taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
7E62E05257DD53F096F0AC92D64CC1B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556044	https://doi.org/10.3897/vz.76.e176241.figure7	Figure 7. Drawings of the heads in dorsal and lateral views of most of the species of Madascincus. A M. arenicola, holotype ZSM 1565 / 2008, Baie des Sakalava. B M. miafina, holotype ZSM 1562 / 2008, Ankarana Special Reserve. C M. stumpffi, holotype SMF 16019, “ Nossibé ” (= Nosy Be). D M. polleni, holotype MNHN 1895.210, “ Mouroundava ” (= Morondava). E M. irery sp. nov., holotype ZSM 135 / 2019, Nosy Be. F Madascincus pyrurus, holotype ZSM 520 / 2001, Mont Ibity. G M. igneocaudatus, ZSM 1600 / 2010, Anakao. H M. mouroundavae, ZSM 13 / 2005, Andasibe. I M. ankodabensis, ZSM 355 / 2006, Ranomafana. J M. melanopleura, ZSM 20 / 2005, Andasibe. K M. minutus, ZSM 400 / 2005, Nosy Mangabe. L M. minotaurus sp. nov., holotype ZSM 116 / 2023, Namoroka. Dwarf species, namely M. nanus and M. macrolepis, not represented. Scale bars = 2 mm.	Figure 7. Drawings of the heads in dorsal and lateral views of most of the species of Madascincus. A M. arenicola, holotype ZSM 1565 / 2008, Baie des Sakalava. B M. miafina, holotype ZSM 1562 / 2008, Ankarana Special Reserve. C M. stumpffi, holotype SMF 16019, “ Nossibé ” (= Nosy Be). D M. polleni, holotype MNHN 1895.210, “ Mouroundava ” (= Morondava). E M. irery sp. nov., holotype ZSM 135 / 2019, Nosy Be. F Madascincus pyrurus, holotype ZSM 520 / 2001, Mont Ibity. G M. igneocaudatus, ZSM 1600 / 2010, Anakao. H M. mouroundavae, ZSM 13 / 2005, Andasibe. I M. ankodabensis, ZSM 355 / 2006, Ranomafana. J M. melanopleura, ZSM 20 / 2005, Andasibe. K M. minutus, ZSM 400 / 2005, Nosy Mangabe. L M. minotaurus sp. nov., holotype ZSM 116 / 2023, Namoroka. Dwarf species, namely M. nanus and M. macrolepis, not represented. Scale bars = 2 mm.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
7E62E05257DD53F096F0AC92D64CC1B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556045	https://doi.org/10.3897/vz.76.e176241.figure8	Figure 8. Plate showing photographs in life of some of the recognized species of Madascincus (not to scale).	Figure 8. Plate showing photographs in life of some of the recognized species of Madascincus (not to scale).	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
7E62E05257DD53F096F0AC92D64CC1B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556046	https://doi.org/10.3897/vz.76.e176241.figure9	Figure 9. Schematic maps for Madagascar showing known distribution for Madascincus species. For reasons of readability, the different species are shown on six separate maps. Colored dots represent localities confirmed by DNA sequences, whereas white dots represent localities of specimens only identified by morphology (more details in Miralles et al. 2016). The supplementary map located at the bottom of the figure, employs minimum convex polygons to compare the distribution ranges of all species.	Figure 9. Schematic maps for Madagascar showing known distribution for Madascincus species. For reasons of readability, the different species are shown on six separate maps. Colored dots represent localities confirmed by DNA sequences, whereas white dots represent localities of specimens only identified by morphology (more details in Miralles et al. 2016). The supplementary map located at the bottom of the figure, employs minimum convex polygons to compare the distribution ranges of all species.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
7E62E05257DD53F096F0AC92D64CC1B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556047	https://doi.org/10.3897/vz.76.e176241.figure10	Figure 10. Alive picture of Madascincus irery sp. nov. A, B, C Holotype ZSM 135 / 2019, from Lokobe National Park, Nosy Be. Photographs by MDS.	Figure 10. Alive picture of Madascincus irery sp. nov. A, B, C Holotype ZSM 135 / 2019, from Lokobe National Park, Nosy Be. Photographs by MDS.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
7E62E05257DD53F096F0AC92D64CC1B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556048	https://doi.org/10.3897/vz.76.e176241.figure11	Figure 11. Preserved specimens of Madascincus irery sp. nov. A Anterior body of the paratype, ZSM 86 / 2015 in dorsal view. B dorsal view of the entire body, C lateral and dorsal view of the head, D drawing of head in lateral and dorsal views of holotype ZSM 135 / 2019 from Lokobe National Park. Abbreviations used in the drawings are explained in the Materials and Methods section.	Figure 11. Preserved specimens of Madascincus irery sp. nov. A Anterior body of the paratype, ZSM 86 / 2015 in dorsal view. B dorsal view of the entire body, C lateral and dorsal view of the head, D drawing of head in lateral and dorsal views of holotype ZSM 135 / 2019 from Lokobe National Park. Abbreviations used in the drawings are explained in the Materials and Methods section.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
7E62E05257DD53F096F0AC92D64CC1B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556049	https://doi.org/10.3897/vz.76.e176241.figure12	Figure 12. A Map indicating known localities of Madascincus irery sp. nov. on Nosy Be (red dots). B Forest at the Lokobe National Park. C Detail of the habitat where the holotype of Madascincus irery sp. nov. was collected.	Figure 12. A Map indicating known localities of Madascincus irery sp. nov. on Nosy Be (red dots). B Forest at the Lokobe National Park. C Detail of the habitat where the holotype of Madascincus irery sp. nov. was collected.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556040	https://doi.org/10.3897/vz.76.e176241.figure3	Figure 3. Photos of Madascincus minotaurus sp. nov. in life. A, B Holotype ZSM 116 / 2023, from Tsingy de Namoroka National Park. C Uncollected specimen from Grand Tsingy. Photographs A and B by AM, C by Ivan Ineich.	Figure 3. Photos of Madascincus minotaurus sp. nov. in life. A, B Holotype ZSM 116 / 2023, from Tsingy de Namoroka National Park. C Uncollected specimen from Grand Tsingy. Photographs A and B by AM, C by Ivan Ineich.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556041	https://doi.org/10.3897/vz.76.e176241.figure4	Figure 4. Preserved holotype of Madascincus minotaurus sp. nov., ZSM 116 / 2023, from Tsingy de Namoroka National Park. A, B photographs and C head drawings in lateral (above) and dorsal view (below). Abbreviations used in the drawings are explained in the Materials and Methods section.	Figure 4. Preserved holotype of Madascincus minotaurus sp. nov., ZSM 116 / 2023, from Tsingy de Namoroka National Park. A, B photographs and C head drawings in lateral (above) and dorsal view (below). Abbreviations used in the drawings are explained in the Materials and Methods section.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556042	https://doi.org/10.3897/vz.76.e176241.figure5	Figure 5. Habitat of Madascincus minotaurus sp. nov. at Tsingy de National Park. A View of the Grand Tsingy from the surrounding savanna. B View from inside a canyon at the Grand Tsingy. C Campsite at the border of Grand Tsingy, showing the leaf litter micro-habitat where the uncollected specimen (sample MIRZC 1217) was found. D Petit Tsingy, exact position of the type locality. E Diagram representing a transverse view of Grand Tsingy during the dry season (during the wet season, the depressed ground is heavily flooded). Specimens of M. minotaurus sp. nov. have been encountered in the leaf-litter located in the border of the karts system (corresponding to the orange surface on the left side of the diagram), although the species is likely present in the corridors too.	Figure 5. Habitat of Madascincus minotaurus sp. nov. at Tsingy de National Park. A View of the Grand Tsingy from the surrounding savanna. B View from inside a canyon at the Grand Tsingy. C Campsite at the border of Grand Tsingy, showing the leaf litter micro-habitat where the uncollected specimen (sample MIRZC 1217) was found. D Petit Tsingy, exact position of the type locality. E Diagram representing a transverse view of Grand Tsingy during the dry season (during the wet season, the depressed ground is heavily flooded). Specimens of M. minotaurus sp. nov. have been encountered in the leaf-litter located in the border of the karts system (corresponding to the orange surface on the left side of the diagram), although the species is likely present in the corridors too.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556043	https://doi.org/10.3897/vz.76.e176241.figure6	Figure 6. Satellite views of the Tsingy de Namoroka. A Namoroka National Park and surrounding areas. B Details of the main karstic area. C, D Details of areas are here interpreted as vestigial karstic fragments located outside of the national park. Karstic fragments (i. e., showing the typical network of canyons / fissures arranged perpendicular to each other and filled with vegetation) are delimited in white, while the perimeter of the national park is delimited in green. Turquoise dots correspond to the known localities of M. minotaurus sp. nov. Satellite imagery from Airbus, via Google Earth.	Figure 6. Satellite views of the Tsingy de Namoroka. A Namoroka National Park and surrounding areas. B Details of the main karstic area. C, D Details of areas are here interpreted as vestigial karstic fragments located outside of the national park. Karstic fragments (i. e., showing the typical network of canyons / fissures arranged perpendicular to each other and filled with vegetation) are delimited in white, while the perimeter of the national park is delimited in green. Turquoise dots correspond to the known localities of M. minotaurus sp. nov. Satellite imagery from Airbus, via Google Earth.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556044	https://doi.org/10.3897/vz.76.e176241.figure7	Figure 7. Drawings of the heads in dorsal and lateral views of most of the species of Madascincus. A M. arenicola, holotype ZSM 1565 / 2008, Baie des Sakalava. B M. miafina, holotype ZSM 1562 / 2008, Ankarana Special Reserve. C M. stumpffi, holotype SMF 16019, “ Nossibé ” (= Nosy Be). D M. polleni, holotype MNHN 1895.210, “ Mouroundava ” (= Morondava). E M. irery sp. nov., holotype ZSM 135 / 2019, Nosy Be. F Madascincus pyrurus, holotype ZSM 520 / 2001, Mont Ibity. G M. igneocaudatus, ZSM 1600 / 2010, Anakao. H M. mouroundavae, ZSM 13 / 2005, Andasibe. I M. ankodabensis, ZSM 355 / 2006, Ranomafana. J M. melanopleura, ZSM 20 / 2005, Andasibe. K M. minutus, ZSM 400 / 2005, Nosy Mangabe. L M. minotaurus sp. nov., holotype ZSM 116 / 2023, Namoroka. Dwarf species, namely M. nanus and M. macrolepis, not represented. Scale bars = 2 mm.	Figure 7. Drawings of the heads in dorsal and lateral views of most of the species of Madascincus. A M. arenicola, holotype ZSM 1565 / 2008, Baie des Sakalava. B M. miafina, holotype ZSM 1562 / 2008, Ankarana Special Reserve. C M. stumpffi, holotype SMF 16019, “ Nossibé ” (= Nosy Be). D M. polleni, holotype MNHN 1895.210, “ Mouroundava ” (= Morondava). E M. irery sp. nov., holotype ZSM 135 / 2019, Nosy Be. F Madascincus pyrurus, holotype ZSM 520 / 2001, Mont Ibity. G M. igneocaudatus, ZSM 1600 / 2010, Anakao. H M. mouroundavae, ZSM 13 / 2005, Andasibe. I M. ankodabensis, ZSM 355 / 2006, Ranomafana. J M. melanopleura, ZSM 20 / 2005, Andasibe. K M. minutus, ZSM 400 / 2005, Nosy Mangabe. L M. minotaurus sp. nov., holotype ZSM 116 / 2023, Namoroka. Dwarf species, namely M. nanus and M. macrolepis, not represented. Scale bars = 2 mm.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556045	https://doi.org/10.3897/vz.76.e176241.figure8	Figure 8. Plate showing photographs in life of some of the recognized species of Madascincus (not to scale).	Figure 8. Plate showing photographs in life of some of the recognized species of Madascincus (not to scale).	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
46AE698614A25638AA4F42F54C767A45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1556046	https://doi.org/10.3897/vz.76.e176241.figure9	Figure 9. Schematic maps for Madagascar showing known distribution for Madascincus species. For reasons of readability, the different species are shown on six separate maps. Colored dots represent localities confirmed by DNA sequences, whereas white dots represent localities of specimens only identified by morphology (more details in Miralles et al. 2016). The supplementary map located at the bottom of the figure, employs minimum convex polygons to compare the distribution ranges of all species.	Figure 9. Schematic maps for Madagascar showing known distribution for Madascincus species. For reasons of readability, the different species are shown on six separate maps. Colored dots represent localities confirmed by DNA sequences, whereas white dots represent localities of specimens only identified by morphology (more details in Miralles et al. 2016). The supplementary map located at the bottom of the figure, employs minimum convex polygons to compare the distribution ranges of all species.	2026-03-06	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Miralles, Aurélien;Scherz, Mark D.;Hyde Roberts, Sam;Rakotoarison, Andolalao;Glaw, Frank;Vences, Miguel			
