identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A82EAD987A465623804692CFA6C73AFB.text	A82EAD987A465623804692CFA6C73AFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tinagma nigripunctum Yagi 2026	<div><p>Tinagma nigripunctum sp. nov.</p><p>Figs 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17</p><p>Japanese name.</p><p>kurohoshi-hime-moguriga.</p><p>Type material.</p><p>Holotype. male. Japan: Okinawa Pref. Itokazu-joato, Itokazu, Nanjo, 24. III. 2021 LT, S. Yagi, J. Oku, HK. Kim, K. Goto, M. Kimura leg., genitalia slide no. SY 1638, Sample ID: SaY 1342 .</p><p>Paratype. 1 female, same data as holotype, genitalia slide no. SY 1639, Sample ID: SaY 1343 .</p><p>Diagnosis.</p><p>Most Tinagma species are difficult to distinguish based on external morphology, as their forewings are greyish brown with white to grey markings. However, this species can be distinguished from others by a combination of characters, particularly an indistinct pale greyish-brown fascia, a fuscous spot at the apex, and small body size. In the male genitalia, this species is similar to T. balteolella (Fischer von Röslerstamm, 1840), T. klimeschi Gaedike, 1987, and T. anchusella (Benander, 1936) in sharing a costal process and an articulated process in the valva. However, it can be distinguished by the acute tip of the valva and the uninterrupted pecten on the outer margin of the valva (roundish tip and interrupted pecten in T. balteolella and T. klimeschi, and roundish tip without pecten in T. anchusella). In the female genitalia, this species is similar to T. balteolella and T. klimeschi in having an elongated sterigma with a U-shaped emargination on the ventral opening. However, it can be distinguished by the straight and uniform width of the sterigma (bulging in T. balteolella and T. klimeschi).</p><p>Description.</p><p>Male (Figs 1, 2, 3, 4, 5, 6). Wingspan: 5.5 mm in holotype. Forewing length: 2.5 mm in holotype. Head: smooth, greyish brown. Frons greyish cream. Ocellus well developed. Antenna greyish brown, filiform but slightly thickened, 0.65 × length of forewing. Proboscis long, without scales. Maxillary palpus reduced. Labial palpus pale brown, short, straight, apically dark greyish-brown. Thorax and tegula: dark greyish brown. Forewing upper side greyish brown with faint pale greyish brown fascia in the middle, anteriorly bordered by dark brown band; apex with fuscous spot; fringe pale brown; underside greyish brown, costa basal half dark brown. Hindwing dull brown; fringe pale brown. Frenulum a single stout seta. Foreleg epiphysis present. Legs greyish brown; each tarsomere dark greyish brown, ringed apically. Abdomen: greyish beige dorsally, beige ventrally. S 2 with slender apodeme, curved inwards; venula present (Fig. 5). S 8 bar-shaped, strongly sclerotised, laterally with a pair of slender rods projected inward. T 8 weakly sclerotised, trapezoid, posteriorly slightly concave (Fig. 6).</p><p>Female. Wingspan: 5.9 mm in paratype. Forewing length: 2.8 mm in paratype. Similar to male but antenna filiform, 0.5 × length of the forewing. Frenulum comprising 2 setae.</p><p>Male genitalia (Figs 7, 8, 9, 10, 11, 12). Uncus and gnathos absent. Tegumen slightly sclerotised, triangular, ventrally with microtrichia and subapically with two pairs of long setae (Fig. 9). Valvae symmetrical; trapezoid valva with short, obliquely upward costal projection; apex of valva with an acuminate articulated process 5 × as long as basal width, medially folded over valval surface and anteriorly projecting to 0.7 × length of valva; ventral margin nearly straight with setose fold; outer margin concave with uninterrupted pecten comprising ca. 18 thickened setae; dorsal margin smooth. Transtilla slender, tapered toward midline. Vinculum a strongly sclerotised band with triangularly projecting lateral lobe; saccus moderately projecting; median membranous area of vinculum with minute microtrichia (Fig. 10). Anellus a sclerotised ring, subapically angular, apically rounded and projecting. Juxta basally flat with broadly sclerotised plate. Phallus long, slender, gently tapered toward tip, basally curved upward; coecum penis small; cornuti absent.</p><p>Female genitalia (Figs 13–16). Papillae anales moderately long. Apophyses posteriores slender, 1.5 × length of apophyses anteriores. Apophyses anteriores connected to a setose triangular lateral sclerite posteriorly. S 9 triangular, with minute spines. Posterior margin of S 8 serrated and almost straight. Ostium bursae arising from the membrane between eighth and ninth sterna, strongly emarginate. Sterigma strongly sclerotised, elongated ring, uniform in width, ventral opening U-shaped and serrated. Ductus bursae slender; slightly constricted posteriorly; posterior 3 / 5 weakly sclerotised, anterior 2 / 5 membranous with minute pectinations; anterior border of sclerotised region with six small sclerotised spinules. Ductus seminalis arising at posterior 3 / 5 of ductus bursae near spinules. Corpus bursae oval, with signum; signum consisting of numerous strong sclerotised slender rods of various sizes, arranged in a radial pattern; minute pectinations around signum.</p><p>DNA data</p><p>(Fig. 17, Table 2). The COI barcode sequences (658 bp) of one male and one female Tinagma nigripunctum sp. nov. 100 % matched. The DNA barcode of Tinagma nigripunctum sp. nov. (BOLD: AGT 4721) is closest to that of T. balteolella (BOLD: AAZ 0894), with a pairwise distance (p-distance) of 9.7 %. The NJ tree also showed similarity to T. anchusella (BOLD: ABW 5879), with a pairwise distance of 10.6 %.</p><p>Biology.</p><p>Unknown (see the discussion below for the suspected host plant family). Adults were collected at the forest edge in late March (Fig. 18).</p><p>Etymology.</p><p>The name of the new species, an adjective, is derived from the Latin nigrum, “black” and punctum, “spot”, referring to the black dot at the apex of the forewing.</p><p>Remarks.</p><p>This species is one of the smallest species of Douglasiidae, along with Tinagma minutissima (Staudinger, 1880) (wingspan 4.5–6.5 mm; Budashkin 2003).</p></div>	https://treatment.plazi.org/id/A82EAD987A465623804692CFA6C73AFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Yagi, Sadahisa	Yagi, Sadahisa (2026): Discovery of a new species of Douglasiidae Heinemann & Wocke, 1876 (Lepidoptera) in Japan. Nota Lepidopterologica 49: 97-108, DOI: 10.3897/nl.49.186673
