identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AC50CD368A1EFF87A2FB7B9BFE89BBFE.text	AC50CD368A1EFF87A2FB7B9BFE89BBFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche agnetae Johanson & Oláh 2008	<div><p>Helicopsyche agnetae, new species</p><p>Fig. 1–9</p><p>Helicopsyche agnetae genitalia are similar to those of H. zhejiangensis . Helicopsyche agnetae is distinguished from H. zhejiangensis by having narrower ventral part of segment IX, broader dorsal branch of gonocoxite and segment X, and much thicker phallus.</p><p>Male head. Cephalic setal warts large, oval to ovoid, about as broad as eyes in dorsal view (Fig. 1); postantennal setal warts absent; interantennal setal warts large, tubular, oriented vertically between antennae. Distal segment of maxillary palps slightly longer than basal segment, shorter than eye diameter, as long as scape. Foreleg anterior spur minute, about one-fifth as long as posterior fore leg spur.</p><p>Male wings: Forewing dark grey, length 2.8 mm; hind wing length 2.2 mm, with 16 hamuli. Venation as in Fig. 2.</p><p>Male abdomen and genitalia (Fig. 3–9): Sternum VI (Fig. 3, 4) with short process, oriented posteroventrally (Fig. 3), triangular in ventral view (Fig. 4). Segment IX, lateral view (Fig. 5), strongly expanded anteriorly, anteroventral margin nearly straight, except concave near anterior apex (Fig. 5); segment about as high as long; in dorsal view (Fig. 6), with inner margin tongue-shaped; in ventral view (Fig. 7) with minute, irregular central posterior process; in lateral view with weak tergal transverse apodeme forming a nearly straight line above superior appendage. Sternite IX with characteristically shaped apodeme along anterior margin and with lateral straight arms oriented posteriorly and converging distally. Tergum X oriented posteroventrad, slightly up-curving along its length, narrow along basal one-fifth in lateral view (Fig. 5), expands into clubshaped distal part. Tergum X with long, posterolaterally pointing lateral spines in central row, sub-apical part slightly expanded into dorsal lobe above spines, apex rounded. Tergum X deeply incised towards start of lateral spines in dorsal view (Fig. 6), slightly narrowing distally, sub-apical part slightly bent laterally before curving posteriorly at apex, with about 3 weak setae on apex. Superior appendage long, club-shaped, slightly curving posteriorly towards apex (Fig. 5). Primary branch of gonocoxite, lateral view (Fig. 5), almost ovoid, oriented dorsally, narrowest at basis, posterior margin strongly undulating due to presence of about 6 produced setal bases; secondary branch of gonocoxite short, smooth, oriented posteriorly in lateral view, laterally in ventral view. Basal plate broad, dorsal margin convex along its length in lateral view (Fig. 5), apex points anteroventrally; in ventral view (Fig. 7) narrow and parallel-sided along its length. Phallus, lateral view (Fig. 8), anterior half nearly uniformly thick; basis with triangular, anteriorly pointing, membranous plate; angled 90° at midlength; distal part thicker and more membranous than basal half; with pair of long, strong, knifeshaped sclerites having 2 ventrally oriented basal teeth; in ventral view (Fig. 9) nearly 3x wider at mid-length than anterior and posterior parts; sperm duct divide into slender anterior and thick posterior part; knife-shaped sclerites run parallel with lateral margin.</p><p>Holotype male: China: Hong Kong, Sai Kung East Country Park, stream 1.2 km E Tin Mei Shan Mt., at Luk Wu, Malaise trap 20–27.v.2005 [A. Olsson &amp; E. Ohlsson] (NRM, alcohol).</p><p>Paratypes: same data as holotype — 3 males .</p><p>Distribution: SE China (Hong Kong) (Fig. 10).</p><p>Etymology: agnetae, named after Ms. Agneta Olsson, one of the collectors of the species.</p><p>Discussion</p><p>Helicopsyche agnetae, new species and H. zhejiangensis share several synapomorphies, like: segment IX strongly produced anteriorly and with a small concavity immediately ventrally to anterior apex; tergite X deeply and widely incised, and has a lateral row of hooks; superior appendage large, oriented posteroventrally and slightly curving posteriorly along its length; gonocoxite oval, oriented dorsally and angled 90° towards the secondary branch of gonocoxite; phallus bent 90° at mid-length, and with a pair of large, knife-shaped sclerites at endotheca.</p><p>The type localities of the two species are situated more than 800 kilometers from each other (Fig. 10) and there are apparent geographical barriers along the shortest line between the distribution areas of the two species that could induce speciation of their common ancestor. Unidentified Helicopsyche specimens, known only from larvae, exist on Taiwan (Fig. 10). Future sampling will hopefully reveal the identity of the Taiwan Helicopsyche fauna, and whether or not the species are closely related to the mainland Chinese Helicopsyche, or other faunas.</p></div>	https://treatment.plazi.org/id/AC50CD368A1EFF87A2FB7B9BFE89BBFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johanson, Kjell Arne;Oláh, János	Johanson, Kjell Arne, Oláh, János (2008): Helicopsyche agnetae, new species (Trichoptera, Helicopsychidae) described from Hong Kong. Zootaxa 1854 (1): 63-68, DOI: 10.11646/zootaxa.1854.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1854.1.6
