tip; evaporatorium with curved border anteriorly, advancing to up to one-fourth of mesopleuron (Figs 16F – 25C).</p><p>Measurements: BL, 2.91 (2.82; 2.82–3.00 xF); BW, 1.49 (1.48; 1.48–1.50 xF); HL, 0.32 (0.29; 0.29–0.36 xF); HW, 0.55 (0.55; 0.55– xF); ID, 0.35 (0.34; 0.34–0.36 xF); PL, 0.57 (0.57; 0.57– xF); PW, 1.17 (1.16; 1.16–1.18 xF); AS, 0.15 (0.15; 0.15– xF); AP, 0.44 (0.42; 0.42–0.46 xF); AB, 0.39 (0.36; 0.36–0.42 xF) and AD, 0.51 (0.51; 0.51 xF).</p><p>Etymology: This species was named after the great American entomologist Dr Randall T. Schuh, who identified these specimens as  Anommatocoris sp. and made important contributions to the study of  Vianaidinae (Schuh et al., 2006); not to mention the countless contributions to  Miridae and Heteroptera in general throughout his fruitful career.</p><p>Distribution: Described from Tungurahua, Ecuador.</p><p>Material examined:   Holotype xF: ECUADOR: Tungurahua Prov., 12.2 km E  Baños, 5000 ft, V-22–93, L. Herman col., #2736, litter near stream [Guidoti PhD —  Vianaidinae 017] (AMNH)  .   Paratype: ECUADOR: Tungurahua Prov., 12.2 km E  Baños, 5000 ft, V-22–93, L. Herman col., #2736, litter near stream. [Guidoti PhD – 1xF,  Vianaidinae 016] (AMNH)  .</p></div>	https://treatment.plazi.org/id/B368F1650778FFF2FED4FF573D64FC32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F165077BFFF2FED3FB893A7AFA65.text	B368F165077BFFF2FED3FB893A7AFA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris serratus Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Anommatocoris serratus</p><p>Guidoti et al., sp. nov. (Fig. 26)</p><p>lsid: urn:lsid:zoobank.org:act: BA12936D-8876- 41FF-9B12-FAA0EE1F66C2</p><p>Diagnosis: This species can be recognized by the serrate aspect of the anterior region of the paranota, which is unique among its congeners. The humeral angle slightly pronounced, the hemelytra carina-like vein reaching the middle of the hemelytra, and the tip of the anterior branch of the peritreme resemble  A. zeteki, but it differs from that species by the larger distance between this tip and the hemelytra border and by the even division of the anterior branch by the posterior branch of the peritreme. From  A. araguanus, it differs by the angled posterior branch of the peritreme and the higher position of the anterior branch tip, which is near the hemelytra.</p><p>Description: Body: Reddish brown, pronotum the darkest; head, antennae, rostrum and legs yellowish (Fig. 26A). Head: Pubescent, with well-spaced hairs; clypeus darker than head; antenniferous process almost half of pedicel length; pedicel subequal to basiflagellomere, both smaller than distiflagellomere and at least twice as big as scape; eyes with few scarcely distributed ommatidia; bucculae rounded, widest anteriorly, with a serrate aspect on its border and few scarcely distributed punctures, slightly concave posteriorly (Fig. 26E); rostrum reaching mostly to the second abdominal segment (Fig. 26B, C). Thorax: Finely punctate, one row at collar and then only on the posterior lobe of pronotum; anterior border usually straight, posterior sinuous; paranota carinate, culminating is a small humeral acute angle (Fig. 26F), with small scale-like projections on its border giving an obvious serrate aspect; scutellum conspicuously large, a little less than one-third the maximum pronotum width (Fig. 26D). Hemelytra: Coriaceous and coleopteroid, evenly pubescent; laterally constricted anteriorly; carina-like vein extending from the anterior border to the middle of hemelytra, fading abruptly; coarsely punctate anteriorly, up to the hemelytra constriction, smooth posteriorly. Scent gland: Anterior branch slightly inclined frontwards, tip with the same width as the body of the anterior branch, tilted horizontally, almost same size of the rest of the anterior branch; posterior branch mostly straight, with slightly enlarged upper edge distally; sulcus prominent on both branches; evaporatorium with curved border anteriorly, advancing to up to one-third of mesopleuron (Figs 16G – 26C).</p><p>Measurements: BL, 1.92; BW, 0.97; HL, 0.21; HW, 0.29; ID, 0.21; PL, 0.38; PW, 0.74; AS, 0.10; AP, 0.24; AB, 0.27 and AD, 0.34.</p><p>Etymology: This species was named after the remarkable serrate aspect of the paranota border, which seems to be unique among its congeners.</p><p>Distribution: Described from Boyacá, Colombia.</p><p>Material examined:   Holotype xF: COLOMBIA: Boyacá: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.29344&amp;materialsCitation.latitude=4.89811" title="Search Plazi for locations around (long -73.29344/lat 4.89811)">Sendero Hyca Quye</a>, ~ 5.5 km NW de Santa Maria, 4.89811ºN 73.29344ºW, 900 m, 7–11 March 2016, D.Forero col. [MPUJ _ ENT 0046245] [Guidoti PhD – Extraction #274] [Guidoti PhD –  Vianaidinae 022] (MPUJ).</p></div>	https://treatment.plazi.org/id/B368F165077BFFF2FED3FB893A7AFA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F165077BFFFCFC27FA293D59FB29.text	B368F165077BFFFCFC27FA293D59FB29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris sucreanus Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Anommatocoris sucreanus</p><p>Guidoti et al., sp. nov. (Fig. 27)</p><p>lsid: urn:lsid:zoobank.org:act: EFD9B78A-86A9- 47FC-BA87-B7DF955CE68F</p><p>Diagnosis: This species can be recognized by the scent gland, which is similar to the one presented by  A. minutissimus . However, it distinguishes from  A. minutissimus by the wider peritreme in both anterior and posterior branches, the longer carina-like vein on the hemelytra and the absence of punctures or punctures marks on the hemelytra.</p><p>Description: Body: Reddish brown; antennae, rostrum and legs lighter; head whitish (Fig. 27A). Head: Pubescent, with small and well-spaced hairs; clypeus darker than head, same colour as antennae; antenniferous process less than half of pedicel length (Fig. 27E); pedicel twice the size of scape; eyes with just a few scarcely distributed ommatidia; bucculae rounded, widest anteriorly, with scarce hairs on its border and punctures organized in a single row in the middle; rostrum reaching at least third abdominal segment (Fig. 27B, C). Thorax: Finely punctate, with tiny punctures on collar and on the posterior lobe of pronotum; anterior border straight, posterior only slightly sinuous; paranota carinate with hairs and few scattered scale-like projections on its border (Fig. 27F); scutellum large, about one-fourth of the maximum pronotum width (Fig. 27D). Hemelytra: Coriaceous and coleopteroid, evenly pubescent; laterally constricted anteriorly; carina-like vein extending from the anterior border to the final third of hemelytra, fading abruptly; finely impressed by punctures anteriorly up to the hemelytra constriction, completely smooth posteriorly to the hemelytra constriction. Scent gland: Anterior branch presenting a lightly cracked texture, tip tilted horizontally, slightly longer than half of the rest of the anterior branch; posterior branch lightly curved, with evenly enlarged edges through its whole extant and the same cracked texture observed on the anterior branch; sulcus more prominent on posterior branch than in anterior branch; evaporatorium with curved border anteriorly, advancing to up to one-third of the mesopleuron on its upper part (Figs 16H – 27C).</p><p>Measurements: BL, 1.59 (1.62; 1.56– xM; 1.62–1.65 xF); BW, 0.72 (0.80; 0.67–0.70 xM; 0.76–0.80 xF); HL, 0.22 (0.19; 0.21–0.25 xM; 0.19–0.25 xF); HW, 0.29 (0.29; 0.27–0.30 xM; 0.29– xF); ID, 0.20 (0.19; 0.19– 0.21 xM; 0.19–0.21 xF); PL, 0.31 (0.30; 0.29–0.32 xM; 0.30–0.32 xF); PW, 0.54 (0.55; 0.51–0.55 xM; 0.55–0.57 xF); AS, 0.10 (0.10; 0.10–0.11 xM; 0.10– xF); AP, 0.21 (0.21; 0.19–0.21 xM; 0.21– xF); AB, 0.20 (0.21; 0.19– 0.21 xM; 0.19–0.21 xF) and AD, 0.31 (0.30; 0.30–0.32 xM; 0.30– xF).</p><p>Etymology: This species was named after the Sucre state, in Venezuela, where the type was collected.</p><p>Distribution: Described from Sucre, Venezuela.</p><p>Material examined: Holotype xF: VENEZUELA: Sucre: 4 m, 7 km S El Pilar, 29 July 1987, S. &amp; J. Peck, rainforest remnant, leaf &amp; log litter [Guidoti PhD –  Vianaidinae 004] (AMNH). Paratypes: VENEZUELA: Sucre: 4 m, 7 km S El Pilar, 29 July 1987, S. &amp; J. Peck, rainforest remnant, leaf &amp; log litter [Guidoti PhD – 3xM,  Vianaidinae 001– 002, 005; 1xF,  Vianaidinae 003] (AMNH).</p></div>	https://treatment.plazi.org/id/B368F165077BFFFCFC27FA293D59FB29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650775FFFDFEA3FB663D63FA0A.text	B368F1650775FFFDFEA3FB663D63FA0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anommatocoris zeteki Drake & Froeschner 1962	<div><p>Anommatocoris zeteki</p><p>Drake &amp; Froeschner, 1962 (Fig. 28)</p><p>Diagnosis:  Anommatocoris zeteki can be identified by the colour of the head, the lightly pronounced humeral angle, the relatively large scutellum, the carina-like vein reaching the middle of the hemelytra and the scent gland unequally divided as in  A. coibensis and  A. schuhi . It differs from these two species in the degree of development of such traits, on the body length and width remarkably smaller, and by the almost straight and with enlarged edges anterior branch of the peritreme.</p><p>Redescription: Body: Brown; head, same as the body; antennae, rostrum and legs lighter (Fig. 28A). Head: Pubescent, with small and well-spaced hairs; clypeus lighter than head, same colour as antennae; antenniferous process about one-third of pedicel length; pedicel more than twice the size of scape; eyes with just a few scarcely distributed ommatidia; bucculae rounded, widest anteriorly, with scarce hairs on its border and punctures organized in a single row in the middle with a couple of extra punctures above, slightly concave posteriorly (Fig. 28E); rostrum reaching at least third abdominal segment (Fig. 28B, C). Thorax: Finely punctate, with fine punctures on collar and on the posterior lobe of pronotum; anterior border straight, posterior sinuous; paranota carinate culminating in a small humeral acute angle (Fig. 28F), with hairs on its border; scutellum large, less than one-third of the maximum pronotum width (Fig. 28D). Hemelytra: Coriaceous and coleopteroid, pubescence apparently concentrated on subcostal and costal area, with some few scattered hairs in the middle of the hemelytra; laterally constricted anteriorly; carina-like vein extending from the anterior border to the middle of hemelytra, fading abruptly; punctures on anterior part up to the hemelytra constriction, completely smooth posteriorly to the hemelytra constriction. Scent gland: Anterior branch with slightly sinuous edges, these considerably enlarged if compared to posterior branch, tip slightly tilted horizontally; posterior straight, edges conspicuously narrow, bottom edge three times as wide as upper edge; sulcus prominent on posterior branch, weakly impressed on anterior branch; evaporatorium with straight border anteriorly, advancing to slightly less than one-third of mesopleuron (Figs 16I – 28C).</p><p>Measurements: BL, 1.85 (1.77; 1.77 xM; 1.94 xF); BW, 0.86 (0.78; 0.78 xM; 0.95 xF); HL, 0.22 (0.23; 0.23 xM; 0.21 xF); HW, 0.32 (0.30; 0.30 xM; 0.34 xF); ID, 0.19 (0.17; 0.17 xM; 0.21 xF); PL, 0.34 (0.32; 0.32 xM; 0.36 xF); PW, 0.58 (0.55; 0.55 xM; 0.61 xF); AS, 0.11 (0.11; 0.11 xM; 0.11 xF) and AP, 0.25 (0.25; 0.25 xM; 0.25 xF).</p><p>Distribution: Known only from the Panama Canal Zone.</p><p>Material examined:   Holotype xM: PANAMA:  Barro Colorado Island: Panama Canal Zone,VII.VIII.42, J. Zetek col. (USNM)  .   Allotype: PANAMA:  Barro Colorado Island: Panama Canal Zone, VII.VIII.42, J. Zetek col., No. 4988. [Guidoti PhD –  Vianaidinae 036] (USNM)  .</p></div>	https://treatment.plazi.org/id/B368F1650775FFFDFEA3FB663D63FA0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650774FFFEFF2DF9873EFFFA6F.text	B368F1650774FFFEFF2DF9873EFFFA6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Henryvianaida Guidoti 2020	<div><p>Henryvianaida Guidoti et al.,  gen. nov.</p><p>lsid: urn:lsid:zoobank.org:act: 9B302C57-62DF-45DB-8BB7-388155AA5787</p><p>Diagnosis: This genus can be easily identified by the constricted mandibular plates, which is unique among the known extant vianaidines. There are only macropterous forms known for this genus but the presence of an anteriorly expanded paranota, the large scutellum, the constriction on the anterior part of the hemelytra, the narrow subcostal area less than half of the width of the discoidal area in its widest part, and the scent gland peritreme clearly Y-shaped and not laterally expanded appears to be diagnostic characters for the genus and possibly not affected by the possible and perhaps expected wing polymorphism.</p><p>Description: Head: Triangular in dorsal view, pubescent, with hairs on vertex; mandibular plates constricted; pedicel three times the length of scape, other segments missing; eyes fully developed, compound; bucculae posterior border straight; rostrum reaching abdomen. Thorax: Posterior lobe raised, barely punctate on most of its extant, but strongly punctate near posterior border; collar somehow distinct, punctate; anterior border convex, passing posterior edge of the eyes; posterior border usually convex; paranota explanate, smooth, expanded anteriorly. Hemelytra: Macropterous, clavus and vein-less membrane well-defined; constricted anteriorly; punctate on coriaceous parts; subcostal area subvertical; costal vein not noticeable; costal area explanate, punctate; hypocosta broad and finely punctate. Scent gland: Anterior branch not laterally projected, strongly inclined forward; posterior branch usually straight; evaporatorium barely projected on mesopleuron, occupying a slightly curved and inclined area. Legs: Femora swollen; tarsi two segment, second segment many times the length of the first. Abdomen: Rectangular, covered with hairs; spiracles located ventrally near lateral margins of abdominal sternites, these straight.</p><p>Etymology: This genus was named after our good friend and American entomologist, Dr Thomas J. Henry. Dr Henry helped the leading author with his PhD by serving as a member on the annual evaluation committee and as his sponsor on a predoctoral fellowship awarded by the Smithsonian Institution. Besides this close relationship with this project, Dr Henry has a productive career, contributing greatly on the study of Heteroptera and specially  Miridae . His enthusiasm and passion are extremely contagious and left a good mark on the leading author after this year working side-by-side at the NMNH. The ending is based on  Vianaida, the type genus of the family.</p><p>Type species:  Henryvianaida colombiensi s sp. nov.</p><p>Distribution: Known from Colombia and Peru (Fig. 18B).</p><p>Discussion:  Henryvianaida is proposed here based on two macropterous singletons, both described as new species in this contribution. The difference in size of these two species is unique among  Vianaidinae genera to this date. The genus presents at least one unique feature among all known extant species of  Vianaidinae, which is the constricted mandibular plate. The general habitus of both species resembles  A. bolivianus, but it is impossible to compare the newly described taxa with this species due to its holotype situation. However, features like the presence of an anteriorly explanate paranota, a subhorizontal subcostal area and an anterior branch of peritreme more inclined forward, together with the aforementioned constricted mandibular plates, put these two species apart from this macropterous species of  Anommatocoris . Following the results of our analysis, these two species are here proposed as a new genus.</p></div>	https://treatment.plazi.org/id/B368F1650774FFFEFF2DF9873EFFFA6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650777FFFEFEF9FA2F3A24F9A2.text	B368F1650777FFFEFEF9FA2F3A24F9A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Henryvianaida colombiensis Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Henryvianaida colombiensis</p><p>Guidoti et al., sp. nov. (Fig. 29)</p><p>lsid: urn:lsid:zoobank.org:act: 4C1E8206-5618-4FFB- B57A-904D59E56BC1</p><p>Diagnosis:  Henryvianaida colombiensis can be easily distinguished from the other  Henryvianaida species by its size: it is almost twice as long and at least twice as large. Additionally, its wider paranota, costal area and consequently hemelytra are also important diagnostic differences among these two species.</p><p>Description: Body: Head, pronotum, hemelytra brown (Fig. 29A); antennae, rostrum, legs and abdomen light brown, yellowish. Head: Pubescent; eyes fully developed, length at least half of length of the head; antenniferous process small; pedicel three times longer than scape, basi- and distiflagellomere missing; bucculae roundish, narrower posteriorly, border pubescent and slightly serrate, few fine punctate present, posterior border straight (Fig. 29E); rostrum reaching first abdominal segment (Fig. 29B, C). Thorax: Pubescent, finely punctate, except for middle of posterior lobe; paranota explanate, narrowing posteriorly, border sinuous, pubescent; scutellum large, less than one-third the maximum width of pronotum (Fig. 29D); sternal laminae narrow, punctate. Hemelytra: Clavus coarsely punctate; discoidal, subcostal and costal area only finely punctate; discoidal area about three times subcostal area at its widest; subcostal area wider at middle, extending to the apex of membrane; costal area broadening posteriorly, widest at middle; membrane without inner row of punctuation (Fig. 29F). Scent gland: Anterior branch strongly inclined forward, tip curved horizontally, short and not swollen, sulcus barely present, fading distally; posterior branch tip swollen and curved, sulcus prominent through its entire length (Figs 16J – 29C).</p><p>Measurements: BL, 4.19; BW, 1.88; HL, 0.27; HW, 0.59; ID, 0.29; PL, 0.70; PW, 1.16; AS, 0.25 and AP, 0.76.</p><p>Etymology: This species was named for the country of its type locality, Colombia, becoming the third  Vianaidinae macropterous species to be named based on its type locality.</p><p>Distribution: Described from Caldas, Colombia.</p><p>Material examined:   Holotype xM: COLOMBIA: Caldas Prov.: Villamaria, 2015, D. Forero col. [MPUJ _ ENT_0046225] [Guidoti PhD – Extraction #374] [Guidoti PhD –  Vianaidinae 059] (MPUJ).</p></div>	https://treatment.plazi.org/id/B368F1650777FFFEFEF9FA2F3A24F9A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650776FFF8FF02FBC13ECAFB0B.text	B368F1650776FFF8FF02FBC13ECAFB0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Henryvianaida machupicchuensis Guidoti 2020	<div><p>Henryvianaida machupicchuensis Guidoti et al.,  sp. nov. (Fig. 30)</p><p>lsid: urn:lsid:zoobank.org:act: 15A7E3EB-F21F-431E-BD14-A5D2E9B620CD</p><p>Diagnosis: This species presents scale-like projections on the anterior part of the hemelytra border, and also differs from  H. colombiensis by the anterior branch of the scent gland peritreme, more curved and shorter, and by the entirely enlarged edges on the posterior branch, and by its subcostal area, only less than twice wider than the costal area in its widest part.</p><p>Description: Body: Head, pronotum, hemelytra light brown (Fig. 30A); antennae, rostrum legs and abdomen even lighter, yellowish. Head: Pubescent; eyes fully developed, length less than half of length of the head; antenniferous process small, less than one-eighth of pedicel length; pedicel less than three times longer than scape, basi- and distiflagellomere missing; bucculae roundish, narrower posteriorly, border pubescent, few coarse punctations near junction with head, a second-row compound by three smaller punctates present, posterior border straight (Fig. 30E); rostrum only reaching first abdominal segment (Fig. 30B, C). Thorax: Hairs mostly anteriorly, finely punctate, except for middle of posterior lobe; paranota explanate, narrowing posteriorly, border sinuous, lightly pubescent (Fig. 30F); scutellum large, less than one-third the maximum width of pronotum (Fig. 30D); sternal laminae narrow, punctate. Hemelytra: Margins with scale-like projections anteriorly; clavus coarsely punctate; discoidal, subcostal areas coarsely punctate anteriorly, then fading to become only finely punctate; discoidal area about two and a half times subcostal area at its widest; subcostal area wider at middle, extending to the apex of membrane; costal area broadening posteriorly, widest at middle; membrane without inner row of punctuation. Scent gland: Anterior branch strongly inclined forward, tip curved horizontally and distant from hemelytra border, short and not swollen, sulcus barely present, fading distally; posterior branch tip curved, sulcus prominent through its entire length (Figs 16K – 30C).</p><p>Measurements: BL, 2.19; BW, 0.78; HL, 0.19; HW, 0.30; ID, 0.17; PL, 0.36; PW, 0.51; AS, 0.13 and AP, 0.32.</p><p>Etymology: This species was collected along the Putucusí trail in Peru from where you can observe Machu Picchu, the famous archeological site of the Incas.</p><p>Distribution: Described from Urubamba, Peru. Material examined: Holotype xM: PERU: Urubamba Prov.: Putucusí trail, 2104 m, 13º09’11.5”S 72º31’38.9”W, 1 January 2010, J. Heraty, cloud forest [H10-178] [Guidoti PhD – Extraction #370] [Guidoti PhD –  Vianaidinae 058] (GC).</p></div>	https://treatment.plazi.org/id/B368F1650776FFF8FF02FBC13ECAFB0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650771FFF8FF47FA8B3A9EF884.text	B368F1650771FFF8FF47FA8B3A9EF884.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterovianaida Montemayor & Carpintero 2007	<div><p>Pterovianaida Montemayor &amp; Carpintero, 2007</p><p>Diagnosis: This genus is characterized by the abundant long hairs on the head, which is remarkably declined, by the presence of paranota constricted in the middle and a conspicuously raised distal part of the pronotum and by the scent gland that is laterally projected with the edges of the tips of both anterior and posterior peritreme branches enlarged.</p><p>Redescription: Head: Triangular in dorsal view, pubescent, with large, long hairs on vertex, once referred as macrochaetae; pedicel subequal to basiflagellomere and to distiflagellomere, each one of these more than twice the length of scape; eyes fully developed, compound; rostrum reaching at least the second abdominal segment. Thorax: Posterior lobe strongly raised, pubescent; entirely punctate; collar distinct, punctate; anterior border straight, posterior convex; paranota explanate, smooth, constricted in the middle, between the anterior and posterior lobes. Hemelytra: Macropterous, clavus and vein-less membrane well-defined; scale-like projections on hemelytra border; punctate on coriaceous parts; membrane with one inner row of punctations; hypocosta broad and finely punctate. Scent gland: Anterior branch gradually projected laterally, tip much more projected than base; tip tilted horizontally, large, conspicuously inclined frontwards; posterior branch usually curved, with enlarged tip. Legs: Femora same width of the other segments; tarsi two segment, second segment remarkably longer and many times the length of the first. Abdomen: Rectangular, covered with hairs; spiracles located ventrally near lateral margins of abdominal sternites, these straight.</p><p>Type species:  Pterovianaida melchiori Montemayor &amp; Carpintero, 2007 .</p><p>Distribution: Known from Brazil and Peru (Fig. 18C).</p><p>Discussion:  Pterovianaida was described by Montemayor &amp; Carpintero (2007) based on a single slide-mounted specimen. Additionally, it is clear the authors were not aware of macropterous species described by Schuh et al. (2006) when proposing this genus. However, the genus gained its second species almost ten years after its original description, based on</p></div>	https://treatment.plazi.org/id/B368F1650771FFF8FF47FA8B3A9EF884	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F1650772FFFBFF37FCA13B2EFA85.text	B368F1650772FFFBFF37FCA13B2EFA85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaumamannia Drake & Davis 1960	<div><p>Thaumamannia Drake &amp; Davis, 1960</p><p>Diagnosis: Characterized by the inclined head, the presence of explanate paranota and costal area, the relatively small scutellum, conspicuously concave hemelytra, and the laterally projected scent gland, strongly Y-shaped with enlarged tips in both anterior and posterior branches of the peritreme.</p><p>Redescription: Head: Declined, usually with most of its length hidden in dorsal view, heavily pubescent; clypeus usually in different colour than head; pedicel subequal to basiflagellomere, both smaller than distiflagellomere and usually twice as big as scape; eyes reduced with none or few ommatidia, triangular in shape; rostrum reaching at least the third abdominal segment. Thorax: Posterior lobe slightly raised; coarsely punctate, mostly on collar and posterior lobe, collar indistinct; anterior border usually concave; posterior usually straight; paranota explanate, smooth, usually projected forwards. Hemelytra: Entirely coriaceous and coleopteroid, ovate, pubescent; remarkably convex, no clavus nor membrane distinct; carina-like vein extending variably from the anterior border, deeply punctate; costal area explanate, with one inner row of punctations only; hypocosta broad and coarsely punctate. Scent gland: Anterior branch gradually projected laterally, tip much more projected than base; varying in height, conspicuously inclined frontwards; tip tilted horizontally, large; posterior branch usually curved, with enlarged tip. Legs: Femora only slightly swollen; tarsi two segment, second segment at least five-times longer.</p><p>Type species:  Thaumamannia manni Drake &amp; Davis, 1960 .</p><p>Distribution: Brazil, Bolivia and Suriname (Fig. 18D).</p><p>Discussion: The genus was described by Drake &amp; Davis (1960) and the second species,  T. vanderdrifti Van Doesburg (1977) was described 17 years later. From these two species, only  T. manni was collected associated with ants (Drake &amp; Davis, 1960). The broadly ovate body, which was considered a strong diagnostic feature of the genus, is now challenged with the description of a new species herein described. However, the conspicuously Y-shaped scent gland remains as an important diagnostic character for the genus, currently composed by a total of four species, including the two species described below.  Thaumamannia vanderdrifti was the first  Vianaidinae reported from Brazil (Guidoti et al., 2014) and now both  T. insolita and  T. urucuana are also reported from that country.  Thaumamannia insolita and one specimen of  T. urucuana had its locality data reported as lost and we only know the state where they came from (Pará State, Brazil). The only species with immature forms described is  T. vanderdrifti, which had its fifth instar described and analysed in SEM (Guidoti et al., 2014).</p></div>	https://treatment.plazi.org/id/B368F1650772FFFBFF37FCA13B2EFA85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F165074DFFC5FED2FF503D84FD83.text	B368F165074DFFC5FED2FF503D84FD83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaumamannia insolita Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Thaumamannia insolita</p><p>Guidoti et al., sp. nov. (Fig. 33)</p><p>lsid: urn:lsid:zoobank.org:act: D94DDB01-3100- 4AF2-A8B1-B8A235C4CF22</p><p>Diagnosis:  Thaumamannia insolita is a new species found to be sister to the other  Thaumamannia species, while presenting a few traits shared with  Anommatocoris . The inclination of the head, the differences in colour between the head and body, the enlarged edges of the tips in the anterior branch of the peritreme and the slender body are characters that distinguish this species from its congeners.</p><p>Description: Body: Brown, with scutellum and paranota borders light brown; head, antennae, rostrum and legs yellowish, except for clypeus, which is as brown as the body (Fig. 33A). Head: Slightly declined, pubescent; antenniferous process almost half the scape length; pedicel subequal to basiflagellomere, these slightly bigger than scape and smaller than distiflagellomere; eyes reduced with few ommatidia, somehow triangular; bucculae roundish, border serrate, widest in the middle, two rows of small punctures, posterior border concave (Fig. 33F); rostrum reaching at least the fourth abdominal segment (Fig. 33B, C). Thorax: Pubescent; coarsely punctate, except for the callus region; anterior border straight; posterior convex; paranota explanate, smooth, not projected frontwards, with scale-like projections on its border giving it a serrate-like aspect; scutellum narrow, less than one-sixth of pronotum maximum width (Fig. 33E); sternal laminae considerably narrow, punctate. Hemelytra: Entirely coriaceous and coleopteroid, ovate, pubescent; remarkably convex; carina-like vein extending from the anterior border to the posterior third of the hemelytra; deeply and entirely punctate, these bigger at anterior region, small at posterior third and even smaller at middle; costal area not punctate, anterior border round. Scent gland: Anterior branch gradually projected laterally, tip tilted horizontally, large, only slightly shorter than the rest of anterior branch, much more projected than base (Fig. 33D), edges equally thick throughout its length; posterior branch straight, with only a slightly enlarged tip with a weak aspect of a cracked texture; sulcus prominent on both branches, fading only distally on the anterior branch tip; evaporatorium curved anteriorly, projected up to half of mesopleuron (Figs 17A – 33C).</p><p>Measurements: BL, 1.52; BW, 0.86; HL, 0.15; HW, 0.29; ID, 0.19; PL, 0.30; PW, 0.62; AS, 0.11; AP, 0.17; AB, 0.21 and AD, 0.30.</p><p>Etymology: This species confused the authors and could only be placed in a genus after the phylogenetic analysis. Due to its puzzled nature we considered this species insolitus, which is Latin for unusual or unaccustomed.</p><p>Distribution: Described from Pará, Brazil.</p><p>Material examined: Holotype xF: BRAZIL: Pará state. [ Hemiptera 6º P8 (E)] [GCLBN27] [Guidoti PhD – FR056] [Guidoti PhD –  Vianaidinae 057] (GC).</p></div>	https://treatment.plazi.org/id/B368F165074DFFC5FED2FF503D84FD83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F165074FFFC7FECCFD113B7FFEF7.text	B368F165074FFFC7FECCFD113B7FFEF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaumamannia urucuana Guidoti & Montemayor & Campos & Guilbert 2020	<div><p>Thaumamannia urucuana</p><p>Guidoti et al., sp. nov. (Fig. 35)</p><p>lsid: urn:lsid:zoobank.org:act: AE4DF1D4-69B9- 48EC-9B79-02A64B87AC08</p><p>Diagnosis: This species resembles the most  T. vanderdrifti, but can be recognized by the straight border of hemelytra on its anterior part at the scent gland level and by the tip of the anterior branch of the peritreme, which does not reach the hemelytra border like in  T. vanderdrifti .</p><p>Description: Body: Dark brown, lighter on paranota and costal area; antennae, rostrum and legs yellowish (Fig. 35A). Head: Strongly declined, pubescent; antenniferous process about one-third of the scape length; pedicel almost three times the scape length, basi- and distiflagellomeres missing; eyes reduced with few ommatidia; bucculae round, smooth and pubescent on its border, widest in the middle, with only few punctures in the middle of the bucculae, posterior border rounded (Fig. 35F); rostrum reaching at least the third abdominal segment (Fig. 35B, C). Thorax: Pubescent; coarsely punctate except for the callus region; anterior and posterior border straight; paranota explanate, not punctate, bearing scale-like projections on its border, projected frontwards surpassing most of the eyes; scutellum large, less than one-eighth of pronotum maximum width (Fig. 35E); sternal laminae narrow, punctate. Hemelytra: Coriaceous, coleopteroid, ovate, densely pubescent and remarkably convex; carina-like vein extending from the anterior border to the posterior third of the hemelytra; deeply and evenly punctate; costal area with scale-like projections on its border, with one row of inner punctures, anterior border straight, revealing scent gland dorsally. Scent gland: Anterior branch gradually projected laterally, tip tilted horizontally, large, same size as the rest of the anterior branch, much more projected than base of anterior branch (Fig. 35D), edges thicker on the tip; posterior branch curved, with considerably enlarged tip with a cracked texture; sulcus prominent on both branches, fading only distally on the anterior branch tip; evaporatorium straighter than curved anteriorly, projected up to half of mesopleuron (Figs 17C – 35C).</p><p>Measurements: BL, 1.77 (1.79; 1.79 xM; 1.75 xF); BW, 1.23 (1.24; 1.24 xM; 1.22 xF); HL, 0.12 (0.10; 0.10 xM; 0.15 xF); HW, 0.34 (0.32; 0.32 xM; 0.36 xF); ID, 0.23 (0.23; 0.23 xM; 0.23 xF); PL, 0.37 (0.40; 0.40 xM; 0.34 xF); PW, 1.03 (1.06; 1.06 xM; 0.99 xF); AS, 0.12 (0.13; 0.13 xM; 0.11 xF) and AP, 0.32 (0.32; 0.32 xM).</p><p>Etymology: This species was named based on Urucu, its type locality.</p><p>Distribution: Described from Brazil; holotype locality is in the State of Amazonas, along the [Urucu-Coari-Manaus gas pipeline]; paratype locality was reported as missing (J. A. M. Fernandes, pers. comm. to the lead author).</p><p>M a t e r i a l e x a m i n e d: H o l o t y p e   x M: B R A Z I L:  Amazonas: Petrobras-Urucu, 25 October 2006, S. Dias col. [clareira 3, método de Winkler]  [SID 103] (MPEG) .   Paratype: BRAZIL:  Pará state. [ Hemiptera 7º P10 (E)]   [GCLBN26] [ Guidoti PhD – FR057] [Guidoti PhD –  Vianaidinae 056]  (GC).</p></div>	https://treatment.plazi.org/id/B368F165074FFFC7FECCFD113B7FFEF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
B368F165074EFFC0FC7AFEC93D09F8B2.text	B368F165074EFFC0FC7AFEC93D09F8B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thaumamannia vanderdrifti Van Doesburg 1977	<div><p>Thaumamannia vanderdrifti</p><p>Van Doesburg, 1977 (Fig. 36)</p><p>Diagnosis: This is the largest species of the genus and it is the only known species of  Thaumamannia to present a laterally projected edge of the anterior branch tip of the peritreme, forming a sulcus that is even noticeable in dorsal view.</p><p>Redescription: Body: Dark brown, slightly lighter on paranota and costal area; antennae, rostrum and legs even lighter (Fig. 36A). Head: Strongly declined, pubescent; antenniferous process about half of the scape length; pedicel mora than two times the scape length, slightly smaller than basiflagellomere, distiflagellomere one and a half times the pedicel length; eyes reduced with few ommatidia; bucculae round, smooth and pubescent on its border, widest anteriorly, with few punctures near insertion with the head, these big and deeply impressed, posterior border concave (Fig. 36F); rostrum reaching at least the third abdominal segment (Fig. 36B, C). Thorax: Pubescent; coarsely punctate including few punctations on the middle of callus region; anterior and posterior border straight; paranota explanate, with few punctations, bearing scale-like projections on its border, projected frontwards surpassing the eyes; scutellum large, almost one-ninth of pronotum maximum width (Fig. 36E); sternal laminae narrow, punctate. Hemelytra: Coriaceous, coleopteroid, ovate, densely pubescent and remarkably convex; carina-like vein extending from the anterior border to the posterior third of the hemelytra; deeply and evenly punctate; costal area with scale-like projections on its border, with one row of inner punctures, anterior border round, revealing scent gland dorsally. Scent gland: Anterior branch gradually projected laterally, tip tilted horizontally, large, almost half the size of the rest of the anterior branch, much more projected than base of anterior branch, edges definitely thicker on the tip, these, unevenly laterally projected to the point of being observable dorsally (Fig. 36D); posterior branch strongly curved distally, with considerably enlarged tip with cracked texture; sulcus prominent on both branches, fading only distally on the anterior branch tip; evaporatorium with straight border anteriorly, only slightly projected on mesopleuron (Figs 17D – 36C).</p><p>Measurements: BL, 2.29 (2.16–2.37 xM; 2.30–2.39 xF); BW, 1.51 (1.37–1.48 xM; 1.56–1.65 xF); HL, 0.12 (0.08–0.17 xM; 0.10–0.15 xF); HW, 0.47 (0.42–0.51 xM; 0.46–0.51 xF); ID, 0.27 (0.25–0.30 xM; 0.25–0.29 xF); PL, 0.53 (0.46–0.55 xM; 0.55–0.57 xF); PW, 1.23 (1.06– 1.22 xM; 1.27–1.37 xF); AS, 0.13 (0.13– xM; 0.11–0.13 xF); AP, 0.30 (0.29–0.30 xM; 0.29–0.30 xF); AB, 0.39 (0.36–0.42 xM; 0.40 xF) and AD, 0.46 (0.46 xM).</p><p>Distribution: Described from Suriname by Van Doesburg (1977) and reported in the Brazilian state of Pará by Guidoti et al. (2014).</p><p>Material examined: Other specimens:   BRAZIL: Pará: Parauapebas, Caverna GEM 1784 (Est. Úmida) [GCLBN5] [ Guidoti PhD – FR036] [Guidoti PhD – 1xM,  Vianaidinae 049] (GC) ;   BRAZIL: Pará: Parauapebas, Gruta S 11 D-081 [Guidoti PhD – Extraction #261] [GCLBN8] [ Guidoti PhD – FR038] [Guidoti PhD – 1xF,  Vianaidinae 050] (GC) ;   BRAZIL: Pará: Parauapebas, Gruta S 11 D-99 [GCLBN10] [ Guidoti PhD – FR042] [Guidoti PhD – 1xF,  Vianaidinae 051] (GC) ;   BRAZIL: Pará: Parauapebas, D-82 [GCLBN11] [Guidoti PhD – FR043] [ Guidoti PhD – 1xF,  Vianaidinae 052] (GC) ;   BRAZIL: Pará: Canaã dos Carajás, Gruta S11 12 [GCLBN9.2] [ Guidoti PhD – FR039] [Guidoti PhD – 1xM,  Vianaidinae 053] (GC) ;   BRAZIL: Pará: Canaã dos Carajás, Gruta S11 12 [Guidoti PhD – Extraction #262] [GCLBN9.3] [ Guidoti PhD – FR041] [Guidoti PhD – 1xM,  Vianaidinae 054] (GC)  .</p></div>	https://treatment.plazi.org/id/B368F165074EFFC0FC7AFEC93D09F8B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Guidoti, Marcus;Montemayor, Sara Itzel;Campos, Luiz Alexandre;Guilbert, Eric	Guidoti, Marcus, Montemayor, Sara Itzel, Campos, Luiz Alexandre, Guilbert, Eric (2020): Phylogenetic analysis and revision of the strangest lace bug subfamily Vianaidinae (Heteroptera: Tingidae). Zoological Journal of the Linnean Society 188 (4): 1172-1212, DOI: 10.1093/zoolinnean/zlz089, URL: https://academic.oup.com/zoolinnean/article/188/4/1172/5680759
