taxonID	type	description	language	source
AB08878A2A34FFB4FF41257D47789859.taxon	materials_examined	Type species Paludina curta Küster, 1853.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A34FFBBFF4125D744B898B4.taxon	description	GenBank sequence numbers: PX 113150 – PX 113167 mOTU D Type locality CROATIA • Šibenicko - Kninska županja, Cviljane, Kotluša spring (locality 4: Fig. 1); 43 ° 56 ’ 57 ” N; 16 ° 23 ’ 56 ” E. 397 a. s. l.; the westernmost major source of the river Cetina, on stones in the spring zone.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A34FFBBFF4125D744B898B4.taxon	materials_examined	Holotype CROATIA • ethanol-fixed specimen, Šibenicko-Kninska županja, Cviljane, Kotluša spring (locality 4: Fig. 1); 43 ° 56 ’ 57 ” N; 16 ° 23 ’ 56 ” E. 397 a. s. l.; 17.03.2017; Jozef Grego leg. NHMW-MO- 113902. (Fig. 4 A). Paratypes CROATIA • 58 dry and 38 ethanol-fixed specimens, same data for holotype, 11 ethanol-fixed specimens; 8 dry and 7 ethanol fixed specimens, GMV 112630, 50 dry and 20 ethanol-fixed specimens, JG F 2583. Description Shell (Figs 4 A – D, 5 A – C and 6 A – F) up to 1.70 (type locality) or 2.26 (locality 8) mm high, ovate-conical, broad or moderately broad, with spire relatively or moderately low, with 4 – 5 whorls, spire height 24 – 26 % (type locality) or 29 – 30 % (locality 8) of the shell height. Teleoconch whorls slightly convex, evenly rounded, growing regularly in diameter, but penultimate whorl disproportionally high in some populations (like at locality 8). Aperture moderately narrow and ovoid, outer lip simple, parietal lip complete, either narrow or broad, umbilicus slit-like. Teleoconch moderately thick-walled, its wall translucent, with delicate growth lines, periostracum yellowish. Shell measurements: Table 3, Fig. 15. Protoconch (Figs 4 E and 5 D) growing abruptly, no sharp border between proto- and teleoconch, its micro sculpture with regular net of depressions. Operculum yellowish, smooth on both surfaces. Radula (Fig. 7 A – B) with central tooth with blunt (Fig. 7 A) or sharp (Fig. 7 B) and massive median cusp and short (less than half of the median cusp length) moderately sharp cusps following formula: 7 – 1 – 7 or (8) 7 – 1 – 7 (8) or 8 – 1 – 8 1 – 1 1 – 1 1 – 1 Lateral tooth with 4 – 1 – 4 moderately sharp cusps, the largest one broad and rounded. Inner marginal tooth with 25 – 27 long and sharp cusps, outer marginal tooth with up to 27 relatively long and sharp cusps on the terminal edge.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A34FFBBFF4125D744B898B4.taxon	diagnosis	Differential diagnosis From probably the most closely related species (not well supported relationship) R. frauenfeldiana sp. nov., it differs in having a bigger distal receptaculum and single outgrowth on the penis. The Molecular Diagnostic Characters in comparison with other new Radomaniola species: binary: 60 (C), 69 (C), 159 (A), 166 (C), 204 (A), 219 (C), 252 (A), 277 (C), 297 (C), 409 (C), 423 (C), asymmetric: 57 (A). In comparison with R. frauenfeldiana: binary: 57 (A), 69 (C), 156 (A), 166 (C), 219 (C), 249 (G), 252 (A), 303 (T), 366 (A), 409 (C), 423 (C), 429 (C) (Supplementary File 2). Etymology Named after Cetina river. The source of the Cetina River is the type locality. Distribution and habitat Found at ten localities: 3 – 12 (Figs 1 and 2 A – B, Table 1): eight in central Croatia in the upper Cetina Basin and two in NW Bosnia and Herzegovina, south Livansko Polje. These are either large or small springs, rivers (Cetina), as well as interstitial habitats. At localities 3 and 10 it occurs in sympatry with R. szarowskae Delicado et Hauffe, 2022, at locality 5 in sympatry with R. curta curta (Küster, 1895), and at locality 11 in sympatry with R. torifera (Schütt, 1961). FIGURE 6. Shell of Radomaniola: A – F — R. cetinensis sp. nov., from locality: A — 3 (2 K 80), B — 5 (1 M 3), C — 6 (1 M 6), D — 7 (1 M 4), E — 10 (2 K 60), F — 11 (2 K 70); G – I — R. testavariabilis sp. nov., from locality: G — 15 (2 L 40), H — 19 (2 K 73), I — 20 (2 K 43); colours of mOTUs as in Figs 1 and 3; bar equals 1 mm. Sequences are specified by extraction numbers. Ecological status The number of known locations is more than eight and they are distributed over 50 km long and 2 – 8 km broad EOO at the Upper Cetina Basin in Croatia; 22 km long and 12 km broad EOO at the lower part of Livansko Polje in Bosnia and Herzegovina and one isolated locality south of Split. The AOO is represented by scattered small local areas around several springs and adjacent small rivulets, which in total represent a much smaller total area of 15 km 2 compared with EOO. All localities are dependent upon the supply of spring water and are threatened by climatic changes resulting in droughts and disappearing or declining of the spring water outlet. Additionally, the localities are endangered by anthropogenic pollution by municipal or chemical waste, as well as by anthropogenic alterations of the springs by construction of small power stations, fountains, picnic zones, concrete-bordered channels and tap water catchments with small or no outlets of the remaining water. Therefore, it is assessed as Vulnerable (VD 2).	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A3BFFBCFF4125BD47489FBF.taxon	description	GenBank sequence numbers: PX 113128 – PX 113141 mOTU A Type locality BOSNIA and HERZEGOVINA • Bosnian Federation, Western Herzegovina Canton, Ljubuški district, Vitina, Vrelo Vrioštica spring, (locality 17: Figs 1 and 2 F); 43 ° 14 ’ 15 ” N; 17 ° 29 ’ 09 ” E; 93 m a. s. l.; on stones in spring and riverbed.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A3BFFBCFF4125BD47489FBF.taxon	materials_examined	Holotype BOSNIA and HERZEGOVINA • Ethanol-fixed specimen (Fig. 8 A); Vrelo Vrioštica spring, Vitina, Ljubuški district, West Herzegovina Canton, Bosnia and Herzegovina, (locality 17: Figs 1 and 2 F); 43 ° 14 ’ 15 ” N; 17 ° 29 ’ 09 ” E; 93 m a. s. l.; 02.04.2018; leg. Jozef Grego, NHMW-MO- 113904. Paratypes BOSNIA and HERZEGOVINA • 148 ethanol-fixed and 62 dry and specimens from type locality, collected with the holotype: 15 ethanol-fixed specimens, ZMUJ / 2024 / 02; 10 ethanol-fixed specimens NHMW- MO- 113905; 123 ethanol fixed and 62 dry specimens, JG F 1219. BOSNIA and HERZEGOVINA • 44 ethanol-fixed and 26 dry, from type locality, 22.04.2019, Jozef Grego leg., coll. JG F 2583; BOSNIA and HERZEGOVINA • 32 ethanol-fixed specimens, from type locality, 03.09.2022, Jozef Grego and Andrzej Falniowski legs., coll. JG F 3735. Description Shell (Figs 6 G – I, 8 A – E, 9 A – B and 10 A – B) up to 2.44 mm high, broadly ovate-conical, nearly trochiform (type locality 11: Fig. 8 A – E and locality 10: Fig. 9 A – B), with spire low, with about three whorls, spire height 11 – 17 % of the shell height. Teleoconch whorls flat, growing rapidly in diameter, penultimate proportionally low. At localities 9, 12, 13 and 14 (Figs 6 G – I and 10 A – B), sharing haplotypes with the other localities, the shell turriform, with about five whorls, spire height 33 % of the shell height, sutures deeper and whorls more convex and growing slower than those at the other localities. Aperture broad, ovoid, outer lip simple, parietal lip complete and broad (except at localities 9, 12, 13 and 14), umbilicus absent or slit-like. Teleoconch moderately thick-walled, its wall slightly translucent, with delicate growth lines, periostracum whitish or yellowish. Shell measurements: Table 3, Fig. 15. Operculum yellowish or brownish, smooth on both surfaces.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A3BFFBCFF4125BD47489FBF.taxon	diagnosis	Differential diagnosis From the most closely related species R. torifera it differs in having somewhat larger double outgrowth on the penis, and from R. mislinensis sp. nov. it differs in having a double instead of single outgrowth. R. testavariabilis sp. nov. differs from R. torifera especially in its female reproductive organs, with ellipsoid, not spherical bursa copulatrix, thicker loop of oviduct, proximal seminal receptacle (rs 2) smaller and distal receptaculum (rs 1) being shorter and broader. The Molecular Diagnostic Characters in comparison with other new Radomaniola species: binary: 183 (A), 357 (A), 429 (A). In comparison with R. mislinensis and R. torifera: binary: 183 (A), 357 (A), 429 (A). Etymology The name reflects variability of the shell, especially of the spire height. Distribution and habitat Found at six localities: 15 – 20 (Figs 1 and 2 D, F, G, Table 1): three in Croatia and three in Bosnia and Herzegovina, five of them in the southern parts of both countries. In springs, rivers (Neretva) and interstitial waters. At locality 14 in sympatry with R. torifera. Ecological status The number of known locations is more than five, and they are distributed over 75 km long and 5 – 20 km broad EOO along the Tihaljina River Basin, Middle Neretva Valley and northern Popovo Polje in Bosnia and Herzegovina, with one elusive locality in Obrovac Sinjski in Croatia. The AOO is represented by scattered local small areas around several springs and adjacent small rivulets, which in total represent a much smaller total area compared to EOO. Each of the localities is dependent upon a supply of spring water and is threatened by climatic changes through droughts and disappearing or declining of the spring water outlet over longer periods. Additionally, the localities are endangered by anthropogenic pollution by municipal or chemical waste, as well as by anthropogenic alterations of the springs by construction of objects as small power stations, houses, fountains, picnic zones, concrete-bordered channels and tap water catchments with small or no outlets of the remaining water. Therefore, it is assessed as Vulnerable (V).	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A3CFFBEFF4126AA41299963.taxon	description	GenBank sequence numbers: PX 113142 – PX 113146 mOTU B Type locality CROATIA • Splitsko-Dalmatinska županja, Metković distr., Neretva River Delta, Mislina, Mislina spring (locality 14: Figs 1 and 2 E, H); 42 ° 58 ’ 56 ” N; 17 ° 36 ’ 24 ” E; 4 m a. s. l.; at the bottom of spring head at the bank of spring lake under the road.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A3CFFBEFF4126AA41299963.taxon	materials_examined	Holotype CROATIA • ethanol-fixed specimen (Fig. 11 A), Mislina spring, Mislina, delta of Neretva River, District Metković, Splitsko-Dalmatinska županja, Croatia (locality 14: Figs 1 and 2 E, H); 42 ° 58 ’ 56 ” N; 17 ° 36 ’ 24 ” E; 04.08.2020; Andrzej Falniowski, Roman Ozimec and Jozef Grego legs.; NHMW-MO- 113906. Paratypes CROATIA • 96 dry and 78 ethanol-fixed specimens; type locality; data as for holotype; 39 ethanol-fixed specimens in ZMUJ / 2024 / 03; 5 dry specimens NHMW-MO- 113907; 91 dry and 46 ethanol-fixed JG F 2583; CROATIA • 38 dry and 72 ethanol-fixed specimens, from type locality, 27.07.2024, Roman Ozimec and Jozef Grego legs., 10 dry and 10 ethanol-fixed specimens in GMV 112631, 28 dry and 55 ethanol-fixes specimens in JG F 3769. Description Shell (Fig. 11 A – D) up to 2.27 mm high, ovate-conical, broad, with spire low or moderately low, with about five whorls, spire height 20 – 24 % of the shell height. Teleoconch whorls slightly convex, evenly rounded, growing regularly and slowly in diameter, penultimate whorl disproportionally low in some specimens. Aperture moderately narrow and ovoid, outer lip simple, parietal lip complete or not, moderately broad, umbilicus slit-like. Teleoconch moderately thick-walled, its wall translucent, growth lines delicate or more visible, periostracum whitish or yellowish. Shell measurements: Table 3, Fig. 15. Protoconch (Fig. 11 E) growing slowly, no sharp border between proto- and teleoconch, its micro sculpture delicate and irregular. Operculum yellowish-brownish, smooth on both surfaces. Soft parts morphology and anatomy Head, mantle and visceral sac intensely pigmented black. Female reproductive organs unknown. Penis (Fig. 11 F – G) long, narrow and bent, proximally broad and ellipsoid in shape, distally sharp, on the left side of the penis is a single narrow and sharp outgrowth, vas deferens visible inside the penis. Differential diagnosis From the most closely related species — R. torifera and R. testavariabilis sp. nov. — it differs in its penis having a single, finger-shaped sharply pointed outgrowth. The Molecular Diagnostic Characters in comparison with all Radomaniola sequences used in phylogenetic analysis: binary 342 (G). In comparison with other new Radomaniola species: binary: 96 (C), 342 (G), 360 (T), 372 (G), asymmetric: 390 (C). In comparison with R. testavariabilis sp. nov. and R. torifera: binary: 96 (C), 342 (G), 360 (T), 372 (G), 390 (C). Etymology The name reflects Mislina spring, its type locality. Distribution and habitat Apart from type locality, found also at spring Studena, locality 13 (Fig. 1, Table 1). Ecological status The AOO and EOO are represented by a spring supplying a small lake Mislina at N edge of swamps adjacent to Kuti Lake and by adjacent parts of the small lake, in area about 50 m 2. Locality in Neretva Delta is dependent upon a supply of fresh spring water and is threatened by climatic changes and declining of the spring water outlet causing eutrophication. Additionally, locality is endangered by anthropogenic pollution by municipal or chemical waste from local settlement and adjacent road, as well as by anthropogenic alternations of the springs by construction activities. Locality in Studena is a small fountain close to a road and heavily affected by road construction and excavation work. Therefore, it is assessed as Endangered (En).	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A21FFA0FF4123F946999A27.taxon	description	GenBank sequence numbers: KC 011744 mOTU F Type locality GREECE • North Peloponnísos Peninsula, Achaia, Province Kalavryta, spring at Bouboukas, near road to Maneli (locality 24: Figs 1 and 2 C); 38 ° 00 ’ 53 ” N; 21 ° 58 ’ 31 ” E; 703 m a. s. l.; on stones and leaves in the spring head.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A21FFA0FF4123F946999A27.taxon	materials_examined	Holotype GREECE • ethanol-fixed specimen (Fig. 12 A); spring at Bouboukas, near road to Maneli, North Peloponnísos Peninsula, Achaia, Province Kalavryta, Greece (locality 24: Figs 1 and 2 C); 38 ° 00 ’ 53 ” N; 21 ° 58 ’ 31 ” E; 703 m a. s. l.; 12.09.2007; Magdalena Szarowska and Andrzej Falniowski legs.; NHMW-MO- 113908. Paratypes GREECE • 18 ethanol-fixed specimens from type locality; 16 ethanol-fixed specimens ZMUJ / 2024 / 04; 2 ethanol-fixed specimens NHMW-MO- 113909. Description Shell (Fig. 12 A – D) up to 2.16 mm high, ovate-conical, broad, spire moderately low, about 5 ½ whorls, spire height 26 – 29 of the shell height. Teleoconch whorls slightly convex, evenly rounded, growing regularly and slowly in diameter. Aperture moderately broad and ovoid, outer lip simple, parietal lip complete, moderately broad or broad, umbilicus in the form of long slit. Teleoconch moderately thick-walled, its wall translucent, growth lines delicate. Periostracum yellowish. Shell measurements: Table 3, Fig. 15. Operculum yellowish, smooth on both surfaces.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A21FFA0FF4123F946999A27.taxon	diagnosis	Differential diagnosis From the most closely related species R. filiola it differs in having a larger, longer and smooth penis with either absent or vestigial outgrowth. The Molecular Diagnostic Characters in comparison with other new Radomaniola species: binary: 39 (A), 63 (T), 75 (T), 76 (A), 138 (G), 147 (C), 171 (C), 213 (A), 235 (T), 411 (G), 426 (T), asymmetric: 57 (G), 249 (C), 390 (G). In comparison with R. filiola: binary: 39 (G), 93 (G), 138 (G), 235 (T). Etymology The name reflects slender and long penis with vestigial outgrowth. Distribution and habitat Known so far only from its type locality. Ecological status The species is known from a single locality at Bouboukas at Northwestern Peloponnese Peninsula. The AOO and EOO are represented by a very small 5 m 2 spring zone already altered by the construction of the fountain. Locality is strongly dependent upon a small supply of water and is threatened by climatic changes and drought. Additionally, locality is endangered by anthropogenic pollution by municipal or chemical waste (detergents) from visitors of the fountain, as well as by further anthropogenic alterations of the fountain by construction activities or tap water catchments. Therefore, it is assessed as Endangered (En).	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A20FFA3FF41273447489C0B.taxon	description	GenBank sequence numbers: MZ 539646, MZ 539647, PX 113147 – PX 113148 mOTU C Type locality CROATIA • Splitsko-Dalamatinska županja, Kostanje-Kučiće, spring Vrutak, a small spring with cisterna at left side of road to Kostanje (locality 1); 43 ° 26 ′ 28 ″ N, 16 ° 49 ′ 21 ″ E.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A20FFA3FF41273447489C0B.taxon	materials_examined	Holotype CROATIA • ethanol-fixed specimen (Fig. 13 A), Splitsko-Dalamatinska županja, Kostanje-Kučiće, Croatia, 43 ° 26 ′ 28 ″ N, 16 ° 49 ′ 21 ″ E, 31.08.2018, leg. Luboš Beran, ZMUJ 2024 / 04. Paratypes CROATIA • 14 ethanol-fixed specimens from type locality, same data as for holotype, two ethanol-fixed specimens ZMUJ 2024 / 05, 12 ethanol-fixed specimens in coll. LB; CROATIA • 16 dry and 36 ethanol-fixed specimens, from type locality, 26.07.2024, Jozef Grego and Roman Ozimec legs., 5 dry specimens in GMV 112631, 11 dry and 36 ethanol-fixed specimens in JG F 3755. Description Shell (Fig. 13 A – D) up to 2.23 mm high, ovate-conical, broad, with spire moderately low, with about 5 ½ whorls, spire height 28 % 26 – 29 of the shell height. Teleoconch whorls slightly convex, evenly rounded, growing regularly and slowly in diameter. Aperture broad and nearly circular or less broad and ovoid, outer lip simple, parietal lip complete, broad in completely grown specimens (Fig. 13 A and D), umbilicus in the form of broad and long slit (Fig. 13 A) or incomplete slit, with parietal lip adjacent to body whorl (Fig. 13 D). Teleoconch moderately thick-walled, its wall translucent, growth lines delicate but well visible, periostracum yellowish. Shell measurements: Table 3, Fig. 15. Operculum yellowish, smooth on both surfaces.	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A20FFA3FF41273447489C0B.taxon	diagnosis	Differential diagnosis From the most closely related species R. cetinensis it differs in smaller distal seminal receptacle and presence of the second, vestigial outgrowth on the penis. The Molecular Diagnostic Characters in comparison with R. cetinensis: binary: 57 (A), 69 (C), 166 (C), 219 (C), 249 (G), 252 (A), 366 (A), 409 (C), 423 (C), 429 (C). Etymology Named in honour of Georg von Frauenfeld, famous Austrian malacologist, devoted also to the study of the Balkan Truncatelloidea (see Eschner 2008). Distribution and habitat The species is known from four springs at lower Cetina Canyon. Ecological status The AOO and EOO are represented by four springs in lower Cetina Canyon, some of them already altered by tap water reservoirs. Further construction activities of tap water supply and road construction are the main treats for the populations, together with increasing rafting tourism in the region and related infrastructure construction. The anthropogenic pollution of the springs is also related to the tourist and camping activities. Therefore, it is assessed as Vulnerable (V).	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
AB08878A2A24FFA7FF41219441EE9F77.taxon	description	Shell (Fig. 14 A – C) up to 2.28 mm high, ovate-conical, with moderately low spire, with about five whorls, spire height 23 – 28 % of the shell height. Teleoconch whorls moderately convex, evenly rounded, growing regularly in diameter. Aperture ovoid, outer lip simple, parietal lip broad and complete, umbilicus slit-like. Teleoconch moderately thick-walled, its wall translucent, with delicate growth lines, periostracum yellowish. Protoconch (Fig. 14 D) initially broad, growing abruptly, no sharp border between proto- and teleoconch, no distinct microsculpture. Operculum brownish, smooth on both surfaces. Soft parts morphology and anatomy Head and mantle intensely pigmented black. Female reproductive organs as described by Delicado & Hauffe (2022). Penis (Fig. 14 E – F) long and narrow, ribbon-shaped, sharply terminated. On the left side of the medial part of the penis there is double outgrowth, consisted of two delicate extensions, vas deferens well visible inside the penis. Distribution and habitat Small springs in Greece, found at localities 17 – 19 (Table 1, Fig. 1).	en	Jaszczyńska, Aleksandra, Grego, Jozef, Beran, Luboš, Hofman, Sebastian, Falniowski, Andrzej, Osikowski, Artur (2025): Five new species of Radomaniola Szarowska, 2006 (Truncatelloidea: Hydrobiidae). Zootaxa 5716 (2): 207-233, DOI: 10.11646/zootaxa.5716.2.2, URL: https://doi.org/10.11646/zootaxa.5716.2.2
