identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9FB85940DD0853AFAB1DD0DD3B2B48C7.text	9FB85940DD0853AFAB1DD0DD3B2B48C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheniella parvispora S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>1. Atheniella parvispora S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 2, 3</p><p>Etymology.</p><p>The specific epithet “parvispora” refers to small size of basidiospores.</p><p>Diagnosis.</p><p>The species is characterized by papillate, umbilicate, orangish yellow pileus with off white margins, lacrymoid, ovoid to subcylindrical, small basidiospores (6.71 × 3.70 µm) and abundant fusiform cheilocystidia.</p><p>Holotype.</p><p>CHINA, Gansu Province, Gannan Tibetan Autonomous Prefecture, Lintan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.5&amp;materialsCitation.latitude=34.95" title="Search Plazi for locations around (long 103.5/lat 34.95)">Lianhuashan National Forest Park</a>, 34°57'N, 103°30'E, on broad-leaved forests land, 2 September 2024, FLF 1713 (GenBank ITS = PX 612304; tef 1 - α = PX 684307)</p><p>Description.</p><p>Basidiomata small-sized. Pileus 1.0– 1.3 cm in diameter, orangish yellow (2.5 R 8 / 4), margins off white (10 R 8 / 1) in circular ring, papillate, umbilicate, dry, smooth at center, fibrillose towards margins, plicate to split, margin decurved to incurved. Context thin, white. Lamellae adnate with decurrent tooth, chalky white (10 R 8 / 1), narrow, subdistant, thin, unequal, even. Lamellulae, in two tiers, alternating with lamellae. Stipe ca. 5.3 cm, chalky white (10 R 8 / 1) to smoky white (2.5 YR 8 / 1), central, equal to slightly flexuous at base, surface scurfy to canescent. Hyphal system composed of generative clamped hyphae, IKI –, inflated in KOH, thin-walled, occasionally branched, subregular to irregularly arranged, 2.74–12.24 µm in diameter.</p><p>Basidiospores 5.32–8.19 × 2.91–4.91 µm, av. L × av. W = 6.71 × 3.70 µm, Q = 1.75–1.81, lacrymoid, ovoid to subcylindrical with an apiculus and oil drop, smooth, hyaline in 5 % KOH, IKI –, guttulated, thin-walled. Probasidia cylindrical to clavate, thin-walled. Basidia 22.56–26.53 × 5.58–7.32 µm, av. L × av. W = 24.82 × 6.34 µm, Q = 3.91, clavate to narrowly clavate, guttulated, hyaline, thin-walled, bi to tetra-sterigmated. Cheilocystidia 46.44–55.01 × 8.81–12.30 µm, av. L × av. W = 51.45 × 9.88 µm, abundant, fusiform, ventricose-rostrate, with an obtuse apex, short- to long-stalked, smooth, hyaline in 5 % KOH, thin – walled. Pleurocystidia not observed. Pileipellis a cutis of inflated, clavate, hyphae, hyphae 10.53–20.11 µm in diameter, thin-walled, with branched excrescences.</p><p>Additional specimen (isotype) examined.</p><p>CHINA, Gansu Province, Gannan Tibetan Autonomous Prefecture, Lintan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.5&amp;materialsCitation.latitude=34.95" title="Search Plazi for locations around (long 103.5/lat 34.95)">Lianhuashan National Forest Park</a>, 34°57'N, 103°30'E on broad-leaved forests land, 2 September 2024, MSX 1713 (GenBank ITS = PX 612305; tef 1 - α = PX 684308) .</p><p>Notes.</p><p>The presence of a brightly colored pileus, an inamyloid hymenophoral trama, and pileipellis covered with simple to branched excrescences suggests that this species belongs to the genus Atheniella . It is characterized by small basidiomata, a pileus with white margins to orangish yellow, thin and white context, a chalky white to smoky white hollow stipe with a scurfy to canescent surface, thin-walled and clamped hyphae inflated in KOH, lacrymoid, ovoid to subcylindrical, small basidiospores (6.71 × 3.70 μm) with an apiculus and oil drop, and abundant fusiform cheilocystidia.</p><p>In the combined ITS + tef 1 - α phylogram, Atheniella parvispora is separated from M. floridula (Fr.) Quél. and A. flavoalba (Fr.) Redhead, Moncalvo, Vilgalys, Desjardin &amp; B. A. Perry. The latter two species were formerly classified in Mycena sect. Adonideae . Redhead (2012) proposed that M. floridula was a synonym of A. adonis (Bull.) Redhead, Moncalvo, Vilgalys, Desjardin &amp; B. A. Perry; however, more recently, M. floridula sensu Kühner has been considered a pink-capped form of M. flavoalba (Kühner and Maire 1938; Aronsen and Læssøe 2016).</p><p>In the phylogeny, the new species forms a sister group to Atheniella adonis (previously named M. floridula). Although morphologically similar in terms of basidiospores, A. parvispora can be differentiated from A. adonis by its papillate, umbilicate, and plicate pileus with a circular ring at the margin, as well as its scurfy to canescent stipe. Anatomically, A. adonis possesses pseudofusiform or lageniform pleurocystidia with a distended venter. Furthermore, A. parvispora has smaller basidiospores (6.71 × 3.70 μm) than the larger ones of A. adonis (7–9.5 × 3.5–5 μm). The close relative is A. flavoalba, which is the most widely distributed species in the Northern Hemisphere and can be easily differentiated by its narrowly ellipsoid basidiospores and larger caulocystidia (up to 60 μm) (Aronsen and Læssøe 2016; Na and Bau 2019).</p></div>	https://treatment.plazi.org/id/9FB85940DD0853AFAB1DD0DD3B2B48C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
95C0829A2CEA542DBE4CA8AA8936E3C8.text	95C0829A2CEA542DBE4CA8AA8936E3C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena albomarginata S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>2. Mycena albomarginata S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 4, 5</p><p>Etymology.</p><p>The specific epithet “albomarginata” refers to the white margins of pileus.</p><p>Diagnosis.</p><p>The species is characterized by virgate to sulcate, brownish gray pileus with white margins, brownish gray stipe with pruinose apex, free to adnexed lamellae, narrowly clavate cheilocystidia with slightly long base, larger (9.08 × 5.51 µm), ellipsoid basidiospores, and bi-sterigmated basidia.</p><p>Holotype.</p><p>CHINA, Gansu Province, Tianzhu Tibetan Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.26667&amp;materialsCitation.latitude=37.466667" title="Search Plazi for locations around (long 103.26667/lat 37.466667)">Xidatan Town</a>, 37°28'N, 103°16'E, in mixed coniferous, broadleaf forest, 19 th September 2024, FLF 1971 (GenBank ITS = PX 612302; tef 1 - α = PX 684305) .</p><p>Description.</p><p>Basidiomata small-sized. Pileus 1.9–2.2 cm in diameter, pigeon gray (N 9 / 0) to gray (7.5 Y 5 / 1), brown, margins white (6.9 GY 7 / 1), in circular ring, conical to parabolic, dry, smooth in center, glabrous, virgate to sulcate, margins smooth, even, entire, regular, slightly recurved to straight. Context thin, gray. Lamellae regular, free to adnexed, fimbriate, gray (7.5 Y 6 / 1), broad, subdistant, thick, wavy. Lamellulae of two different size, in two tiers, alternating with lamellae. Stipe approximately 4.5 cm long, brownish gray (10 Y 6 / 1) with white pruinose apex, upper 3 rd part of stipe is white, central, cylindrical to flexuous, only apex floccose, smooth towards base, base white and slightly bulbous. Hyphal system composed of hyphae without clamp connections, dextronoid, inflated in KOH, thin-walled, branched, septate, regular to subregular arrangement, 4.84–8.29 µm in diameter.</p><p>Basidiospores 7.56–10.43 × 5.5–6.90 µm, av. L × av. W = 9.08 × 5.51 µm, Q = 1.54–1.66, broadly ellipsoid with apiculus and oil drop, smooth, hyaline in 5 % KOH, amiloid, thin-walled. Probasidia cylindrical to clavate, thin-walled. Basidia 24.49–37.44 × 6.18–8.67 µm, av. L × av. W = 32.30 × 7.39 µm, Q = 4.25–4.36, clavate, guttulated, hyaline, thin-walled, two-spored. Cheilocystidia 27.84–33.63 × 5.28–6.67 µm, av. L × av. W = 31.84 × 5.99 µm, abundant, narrowly clavate with slightly long base, smooth, hyaline in 5 % KOH, thin-walled, in groups. Pileipellis a cutis; hyphae 11.4–20.1 µm in diam., inflated, thin-walled, predominantly smooth but with scattered excrescences.</p><p>Additional specimens examined.</p><p>CHINA, Gansu Province, Tianzhu Tibetan Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.96667&amp;materialsCitation.latitude=37.416668" title="Search Plazi for locations around (long 102.96667/lat 37.416668)">Xidatan Town</a>, 37°25'N, 102°58'E, in mixed coniferous-broadleaf forests, 29 September 2024, MSX 1971 (GenBank ITS = PX 612303; tef 1 - α = PX 684306) .</p><p>Notes.</p><p>Mycena albomarginata belongs to Mycena sect. Mycena . The species is characterized by a virgate to sulcate, brownish-gray pileus with white margins, a brownish-gray stipe with a pruinose apex, free to adnexed gray lamellae, narrowly clavate cheilocystidia with a slightly elongated base, larger, ellipsoid basidiospores (9.08 × 5.51 μm), and 2 - st erigmated basidia. In the phylogenetic analysis (Fig. 1), M. albomarginata forms a sister group with M. semivestipes (Peck) A. H. Sm., sharing similar pileus and stipe morphology as well as spore shape. However, M. albomarginata is distinguished by a smaller (2.2 cm), brownish-gray pileus with white margins and gray lamellae, whereas M. semivestipes possesses a dark brown to nearly black pileus with darker margins and white to pink lamellae. Anatomically, M. albomarginata has larger basidiospores (9.08 × 5.51 μm) and larger, 2 - spored basidia (24.5–37.4 × 6.2–8.7 μm), whereas M. semivestipes has smaller, 4 - spored basidia (21–30 × 4.5–5.4 μm) (Bessette et al. 1995).</p><p>Another close relative is M. flos-nivium Kühner, which can be easily distinguished in the field by the presence of a stipe base densely and conspicuously covered with long, coarse, flexuous, whitish fibrils. Both species share similarities in pileus morphology and coloration, exhibiting very dark sepia brown to dark brown tones that become paler toward the margins. Mycena albomarginata has larger basidiospores (9.08 × 5.51 μm) and smaller, smooth, narrowly clavate cheilocystidia (27.8–33.6 × 5.3–6.7 μm) with a slightly elongated base. In contrast, M. flos-nivium possesses smaller basidiospores (8–11.5 × 4–5 μm) and larger cheilocystidia (20–80 × 6–12 μm) that are clavate, with some very long-stalked and covered with fairly few, unevenly spaced, usually rather coarse, simple to branched, straight to curved excrescences (Na and Bau 2020).</p></div>	https://treatment.plazi.org/id/95C0829A2CEA542DBE4CA8AA8936E3C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
8D4BAEF2AB615AEDA66A9F6F69369DC5.text	8D4BAEF2AB615AEDA66A9F6F69369DC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena lavendula S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>4. Mycena lavendula S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 8, 9</p><p>Etymology.</p><p>The specific epithet “lavendula” refers to the lilac color of basidiomata.</p><p>Diagnosis.</p><p>Mycena lavendula is characterized by a subcylindrical to ellipsoid basidiospores (6.68 × 2.56 µm), polymorphic cheilocystidia with rounded apex and generative hyphae with clamp connections.</p><p>Holotype.</p><p>CHINA, Gansu Province, Tibetan Autonomous of Gannan, Taohe National Nature Reserve, 34°40'70"N, 103°53'26"E, in mixed coniferous and broadleaf forests, 3 rd September 2024 FLF 1785 (GenBank ITS = PX 612289; tef 1 - α = PX 684295)</p><p>Description.</p><p>Basidiomata medium-sized. Pileus 2.1–2.4 cm in diameter, lilac (10 P 8 / 2), hemispherical to convex, becoming plano-convex, sticky, glabrous to sericeous, margin incurved, with purple (10 P 6 / 10) fibrills, and smooth, entire. Context thin, lilac. Lamellae subdecurrent, lilac (10 P 8 / 2), narrow, crowded, thick, edges even, entire to eroded. Lamellulae abundant, irregular, of varying lengths, in more than two tiers, alternating with lamellae. Stipe 3.4–3.8 cm long, reddish purple (2.5 RP 2 / 2) at the apex and darker than the base, lilac (10 P 8 / 2) at the base, central, cylindrical to flexuous, slightly pruinose, longitudinally striate, hollow, context lilac (10 P 8 / 2). Hyphal system composed of generative hyphae with clamp connections, dextrinoid, inflated in KOH, thin-walled, occasionally branched, regular to subregularly arranged, 4.1–10.8 µm in diameter.</p><p>Basidiospores 4.95–8.31 × 2.56–5 µm, av. L × av. W = 6.68 × 2.56 µm, Q = 1.76–1.88, subcylindrical to ellipsoid with an apiculus and oil drop, amyloid, smooth, hyaline in 5 % KOH, amyloid, thin-walled. Probasidia subclavate to clavate, thin-walled. Basidia 22.36–35.12 × 7.64–8.73 µm, av. L × av. W = 30.60 × 8.05 µm, Q = 3.58–3.79, broadly clavate to slightly pyriform, smooth, hyaline in 5 % KOH, thin-walled, four-spored. Cheilocystidia 28.32–44.76 × 8.56–13.75, av. L × av. W = 35.99 × 11.81 µm, abundant, polymorphic, clavate, fusiform, pyriform, some with a rounded apex, smooth, hyaline in 5 % KOH, thin-walled. Pleurocystidia not observed. Pileipellis a cutis of irregular and inflated smooth hyphae, thin-walled, 17.32–23.93 µm in diameter.</p><p>Additional specimen (paratype) examined.</p><p>CHINA. Gansu Province, Tibetan Autonomous of Gannan, Taohe National Nature Reserve, 34°40'70"N, 103°53'26"E, mossy ground in mixed forests, 4 September 2024, MSX 1785. (GenBank ITS = PX 612290; tef 1 - α = PX 684296) .</p><p>Notes.</p><p>Species of Mycena sect. Calodontes are easily recognized by the pinkish, reddish, purplish to brownish pileus and large-sized basidiomata. Mycena lavendula is characterized by its lilac pileus and reddish-purple stipe that becomes lilac toward the base, is longitudinally striated, and has a pruinose apex. Microscopically, it is distinguished by small, subcylindrical to ellipsoid basidiospores (av. 6.68 × 2.56 μm) with an oil drop, polymorphic cheilocystidia, and clamped generative hyphae.</p><p>Mycena lavendula resembles M. luciferina Cortés-Pérez, Guzm. - Dáv. &amp; Ram. - Cruz., a recently described bioluminescent species from Mexico, in the shape and size of its basidiospores and cheilocystidia with rounded apices. However, M. lavendula can be differentiated from M. luciferina by its unique pileus and stipe morphology. Mycena luciferina has a reddish-pink pileus and white stipe, whereas M. lavendula has a lilac pileus and a reddish-purple stipe with a pruinose apex. Anatomically, M. lavendula differs from M. luciferina by its larger basidia (22.36–35.12 × 7.64–8.73 μm vs. 15.2–27.2 × 4.5–5.6 μm). Furthermore, M. luciferina has larger basidiospores (7.4 × 3.8 μm), whereas M. lavendula possesses smaller ones (6.68 × 2.56 μm) (Cortés-Pérez et al. 2023). In the phylogenetic tree, Mycena lavendula is separated from M. oblongispora, another species described in this study, and forms a sister clade with it. Mycena oblongispora can be differentiated from M. lavendula by its grayish-brown pileus with brownish-black fibrils, clavate cheilocystidia, and larger basidiospores, whereas M. lavendula has a lilac pileus, polymorphic cheilocystidia, and smaller basidiospores (6.68 × 2.56 μm vs. 7.25 × 4.96 μm).</p></div>	https://treatment.plazi.org/id/8D4BAEF2AB615AEDA66A9F6F69369DC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
E31A1660371D52C49AABFDEE389D14B0.text	E31A1660371D52C49AABFDEE389D14B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena longnanensis S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>5. Mycena longnanensis S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 10, 11</p><p>Etymology.</p><p>The specific epithet “longnanensis” refers to type locality Longnan.</p><p>Diagnosis.</p><p>The species is characterized by an umbonate, radially fibrillose, lilac to dark purple pileus, decurrent and lilac lamellae, ellipsoid to subcylindrical basidiospores (6.75 × 3.45 µm), and spheropedunculate, small cheilocystidia (39 × 14.21 µm), with a slightly long tail and a round base.</p><p>Holotype.</p><p>CHINA, Gansu Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.13333&amp;materialsCitation.latitude=34.0" title="Search Plazi for locations around (long 103.13333/lat 34.0)">Longnan City</a>, Dangchang County, 34°N, 103°8'E, in mixed coniferous-broadleaf forests, 4 September 2024, FLF 1804 (GenBank ITS = PX 612291; tef 1 - α = PX 684297) .</p><p>Description.</p><p>Basidiomata medium-sized. Pileus 2.2–2.4 cm in diameter, grayish lilac (5 P 18) to dark purple (7.5 P 5 / 10), campanulate, with a round and prominent lilac (10 P 8 / 2) umbo. Surface radially fibrillose and sticky. Margin decurved to incurved, sulcate to slightly undulating, dark purple (7.5 P 5 / 10) in a ring, followed by a lilac (10 P 8 / 2) circle. Context thin, grayish lilac. Lamellae uncinate, deeply decurrent, lilac (10 P 8 / 2), broad, crowded, thick, even, crenate, forking. Lamellulae frequent, irregular, in more than two tiers, alternating with lamellae. Stipe c. 4.3 cm long, reddish purple (2.5 RP 2 / 2) to dark purple brown (10 YR 2 / 3) with lilac (10 P 8 / 2) ting at base, central, flexuous, cylindrical, tapering towards the base, fibrillose, fine appressed longitudinal fibrils, solid.</p><p>Hyphal system composed of generative clamped hyphae, dextrinoid, inflated in KOH, thin-walled, occasionally branched, regularly to subregularly arranged, 5.87–14.94 µm in diameter.</p><p>Basidiospores 5.73–8.28 × 2.75–4.39 µm, av. L × av. W = 6.75 × 3.45 µm, Q = 1.85–1.96, long-ellipsoid to subcylindrical with an apiculus and oil drop, smooth, hyaline in 5 % KOH, amyloid, thin-walled. Probasidia cylindrical to clavate, thick-walled. Basidia 22.83–31.92 × 4.68–6.75 µm, av. L × av. W = 27.88 × 5.92 µm, Q = 4.95, narrowly clavate, smooth, hyaline in 5 % KOH, thin-walled, two or four-spored. Cheilocystidia 31.61–47.81 × 10.79–17.70 µm, av. L × av. W = 39.00 × 14.21 µm, abundant, spheropedunculate with slightly long tail, round apex, smooth, hyaline in 5 % KOH, thin-walled. Pleurocystidia not observed. Pileipellis a cutis of smooth, thin-walled and clamped hyphae of 12.15–18.21 µm in diameter.</p><p>Additional specimen (isotype) examined.</p><p>CHINA. Gansu Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.13333&amp;materialsCitation.latitude=34.0" title="Search Plazi for locations around (long 103.13333/lat 34.0)">Longnan City</a>, Dangchang County, 34°N, 103°8'E, in mixed coniferous-broadleaf forests, 4 September 2024, MSX 1804. (GenBank ITS = PX 612292; tef 1 - α = PX 684298) .</p><p>Notes.</p><p>The new species is characterized by an umbonate, radially fibrillose, lilac to dark purple pileus, decurrent and lilac lamellae, ellipsoid to subcylindrical basidiospores (6.75 × 3.45 μm), and smaller, spheropedunculate cheilocystidia (39 × 14.21 μm) with a slightly elongated tail and rounded apex. It resembles M. brunneoviolacea A. C. Cooper, Desjardin &amp; B. A. Perry in its pileus and stipe morphology. However, the new species is distinguished by a grayish-lilac umbo and a radially fibrillose pileus. Furthermore, M. brunneoviolacea possesses ascending to adnate, white to pale gray lamellae, whereas M. longnanensis features uncinate, deeply decurrent, lilac lamellae. Anatomically, M. longnanensis can be differentiated by its unique, smaller, spheropedunculate cheilocystidia (av. 39.0 × 14.2 μm), whereas Mycena brunneoviolacea possesses larger, lageniform cheilocystidia (32–72 × 9.6–18.4 μm) (Cooper et al. 2018). Another similar species is M. diosma Krieglst. &amp; Schwöbel, which features basidiomata with a dark violet-brown or reddish-purple, hygrophanous pileus. Furthermore, M. diosma is distinguished by its dark brownish-violet to dark violet lamellae and larger cheilocystidia [20–60 (– 80) × 3.5–20 μm] (Aronsen and Læssøe 2016). Another species in the same clade is M. cahaya A. L. C. Chew &amp; Desjardin, which has a brown pileus, elongated basidiospores (av. 7.1 × 3.8 μm), and abundant clavate to ventricose cheilocystidia (Chew et al. 2014). Mycena longnanensis resembles M. glabra in having long-ellipsoid to subcylindrical basidiospores and bi- or tetrasporic basidia. However, M. longnanensis is distinguished by its dark purple pileus, lilac lamellae, and reddish-purple to dark purple-brown stipe, whereas M. glabra has white to cream basidiomata, white to pale pink-yellow lamellae, and a white to pale wax-yellow stipe. Anatomically, M. longnanensis is distinguished from M. glabra by its narrower basidiospores (2.75–4.39 μm vs. 3.7–5.5 μm), significantly smaller basidia (22.83–31.92 × 4.68–6.75 μm vs. 29.0–45.0 × 6.3–12.0 μm), and shorter cheilocystidia (31.61–47.81 × 10.79–17.70 μm vs. 48.0–77.4 × 10.0–16.4 μm) (Fan et al. 2024).</p></div>	https://treatment.plazi.org/id/E31A1660371D52C49AABFDEE389D14B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
CD21CCB0AA89520BBE1C426C2E20F8E8.text	CD21CCB0AA89520BBE1C426C2E20F8E8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena minispora S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>6. Mycena minispora S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 12, 13</p><p>Etymology.</p><p>The specific epithet minispora ” refers to the small-sized basidiospores.</p><p>Diagnosis.</p><p>Mycena minispora is characterized by its glutinous, fibrillose golden-brown pileus with a low obtuse umbo, milky-yellow lamellae, smaller (6.04 × 3.13 µm) amyloid basidiospores and fusiform to subclavate cheilocystidia with blunt and rounded apices.</p><p>Holotype.</p><p>CHINA, Gansu Province, Gannan Tibetan Autonomous Prefecture, Yeliguan National Forest Park, 34°95'N, 103°60'E, in mixed coniferous, broadleaf forests, 5 July 2024, FLF 1223 (GenBank ITS = PX 612293) .</p><p>Description.</p><p>Basidiomata large-sized. Pileus 3.5–3.7 cm in diameter, brown (7.5 YR 6 / 8) with a light golden-brown (2.5 Y 7 / 8) umbo and margins; plane to slightly convex with a low, obtuse, non-prominent umbo; surface glutinous to viscid, hygrophanous, and fibrillose; margin straight to very slightly uplifted, plicate. Context thin, white to gray. Lamellae sinuate, edges eroded to crenate, milky-yellow (5 Y 8 / 1), broad, close, thick, even, unequal. Lamellulae frequent, intervenose, irregular, arranged in more than two tiers and alternating with the lamellae. Stipe approximately 4.7 cm long, eccentric, terete and cylindrical, firm, ribbed or costate, pale yellow (5 Y 8 / 4) with a light green (7.5 GY 8 / 2) tinge at the base; apex slightly pruinose.</p><p>Hyphal system composed of generative clamped hyphae, dextrinoid, inflated in KOH, thin-walled, septate, regular to subregularly arranged, 6.28–10.59 µm in diameter.</p><p>Basidiospores 4.61–6.92 × 2.28–4.04 µm, av. L × av. W = 6.04 × 3.13 µm, Q = 1.89–1.95, ellipsoid to subcylindrical with a distinct apiculus and oil drop, smooth, pale yellow in 5 % KOH, amyloid, thin-walled, guttulated. Probasidia subclavate to clavate, thin-walled. Basidia 17.28–26.82 × 4.67–6.46 µm, av. L × av. W = 23.96 × 5.62 µm, Q = 4.21–4.25, narrowly clavate, pale yellow in 5 % KOH, thin-walled, four-spored. Cheilocystidia 35.82–59.93 × 12.86–20.37 µm, av. L × av. W = 48.76 × 16.18 µm, abundant fusiform to subclavate with a blunt, rounded apex, smooth, pale yellow in 5 % KOH, thin-walled. Pleurocystidia not observed. Pileipellis a cutis, made up smooth, thin-walled and clamped hyphae of 13.62–21.32 µm in diameter.</p><p>Additional specimen examined.</p><p>CHINA. Gansu Province, Gannan Tibetan Autonomous Prefecture, Yeliguan National Forest Park, 34°95'N, 103°50'E, in mixed coniferous-broadleaf forests, 5 July 2024, FLF 1244. (GenBank ITS = PX 612294; tef 1 - α = PX 684299) .</p><p>Notes.</p><p>The presence of a hygrophanous, brownish pileus, intervenose lamellae, smooth cheilocystidia, and amyloid basidiospores confirms the placement of M. minispora within Mycena sect. Calodontes . Phylogenetic analyses based on ITS and tef 1 - α further support this taxonomic position.</p><p>Mycena minispora is characterized by a brown pileus with a light golden-brown umbo, milky yellow, sinuate lamellae with eroded to crenate edges, and a pale-yellow stipe exhibiting a light green tinge at the base. Microscopically, the species possesses small, ellipsoid to subcylindrical basidiospores (6.04 × 3.13 μm) and fusiform or pyriform cheilocystidia. Mycena minispora can be readily distinguished from M. pura (Pers.) P. Kumm, which has larger basidiospores (6–9.5 × 4–5.2 μm) and pink to violet lamellae. Mycena minispora is separated from M. luciferina, which is characterized by a campanulate, reddish-pink to pale pink pileus and white intervenose lamellae. In contrast, M. minispora possesses a hygrophanous, brownish pileus, a yellow stipe with a light green base, and smaller basidiospores (6.04 × 3.13 μm vs. 7.4 × 3.8 μm; Cortés-Pérez et al. 2023).</p><p>Another morphologically similar species is M. shengshanensis, which shares several features with M. minispora, including a convex pileus with an obtuse, disc-like umbo, a paler striate margin, golden-brown to brown coloration, sinuate and irregularly serrulate lamellae, and a central, cylindrical stipe. However, M. minispora differs by having a larger pileus (3.5–3.7 cm vs. 1.3–2.6 cm) and a pale-yellow stipe with a light green tinge at the base, whereas M. shengshanensis possesses a reddish-brown to brownish-gray stipe. Anatomically, M. minispora is further distinguished by its smaller basidiospores and basidia (4.61–6.92 × 2.28–4.04 μm vs. 6.10–8.10 × 3.40–4.40 μm for basidiospores; 17.28–26.82 × 4.67–6.46 μm vs. 22–32 × 6–8 μm for basidia; Liu et al. 2022).</p></div>	https://treatment.plazi.org/id/CD21CCB0AA89520BBE1C426C2E20F8E8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
78D64CFC46F856D2B8C172101EC001AB.text	78D64CFC46F856D2B8C172101EC001AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena oblongispora S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>3. Mycena oblongispora S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 6, 7</p><p>Etymology.</p><p>The specific epithet “oblongispora” refers to the oblong shape of spores.</p><p>Diagnosis.</p><p>It is characterized by grayish brown pileus having brownish black fibrils, gray, free to remote lamellae, with rhizomorphs at the base stipe, growing on decaying logs, smaller (7.25 × 4.96 µm), subglobose to ellipsoid to oblong basidiospores, smooth, smaller (35.27 × 12.40 µm) cheliocystidia and with scattered excrescences in pileipellis.</p><p>Holotype.</p><p>CHINA, Gansu Province, Longnan City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.03333&amp;materialsCitation.latitude=33.15" title="Search Plazi for locations around (long 104.03333/lat 33.15)">Guan’e gully national forest park</a>, 33°9'N, 104°2'E, grow on decaying logs in mixed coniferous-broadleaf forests, 5 September 2024, FLF 1861 (GenBank ITS = PX 612295; tef 1 - α = PX 684300) .</p><p>Description.</p><p>Basidiomata small-sized. Pileus 1.2–1.9 cm in diameter, with brownish black (10 YR 2 / 2) fibrils on a grayish brown (10 YR 4 / 2) context, parabolic, dry, sulcate to plicate, striated, striation expands towards margins, margins slightly uplifted to straight, even, entire. Context thin, white to grayish brown (10 YR 4 / 2). Lamellae free to remote, even, entire, gray (2.5 GY 7 / 1), broad, subdistant, average, unequal to forked. Lamellulae regular, of two different sizes, alternating with lamellae, in two tiers. Stipe 2.5–3.9 cm long, central, cylindrical to flexuous, smooth to virgate with maturity, apex gray (2.5 GY 7 / 1), becoming black (10 YR 3 / 2) to dark brown (10 YR 2 / 3) towards the base whitish (10 R 8 / 1) with distinct rhizomorphs. Hyphal system composed of generative hyphae with clamp connections, dextrinoid, inflated in KOH, thin-walled, branched, regultar to subregularly arranged, 1.94–6.32 µm in diameter.</p><p>Basidiospores 5.94–8.94 × 3.38–7.06 µm, av. L × av. W = 7.25 × 4.96 µm, Q = 1.34–1.46, subglobose to oblong ellipsoid or slightly ovoid with an apiculus and oil drop, smooth, hyaline in 5 % KOH, amyloid, thin-walled. Probasidia cylindrical to clavate, elongated and slender at the base, thin-walled. Basidia 23.98–34.59 × 5.78–9.88 µm, av. L × av. W = 28.83 × 8.76 µm, Q = 2.28, clavate, guttulated, hyaline, thin-walled, two to four spored. Cheilocystidia 29.60–43.74 × 8.46–14.87 µm, av. L × av. W = 35.27 × 12.40 µm, abundant, clavate with capitate apex, smooth, hyaline in 5 % KOH, thin-walled. Pleurocystidia not observed. Hyphae of the pileipellis 5.87–16.77 µm, inflated, thin-walled, with scattered excrescences.</p><p>Additional specimen (isotype) examined.</p><p>CHINA. Gansu Province, Longnan City, Guan’e gully national forest park, 33°93'N, 104°2'E, grow on decaying logs in mixed coniferous-broadleaf forests, 5 September 2024, MSX 1861. (GenBank ITS = PX 612296; tef 1 - α = PX 684301) .</p><p>Notes.</p><p>This new species belongs to Mycena sect. Fragilipedes . It is characterized by a grayish-brown pileus with brownish-black fibrils, gray, free to remote lamellae, and a stipe base with distinct rhizomorphs. Ecologically, it grows on decaying logs. Microscopically, it is distinguished by its smaller, subglobose to oblong-ellipsoid basidiospores (av. 7.25 × 4.96 μm) and hyphae of the pileipellis with scattered excrescences. Mycena stipata Maas Geest. &amp; Schwöbel shares similarities with M. oblongispora, as both possess a brownish-beige to blackish-brown pileus. However, M. oblongispora can be distinguished by its fibrillose pileus (smooth in M. stipata), gray, free to remote lamellae (adnate to adnexed in M. stipata), clavate cheilocystidia with a capitate apex (fusiform to lageniform in M. stipata), and smaller basidiospores (7.25 × 4.96 μm).</p><p>Another close relative, Mycena silvae-nigrae Maas Geest. &amp; Schwöbel, shares several similarities, such as a blackish-brown pileus, whitish to grayish lamellae, and the presence of excrescences in the pileipellis. However, M. oblongispora has smaller basidiospores (7.25 × 4.96 μm) and smaller cheilocystidia (29.60–43.74 × 8.46–14.87 μm) with a rounded apex, while M. silvae-nigrae has larger basidiospores (9.5 × 9 μm) and larger cheilocystidia (20–73 × 7–20 μm) (Osmundson et al. 2013).</p></div>	https://treatment.plazi.org/id/78D64CFC46F856D2B8C172101EC001AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
312664694D51548AB9C0BD61FFEA5B60.text	312664694D51548AB9C0BD61FFEA5B60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena pakistanica A. Naseer, A. Khalid, A. N. Khalid, S. X. Ma & L. F. Fan 2026	<div><p>7. Mycena pakistanica A. Naseer, A. Khalid, A. N. Khalid, S. X. Ma &amp; L. F. Fan sp. nov.</p><p>Figs 14, 15</p><p>Etymology.</p><p>The specific epithet “pakistanica” refers to the country name, Pakistan.</p><p>Diagnosis.</p><p>The species is characterized by a lavender pileus with grape wine fibrils, larger (12 cm) stipe, lacrymoid subcylindrical to ellipsoid, smaller (5.9 × 2.6 µm) basidiospores, spathulate pleurocystidia, and fusiform to sublageniform cheilocystidia with both sharp ends.</p><p>Holotype.</p><p>PAKISTAN, Khyber Pakhtunkhwa Province, Abbottabad district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.416664&amp;materialsCitation.latitude=34.033333" title="Search Plazi for locations around (long 73.416664/lat 34.033333)">Kala Bagh</a>, 34°2'0"N, 73°25'0"E, in mixed coniferous forest, found on wood, 11 August 2024, Arooj Naseer, KB – 16 (GenBank ITS = PX 612297) .</p><p>Description.</p><p>Basidiomata large-sized, solitary, or scattered. Pileus 2.0– 2.9 cm in diameter, lavender (7.5 R 5 / 3) with grape-wine (7.5 R 6 / 2) fibrils; shape conical to parabolic, becoming planate when mature; surface dry, fibrillose, with distinct veil remnants; margin decurved, becoming slightly incurved with age, varying from plicate or undulating when young to sulcate and entire at maturity. Context thin, lavender. Lamellae emerginate, light grayish-purple (N 7 / 0), ventricose, thick, and subdistant, unequal to forked; Lamellulae frequent, irregular, arranged in one to two tiers alternating with the lamellae. Stipe 8.0–12.0 × 0.1–0.3 cm, central, equal to slightly flexuous, fragile; surface smooth to fibrillose towards the base, light grayish-purple (N 7 / 0) at the apex, becoming dark purplish-brown (7.5 YR 3 / 4) towards the base.</p><p>Hyphal system composed of generative clamped hyphae, dextrinoid, inflated in KOH. generative hyphae in context brown to reddish brown, thin-walled, occasionally branched, septate, irregularly arranged, 2.2–11.8 µm in diameter. Generative hyphae in lamella hyaline in KOH, thin-walled, occasionally branched, irregularly arranged, 2–8 µm in diameter.</p><p>Basidiospores (4.4 –) 4.9 – 7.3 (– 7.4) × (2.2 –) 2.4 – 2.8 (– 3) µm, av. L × av. W = 5.9 × 2.6 µm, Q = 1.8 – 3.3, av. Q. = 2.3, lacrymoid, subcylindrical to ellipsoid, with suprahilar depression, phaseoliform in side view, oblong in frontal view, apiculate, smooth, hyaline to light yellow in 5 % KOH, amyloid, thin-walled. Basidia (18.6 –) 18.8 – 22.3 (– 22.9) × (4.5 –) 4.6 – 5.7 (– 6) µm, av. L × av. W = 20.1 × 5.4 µm, clavate to slightly pyriform, smooth, hyaline in 5 % KOH, thick-walled, mostly four-spored. Cheilocystidia (57.8 –) 58.2 – 78.4 (– 91) × (11.2 –) 12.7 – 16.2 (– 16.4) µm, av. L × av. W = 71.6 × 13.9 µm, frequently present, mostly clavate to broadly clavate with acute apices, fusiform to sublageniform with both sharp ends, rarely lecythiform, mucronate, smooth, hyaline to light brown in 5 % KOH, thick-walled. Pleurocystidia (49.5 –) 54.2 – 72 (– 74) × (9.5 –) 10 – 17.6 (– 19.5) µm, av. L × av. W = 62.3 × 14.3 µm, similar in shape to cheilocystidia but smaller, mostly clavate to spathulate with knob-like outgrowth, narrowly lageniform to mucronate or with cylindrical outgrowth, smooth, hyaline to light brown in 5 % KOH, thin to rarely thick-walled. Pileipellis a cutis, made up of irregular, branched 1.6–7.6 µm, average width 4.02 µm, inflated, hyaline to pale yellow in 5 % KOH, septate, thick-walled.</p><p>Notes.</p><p>The presence of a pinkish, reddish, purplish to brownish pileus, large-sized basidiomata, and smooth hymenial cystidia supports the placement of M. pakistanica within Mycena sect. Calodontes . Mycena pakistanica is characterized by a lavender pileus with grape-wine fibrils, a long stipe (up to 12 cm), and small, lacrymoid, subcylindrical to ellipsoid basidiospores (av. 5.9 × 2.6 μm). Additionally, it features spathulate pleurocystidia and fusiform to sublageniform cheilocystidia with acute apices.</p><p>Mycena pakistanica shares morphological similarities with Mycena pura, including light purplish-gray lamellae, a dark purplish-brown, smooth, and equal stipe, and clavate to fusiform cheilocystidia. However, M. pakistanica can be distinguished from M. pura by its smaller (2.9 cm in diameter), conical to flat, fibrillose pileus with plicate to sulcate margins and emarginate lamellae. Additionally, M. pakistanica possesses a longer (12 cm in length) stipe that is fibrillose toward the base, whereas M. pura has a relatively shorter stipe (up to 11 cm) that is pruinose to glabrescent. Anatomically, M. pakistanica is distinguished by its smaller, lacrymoid, subcylindrical to ellipsoid basidiospores (av. 5.9 × 2.6 μm), whereas M. pura possesses larger, narrowly ellipsoid to cylindrical basidiospores (av. 8.4 × 4 μm) (Perry 2002). Moreover, M. pakistanica has larger, sublageniform cheilocystidia with both ends sharp and mucronate (up to 91 μm) and spathulate pleurocystidia (up to 74 μm).</p><p>Mycena pakistanica is also comparable with M. pelianthina (Fr.) Quél. (1872) . Both share a dark brown pileus, purple lamellae, an equal and fibrillose stipe, thin-walled clavate basidia, and fusiform cheilocystidia. However, M. pelianthina differs from M. pakistanica mainly in its larger (up to 50 mm), hemispherical to campanulate pileus with a glabrous surface that is slightly lubricous when moist. Furthermore, M. pelianthina features adnate to decurrent lamellae and a longer (up to 120 mm), whitish to lilaceous stipe that is striate with dark fibrils. Microscopically, M. pelianthina is distinguished by larger pip-shaped basidiospores (av. 7 × 3.8 μm), smaller cheilo- and pleurocystidia (up to 70 μm), and smooth cortical hyphae in the pileipellis (Harder et al. 2010, 2012).</p></div>	https://treatment.plazi.org/id/312664694D51548AB9C0BD61FFEA5B60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
8899FD5BD59C5D5C885F3F42F483F6A6.text	8899FD5BD59C5D5C885F3F42F483F6A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena tephroleuca S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>8. Mycena tephroleuca S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 16, 17</p><p>Etymology.</p><p>The specific epithet “tephroleuca” refers to color of the basidiomata.</p><p>Diagnosis.</p><p>The species is characterized by white to grayish-white pileus, adnate lamellae with a decurrent tooth, a white stipe with brown tinge at base, thin-walled hyphae and clamped, and the presence of clavate to narrowly fusiform or utriform cheilocystidia with cylindrical outgrowths.</p><p>Holotype.</p><p>CHINA, Gansu Province, Gannan Tibetan Autonomous Prefecture, Diebu County, Lazikou Forest Protection Station, 33°85'N, 104°55'E, in broad-leaved forests, 4 th September 2024, FLF 1816 (GenBank ITS = PX 612306; Tef 1 - α = PX 684309) .</p><p>Description.</p><p>Basidiomata small-sized. Pileus 0.7–1.5 cm in diameter, chalky white (10 R 8 / 1) to chalky white (10 R 8 / 1), pulvinate, slightly sticky, glabrous in center, marginally fibrillose, even, entire to plicate, margins incurved to straight. Context thin, white. Lamellae adnate with decurrent tooth, fimbriate, chalky white (10 R 8 / 1), broad, subdistant, thick, unequal, wavy. Lamellulae are of two different sizes, in two tiers, alternating with lamellae. Stipe 3.8–4.3 cm long, chalky white (10 R 8 / 1) to smoky white (2.5 YR 8 / 1), flexuous, equal, surface smooth, with some brown tinge at base, white rhizomorphs, central.</p><p>Hyphal system composed of generative hyphae with clamp connections, dextrinoid, inflated in KOH, thin-walled, regular to subregularly arranged, 1.26–9.20 µm in diameter.</p><p>Basidiospores 7–8.77 × 3.8–5.9 µm, av. L × av. W = 7.73 × 4.79 µm, Q = 1.61–1.73, oblong to elliptical, with an apiculus and oil drop, smooth, hyaline in 5 % KOH, amyloid, thin-walled. Basidia 24.97–32.57 × 7.38–9.16 µm, av. L × av. W = 29.45 × 8.19 µm, clavate, guttulated, hyaline, thin-walled, two to four-spored. Cheilocystidia 48.22–54.80 × 13.04–18.80 µm, av. L × av. W = 14.26 × 14.7 µm, clavate, narrowly fusiform, utriform, mucronate or with a cylindrical outgrowth, smooth, hyaline in 5 % KOH, thin-walled. Pleurocystidia not observed. Pileipellis a cutis made up clamped hyphae of 10.08–24.14 µm in diameter, smooth, thin-walled.</p><p>Additional specimen (paratype) examined.</p><p>CHINA. Gansu Province, Gannan Tibetan Autonomous Prefecture, Lazikou Forest Protection Station, Laolong Gully, 33°85'N, 104°55'E, in broad-leaved forests, 4 September 2024, MSX 1816. (GenBank ITS = PX 612307; tef 1 - α = PX 684310) .</p><p>Notes.</p><p>The species is characterized by small-sized basidiomata, a chalky-white to grayish-white pileus, adnate lamellae with a decurrent tooth, a chalky-white to smoky-white stipe with some brown tinge at the base, a pileipellis as a cutis made up of clamped hyphae 10.08–24.14 μm in diameter, smooth, thin-walled hyphae, and abundant, smooth, clavate to narrowly fusiform or utriform cheilocystidia with cylindrical outgrowths.</p><p>Mycena tephroleuca shows similarity to M. cahaya in the size of its basidiospores and two- or four-spored basidia. However, M. tephroleuca possesses a pulvinate, marginally fibrillose pileus with entire to plicate margins and a brown tinge at the stipe base with white rhizomorphs, whereas M. cahaya possesses a parabolic to convex or campanulate, uneven to slightly wrinkled pileus and a yellowish-gray to beige, glabrous stipe. Anatomically, M. tephroleuca is distinguished by spherical to phaseoliform basidiospores and broader basidia, whereas M. cahaya has elongate to cylindrical basidiospores and narrower basidia (7.38–9.16 μm vs. 4.0–6.4 μm) (Chew et al. 2014).</p><p>Mycena tephroleuca the novel species described in this study, forms a sister group to M. glabra B. Y. Wang, T. F. Ma &amp; L. F. Fan in the phylogenetic analysis, sharing similarities in its long ellipsoid basidiospores and comparable basidial size and shape. Morphologically, M. tephroleuca possesses a chalky-white to grayish-white, pulvinate pileus that is glabrous at the center and fibrillose at the margin, with fimbriate lamellae, whereas M. glabra features a white to cream pileus with vertical stripes or grooves and subdecurrent lamellae. In M. tephroleuca, the stipe is flexuous, chalky-white to smoky-white with brown tinges at the base, and covered with white rhizomorphs, whereas M. glabra has a smooth, white to cream stipe that is pale wax-yellow at the base. Anatomically, M. tephroleuca has broader cheilocystidia (13.04–18.80 μm), whereas M. glabra has narrower cheilocystidia (10.0–16.4 μm).</p><p>Mycena rufobrunnea shares morphological affinities with Mycena tephroleuca, particularly in small-sized basidiomata with a relatively thin, white pileus context, basidiospore dimensions, and the absence of pleurocystidia. However, M. rufobrunnea is readily distinguished by its vividly colored stipe, ranging from grayish-magenta to dark purple, in contrast to the entirely chalky-white to smoky-white stipe of M. tephroleuca . Anatomically, M. tephroleuca possesses smaller basidiospores (7.73 × 4.79 μm) and larger cheilocystidia (48.22–54.80 × 13.04–18.80 μm) that are more morphologically diverse, including mucronate forms and cylindrical outgrowths. In contrast, M. rufobrunnea has larger basidiospores (8.3 × 4.4 μm) and smaller, utriform to clavate cheilocystidia (23–44 × 7–17 μm) (Wei et al. 2024). This species also shares similarities with another taxon described in this study, Mycena lavendula, which is characterized by a lilac pileus and reddish-purple stipe with a lilac base, longitudinal striations, and a pruinose apex. Anatomically, M. lavendula has subcylindrical to ellipsoid basidiospores (6.68 × 2.56 μm), polymorphic cheilocystidia with rounded apices, and clamped generative hyphae.</p></div>	https://treatment.plazi.org/id/8899FD5BD59C5D5C885F3F42F483F6A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
D767F70A45515F749344561B1D007542.text	D767F70A45515F749344561B1D007542.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phloeomana qilianensis S. X. Ma, A. Naseer, A. N. Khalid & L. F. Fan 2026	<div><p>9. Phloeomana qilianensis S. X. Ma, A. Naseer, A. N. Khalid &amp; L. F. Fan sp. nov.</p><p>Figs 18, 19</p><p>Etymology.</p><p>The specific epithet “ qilianensis ” refers to the type locality, Qilian.</p><p>Diagnosis.</p><p>Phloeomana qilianensis is characterized by small (up to 1 cm) basidiomata, an off-white to pale yellow pileus, fimbriate lamellae, a furfuraceous stipe, lacrymoid basidiospores (7.39 × 3.99 µm), and abundant, fusiform to ventricose-rostrate cheilocystidia with an obtuse apex and tapered base.</p><p>Holotype.</p><p>CHINA, Gansu Province, Zhangye City, Qilian Mountains National Nature Reserve, Qinglong Resource Conservation Station, 38°68'N, 99°92'E, in mixed coniferous-broadleaf forests, 3 August 2024, FLF 1582. (GenBank ITS = PX 612301) .</p><p>Description.</p><p>Basidiomata small, in cluster. Pileus 0.4–1.0 cm in diameter, off-white (10 R 8 / 1) when young, becoming pale yellow (2.5 R 8 / 4) with age, campanulate to parabolic, dry, sometimes viscid, smooth at the center, fibrillose towards margins, plicate to split, margin incurved to slightly uplifted. Context thin, white. Lamellae regular, fimbriate, off-white (10 R 8 / 1), narrow, subdistant, thick, unequal, even. Lamellulae of two lengths, arranged in two tiers, alternating with the lamellae. Stipe central, flexuous, cylindrical to slightly bulbous towards base 2.1–3.4 cm, smoky-white (2.5 YR 8 / 1) with a white pruinose cpating, surface scurfy, branny, furfuraceous.</p><p>Hyphal system composed of generative hyphae with clamp connections, IKI –, inflated in KOH, thin-walled, occasionally branched, septate, regular to subregularly arranged, 3.31–3.94 µm in diameter.</p><p>Basidiospores 6–8.5 × 3.43–4.72 µm, av. L × av. W = 7.39 × 3.99 µm, Q = 1.72–1.85, lacrymoid, oblong ellipsoid to subcylindrical with an apiculus and oil drop, smooth, hyaline in 5 % KOH, IKI –, guttulated, thin-walled. Probasidia cylindrical to clavate, thin-walled. Basidia 21.01–33.21 × 5.21–6.56 cm, av. L × av. W = 28.16 × 5.92 µm, Q = 4.75, narrowly clavate, guttulated, hyaline, thin – walled, two or four – spored. Cheilocystidia 24.65–28.56 × 3.13–3.87 µm, av. L × av. W = 27.04 × 3.37 µm, abundant, fusiform, ventricose-rostrate, with obtuse apex, base tapered, with short to long stalk, smooth, hyaline in 5 % KOH, thin-walled. Pleurocystidia not observed. Pileipellis a cutis of smooth, thin-walled hyphae of 3.64–11.55 µm in diameter, with clavate terminal cells.</p><p>Additional specimen examined.</p><p>CHINA. Gansu Province, Qilian Mountains National Nature Reserve, Tianzhu Tibetan Autonomous County, Dongdatan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.123055&amp;materialsCitation.latitude=37.533333" title="Search Plazi for locations around (long 102.123055/lat 37.533333)">Xichakou Village</a>, 37°32'N, 102°7'23"E, in mixed coniferous-broadleaf forests, 28 July 2024, FLF 1269. (GenBank ITS = PX 612300; tef 1 - α = PX 684304) .</p><p>Notes.</p><p>The presence of horizontal to arcuate lamellae with straight or concave edges, smooth cheilocystidia, and inamyloid spores confirms that the new species belongs to the genus Phloeomana Redhead. The new species is characterized by small, clustered basidiomata, an off-white to pale yellow pileus, fimbriate lamellae, a scurfy, furfuraceous stipe, lacrymoid to oblong-ellipsoid or subcylindrical basidiospores, and abundant, fusiform, ventricose-rostrate cheilocystidia with an obtuse apex and a tapered base.</p><p>Phloeomana qilianensis resembles P. speirea (Fr.) Redhead in having small basidiomata with furfuraceous surfaces and ellipsoid basidiospores. However, P. qilianensis can be distinguished from P. speirea by its off-white to pale yellow pileus and smoky-white stipe covered with white dust, whereas P. speirea displays a pale gray-brown to pale yellowish-brown pileus and a stipe that is densely but minutely puberulous throughout. Both species share similar cheilocystidia in terms of abundance and general fusiform shape, but they differ in apex morphology — P. qilianensis has ventricose-rostrate cheilocystidia with tapered bases and obtuse apices, whereas P. speirea features utriform or somewhat irregularly shaped cheilocystidia, usually simple but sometimes furcated, lobed, or bearing a few excrescences, and often curved to somewhat flexuous.</p></div>	https://treatment.plazi.org/id/D767F70A45515F749344561B1D007542	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Fan, Long Fei;Naseer, Arooj;Khalid, Ammara;Khalid, Abdul Nasir;Deng, Yan;Wang, Chen Yan;Wang, Yuan;Ma, Si Xuan	Fan, Long Fei, Naseer, Arooj, Khalid, Ammara, Khalid, Abdul Nasir, Deng, Yan, Wang, Chen Yan, Wang, Yuan, Ma, Si Xuan (2026): Unveiling the diversity of the families Cyphellaceae and Mycenaceae from Southeast Asia. IMA Fungus 17: e 182174, DOI: 10.3897/imafungus.17.182174
