identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5081F89D1CBA562790AFE373CCB9E305.text	5081F89D1CBA562790AFE373CCB9E305.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplazium velutinum Holttum	<div><p>Diplazium velutinum Holttum, Gardens’ Bulletin Straits Settlements 9: 126. 1937.</p><p>Lectotype (designated here).</p><p>• R. E. Holttum S. F. N. 31221 (SING!); K [K 000492397, K 000451191, K 000492398]! and L [L. 3514169]!.</p><p>Syntypes.</p><p>• R. E. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.38333&amp;materialsCitation.latitude=4.4666667" title="Search Plazi for locations around (long 101.38333/lat 4.4666667)">Holttum</a> S. F. N. 31221 (SING!, K [K 000492397, K 000451191, K 000492398]!, L [L. 3514169]!), Pahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.38333&amp;materialsCitation.latitude=4.4666667" title="Search Plazi for locations around (long 101.38333/lat 4.4666667)">Cameron Highlands</a>, Jalan Telom, 47 th mile, 4 ° 28 ' N, 101 ° 23 ' E, 1,463 m elev., Malaysia, 14 May 1936 ; • R. E. Holttum S. F. N. 23428 (K!), Pahang, Cameron Highlands, 4,800 ft., MALAYSIA, 5 Apr 1930 .</p><p>In his original treatment, Holttum listed two specimens under a single “ Type ” heading — S. F. N. 31221 and S. F. N. 23428 — without designating a holotype and cited no additional material under a separate “ Other specimens ” entry, as he did for other species in the same paper. These two specimens therefore constitute syntypes, and lectotypification is required to fix the nomenclatural type.</p><p>S. F. N. 31221 is selected as the lectotype over S. F. N. 23428 for the following reasons. First, S. F. N. 31221 is the more complete and diagnostically informative specimen, collected from the same locality (Cameron Highlands, 47 th mile, Telom Road, 4,800 ft) explicitly cited in Holttum’s (1937) morphological description and ecological note that the species grows in “ the wettest and shadiest mountain forest, near streams. ” Second, S. F. N. 31221 is represented by five sheets across three herbaria — three sheets at Kew (K 000492397, K 000451191, K 000492398), one at SING, and one at Naturalis (L) — providing exceptional accessibility for future workers, further supporting its selection over S. F. N. 23428, which is held at a single institution. Third, S. F. N. 23428, collected earlier in 1930 from a less precisely localized Cameron Highlands site, appears to have been cited by Holttum as a secondary supporting record. Designation of S. F. N. 31221 therefore best serves nomenclatural stability and is consistent with the intent of Art. 9.12 of the International Code of Nomenclature for algae, fungi, and plants (Turland et al. 2018), which recommends selecting the lectotype that most completely represents the protologue.</p><p>Description.</p><p>Rhizome short, erect. Stipe to 40 cm long, densely clothed with multicellular brown hairs; scales nearly black, tapering, entire, to 10 × 0.8 mm. Lamina bipinnatifid – tripinnatifid, to 25 × 15 cm; rachis and costa with dense brown indument. Pinnae 12–15 pairs; sub-basal pinnae largest, to 8 × 3 cm. Pinnules to 12 × 4 mm, margin ciliate or lobed to halfway to costule. Venation free, 7–10 – jugate. Sori diplazioid; indusium thin, broad. Stele in transverse section at upper stipe.</p><p>Differential diagnosis.</p><p>Diplazium velutinum differs from D. tomentosum (Table 1) in its bipinnatifid – tripinnatifid lamina (vs. pinnate – bipinnatifid), nearly black stipe scales to 10 mm long (vs. pale brown, ca. 4 mm), indument covering the entire stipe and abaxial lamina (vs. restricted to costa and rachis), and restriction to upper montane forest above 1,400 m.</p><p>* Data based on fresh material and UKMB specimens collected in 2018 (NA 2018-49, NA 2018-52).</p></div>	https://treatment.plazi.org/id/5081F89D1CBA562790AFE373CCB9E305	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aliah, Nur;Nadhirah, Nurul;Maideen, Haja;Norhazrina, Nik;Khamis, Shamsul;Mohd Yusop, Syazwani;Basir, Syazwani;Syazwana, Nur;Zainal Abidin, Danial Hariz;Zainee, Nur Farah Ain	Aliah, Nur, Nadhirah, Nurul, Maideen, Haja, Norhazrina, Nik, Khamis, Shamsul, Mohd Yusop, Syazwani, Basir, Syazwani, Syazwana, Nur, Zainal Abidin, Danial Hariz, Zainee, Nur Farah Ain (2026): Rediscovery and lectotypification of Diplazium velutinum Holttum (Athyriaceae), a rare endemic fern from Peninsular Malaysia. PhytoKeys 276: 139-149, DOI: 10.3897/phytokeys.276.197738
