identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AD691A0AB624FFA982E731E4FA6FFCCA.text	AD691A0AB624FFA982E731E4FA6FFCCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bhatia Distant 1908	<div><p>Bhatia Distant, 1908</p><p>Bhatia Distant, 1908: 357; Webb, 1994: 116; Zhang &amp; Webb, 1996: 12; Viraktamath &amp; Murthy, 2009: 47; Shang, Shen, Zhang &amp; Li, 2006: 565; Lu &amp; Zhang, 2015: 146; Yu, Qu, Dai &amp; Yang, 2019: 143; Lu &amp; Zhang, 2024: 278.</p><p>Type species: Eutettix? olivacea Melichar, 1903 .</p><p>Diagnosis. Head short, crown broadly rounded anteriorly, with transverse carina; median length slightly greater than length next to eye; transverse groove near anterior margin; head slightly wider than pronotum. Ocelli marginal, distant from eye. Antennae long, inserted at upper angle of eye. Hind femur apical setal formula 2+2+1.</p><p>Male pygofer with numerous macrosetae on lateral lobe, with or without an inner marginal ridge; if present, the ridge runs obliquely from inner margin to ventroposterior corner. Subgenital plate nearly triangular, inner margin nearly straight. Valve nearly quadrate. Connective Y-shaped or H-shaped; in some species, with a sclerotized plate or an elongate process between connective and aedeagus. Aedeagus with one or more pairs of basal processes.</p><p>Remarks. Species of Bhatia are similar in external morphology and male genitalia, but can be immediately separated from Bhatiahamus by the absence of a pair of stout apical processes on the male tenth abdominal segment. Furthermore, this genus is also similar to Fistulatus, from which it differs most notably in lacking posterior elongations on the pygofer lobe.</p><p>Distribution. Oriental Region and Pacific areas.</p><p>Checklist of Chinese species of Bhatia</p><p>B. biconjugara Zhang &amp; Zhang, 1998</p><p>B. biconjugara Zhang &amp; Zhang, 1998: 178; Shang, Shen, Zhang &amp; Li, 2006: 566; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Sichuan, Guangxi, Guangdong, Fujiang).</p><p>B. bilateritra sp. nov.</p><p>Distribution: China (Hainan).</p><p>B. digitata Shang &amp; Shen, 2006</p><p>B. digitata Shang &amp; Shen, 2006: 568; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Anhui, Henan, Guangxi).</p><p>B. furcata sp. nov.</p><p>Distribution: China (Yunnan).</p><p>B. hastata Shang &amp; Shen, 2006</p><p>B. hastata Shang &amp; Shen, 2006: 567; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Guangxi).</p><p>B. holodonta sp. nov.</p><p>Distribution: China (Yunnan).</p><p>B. koreana (Kwon &amp; Lee, 1979)</p><p>Koreanopsis koreana Kwon &amp; Lee, 1979: 50 .</p><p>B. koreana Zhang &amp; Webb, 1996: 12 .</p><p>Distribution: China (Guangxi, Shaanxi); South Korea.</p><p>B. affinikoreana sp. nov.</p><p>Distribution: China (Hainan).</p><p>B. loba Lu &amp; Zhang, 2024</p><p>B. loba Lu &amp; Zhang, 2024: 283 .</p><p>Distribution: China (Tibet).</p><p>B. multispinosa Lu &amp; Zhang, 2015</p><p>B. multispinosa Lu &amp; Zhang, 2015: 148; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Sichuan).</p><p>B. olivacea (Melichar, 1903)</p><p>Eutettix? olivaceus Melichar, 1903: 191 .</p><p>B. olivacea (Melichar, 1903), Viraktamath, 1998: 171; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Melichariella olivacea Matsumura, 1914: 237 .</p><p>B. divacea Evans, 1947: 230 .</p><p>Distribution: China (Hainan); India; Sri Lanka; Thailand; Java; Indonesia.</p><p>B. parasagittata Lu &amp; Zhang, 2024</p><p>B. parasagittata Lu &amp; Zhang, 2024: 281 .</p><p>Distribution: China (Guizhou).</p><p>B. quadrispinosa Shang &amp; Zhang, 2006</p><p>B. quadrispinosa Shang &amp; Zhang, 2006: 571; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Sichuan).</p><p>B. sagittata Cai &amp; Shen, 1999</p><p>B. sagittata Cai &amp; Shen, 1999: 38 .</p><p>Distribution: China (Henan, Jiangxi).</p><p>B. satsumensis (Matsumura, 1914)</p><p>Melichariella satsumensis Matsumura, 1914: 237 .</p><p>B. satsumensis Esaki &amp; Ito, 1954: 94: Zhang &amp; Webb, 1996: 12; Zhang &amp; Zhang, 1998: 179; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Guangdong); Japan.</p><p>B. unicornis Shang &amp; Li, 2006</p><p>B. unicornis Shang &amp; Li, 2006: 573; Yu, Qu, Dai &amp; Yang, 2019: 143.</p><p>Distribution: China (Guangdong).</p><p>Key to Chinese species of Bhatia (males)</p><p>1 Aedeagus with single basal process.............................................. B. unicornis Shang &amp; Li, 2006</p><p>– Aedeagus with two or more basal processes................................................................ 2</p><p>2 Aedeagal shaft with processes........................................................................... 3</p><p>– Aedeagal shaft without processes........................................................................ 8</p><p>3 Aedeagal shaft with denticulate lateral flanges.............................................................. 4</p><p>– Aedeagal shaft without denticulate lateral flanges........................................................... 5</p><p>4 Style apex rounded, subapex acute (Fig. 48)............................................... B. holodonta sp. nov.</p><p>– Style apex acute, subapex rounded................................................ B. koreana Kwon &amp; Li, 1979</p><p>5 Aedeagal shaft with two pairs of processes (Figs. 13, 19).......................... B. satsumensis (Matsumura, 1914)</p><p>– Aedeagal shaft with only one pair of processes............................................................. 6</p><p>6 Processes at apex of aedeagal shaft arrow-shaped in ventral view............................................... 7</p><p>– Processes at apex of aedeagal shaft not above (Figs. 4, 10).............................. B. olivacea (Melichar, 1903)</p><p>7 Aedeagal shaft gradually tapered subapically (Figs. 18, 24)........................ B. parasagittata Lu &amp; Zhang, 2024</p><p>– Aedeagal shaft distinctly inflated subapically (Figs. 3, 9).............................. B. sagittata Cai &amp; Shen, 1999</p><p>8 Aedeagus with one pair of basal processes................................................................. 9</p><p>– Aedeagus with two or more pairs of basal processes........................................................ 14</p><p>9 Leaf-shaped sclerite articulated between aedeagus and connective (Figs. 6, 12)............... B. loba Lu &amp; Zhang, 2024</p><p>– Leaf-shaped sclerite articulated absent between aedeagus and connective........................................ 10</p><p>10 Aedeagal shaft with basal processes crossing ventrally (Figs. 41–42).............................. B. furcata sp. nov.</p><p>– Aedeagal shaft with basal processes not crossing ventrally................................................... 11</p><p>11 Aedeagal shaft medially inflated, not inflated apically, in ventral view (Fig. 32).................. B. bilateritra sp. nov.</p><p>– Aedeagal shaft apically inflated, not inflated medially, in ventral view (Fig. 22).................................. 12</p><p>12 Style with apical process long and straight........................................ B. hastata Shang &amp; Shen, 2006</p><p>– Style with apical process short and hooklike.............................................................. 13</p><p>13 Margins of basal aedeagal processes smooth (Fig. 22)............................... B. digitata Shang &amp; Shen, 2006</p><p>– Margins of basal aedeagal processes undulate (Fig. 60).................................... B. affinikoreana sp. nov.</p><p>14 Style apex rounded, subapex acute......................................... B. biconjugara Zhang &amp; Zhang, 1998</p><p>– Style apex acute, subapex rounded...................................................................... 15</p><p>15 Single lateral process present between aedeagus and connective (Fig. 23)............. B. multispinosa Lu &amp; Zhang, 2015</p><p>– No process between aedeagus and connective (Fig. 7)......................... B. quadrispinosa Shang &amp; Zhang, 2006</p></div>	https://treatment.plazi.org/id/AD691A0AB624FFA982E731E4FA6FFCCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Qu, Ling;Yang, Yan-Qiong;Yu, Zhou;Dai, Ren-Huai	Qu, Ling, Yang, Yan-Qiong, Yu, Zhou, Dai, Ren-Huai (2025): Four new species of leafhopper genus Bhatia Distant (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) from southern China with an updated checklist and key to Chinese fauna. Zootaxa 5737 (2): 277-285, DOI: 10.11646/zootaxa.5737.2.7, URL: https://doi.org/10.11646/zootaxa.5737.2.7
AD691A0AB621FFA882E73030FE1AFDC7.text	AD691A0AB621FFA882E73030FE1AFDC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bhatia bilateritra Qu & Yang & Yu & Dai 2025	<div><p>Bhatia bilateritra sp. nov.</p><p>(FIGURES 25–33)</p><p>Description. Length (including tegmen): (holotype), 6.9 mm (Fig. 25).</p><p>Body brown; face bright yellow; eyes brown (Figs. 25–27). Crown with two black transverse bands. Crown evenly arcuate, median length subequal to length at level of the eye. Ocellus separated from the corresponding eye by a distance equal to its own diameter. Antennal pit above midline of eye; antenna nearly more than half body length. Clypeus apically expanded. Lateral frontal sutures extending to level of respective ocellus. Hind femoral macrosetal formula 2+2+1.</p><p>Male genitalia. Pygofer side with numerous macrosetae (Fig. 28). Subgenital plate nearly triangular, apical portion digitately curved laterally (Fig. 29). Style with apophysis short and stout, apex bluntly rounded (Fig. 30). Connective Y-shaped, stem markedly longer than arms (Fig. 31). Aedeagal shaft cylindrical, slightly expanded subapically with expansion finely serrate along margin in ventral view; shaft weakly sinuated in lateral view; aedeagus with a pair of elongate laterobase processes arising between aedeagus and connective, the procecess extended beyond approximately 1.5 times the length of aedeagal shaft, internally spiral-like (Figs. 32–33).</p><p>Material examined. HOLOTYPE, ♂, China: Hainan Province, Jianfengling Mountains, 17.IX.2009, collected by Xinfeng Zhang.</p><p>Distribution. China (Hainan).</p><p>Etymology. The new species name is derived from the Latin word “ bilateritra ”, referring to the paired processes of the aedeagus connecting to the connective.</p><p>Remarks. The new species is morphologically similar to B. unicornis in having a columnar aedeagal shaft that is subapically inflated, with the inflated portion exhibiting denticulate margins in ventral view. However, the two species can be consistently distinguished by the following characters: presence of paired of processes linking the aedeagus to the connective (vs. a single process in B. unicornis); absence of a baso-posterior process between the aedeagus and connective (vs. present in B. unicornis); and apical apophysis of the style blunt and rounded (vs. sharply acute in B. unicornis).</p></div>	https://treatment.plazi.org/id/AD691A0AB621FFA882E73030FE1AFDC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Qu, Ling;Yang, Yan-Qiong;Yu, Zhou;Dai, Ren-Huai	Qu, Ling, Yang, Yan-Qiong, Yu, Zhou, Dai, Ren-Huai (2025): Four new species of leafhopper genus Bhatia Distant (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) from southern China with an updated checklist and key to Chinese fauna. Zootaxa 5737 (2): 277-285, DOI: 10.11646/zootaxa.5737.2.7, URL: https://doi.org/10.11646/zootaxa.5737.2.7
AD691A0AB620FFAB82E7313DFD03FEC3.text	AD691A0AB620FFAB82E7313DFD03FEC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bhatia furcata Qu & Yang & Yu & Dai 2025	<div><p>Bhatia furcata sp. nov.</p><p>(FIGURES 34–42)</p><p>Description. Length (including tegmen): (holotype), 7.2 mm (Fig. 34).</p><p>Body dark brown. Crown anterior margin rounded and protruding, with distinct dark patches. Eyes deep brown. Pronotum with black patches and transverse striations. Scutellum yellowish-brown, with paired symmetrical black patches near midlength; transverse impression arcuate and deeply depressed. Face yellowish-brown; frontoclypeus to anteclypeus dark brown, with paired symmetrical striae laterally (Fig. 36). Hind femoral macrosetal formula 2+2+1.</p><p>Male genitalia. Pygofer side sub-trapezoidal, broader basally, narrowing apically, with numerous basal setae (Fig. 37). Valve semicircular. Subgenital plate basally inflated, forming semicircular lobe, apically constricted into slender, finger-like process (Fig. 38). Style with apical apophysis prolonged and beak-like distally (Fig. 39). Connective short, H-shaped; arms slightly extended apically (Fig. 40). Aedeagal shaft columnar and short, with paired basal processes extending ventrally, bent at a right angle and crossing distally, process apices serrate in ventral view; aedeagus inflated, short, weakly curved dorsad in lateral view (Figs. 41–42).</p><p>Material examined. HOLOTYPE: ♂, China: Yunnan, Nannuo Mountain, Xishuangbanna, 9.VI.2023, collected by Yanqiong Yang. PARATYPES: 2♂♂, same collection data as holotype .</p><p>Distribution. China (Yunnan).</p><p>Etymology. The new species name is derived from the Latin meaning "forked", referring to the crossed configuration of the aedeagal processes.</p><p>Remarks. The new species resembles B. digitata in possessing long aedeagal processes but differs as follows: processes of the new species extend in a crossed arrangement ventrally (vs. uncrossed in B. digitata); aedeagal shaft truncate and non-dilated in ventral view (vs. arcuate and dilated in B. digitata); and aedeagus inflated and shortened in lateral view (vs. lacking this feature in B. digitata).</p></div>	https://treatment.plazi.org/id/AD691A0AB620FFAB82E7313DFD03FEC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Qu, Ling;Yang, Yan-Qiong;Yu, Zhou;Dai, Ren-Huai	Qu, Ling, Yang, Yan-Qiong, Yu, Zhou, Dai, Ren-Huai (2025): Four new species of leafhopper genus Bhatia Distant (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) from southern China with an updated checklist and key to Chinese fauna. Zootaxa 5737 (2): 277-285, DOI: 10.11646/zootaxa.5737.2.7, URL: https://doi.org/10.11646/zootaxa.5737.2.7
AD691A0AB623FFAA82E7323DFD70FE72.text	AD691A0AB623FFAA82E7323DFD70FE72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bhatia holodonta Qu & Yang & Yu & Dai 2025	<div><p>Bhatia holodonta sp. nov.</p><p>(FIGURES 43–51)</p><p>Description. Length (including tegmen): (holotype), 6.2 mm (Fig. 43).</p><p>Body yellowish-brown (Figs. 43–45). Crown angularly produced, subapically depressed with two black transverse bands (one discontinuous) and sparse spots along anterior margin; anterior and posterior margins nearly parallel; eyes dark red. Ocellus marginal, distance between ocellus and adjacent eye 4 times ocellar diameter; antennal located just above the eye, antennal pits strongly developed; frontoclypeus narrowing from apex to base, medially convex, with fine black striae laterally; postclypeus rectangular, apex slightly expanded. Face densely mottled with minute brown dots (Fig. 45). Pronotum with straight lateral margins, posterior margin truncate; surface with dense transverse striations and scattered black patches along anterior margin (Fig. 43). Hind femur with macrosetal formula 2+2+1.</p><p>Male genitalia. Pygofer side subcircular in lateral view, posterodorsal angle right-angled; apex bearing numerous long setae (Fig. 46). Valve broadly rounded, subgenital plate relatively short, covered with minute setae, apex extended into a slender process (Fig. 47). Style slender, apex acuminate, subapically bearing a distinct spiniform projection directed laterally (Fig. 48). Connective Y-shaped, arms approximately twice as long as stem (Fig. 49). Aedeagus tubular in dorsal/ventral aspect, lateral margins finely denticulate, with a pair of elongate processes at base, 1.5 times the length of aedeagal shaft. Shaft ventrally arcuate in lateral view, dorsal margin with a serrated, lamellate tooth (Figs. 50–51). Genital pore positioned on ventral surface of shaft slightly proximal to apex (Figs. 50–51).</p><p>Material examined.</p><p>HOLOTYPE, ♂, China: Yunnan Province, Baoshan City, Baihualing, 05.VIII.2018, collected by Lu Yang.</p><p>Distribution. China (Yunnan).</p><p>Etymology. The new species name is derived from the Latin word " holodontus ", referring to the serrated lamellar processes on the aedeagal shaft.</p><p>Remarks. This new species was previously misidentified and reported as B. koreana (Kwon &amp; Lee) by Yu et al. (2019, Figs. 2i–j). However, a detailed re-examination of the male genitalia (including the aedeagal shaft, connective, style, and pygofer lobe) reveals consistent and significant differences that warrant its recognition as a distinct species. B. holodonta sp. nov. can be distinguished from B. koreana by the following combination of characters: style with acute subapex and rounded apex (vs. rounded subapex and acute apex in B. koreana); aedeagal shaft acutely pointed apically in ventral view (vs. truncate in B. koreana); and aedeagal shaft distinctly wider than basal processes in lateral view (vs. subequal in width in B. koreana).</p><p>The new species is comparable to B. digitata in having the aedeagal shaft with a pair of processes. It differs by: the serrated margin of the aedeagal shaft in lateral view (smooth in B. digitata); the serrated margins of both the shaft and its processes in ventral view (smooth in B. digitata); and the style with the subapical process elongated (vs. apical process elongated in B. digitata).</p></div>	https://treatment.plazi.org/id/AD691A0AB623FFAA82E7323DFD70FE72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Qu, Ling;Yang, Yan-Qiong;Yu, Zhou;Dai, Ren-Huai	Qu, Ling, Yang, Yan-Qiong, Yu, Zhou, Dai, Ren-Huai (2025): Four new species of leafhopper genus Bhatia Distant (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) from southern China with an updated checklist and key to Chinese fauna. Zootaxa 5737 (2): 277-285, DOI: 10.11646/zootaxa.5737.2.7, URL: https://doi.org/10.11646/zootaxa.5737.2.7
AD691A0AB622FFAA82E732A9FB17FA8A.text	AD691A0AB622FFAA82E732A9FB17FA8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bhatia affinikoreana Qu & Yang & Yu & Dai 2025	<div><p>Bhatia affinikoreana sp. nov.</p><p>(FIGURE 52–60)</p><p>Description. Length (including tegmen): (holotype), 6.52 mm (Fig. 52).</p><p>Body overall brownish-yellow (Figs. 52–54). Crown anterior margin rounded and prominent. Pronotum densely covered with fine transverse striations, lateral margins with irregular patches (Fig. 52). Forewings yellowish-brown, hyaline with pale venation. Face yellowish brown, frontoclypeal margins with pale markings (Fig. 53). Hind femoral macrosetal formula 2+2+1.</p><p>Male genitalia. Pygofer side subtriangular and densely setose in lateral view (Fig. 55). Subgenital plate basally inflated, forming a subcircular lobe, constricted subapically, terminating in digitate process with irregular marginal folds distally (Fig. 56). Style with apex apophysis tapered into a sharp, laterally directed spine (Fig. 57). Connective H-shaped, both basal and apical portions broad, medial portion constricted and inwardly curved (Fig. 58). Aedeagal shaft smooth and columnar, acute distally, curved C-shaped with paired laterobasal processes extending beyond the length of aedeagal shaft arising from between connective and shaft; the processes rough with apex acuminate and subapically serrate in ventral view; appendage between aedeagus and connective absent (Figs. 59–60).</p><p>Material examined. HOLOTYPE, ♂, China: Hainan, Bawangling Nature Reserve, 25.VIII.2015, collected by Yingjian Wang.</p><p>Distribution. China (Hainan).</p><p>Etymology. The specific name refers to its morphological similarity to B. koreana .</p><p>Remarks. The new species resembles B. koreana but differs markedly in male genital structures: subgenital plate constricted subapically with semicircular basal dilation (vs. evenly tapered without basal dilation in B. koreana); aedeagal shaft lacking dentate processes (vs. with distinct dentiform projections in B. koreana); aedeagus acutely pointed in lateral view (vs. truncate apically in B. koreana); and style terminating directly in an acute projection without distal elongation (vs. distally extended before terminating in an acute apex in B. koreana).</p></div>	https://treatment.plazi.org/id/AD691A0AB622FFAA82E732A9FB17FA8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Qu, Ling;Yang, Yan-Qiong;Yu, Zhou;Dai, Ren-Huai	Qu, Ling, Yang, Yan-Qiong, Yu, Zhou, Dai, Ren-Huai (2025): Four new species of leafhopper genus Bhatia Distant (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) from southern China with an updated checklist and key to Chinese fauna. Zootaxa 5737 (2): 277-285, DOI: 10.11646/zootaxa.5737.2.7, URL: https://doi.org/10.11646/zootaxa.5737.2.7
