taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
A67987E6FFA76765F2BD002FFD1CF86D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFA76765F2BD002FFD1CF86D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFA96766F2BD0275FC28FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFA96766F2BD0275FC28FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFA96766F2BD0275FC28FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAB6761F2BD0636FDA5FDEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166181/files/figure.png	https://doi.org/10.5281/zenodo.19166181	FIGURE 4. Acteon romanicus sp. nov. A1–A2. Holotype, NHMW 2016/0177/1298, Lăpugiu de Sus (Romania).B1–B2. Paratype, NHMW 2016/0177/1298a, Lăpugiu de Sus (Romania). C. Paratype, NHMW 2016/0177/1298b, Lăpugiu de Sus (Romania).	FIGURE 4. Acteon romanicus sp. nov. A1–A2. Holotype, NHMW 2016/0177/1298, Lăpugiu de Sus (Romania).B1–B2. Paratype, NHMW 2016/0177/1298a, Lăpugiu de Sus (Romania). C. Paratype, NHMW 2016/0177/1298b, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAB6761F2BD0636FDA5FDEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAB6761F2BD0636FDA5FDEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAC6760F2BD059EFD52F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166185/files/figure.png	https://doi.org/10.5281/zenodo.19166185	FIGURE 5. Acteon semistriatus (Férussac, 1822). A1–A2. NHMW 2018/0248/0484, Coşteiu de Sus (Romania). B1–B2. NHMW2018/0248/0484a, Coşteiu de Sus (Romania). C1–C2. NHMW 1859/0038/0121, Grund (Austria).	FIGURE 5. Acteon semistriatus (Férussac, 1822). A1–A2. NHMW 2018/0248/0484, Coşteiu de Sus (Romania). B1–B2. NHMW2018/0248/0484a, Coşteiu de Sus (Romania). C1–C2. NHMW 1859/0038/0121, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAC6760F2BD059EFD52F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAC6760F2BD059EFD52F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAC6760F2BD059EFD52F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAE6763F2BD0786FCEEF845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAE6763F2BD0786FCEEF845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAE6763F2BD0786FCEEF845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAF6762F2BD0786FBD2F830.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166188/files/figure.png	https://doi.org/10.5281/zenodo.19166188	FIGURE 6. A–F. Acteon trifasciatus sp. nov. A1–A2. Paratype, NHMW 2010/0004/1662, Bad Vöslau (Austria). B1–B2. Paratype, NHMW 2010/0004/1662a, Bad Vöslau (Austria). C1–C2. Paratype, NHMW 2010/0004/1662b, Bad Vöslau (Austria). D1–D3. Holotype, NHMW 1851/0002/0071, Grund (Austria). E1–E2. Paratype, NHMW 1864/0001/0619, Grund (Austria). F1– F2. Paratype, NHMW 1851/0002/0071, Grund (Austria). G. Rictaxis ursulae sp. nov. Paratype, NHMW 2007z0086/0000d, Grund (Austria).	FIGURE 6. A–F. Acteon trifasciatus sp. nov. A1–A2. Paratype, NHMW 2010/0004/1662, Bad Vöslau (Austria). B1–B2. Paratype, NHMW 2010/0004/1662a, Bad Vöslau (Austria). C1–C2. Paratype, NHMW 2010/0004/1662b, Bad Vöslau (Austria). D1–D3. Holotype, NHMW 1851/0002/0071, Grund (Austria). E1–E2. Paratype, NHMW 1864/0001/0619, Grund (Austria). F1– F2. Paratype, NHMW 1851/0002/0071, Grund (Austria). G. Rictaxis ursulae sp. nov. Paratype, NHMW 2007z0086/0000d, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAF6762F2BD0786FBD2F830.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAF6762F2BD0786FBD2F830.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFAF6762F2BD0786FBD2F830.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB0677CF2BD016AFE28F845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166190/files/figure.png	https://doi.org/10.5281/zenodo.19166190	FIGURE 7. A1–A2. Acteon viciani sp. nov., Holotype, HNHM INV 2025.97, Parádfürdő (Hungary).	FIGURE 7. A1–A2. Acteon viciani sp. nov., Holotype, HNHM INV 2025.97, Parádfürdő (Hungary).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB0677CF2BD016AFE28F845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB0677CF2BD016AFE28F845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB2677EF2BD0786FE0FFCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166192/files/figure.png	https://doi.org/10.5281/zenodo.19166192	FIGURE 8. Acteon vindobonensis sp. nov. A1–A2. Paratype, NHMW 1866/0040/0219, Baden Sooss (Austria). B1–B2. Holotype, NHMW 1855/0045/0523, Baden (Austria). C1–C2. Paratype, NHMW 1855/0045/0523a, Baden (Austria). D1–D2. Paratype, NHMW 1861/0001/0469, Bad Vöslau (Austria).	FIGURE 8. Acteon vindobonensis sp. nov. A1–A2. Paratype, NHMW 1866/0040/0219, Baden Sooss (Austria). B1–B2. Holotype, NHMW 1855/0045/0523, Baden (Austria). C1–C2. Paratype, NHMW 1855/0045/0523a, Baden (Austria). D1–D2. Paratype, NHMW 1861/0001/0469, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB2677EF2BD0786FE0FFCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB2677EF2BD0786FE0FFCC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB36779F2BD032FFBD4FA99.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166194/files/figure.png	https://doi.org/10.5281/zenodo.19166194	FIGURE 9. Crenilabium reussi (Boettger, 1904). A. Lectotype, SMF 363156, Coșteiu de Sus (Romania). B1–B2. NHMW 2020/0004/1666, Bad Vöslau (Austria). C1–C2. NHMW 2020/0004/1666a, Bad Vöslau (Austria).	FIGURE 9. Crenilabium reussi (Boettger, 1904). A. Lectotype, SMF 363156, Coșteiu de Sus (Romania). B1–B2. NHMW 2020/0004/1666, Bad Vöslau (Austria). C1–C2. NHMW 2020/0004/1666a, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB36779F2BD032FFBD4FA99.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB36779F2BD032FFBD4FA99.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB4677BF2BD0169FE56FD53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166196/files/figure.png	https://doi.org/10.5281/zenodo.19166196	FIGURE 10. A–B. Rictaxis austriacus sp. nov. A1–A2. Holotype, NHMW 2007z0086/0000e, Grund (Austria). B1–B2. Paratype, NHMW 2007z0086/0000f, Grund (Austria). C1–C2. Paratype, NHMW 2007z0086/0000g, Grund (Austria). D–E. Rictaxis siedli sp. nov. D1–D2. Holotype, NHMW 1997z0178/1214, Bad Vöslau (Austria). E1–E2. Paratype, NHMW 2010/0004/2663a, Bad Vöslau (Austria).	FIGURE 10. A–B. Rictaxis austriacus sp. nov. A1–A2. Holotype, NHMW 2007z0086/0000e, Grund (Austria). B1–B2. Paratype, NHMW 2007z0086/0000f, Grund (Austria). C1–C2. Paratype, NHMW 2007z0086/0000g, Grund (Austria). D–E. Rictaxis siedli sp. nov. D1–D2. Holotype, NHMW 1997z0178/1214, Bad Vöslau (Austria). E1–E2. Paratype, NHMW 2010/0004/2663a, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB4677BF2BD0169FE56FD53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB4677BF2BD0169FE56FD53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB6677AF2BD05C6FF0BFD53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166196/files/figure.png	https://doi.org/10.5281/zenodo.19166196	FIGURE 10. A–B. Rictaxis austriacus sp. nov. A1–A2. Holotype, NHMW 2007z0086/0000e, Grund (Austria). B1–B2. Paratype, NHMW 2007z0086/0000f, Grund (Austria). C1–C2. Paratype, NHMW 2007z0086/0000g, Grund (Austria). D–E. Rictaxis siedli sp. nov. D1–D2. Holotype, NHMW 1997z0178/1214, Bad Vöslau (Austria). E1–E2. Paratype, NHMW 2010/0004/2663a, Bad Vöslau (Austria).	FIGURE 10. A–B. Rictaxis austriacus sp. nov. A1–A2. Holotype, NHMW 2007z0086/0000e, Grund (Austria). B1–B2. Paratype, NHMW 2007z0086/0000f, Grund (Austria). C1–C2. Paratype, NHMW 2007z0086/0000g, Grund (Austria). D–E. Rictaxis siedli sp. nov. D1–D2. Holotype, NHMW 1997z0178/1214, Bad Vöslau (Austria). E1–E2. Paratype, NHMW 2010/0004/2663a, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB6677AF2BD05C6FF0BFD53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB6677AF2BD05C6FF0BFD53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB7677AF2BD05C6FE56F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166188/files/figure.png	https://doi.org/10.5281/zenodo.19166188	FIGURE 6. A–F. Acteon trifasciatus sp. nov. A1–A2. Paratype, NHMW 2010/0004/1662, Bad Vöslau (Austria). B1–B2. Paratype, NHMW 2010/0004/1662a, Bad Vöslau (Austria). C1–C2. Paratype, NHMW 2010/0004/1662b, Bad Vöslau (Austria). D1–D3. Holotype, NHMW 1851/0002/0071, Grund (Austria). E1–E2. Paratype, NHMW 1864/0001/0619, Grund (Austria). F1– F2. Paratype, NHMW 1851/0002/0071, Grund (Austria). G. Rictaxis ursulae sp. nov. Paratype, NHMW 2007z0086/0000d, Grund (Austria).	FIGURE 6. A–F. Acteon trifasciatus sp. nov. A1–A2. Paratype, NHMW 2010/0004/1662, Bad Vöslau (Austria). B1–B2. Paratype, NHMW 2010/0004/1662a, Bad Vöslau (Austria). C1–C2. Paratype, NHMW 2010/0004/1662b, Bad Vöslau (Austria). D1–D3. Holotype, NHMW 1851/0002/0071, Grund (Austria). E1–E2. Paratype, NHMW 1864/0001/0619, Grund (Austria). F1– F2. Paratype, NHMW 1851/0002/0071, Grund (Austria). G. Rictaxis ursulae sp. nov. Paratype, NHMW 2007z0086/0000d, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB7677AF2BD05C6FE56F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166198/files/figure.png	https://doi.org/10.5281/zenodo.19166198	FIGURE 11. Rictaxis ursulae sp. nov. A1–A2. Holotype, NHMW 2007z0086/0000, Grund (Austria). B1–B2. Paratype, NHMW 2007z0086/0000a, Grund (Austria). C1–C2. Paratype, NHMW 2007z0086/0000b, Grund (Austria). D1–D2. Paratype, NHMW 2007z00860/000c, Grund (Austria). E1–E2. Paratype, NHMW 2007z0086/0000d, Grund (Austria).	FIGURE 11. Rictaxis ursulae sp. nov. A1–A2. Holotype, NHMW 2007z0086/0000, Grund (Austria). B1–B2. Paratype, NHMW 2007z0086/0000a, Grund (Austria). C1–C2. Paratype, NHMW 2007z0086/0000b, Grund (Austria). D1–D2. Paratype, NHMW 2007z00860/000c, Grund (Austria). E1–E2. Paratype, NHMW 2007z0086/0000d, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB7677AF2BD05C6FE56F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB7677AF2BD05C6FE56F82C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB96776F2BD057BFAA8FE23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166202/files/figure.png	https://doi.org/10.5281/zenodo.19166202	FIGURE 12. A1–A2. Ringiculocosta costata (Eichwald, 1830), NHMW 1855/0045/0036, Steinebrunn (Austria). B1–B2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C1–C2, Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). D1–D2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Buituri (Romania). E1–E2. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO- 75000-E-57959, Grund (Austria).	FIGURE 12. A1–A2. Ringiculocosta costata (Eichwald, 1830), NHMW 1855/0045/0036, Steinebrunn (Austria). B1–B2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C1–C2, Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). D1–D2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Buituri (Romania). E1–E2. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO- 75000-E-57959, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB96776F2BD057BFAA8FE23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166204/files/figure.png	https://doi.org/10.5281/zenodo.19166204	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB96776F2BD057BFAA8FE23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFB96776F2BD057BFAA8FE23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBB6771F2BD052EFB60FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166202/files/figure.png	https://doi.org/10.5281/zenodo.19166202	FIGURE 12. A1–A2. Ringiculocosta costata (Eichwald, 1830), NHMW 1855/0045/0036, Steinebrunn (Austria). B1–B2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C1–C2, Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). D1–D2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Buituri (Romania). E1–E2. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO- 75000-E-57959, Grund (Austria).	FIGURE 12. A1–A2. Ringiculocosta costata (Eichwald, 1830), NHMW 1855/0045/0036, Steinebrunn (Austria). B1–B2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C1–C2, Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). D1–D2. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Buituri (Romania). E1–E2. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO- 75000-E-57959, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBB6771F2BD052EFB60FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166204/files/figure.png	https://doi.org/10.5281/zenodo.19166204	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBB6771F2BD052EFB60FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBB6771F2BD052EFB60FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBC6773F2BD01FBFF0BFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166206/files/figure.png	https://doi.org/10.5281/zenodo.19166206	FIGURE 14. Ringicula minor (Grateloup, 1838). A1–A2. NHMW 1862/0001/0521, Buituri (Romania). B1–B2. NHMW 1862/0001/0521a, Buituri (Romania). C1–C2. NHMW 1850/0002/0012, Grund (Austria). D1–D2. NHMW 1862/0001/0521b, Buituri (Romania). E1–E2. NHMW-ZOO-MO-75000-E-57962, Bad Vöslau (Austria). F–J. Ringicula daisyae sp. nov. F1–F2. Holotype, NHMW 2025/0245/0001, Nemeşeşti (Romania). G1–G2. Paratype, NHMW 2025/0245/0002, Nemeşeşti (Romania). H1–H2. Paratype, NHMW 2025/0245/0003, Nemeşeşti (Romania). I1–I2. Paratype, NHMW 2025/0245/0004, Nemeşeşti (Romania). J1–J2. Paratype, NHMW 2025/0245/0005, Nemeşeşti (Romania).	FIGURE 14. Ringicula minor (Grateloup, 1838). A1–A2. NHMW 1862/0001/0521, Buituri (Romania). B1–B2. NHMW 1862/0001/0521a, Buituri (Romania). C1–C2. NHMW 1850/0002/0012, Grund (Austria). D1–D2. NHMW 1862/0001/0521b, Buituri (Romania). E1–E2. NHMW-ZOO-MO-75000-E-57962, Bad Vöslau (Austria). F–J. Ringicula daisyae sp. nov. F1–F2. Holotype, NHMW 2025/0245/0001, Nemeşeşti (Romania). G1–G2. Paratype, NHMW 2025/0245/0002, Nemeşeşti (Romania). H1–H2. Paratype, NHMW 2025/0245/0003, Nemeşeşti (Romania). I1–I2. Paratype, NHMW 2025/0245/0004, Nemeşeşti (Romania). J1–J2. Paratype, NHMW 2025/0245/0005, Nemeşeşti (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBC6773F2BD01FBFF0BFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBC6773F2BD01FBFF0BFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBE674DF2BD0397FF37FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166204/files/figure.png	https://doi.org/10.5281/zenodo.19166204	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBE674DF2BD0397FF37FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166208/files/figure.png	https://doi.org/10.5281/zenodo.19166208	FIGURE 15. A–D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023. A1–A2. NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). B1–B2. NHMW 1870/0014/0001a, Golubytsya (Hołubica) (Ukraine). C1–C2. NHMW 1870/0014/0001b, Golubytsya (Hołubica) (Ukraine). D1–D2. NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E1–E3. NHMW-ZOO- MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria). F1–F3. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria).	FIGURE 15. A–D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023. A1–A2. NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). B1–B2. NHMW 1870/0014/0001a, Golubytsya (Hołubica) (Ukraine). C1–C2. NHMW 1870/0014/0001b, Golubytsya (Hołubica) (Ukraine). D1–D2. NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E1–E3. NHMW-ZOO- MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria). F1–F3. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBE674DF2BD0397FF37FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFBE674DF2BD0397FF37FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF80674CF2BD0637FE3DFAF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166204/files/figure.png	https://doi.org/10.5281/zenodo.19166204	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF80674CF2BD0637FE3DFAF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166208/files/figure.png	https://doi.org/10.5281/zenodo.19166208	FIGURE 15. A–D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023. A1–A2. NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). B1–B2. NHMW 1870/0014/0001a, Golubytsya (Hołubica) (Ukraine). C1–C2. NHMW 1870/0014/0001b, Golubytsya (Hołubica) (Ukraine). D1–D2. NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E1–E3. NHMW-ZOO- MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria). F1–F3. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria).	FIGURE 15. A–D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023. A1–A2. NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). B1–B2. NHMW 1870/0014/0001a, Golubytsya (Hołubica) (Ukraine). C1–C2. NHMW 1870/0014/0001b, Golubytsya (Hołubica) (Ukraine). D1–D2. NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E1–E3. NHMW-ZOO- MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria). F1–F3. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF80674CF2BD0637FE3DFAF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF80674CF2BD0637FE3DFAF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF81674EF2BD0263FE13FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166210/files/figure.png	https://doi.org/10.5281/zenodo.19166210	FIGURE 16. Ringicula hoernesii Seguenza, 1881. A1–A2. NHMW 1846/0037/0090, Baden (Austria). B1–B2. NHMW 1846/0037/0090a, Baden (Austria). C1–C2. NHMW 1855/0045/0866, Bad Vöslau (Austria). D1–D2. NHMW 1862/0001/0195, Baden (Austria).	FIGURE 16. Ringicula hoernesii Seguenza, 1881. A1–A2. NHMW 1846/0037/0090, Baden (Austria). B1–B2. NHMW 1846/0037/0090a, Baden (Austria). C1–C2. NHMW 1855/0045/0866, Bad Vöslau (Austria). D1–D2. NHMW 1862/0001/0195, Baden (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF81674EF2BD0263FE13FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF81674EF2BD0263FE13FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF836748F2BD05A3FF37FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166204/files/figure.png	https://doi.org/10.5281/zenodo.19166204	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	FIGURE 13. A. Ringiculocosta cancellarioides (Seguenza, 1881), NHMW-ZOO-MO-75000-E-3283, Bad Vöslau (Austria). B. Ringiculocosta costata (Eichwald, 1830), NHMW 1863/0015/0616, Niederleis (Austria). C. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023, NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria). F. Ringicula laevigata (Eichwald, 1830), NHMW 1949/0005/0009, Steinebrunn (Austria). G. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57964, Steinebrunn (Austria). H. Ringicula laevigata (Eichwald, 1830), NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF836748F2BD05A3FF37FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166212/files/figure.png	https://doi.org/10.5281/zenodo.19166212	FIGURE 17. Ringicula laevigata (Eichwald, 1830). A1–A2. NHMW 1949/0005/0009, Steinebrunn (Austria). B1–B2. NHMW 1949/0005/0009, Steinebrunn (Austria). C1–C2. NHMW-ZOO-MO-75000-E-57969, Grund (Austria). D1–D2. NHMW-ZOO- MO-75000-E-57964, Steinebrunn (Austria). E1–E2. NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria). F1–F2. NHMW 2018/0248/0490, Coşteiu de Sus (Romania). G1–G2. NHMW 2018/0248/0490, Coşteiu de Sus (Romania). H. NHMW 2018/0248/0490, Coşteiu de Sus (Romania).	FIGURE 17. Ringicula laevigata (Eichwald, 1830). A1–A2. NHMW 1949/0005/0009, Steinebrunn (Austria). B1–B2. NHMW 1949/0005/0009, Steinebrunn (Austria). C1–C2. NHMW-ZOO-MO-75000-E-57969, Grund (Austria). D1–D2. NHMW-ZOO- MO-75000-E-57964, Steinebrunn (Austria). E1–E2. NHMW-ZOO-MO-75000-E-57966, Ritzing (Austria). F1–F2. NHMW 2018/0248/0490, Coşteiu de Sus (Romania). G1–G2. NHMW 2018/0248/0490, Coşteiu de Sus (Romania). H. NHMW 2018/0248/0490, Coşteiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF836748F2BD05A3FF37FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF836748F2BD05A3FF37FA17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF85674AF2BD0103FDE7FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166206/files/figure.png	https://doi.org/10.5281/zenodo.19166206	FIGURE 14. Ringicula minor (Grateloup, 1838). A1–A2. NHMW 1862/0001/0521, Buituri (Romania). B1–B2. NHMW 1862/0001/0521a, Buituri (Romania). C1–C2. NHMW 1850/0002/0012, Grund (Austria). D1–D2. NHMW 1862/0001/0521b, Buituri (Romania). E1–E2. NHMW-ZOO-MO-75000-E-57962, Bad Vöslau (Austria). F–J. Ringicula daisyae sp. nov. F1–F2. Holotype, NHMW 2025/0245/0001, Nemeşeşti (Romania). G1–G2. Paratype, NHMW 2025/0245/0002, Nemeşeşti (Romania). H1–H2. Paratype, NHMW 2025/0245/0003, Nemeşeşti (Romania). I1–I2. Paratype, NHMW 2025/0245/0004, Nemeşeşti (Romania). J1–J2. Paratype, NHMW 2025/0245/0005, Nemeşeşti (Romania).	FIGURE 14. Ringicula minor (Grateloup, 1838). A1–A2. NHMW 1862/0001/0521, Buituri (Romania). B1–B2. NHMW 1862/0001/0521a, Buituri (Romania). C1–C2. NHMW 1850/0002/0012, Grund (Austria). D1–D2. NHMW 1862/0001/0521b, Buituri (Romania). E1–E2. NHMW-ZOO-MO-75000-E-57962, Bad Vöslau (Austria). F–J. Ringicula daisyae sp. nov. F1–F2. Holotype, NHMW 2025/0245/0001, Nemeşeşti (Romania). G1–G2. Paratype, NHMW 2025/0245/0002, Nemeşeşti (Romania). H1–H2. Paratype, NHMW 2025/0245/0003, Nemeşeşti (Romania). I1–I2. Paratype, NHMW 2025/0245/0004, Nemeşeşti (Romania). J1–J2. Paratype, NHMW 2025/0245/0005, Nemeşeşti (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF85674AF2BD0103FDE7FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166208/files/figure.png	https://doi.org/10.5281/zenodo.19166208	FIGURE 15. A–D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023. A1–A2. NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). B1–B2. NHMW 1870/0014/0001a, Golubytsya (Hołubica) (Ukraine). C1–C2. NHMW 1870/0014/0001b, Golubytsya (Hołubica) (Ukraine). D1–D2. NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E1–E3. NHMW-ZOO- MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria). F1–F3. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria).	FIGURE 15. A–D. Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023. A1–A2. NHMW 1870/0014/0001, Golubytsya (Hołubica) (Ukraine). B1–B2. NHMW 1870/0014/0001a, Golubytsya (Hołubica) (Ukraine). C1–C2. NHMW 1870/0014/0001b, Golubytsya (Hołubica) (Ukraine). D1–D2. NHMW-ZOO-MO-75000-E-57957, Steinebrunn (Austria). E1–E3. NHMW-ZOO- MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria). F1–F3. Ringicula dentata Berger, 1954, Holotype, NHMW-ZOO-MO-75000-E-57968, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF85674AF2BD0103FDE7FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF85674AF2BD0103FDE7FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF886745F2BD0608FEB6FABD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166216/files/figure.png	https://doi.org/10.5281/zenodo.19166216	FIGURE 18. A. Bulla hungarica Boettger, 1907. lectotype, SMF 363157, Coșteiu de Sus (Romania). B1–B3. Bulla? elalbanii sp. nov., holotype, NHMW 1859/0045/0420, Oles’ko (Lviv Region, Ukraine).	FIGURE 18. A. Bulla hungarica Boettger, 1907. lectotype, SMF 363157, Coșteiu de Sus (Romania). B1–B3. Bulla? elalbanii sp. nov., holotype, NHMW 1859/0045/0420, Oles’ko (Lviv Region, Ukraine).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF886745F2BD0608FEB6FABD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF886745F2BD0608FEB6FABD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF896744F2BD0786FE86FBBE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166216/files/figure.png	https://doi.org/10.5281/zenodo.19166216	FIGURE 18. A. Bulla hungarica Boettger, 1907. lectotype, SMF 363157, Coșteiu de Sus (Romania). B1–B3. Bulla? elalbanii sp. nov., holotype, NHMW 1859/0045/0420, Oles’ko (Lviv Region, Ukraine).	FIGURE 18. A. Bulla hungarica Boettger, 1907. lectotype, SMF 363157, Coșteiu de Sus (Romania). B1–B3. Bulla? elalbanii sp. nov., holotype, NHMW 1859/0045/0420, Oles’ko (Lviv Region, Ukraine).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF896741F2BD01A0FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166218/files/figure.png	https://doi.org/10.5281/zenodo.19166218	FIGURE 19. A–E. Pyrunculus elongatus (Eichwald, 1830). A1–A3. NHMW 2010/0004/1656, Bad Vöslau (Austria). B1–B3. NHMW 2010/0004/1656a, Bad Vöslau (Austria). C1–C2. NHMW 1869/0001/0775, Coşteiu de Sus (Romania). D1–D2. NHMW 2010/0004/1656b, Bad Vöslau (Austria).E1–E3. NHMW-ZOO-MO-75000-E-58763, Gainfarn (Austria). F. Pyrunculus frequens (Boettger, 1902), NHMW 1865/0015/0155, Lysice (Czech Republic).	FIGURE 19. A–E. Pyrunculus elongatus (Eichwald, 1830). A1–A3. NHMW 2010/0004/1656, Bad Vöslau (Austria). B1–B3. NHMW 2010/0004/1656a, Bad Vöslau (Austria). C1–C2. NHMW 1869/0001/0775, Coşteiu de Sus (Romania). D1–D2. NHMW 2010/0004/1656b, Bad Vöslau (Austria).E1–E3. NHMW-ZOO-MO-75000-E-58763, Gainfarn (Austria). F. Pyrunculus frequens (Boettger, 1902), NHMW 1865/0015/0155, Lysice (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF896741F2BD01A0FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF896741F2BD01A0FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF896741F2BD01A0FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8C6740F2BD046CFBD8FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166218/files/figure.png	https://doi.org/10.5281/zenodo.19166218	FIGURE 19. A–E. Pyrunculus elongatus (Eichwald, 1830). A1–A3. NHMW 2010/0004/1656, Bad Vöslau (Austria). B1–B3. NHMW 2010/0004/1656a, Bad Vöslau (Austria). C1–C2. NHMW 1869/0001/0775, Coşteiu de Sus (Romania). D1–D2. NHMW 2010/0004/1656b, Bad Vöslau (Austria).E1–E3. NHMW-ZOO-MO-75000-E-58763, Gainfarn (Austria). F. Pyrunculus frequens (Boettger, 1902), NHMW 1865/0015/0155, Lysice (Czech Republic).	FIGURE 19. A–E. Pyrunculus elongatus (Eichwald, 1830). A1–A3. NHMW 2010/0004/1656, Bad Vöslau (Austria). B1–B3. NHMW 2010/0004/1656a, Bad Vöslau (Austria). C1–C2. NHMW 1869/0001/0775, Coşteiu de Sus (Romania). D1–D2. NHMW 2010/0004/1656b, Bad Vöslau (Austria).E1–E3. NHMW-ZOO-MO-75000-E-58763, Gainfarn (Austria). F. Pyrunculus frequens (Boettger, 1902), NHMW 1865/0015/0155, Lysice (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8C6740F2BD046CFBD8FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166220/files/figure.png	https://doi.org/10.5281/zenodo.19166220	FIGURE 20. Pyrunculus frequens (Boettger, 1902). A1–A2. NHMW 1865/0015/0155, Lysice (Czech Republic). B1–B2. NHMW- ZOO-MO-75000-E-47049, Lysice (Czech Republic). C. Lectotype. SMF 363160, Coşteiu de Sus (Romania).	FIGURE 20. Pyrunculus frequens (Boettger, 1902). A1–A2. NHMW 1865/0015/0155, Lysice (Czech Republic). B1–B2. NHMW- ZOO-MO-75000-E-47049, Lysice (Czech Republic). C. Lectotype. SMF 363160, Coşteiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8C6740F2BD046CFBD8FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8C6740F2BD046CFBD8FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8D6742F2BD0345FD60FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166222/files/figure.png	https://doi.org/10.5281/zenodo.19166222	FIGURE 21. Pyrunculus latesulcatus (Boettger, 1902). A1–A3. NHMW 1855/0045/0258, Steinebrunn (Austria). B1–B3. NHMW 1855/0045/0258a, Steinebrunn (Austria). C. Lectotype, SMF 363159, Coşteiu de Sus (Romania). D1–D3. NHMW- ZOO-MO-75000-E-47032, Lăpugiu de Sus (Romania). E1–E2. NHMW 2010/0004/1655, Bad Vöslau (Austria). F1–F2. NHMW 1855/0045/0258a, Steinebrunn (Austria).	FIGURE 21. Pyrunculus latesulcatus (Boettger, 1902). A1–A3. NHMW 1855/0045/0258, Steinebrunn (Austria). B1–B3. NHMW 1855/0045/0258a, Steinebrunn (Austria). C. Lectotype, SMF 363159, Coşteiu de Sus (Romania). D1–D3. NHMW- ZOO-MO-75000-E-47032, Lăpugiu de Sus (Romania). E1–E2. NHMW 2010/0004/1655, Bad Vöslau (Austria). F1–F2. NHMW 1855/0045/0258a, Steinebrunn (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8D6742F2BD0345FD60FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8D6742F2BD0345FD60FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8D6742F2BD0345FD60FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F6742F2BD0683FD2DF9DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166224/files/figure.png	https://doi.org/10.5281/zenodo.19166224	FIGURE 22. Pyrunculus leuchsi (Berger, 1949). A1–A2. Holotype, NHMW-ZOO-MO-75000-E-11815A, Bad Vöslau (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-11815, Bad Vöslau (Austria). C1–C3. Paratype, NHMW-ZOO-MO-75000-E- 11815, Bad Vöslau (Austria). D1–D3. NHMW-ZOO-MO-75000-E-3308, Bad Vöslau (Austria). E1–E3. NHMW-ZOO-MO- 75000-E-3308, Bad Vöslau (Austria).	FIGURE 22. Pyrunculus leuchsi (Berger, 1949). A1–A2. Holotype, NHMW-ZOO-MO-75000-E-11815A, Bad Vöslau (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-11815, Bad Vöslau (Austria). C1–C3. Paratype, NHMW-ZOO-MO-75000-E- 11815, Bad Vöslau (Austria). D1–D3. NHMW-ZOO-MO-75000-E-3308, Bad Vöslau (Austria). E1–E3. NHMW-ZOO-MO- 75000-E-3308, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F6742F2BD0683FD2DF9DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F6742F2BD0683FD2DF9DB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F675CF2BD014FFE28FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166226/files/figure.png	https://doi.org/10.5281/zenodo.19166226	FIGURE 23. Pyrunculus microromanicus sp. nov. A1–A2. Holotype, NHMW 2018/0248/0475, Coşteiu de Sus (Romania). B1– B2. Paratype, NHMW 2018/0248/0475a, Coşteiu de Sus (Romania). C. Paratype, NHMW-ZOO-MO-75000-E-147038, Lăpugiu de Sus (Romania).	FIGURE 23. Pyrunculus microromanicus sp. nov. A1–A2. Holotype, NHMW 2018/0248/0475, Coşteiu de Sus (Romania). B1– B2. Paratype, NHMW 2018/0248/0475a, Coşteiu de Sus (Romania). C. Paratype, NHMW-ZOO-MO-75000-E-147038, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F675CF2BD014FFE28FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166228/files/figure.png	https://doi.org/10.5281/zenodo.19166228	FIGURE 24. A–F. Pyrunculus parangistoma (Boettger, 1902). A1–A2. NHMW 1855/0045/0258, Steinebrunn (Austria). B1–B3. NHMW 1855/0045/0258a, Steinebrunn (Austria). C1–C2. NHMW 1855/0045/0258b, Steinebrunn (Austria). D1–D3. NHMW- ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E1–E2. RGM 776934, Seyithasan (Turkey), Serravallian). F. Lectotype, SMF 363161 - 12.2551a, Coşteiu de Sus (Romania). G. Pyrunculus microromanicus sp. nov., holotype, NHMW 2018/0248/0475, Coşteiu de Sus (Romania). H. Pyrunculus microromanicus sp. nov., paratype, NHMW 2018/0248/0475a, Coşteiu de Sus (Romania).	FIGURE 24. A–F. Pyrunculus parangistoma (Boettger, 1902). A1–A2. NHMW 1855/0045/0258, Steinebrunn (Austria). B1–B3. NHMW 1855/0045/0258a, Steinebrunn (Austria). C1–C2. NHMW 1855/0045/0258b, Steinebrunn (Austria). D1–D3. NHMW- ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E1–E2. RGM 776934, Seyithasan (Turkey), Serravallian). F. Lectotype, SMF 363161 - 12.2551a, Coşteiu de Sus (Romania). G. Pyrunculus microromanicus sp. nov., holotype, NHMW 2018/0248/0475, Coşteiu de Sus (Romania). H. Pyrunculus microromanicus sp. nov., paratype, NHMW 2018/0248/0475a, Coşteiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F675CF2BD014FFE28FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF8F675CF2BD014FFE28FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF91675EF2BD0246FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166228/files/figure.png	https://doi.org/10.5281/zenodo.19166228	FIGURE 24. A–F. Pyrunculus parangistoma (Boettger, 1902). A1–A2. NHMW 1855/0045/0258, Steinebrunn (Austria). B1–B3. NHMW 1855/0045/0258a, Steinebrunn (Austria). C1–C2. NHMW 1855/0045/0258b, Steinebrunn (Austria). D1–D3. NHMW- ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E1–E2. RGM 776934, Seyithasan (Turkey), Serravallian). F. Lectotype, SMF 363161 - 12.2551a, Coşteiu de Sus (Romania). G. Pyrunculus microromanicus sp. nov., holotype, NHMW 2018/0248/0475, Coşteiu de Sus (Romania). H. Pyrunculus microromanicus sp. nov., paratype, NHMW 2018/0248/0475a, Coşteiu de Sus (Romania).	FIGURE 24. A–F. Pyrunculus parangistoma (Boettger, 1902). A1–A2. NHMW 1855/0045/0258, Steinebrunn (Austria). B1–B3. NHMW 1855/0045/0258a, Steinebrunn (Austria). C1–C2. NHMW 1855/0045/0258b, Steinebrunn (Austria). D1–D3. NHMW- ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E1–E2. RGM 776934, Seyithasan (Turkey), Serravallian). F. Lectotype, SMF 363161 - 12.2551a, Coşteiu de Sus (Romania). G. Pyrunculus microromanicus sp. nov., holotype, NHMW 2018/0248/0475, Coşteiu de Sus (Romania). H. Pyrunculus microromanicus sp. nov., paratype, NHMW 2018/0248/0475a, Coşteiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF91675EF2BD0246FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF91675EF2BD0246FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF91675EF2BD0246FED3FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF936759F2BD049BFCCCFEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166230/files/figure.png	https://doi.org/10.5281/zenodo.19166230	FIGURE 25.A–D. Pyrunculus szobiensis sp. nov. A1–A3. Holotype, NHMW 186100010507, Szob (Hungary). B1–B2. Paratype, NHMW 186100010507a, Szob (Hungary). C1–C2. Paratype, NHMW 186100010507b, Szob (Hungary). D1–D3. Paratype, NHMW-ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E–F. Pyrunculus zuschini sp. nov. E. Holotype, NHMW-ZOO-MO- 75000-E-42898A, Vienna/Ottakring (Austria). F. Paratype, NHMW 1859/0045/0604, Lăpugiu de Sus (Romania).	FIGURE 25.A–D. Pyrunculus szobiensis sp. nov. A1–A3. Holotype, NHMW 186100010507, Szob (Hungary). B1–B2. Paratype, NHMW 186100010507a, Szob (Hungary). C1–C2. Paratype, NHMW 186100010507b, Szob (Hungary). D1–D3. Paratype, NHMW-ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E–F. Pyrunculus zuschini sp. nov. E. Holotype, NHMW-ZOO-MO- 75000-E-42898A, Vienna/Ottakring (Austria). F. Paratype, NHMW 1859/0045/0604, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF936759F2BD049BFCCCFEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF936759F2BD049BFCCCFEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF936759F2BD049BFCCCFEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF94675BF2BD0196FCB1FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166230/files/figure.png	https://doi.org/10.5281/zenodo.19166230	FIGURE 25.A–D. Pyrunculus szobiensis sp. nov. A1–A3. Holotype, NHMW 186100010507, Szob (Hungary). B1–B2. Paratype, NHMW 186100010507a, Szob (Hungary). C1–C2. Paratype, NHMW 186100010507b, Szob (Hungary). D1–D3. Paratype, NHMW-ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E–F. Pyrunculus zuschini sp. nov. E. Holotype, NHMW-ZOO-MO- 75000-E-42898A, Vienna/Ottakring (Austria). F. Paratype, NHMW 1859/0045/0604, Lăpugiu de Sus (Romania).	FIGURE 25.A–D. Pyrunculus szobiensis sp. nov. A1–A3. Holotype, NHMW 186100010507, Szob (Hungary). B1–B2. Paratype, NHMW 186100010507a, Szob (Hungary). C1–C2. Paratype, NHMW 186100010507b, Szob (Hungary). D1–D3. Paratype, NHMW-ZOO-MO-75000-E-42592, Bad Vöslau (Austria). E–F. Pyrunculus zuschini sp. nov. E. Holotype, NHMW-ZOO-MO- 75000-E-42898A, Vienna/Ottakring (Austria). F. Paratype, NHMW 1859/0045/0604, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF94675BF2BD0196FCB1FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166232/files/figure.png	https://doi.org/10.5281/zenodo.19166232	FIGURE 26. Pyrunculus zuschini sp. nov. A1–A2. Holotype, NHMW-ZOO-MO-75000-E-42898A, Vienna/Ottakring (Austria). B1–B2. Paratype, NHMW 1859/0045/0604, Lăpugiu de Sus (Romania).	FIGURE 26. Pyrunculus zuschini sp. nov. A1–A2. Holotype, NHMW-ZOO-MO-75000-E-42898A, Vienna/Ottakring (Austria). B1–B2. Paratype, NHMW 1859/0045/0604, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF94675BF2BD0196FCB1FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF94675BF2BD0196FCB1FF03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF96675BF2BD06E3FB53F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166234/files/figure.png	https://doi.org/10.5281/zenodo.19166234	FIGURE 27. Retusa mystica sp. nov. A1–A3. Paratype, NHMW 1855/0045/0260c, Steinebrunn (Austria). B1–B3. Paratype, NHMW 1973/1615/0124, Lăpugiu de Sus (Romania). C1–C3. Paratype, NHMW 1855/0045/0260d, Steinebrunn (Austria). D1– D3. Holotype, NHMW 1855/0043/0086, Lăpugiu de Sus (Romania). E1–E2. Paratype, NHMW 1997z0178/1212, Bad Vöslau (Austria).	FIGURE 27. Retusa mystica sp. nov. A1–A3. Paratype, NHMW 1855/0045/0260c, Steinebrunn (Austria). B1–B3. Paratype, NHMW 1973/1615/0124, Lăpugiu de Sus (Romania). C1–C3. Paratype, NHMW 1855/0045/0260d, Steinebrunn (Austria). D1– D3. Holotype, NHMW 1855/0043/0086, Lăpugiu de Sus (Romania). E1–E2. Paratype, NHMW 1997z0178/1212, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF96675BF2BD06E3FB53F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF96675BF2BD06E3FB53F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF96675BF2BD06E3FB53F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF976754F2BD0185FF37FC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166236/files/figure.png	https://doi.org/10.5281/zenodo.19166236	FIGURE 28. Retusa promamillata Boettger, 1907. A1–A3. NHMW-ZOO-MO-75000-E-47025, Guntersdorf (Austria). B1– B3. NHMW-ZOO-MO-75000-E-47026, Grund (Austria). C. Lectotype, SMF 363158, Coșteiu de Sus (Romania). D1–D2. NHMW 2018/0248/0477, Coşteiu de Sus (Romania). E1–E2. NHMW 1864/0001/0224, Gainfarn (Austria). F1–F3. NHMW 1864/0001/0224a, Gainfarn (Austria).	FIGURE 28. Retusa promamillata Boettger, 1907. A1–A3. NHMW-ZOO-MO-75000-E-47025, Guntersdorf (Austria). B1– B3. NHMW-ZOO-MO-75000-E-47026, Grund (Austria). C. Lectotype, SMF 363158, Coșteiu de Sus (Romania). D1–D2. NHMW 2018/0248/0477, Coşteiu de Sus (Romania). E1–E2. NHMW 1864/0001/0224, Gainfarn (Austria). F1–F3. NHMW 1864/0001/0224a, Gainfarn (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF976754F2BD0185FF37FC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF976754F2BD0185FF37FC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF976754F2BD0185FF37FC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF996756F2BD04E6FB44FEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166238/files/figure.png	https://doi.org/10.5281/zenodo.19166238	FIGURE 29. A–D. Retusa sarmatica Berger, 1953. A1–A3. NHMW 1855/0045/0260, Steinebrunn (Austria), Badenian. B1–B3. NHMW 1870/0037/0063, Ritzing (Austria). C1–C2. Holotype, NHMW-ZOO-MO-75000-E-47021A, Vienna/Hernals (Austria), Sarmatian. NHMW 1870/0037/0063, Ritzing (Austria), Badenian. D. NHMW-ZOO-MO-75000-E-47018, Hölles (Austria), Sarmatian. E–F. Retusa sinzovi (Kolesnikov, 1935). E1–E3. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic). F1–F2. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic).	FIGURE 29. A–D. Retusa sarmatica Berger, 1953. A1–A3. NHMW 1855/0045/0260, Steinebrunn (Austria), Badenian. B1–B3. NHMW 1870/0037/0063, Ritzing (Austria). C1–C2. Holotype, NHMW-ZOO-MO-75000-E-47021A, Vienna/Hernals (Austria), Sarmatian. NHMW 1870/0037/0063, Ritzing (Austria), Badenian. D. NHMW-ZOO-MO-75000-E-47018, Hölles (Austria), Sarmatian. E–F. Retusa sinzovi (Kolesnikov, 1935). E1–E3. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic). F1–F2. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF996756F2BD04E6FB44FEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF996756F2BD04E6FB44FEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF996756F2BD04E6FB44FEB3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9B6751F2BD06A6FA5BFF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166238/files/figure.png	https://doi.org/10.5281/zenodo.19166238	FIGURE 29. A–D. Retusa sarmatica Berger, 1953. A1–A3. NHMW 1855/0045/0260, Steinebrunn (Austria), Badenian. B1–B3. NHMW 1870/0037/0063, Ritzing (Austria). C1–C2. Holotype, NHMW-ZOO-MO-75000-E-47021A, Vienna/Hernals (Austria), Sarmatian. NHMW 1870/0037/0063, Ritzing (Austria), Badenian. D. NHMW-ZOO-MO-75000-E-47018, Hölles (Austria), Sarmatian. E–F. Retusa sinzovi (Kolesnikov, 1935). E1–E3. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic). F1–F2. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic).	FIGURE 29. A–D. Retusa sarmatica Berger, 1953. A1–A3. NHMW 1855/0045/0260, Steinebrunn (Austria), Badenian. B1–B3. NHMW 1870/0037/0063, Ritzing (Austria). C1–C2. Holotype, NHMW-ZOO-MO-75000-E-47021A, Vienna/Hernals (Austria), Sarmatian. NHMW 1870/0037/0063, Ritzing (Austria), Badenian. D. NHMW-ZOO-MO-75000-E-47018, Hölles (Austria), Sarmatian. E–F. Retusa sinzovi (Kolesnikov, 1935). E1–E3. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic). F1–F2. NHMW-ZOO-MO-75000-E-47058, Bílovice u Uherského Hradiště (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9B6751F2BD06A6FA5BFF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9B6751F2BD06A6FA5BFF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9B6751F2BD06A6FA5BFF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9C6753F2BD065AFF26FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166240/files/figure.png	https://doi.org/10.5281/zenodo.19166240	FIGURE 30. Retusa truncatula (Bruguière, 1792).A1–A3. NHMW 2018/0248/0476, Coșteiu de Sus (Romania). B1–B2. NHMW- ZOO-MO-75000-E-41686, Steinebrunn (Austria). C1–C3. NHMW-ZOO-MO-75000-E-47002, Grund (Austria), holotype of Retusa grundensis Berger, 1953. D1–D3. NHMW 2025/0152/0001, Bad Vöslau (Austria). E1–E3. NHMW 2025/0152/0002, Bad Vöslau (Austria).	FIGURE 30. Retusa truncatula (Bruguière, 1792).A1–A3. NHMW 2018/0248/0476, Coșteiu de Sus (Romania). B1–B2. NHMW- ZOO-MO-75000-E-41686, Steinebrunn (Austria). C1–C3. NHMW-ZOO-MO-75000-E-47002, Grund (Austria), holotype of Retusa grundensis Berger, 1953. D1–D3. NHMW 2025/0152/0001, Bad Vöslau (Austria). E1–E3. NHMW 2025/0152/0002, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9C6753F2BD065AFF26FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9C6753F2BD065AFF26FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9C6753F2BD065AFF26FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9E6752F2BD0372FDF1F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166242/files/figure.png	https://doi.org/10.5281/zenodo.19166242	FIGURE 31. Retusa vindobonensis (Berger,1949).A1–A3. Holotype,NHMW-ZOO-MO-75000-E-3311A,Bad Vöslau(Austria). B1–B3. NHMW-ZOO-MO-75000-E-7041, Grund (Austria). C1–C2. NHMW-ZOO-MO-75000-E-7041, Grund (Austria).	FIGURE 31. Retusa vindobonensis (Berger,1949).A1–A3. Holotype,NHMW-ZOO-MO-75000-E-3311A,Bad Vöslau(Austria). B1–B3. NHMW-ZOO-MO-75000-E-7041, Grund (Austria). C1–C2. NHMW-ZOO-MO-75000-E-7041, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9E6752F2BD0372FDF1F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9E6752F2BD0372FDF1F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9F672DF2BD01E7FE54F9B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166244/files/figure.png	https://doi.org/10.5281/zenodo.19166244	FIGURE 32. Retusa voeslauensis (Berger, 1953). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-3318A, Bad Vöslau (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3318, Bad Vöslau (Austria). C1–C3. NHMW-ZOO-MO-75000-E-3318, Bad Vöslau (Austria). D1–D2. NHMW-ZOO-MO-75000-E-3318, Steinebrunn (Austria). E1–E3. NHMW-ZOO-MO-75000-E-3318, Steinebrunn (Austria).	FIGURE 32. Retusa voeslauensis (Berger, 1953). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-3318A, Bad Vöslau (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3318, Bad Vöslau (Austria). C1–C3. NHMW-ZOO-MO-75000-E-3318, Bad Vöslau (Austria). D1–D2. NHMW-ZOO-MO-75000-E-3318, Steinebrunn (Austria). E1–E3. NHMW-ZOO-MO-75000-E-3318, Steinebrunn (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9F672DF2BD01E7FE54F9B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9F672DF2BD01E7FE54F9B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FF9F672DF2BD01E7FE54F9B1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE1672FF2BD01CEFB89F845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166246/files/figure.png	https://doi.org/10.5281/zenodo.19166246	FIGURE 33. Volvulella acuminata (Bruguière, 1792). A1–A2. NHMW 2010/0004/1651, Bad Vöslau (Austria). B1–B2. NHMW 1876/0011/0171, Lăpugiu de Sus (Romania). C1–C2. NHMW 2010/0004/1652, Bad Vöslau (Austria).	FIGURE 33. Volvulella acuminata (Bruguière, 1792). A1–A2. NHMW 2010/0004/1651, Bad Vöslau (Austria). B1–B2. NHMW 1876/0011/0171, Lăpugiu de Sus (Romania). C1–C2. NHMW 2010/0004/1652, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE1672FF2BD01CEFB89F845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE1672FF2BD01CEFB89F845.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE36728F2BD049AFED3F89B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166250/files/figure.png	https://doi.org/10.5281/zenodo.19166250	FIGURE 34. Acteocina fusiformis (Boettger, 1907). A1–A2. NHMW 1876/0011/0170, Lăpugiu de Sus (Romania). B1– B2.NHMW 2020/0004/1661, Bad Vöslau (Austria). C1–C2. NHMW 2020/0004/1661a, Bad Vöslau (Austria). D1–D2. NHMW 1867/0019/0322, Coșteiu de Sus (Romania).E1–E3. NHMW 1997z0178/1210, BadVöslau (Austria). F1–F2. NHMW 1997z0178/ 1210a, Bad Vöslau (Austria). G1–G3. NHMW 1876/0011/0170a, Lăpugiu de Sus (Romania). H1–H3. NHMW 2008/0248/0479, Coșteiu de Sus (Romania). I. NHMW 1859/0037/0570, Vienna/Pötzleinsdorf (Austria). J. NHMW-ZOO-MO-75000-E-3317A, Bad Vöslau (Austria), holotype of Acteocina heraclitica Berger, 1949.	FIGURE 34. Acteocina fusiformis (Boettger, 1907). A1–A2. NHMW 1876/0011/0170, Lăpugiu de Sus (Romania). B1– B2.NHMW 2020/0004/1661, Bad Vöslau (Austria). C1–C2. NHMW 2020/0004/1661a, Bad Vöslau (Austria). D1–D2. NHMW 1867/0019/0322, Coșteiu de Sus (Romania).E1–E3. NHMW 1997z0178/1210, BadVöslau (Austria). F1–F2. NHMW 1997z0178/ 1210a, Bad Vöslau (Austria). G1–G3. NHMW 1876/0011/0170a, Lăpugiu de Sus (Romania). H1–H3. NHMW 2008/0248/0479, Coșteiu de Sus (Romania). I. NHMW 1859/0037/0570, Vienna/Pötzleinsdorf (Austria). J. NHMW-ZOO-MO-75000-E-3317A, Bad Vöslau (Austria), holotype of Acteocina heraclitica Berger, 1949.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE36728F2BD049AFED3F89B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE36728F2BD049AFED3F89B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE36728F2BD049AFED3F89B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE36728F2BD049AFED3F89B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166257/files/figure.png	https://doi.org/10.5281/zenodo.19166257	FIGURE 37. A–G. Acteocina volhynica (Eichwald, 1830). A1–A3. NHMW 1870/0013/0012, Vienna/Heiligenstadt (Austria). B1–B2. NHMW 1870/0013/0012a, Vienna/Heiligenstadt (Austria). C1–C3. NHMW 1860/0001/0346a, Lednice (= Eisgrub) (Czech Republic). D1–D2. NHMW 1861/0001/0355, Podivín (= Kostel) (Czech Republic). E1–E2. NHMW 1861/0001/0355a, Podivín (= Kostel) (Czech Republic). F1–F2. NHMW 1861/0001/0355b, Podivín (= Kostel) (Czech Republic). G1–G2. NHMW 1872/0003/00o6,Vienna/Hernals(Austria).H1–H3. Acteocina lajonkaireana (de Basterot,1825),MNHN.F.A71213,Burdigalian, Saucats (France).	FIGURE 37. A–G. Acteocina volhynica (Eichwald, 1830). A1–A3. NHMW 1870/0013/0012, Vienna/Heiligenstadt (Austria). B1–B2. NHMW 1870/0013/0012a, Vienna/Heiligenstadt (Austria). C1–C3. NHMW 1860/0001/0346a, Lednice (= Eisgrub) (Czech Republic). D1–D2. NHMW 1861/0001/0355, Podivín (= Kostel) (Czech Republic). E1–E2. NHMW 1861/0001/0355a, Podivín (= Kostel) (Czech Republic). F1–F2. NHMW 1861/0001/0355b, Podivín (= Kostel) (Czech Republic). G1–G2. NHMW 1872/0003/00o6,Vienna/Hernals(Austria).H1–H3. Acteocina lajonkaireana (de Basterot,1825),MNHN.F.A71213,Burdigalian, Saucats (France).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE6672AF2BD0786FDEEFB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166253/files/figure.png	https://doi.org/10.5281/zenodo.19166253	FIGURE 35. Acteocina lichtensteinii (Eichwald, 1830). A1–A3. NHMW-ZOO-MO-75000-E-47062, Żukowce (Zhukivtsi) (Ukraine). B1–B2. NHMW 1859/9945/0507, Golubytsya (Hołubica), (Ukraine). C1–C4. NHMW 1843/0009/0043, Żukowce (Zhukivtsi) (Ukraine). D1–D3. NHMW-ZOO-MO-75000-E-47067, Golubytsya (Hołubica) (Ukraine).	FIGURE 35. Acteocina lichtensteinii (Eichwald, 1830). A1–A3. NHMW-ZOO-MO-75000-E-47062, Żukowce (Zhukivtsi) (Ukraine). B1–B2. NHMW 1859/9945/0507, Golubytsya (Hołubica), (Ukraine). C1–C4. NHMW 1843/0009/0043, Żukowce (Zhukivtsi) (Ukraine). D1–D3. NHMW-ZOO-MO-75000-E-47067, Golubytsya (Hołubica) (Ukraine).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE6672AF2BD0786FDEEFB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE6672AF2BD0786FDEEFB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE6672AF2BD0786FDEEFB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE6672AF2BD0786FDEEFB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166181/files/figure.png	https://doi.org/10.5281/zenodo.19166181	FIGURE 4. Acteon romanicus sp. nov. A1–A2. Holotype, NHMW 2016/0177/1298, Lăpugiu de Sus (Romania).B1–B2. Paratype, NHMW 2016/0177/1298a, Lăpugiu de Sus (Romania). C. Paratype, NHMW 2016/0177/1298b, Lăpugiu de Sus (Romania).	FIGURE 4. Acteon romanicus sp. nov. A1–A2. Holotype, NHMW 2016/0177/1298, Lăpugiu de Sus (Romania).B1–B2. Paratype, NHMW 2016/0177/1298a, Lăpugiu de Sus (Romania). C. Paratype, NHMW 2016/0177/1298b, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE76724F2BD03BAFCEBF91B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166255/files/figure.png	https://doi.org/10.5281/zenodo.19166255	FIGURE 36. Acteocina melinda (De Gregorio, 1885). A1–A2. NHMW 1846/0037/0526a, Vienna/Gaudenzdorf (Austria). B1–B3. Lectotype, NHMW 1846/0037/0526b, Vienna/Gaudenzdorf (Austria). C1–C2. NHMW 1846/0037/0527, Vienna/Gaudenzdorf (Austria). D1–D2. NHMW-ZOO-MO-75000-E-42947A, Vienna/Ottakring (Austria). E1–E2. NHMW-ZOO-MO-75000-E-42947, Vienna/Ottakring (Austria). F1–F3. NHMW-ZOO-MO-75000-E-42947, Vienna/Ottakring (Austria). G1–G3. NHMW-ZOO-MO- 75000-E-47061, Vienna/Hundsturm (Austria). H1–H2. NHMW-ZOO-MO-75000-E-47061A, Vienna/Hundsturm (Austria).	FIGURE 36. Acteocina melinda (De Gregorio, 1885). A1–A2. NHMW 1846/0037/0526a, Vienna/Gaudenzdorf (Austria). B1–B3. Lectotype, NHMW 1846/0037/0526b, Vienna/Gaudenzdorf (Austria). C1–C2. NHMW 1846/0037/0527, Vienna/Gaudenzdorf (Austria). D1–D2. NHMW-ZOO-MO-75000-E-42947A, Vienna/Ottakring (Austria). E1–E2. NHMW-ZOO-MO-75000-E-42947, Vienna/Ottakring (Austria). F1–F3. NHMW-ZOO-MO-75000-E-42947, Vienna/Ottakring (Austria). G1–G3. NHMW-ZOO-MO- 75000-E-47061, Vienna/Hundsturm (Austria). H1–H2. NHMW-ZOO-MO-75000-E-47061A, Vienna/Hundsturm (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE76724F2BD03BAFCEBF91B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE76724F2BD03BAFCEBF91B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFE76724F2BD03BAFCEBF91B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEA6723F2BD018FFF39FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166257/files/figure.png	https://doi.org/10.5281/zenodo.19166257	FIGURE 37. A–G. Acteocina volhynica (Eichwald, 1830). A1–A3. NHMW 1870/0013/0012, Vienna/Heiligenstadt (Austria). B1–B2. NHMW 1870/0013/0012a, Vienna/Heiligenstadt (Austria). C1–C3. NHMW 1860/0001/0346a, Lednice (= Eisgrub) (Czech Republic). D1–D2. NHMW 1861/0001/0355, Podivín (= Kostel) (Czech Republic). E1–E2. NHMW 1861/0001/0355a, Podivín (= Kostel) (Czech Republic). F1–F2. NHMW 1861/0001/0355b, Podivín (= Kostel) (Czech Republic). G1–G2. NHMW 1872/0003/00o6,Vienna/Hernals(Austria).H1–H3. Acteocina lajonkaireana (de Basterot,1825),MNHN.F.A71213,Burdigalian, Saucats (France).	FIGURE 37. A–G. Acteocina volhynica (Eichwald, 1830). A1–A3. NHMW 1870/0013/0012, Vienna/Heiligenstadt (Austria). B1–B2. NHMW 1870/0013/0012a, Vienna/Heiligenstadt (Austria). C1–C3. NHMW 1860/0001/0346a, Lednice (= Eisgrub) (Czech Republic). D1–D2. NHMW 1861/0001/0355, Podivín (= Kostel) (Czech Republic). E1–E2. NHMW 1861/0001/0355a, Podivín (= Kostel) (Czech Republic). F1–F2. NHMW 1861/0001/0355b, Podivín (= Kostel) (Czech Republic). G1–G2. NHMW 1872/0003/00o6,Vienna/Hernals(Austria).H1–H3. Acteocina lajonkaireana (de Basterot,1825),MNHN.F.A71213,Burdigalian, Saucats (France).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEA6723F2BD018FFF39FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEA6723F2BD018FFF39FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEA6723F2BD018FFF39FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEF673CF2BD02BBFF37FE07.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166261/files/figure.png	https://doi.org/10.5281/zenodo.19166261	FIGURE 39. A1–A3. Acrostemma kuehnelti (Berger, 1953), holotype, NHMW-ZOO-MO-75000-E-42897, Vienna/Ottakring (Austria). B–C. Acrostemma edlaueri (Berger, 1953), B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3310, Bad Vöslau (Austria), Bad Vöslau (Austria). C1–C3. Holotype, NHMW-ZOO-MO-75000-E-3310A, Bad Vöslau (Austria). D1–D2. Acrostemma anatolica sp. nov., holotype, RGM 776925, Seyithasan (Turkey).	FIGURE 39. A1–A3. Acrostemma kuehnelti (Berger, 1953), holotype, NHMW-ZOO-MO-75000-E-42897, Vienna/Ottakring (Austria). B–C. Acrostemma edlaueri (Berger, 1953), B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3310, Bad Vöslau (Austria), Bad Vöslau (Austria). C1–C3. Holotype, NHMW-ZOO-MO-75000-E-3310A, Bad Vöslau (Austria). D1–D2. Acrostemma anatolica sp. nov., holotype, RGM 776925, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEF673CF2BD02BBFF37FE07.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFEF673CF2BD02BBFF37FE07.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF1673CF2BD0532FEEBFA5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166261/files/figure.png	https://doi.org/10.5281/zenodo.19166261	FIGURE 39. A1–A3. Acrostemma kuehnelti (Berger, 1953), holotype, NHMW-ZOO-MO-75000-E-42897, Vienna/Ottakring (Austria). B–C. Acrostemma edlaueri (Berger, 1953), B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3310, Bad Vöslau (Austria), Bad Vöslau (Austria). C1–C3. Holotype, NHMW-ZOO-MO-75000-E-3310A, Bad Vöslau (Austria). D1–D2. Acrostemma anatolica sp. nov., holotype, RGM 776925, Seyithasan (Turkey).	FIGURE 39. A1–A3. Acrostemma kuehnelti (Berger, 1953), holotype, NHMW-ZOO-MO-75000-E-42897, Vienna/Ottakring (Austria). B–C. Acrostemma edlaueri (Berger, 1953), B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3310, Bad Vöslau (Austria), Bad Vöslau (Austria). C1–C3. Holotype, NHMW-ZOO-MO-75000-E-3310A, Bad Vöslau (Austria). D1–D2. Acrostemma anatolica sp. nov., holotype, RGM 776925, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF1673CF2BD0532FEEBFA5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF1673CF2BD0532FEEBFA5B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF1673FF2BD02CEFED3FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166261/files/figure.png	https://doi.org/10.5281/zenodo.19166261	FIGURE 39. A1–A3. Acrostemma kuehnelti (Berger, 1953), holotype, NHMW-ZOO-MO-75000-E-42897, Vienna/Ottakring (Austria). B–C. Acrostemma edlaueri (Berger, 1953), B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3310, Bad Vöslau (Austria), Bad Vöslau (Austria). C1–C3. Holotype, NHMW-ZOO-MO-75000-E-3310A, Bad Vöslau (Austria). D1–D2. Acrostemma anatolica sp. nov., holotype, RGM 776925, Seyithasan (Turkey).	FIGURE 39. A1–A3. Acrostemma kuehnelti (Berger, 1953), holotype, NHMW-ZOO-MO-75000-E-42897, Vienna/Ottakring (Austria). B–C. Acrostemma edlaueri (Berger, 1953), B1–B3. Paratype, NHMW-ZOO-MO-75000-E-3310, Bad Vöslau (Austria), Bad Vöslau (Austria). C1–C3. Holotype, NHMW-ZOO-MO-75000-E-3310A, Bad Vöslau (Austria). D1–D2. Acrostemma anatolica sp. nov., holotype, RGM 776925, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF1673FF2BD02CEFED3FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF1673FF2BD02CEFED3FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF26739F2BD049BFDB5FB77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166263/files/figure.png	https://doi.org/10.5281/zenodo.19166263	FIGURE 40. Cylichna caudex sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-18735, Guntersdorf (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-18735, Guntersdorf (Austria). C1–C3. Paratype, NHMW-ZOO-MO-75000-E-18735, Guntersdorf (Austria). D1–D3. Paratype, NHMW 1859/0038/0135, Grund (Austria). E1–E2. NHMW 2025/0153/0001, Stetten (Austria).	FIGURE 40. Cylichna caudex sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-18735, Guntersdorf (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-18735, Guntersdorf (Austria). C1–C3. Paratype, NHMW-ZOO-MO-75000-E-18735, Guntersdorf (Austria). D1–D3. Paratype, NHMW 1859/0038/0135, Grund (Austria). E1–E2. NHMW 2025/0153/0001, Stetten (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF26739F2BD049BFDB5FB77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF26739F2BD049BFDB5FB77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF26739F2BD049BFDB5FB77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF4673BF2BD03E3FECEFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166267/files/figure.png	https://doi.org/10.5281/zenodo.19166267	FIGURE 41. Cylichna cylindracea (Pennant, 1777). A1–A3. NHMW 1846/0037/0525, Steinebrunn (Austria). B1–B3. NHMW 2013/0078/06554, Baden (Austria).C1–C3. NHMW 1997z0178/1209, Bad Vöslau (Austria).D1–D2. NHMW 2013/0078/06554a, Baden (Austria). E1–E3. NHMW 2013/0078/06554d, Baden (Austria).	FIGURE 41. Cylichna cylindracea (Pennant, 1777). A1–A3. NHMW 1846/0037/0525, Steinebrunn (Austria). B1–B3. NHMW 2013/0078/06554, Baden (Austria).C1–C3. NHMW 1997z0178/1209, Bad Vöslau (Austria).D1–D2. NHMW 2013/0078/06554a, Baden (Austria). E1–E3. NHMW 2013/0078/06554d, Baden (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF4673BF2BD03E3FECEFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF4673BF2BD03E3FECEFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF4673BF2BD03E3FECEFBE3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF66735F2BD0397FD74FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166269/files/figure.png	https://doi.org/10.5281/zenodo.19166269	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF66735F2BD0397FD74FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF66735F2BD0397FD74FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF86735F2BD06DCFED3F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166271/files/figure.png	https://doi.org/10.5281/zenodo.19166271	FIGURE 43. Cylichna intermedia (Sacco, 1897). A1–A3. NHMW-ZOO-MO-75000-E-7083, Gainfarn (Austria). B1–B3. NHMW-ZOO-MO-75000-E-47087, Steinebrunn (Austria). C1–C2. NHMW-ZOO-MO-75000-E-33312, Guntersdorf (Austria). D1–D3. NHMW 1854/0035/0386, Lăpugiu de Sus (Romania). E1–E3. NHMW 1997z0178/1884, Gainfarn (Austria).	FIGURE 43. Cylichna intermedia (Sacco, 1897). A1–A3. NHMW-ZOO-MO-75000-E-7083, Gainfarn (Austria). B1–B3. NHMW-ZOO-MO-75000-E-47087, Steinebrunn (Austria). C1–C2. NHMW-ZOO-MO-75000-E-33312, Guntersdorf (Austria). D1–D3. NHMW 1854/0035/0386, Lăpugiu de Sus (Romania). E1–E3. NHMW 1997z0178/1884, Gainfarn (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF86735F2BD06DCFED3F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF86735F2BD06DCFED3F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF86735F2BD06DCFED3F821.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF96737F2BD0040FB64FB15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166269/files/figure.png	https://doi.org/10.5281/zenodo.19166269	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF96737F2BD0040FB64FB15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFF96737F2BD0040FB64FB15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFA6736F2BD0205FDB0FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166269/files/figure.png	https://doi.org/10.5281/zenodo.19166269	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFA6736F2BD0205FDB0FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFA6736F2BD0205FDB0FCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFB6731F2BD049BFDE2FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166269/files/figure.png	https://doi.org/10.5281/zenodo.19166269	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFB6731F2BD049BFDE2FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFB6731F2BD049BFDE2FE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFC6730F2BD0683FA81FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166269/files/figure.png	https://doi.org/10.5281/zenodo.19166269	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	FIGURE 42. A1–A3. Cylichna dolium sp. nov., holotype, NHMW 1847/0037/0130, Mikulov (Czech Republic).B1–B3. Cylichna miosimplex sp. nov., holotype, NHMW-ZOO-MO-75000-E-19695, Poysdorf (Austria). C1–C3. Cylichna eschnerae sp. nov., holotype, NHMW-ZOO-MO-75000-E-20707, Žabčice (Czech Republic). D1–D3. Cylichna steiningeri sp. nov., Institute for Paleontology, University Vienna, no. nr. E1–E2. Cylichna subbacillus Berger, 1953, holotype, NHMW-ZOO-MO-75000-E- 2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFC6730F2BD0683FA81FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFC6730F2BD0683FA81FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFC6730F2BD0683FA81FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166275/files/figure.png	https://doi.org/10.5281/zenodo.19166275	FIGURE 44. A1–A3. Cylichna straussi sp. nov., Institute for Paleontology, University Vienna, no. nr. B1–B3. Cylichna sp. 1, NHMW-ZOO-MO-75000-E-3324, Bad Vöslau (Austria). C. Cylichna subbacillus Berger, 1953, Holotype, NHMW-ZOO-MO- 75000-E-2201, Bad Vöslau (Austria).	FIGURE 44. A1–A3. Cylichna straussi sp. nov., Institute for Paleontology, University Vienna, no. nr. B1–B3. Cylichna sp. 1, NHMW-ZOO-MO-75000-E-3324, Bad Vöslau (Austria). C. Cylichna subbacillus Berger, 1953, Holotype, NHMW-ZOO-MO- 75000-E-2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFD6730F2BD057BFDE2F8F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166275/files/figure.png	https://doi.org/10.5281/zenodo.19166275	FIGURE 44. A1–A3. Cylichna straussi sp. nov., Institute for Paleontology, University Vienna, no. nr. B1–B3. Cylichna sp. 1, NHMW-ZOO-MO-75000-E-3324, Bad Vöslau (Austria). C. Cylichna subbacillus Berger, 1953, Holotype, NHMW-ZOO-MO- 75000-E-2201, Bad Vöslau (Austria).	FIGURE 44. A1–A3. Cylichna straussi sp. nov., Institute for Paleontology, University Vienna, no. nr. B1–B3. Cylichna sp. 1, NHMW-ZOO-MO-75000-E-3324, Bad Vöslau (Austria). C. Cylichna subbacillus Berger, 1953, Holotype, NHMW-ZOO-MO- 75000-E-2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFD6730F2BD057BFDE2F8F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFD6730F2BD057BFDE2F8F5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFE6733F2BD0786FF0BFCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166275/files/figure.png	https://doi.org/10.5281/zenodo.19166275	FIGURE 44. A1–A3. Cylichna straussi sp. nov., Institute for Paleontology, University Vienna, no. nr. B1–B3. Cylichna sp. 1, NHMW-ZOO-MO-75000-E-3324, Bad Vöslau (Austria). C. Cylichna subbacillus Berger, 1953, Holotype, NHMW-ZOO-MO- 75000-E-2201, Bad Vöslau (Austria).	FIGURE 44. A1–A3. Cylichna straussi sp. nov., Institute for Paleontology, University Vienna, no. nr. B1–B3. Cylichna sp. 1, NHMW-ZOO-MO-75000-E-3324, Bad Vöslau (Austria). C. Cylichna subbacillus Berger, 1953, Holotype, NHMW-ZOO-MO- 75000-E-2201, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFE6733F2BD0786FF0BFCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFE6733F2BD0786FF0BFCEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFE670DF2BD032EFCF9FB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166277/files/figure.png	https://doi.org/10.5281/zenodo.19166277	FIGURE 45. Cylichnella bitaeniata (Boettger, 1902). A1–A3. NHMW 1997z9178/1903, Bad Vöslau (Austria). B1–B3. NHMW 1997z9178/1903a, Bad Vöslau (Austria). C1–C3. NHMW 187600110169, Lăpugiu de Sus (Romania). D1–D3. NHMW 185900190131, Porzteich (Czech Republic).	FIGURE 45. Cylichnella bitaeniata (Boettger, 1902). A1–A3. NHMW 1997z9178/1903, Bad Vöslau (Austria). B1–B3. NHMW 1997z9178/1903a, Bad Vöslau (Austria). C1–C3. NHMW 187600110169, Lăpugiu de Sus (Romania). D1–D3. NHMW 185900190131, Porzteich (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFE670DF2BD032EFCF9FB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFFE670DF2BD032EFCF9FB8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC1670FF2BD0786FDA7FF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166279/files/figure.png	https://doi.org/10.5281/zenodo.19166279	FIGURE 46. Neubaueriella lapugyensis (Berger, 1949). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24786A, Lăpugiu (Romania). B1–B3. NHMW 1862/0029/0920, Rudice (Czech Republic).	FIGURE 46. Neubaueriella lapugyensis (Berger, 1949). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24786A, Lăpugiu (Romania). B1–B3. NHMW 1862/0029/0920, Rudice (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC1670FF2BD0786FDA7FF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC1670FF2BD0786FDA7FF2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC3670EF2BD067EFED3F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166283/files/figure.png	https://doi.org/10.5281/zenodo.19166283	FIGURE 48. Haminoea miocaenica (Berger, 1949). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-3290, Bad Vöslau (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-36704. Guntersdorf (Austria). C1–C3. NHMW 1997z01781213, Bad Vöslau (Austria).	FIGURE 48. Haminoea miocaenica (Berger, 1949). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-3290, Bad Vöslau (Austria). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-36704. Guntersdorf (Austria). C1–C3. NHMW 1997z01781213, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC3670EF2BD067EFED3F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC3670EF2BD067EFED3F823.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC4670BF2BD0076FCC9FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166285/files/figure.png	https://doi.org/10.5281/zenodo.19166285	FIGURE 49. A–C. Vellicolla postneglecta (Berger, 1953). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-47055A, Lăpugiu (Romania). B1–B2. Paratype, NHMW-ZOO-MO-75000-E-11797, Bad Vöslau (Austria). C1–C3. Paratype, NHMW-ZOO-MO- 75000-E-11797, Bad Vöslau (Austria). D–E. Vellicolla pannonica (Csepreghy-Meznerics, 1952). D1–D2. Holotype, M.61.4301, Szob (Hungary). E1–E3. NHMW-ZOO-MO-75000-E-3330, Bad Vöslau (Austria).	FIGURE 49. A–C. Vellicolla postneglecta (Berger, 1953). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-47055A, Lăpugiu (Romania). B1–B2. Paratype, NHMW-ZOO-MO-75000-E-11797, Bad Vöslau (Austria). C1–C3. Paratype, NHMW-ZOO-MO- 75000-E-11797, Bad Vöslau (Austria). D–E. Vellicolla pannonica (Csepreghy-Meznerics, 1952). D1–D2. Holotype, M.61.4301, Szob (Hungary). E1–E3. NHMW-ZOO-MO-75000-E-3330, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC4670BF2BD0076FCC9FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC4670BF2BD0076FCC9FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC4670BF2BD0076FCC9FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC6670BF2BD04C3FC80F827.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166285/files/figure.png	https://doi.org/10.5281/zenodo.19166285	FIGURE 49. A–C. Vellicolla postneglecta (Berger, 1953). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-47055A, Lăpugiu (Romania). B1–B2. Paratype, NHMW-ZOO-MO-75000-E-11797, Bad Vöslau (Austria). C1–C3. Paratype, NHMW-ZOO-MO- 75000-E-11797, Bad Vöslau (Austria). D–E. Vellicolla pannonica (Csepreghy-Meznerics, 1952). D1–D2. Holotype, M.61.4301, Szob (Hungary). E1–E3. NHMW-ZOO-MO-75000-E-3330, Bad Vöslau (Austria).	FIGURE 49. A–C. Vellicolla postneglecta (Berger, 1953). A1–A3. Holotype, NHMW-ZOO-MO-75000-E-47055A, Lăpugiu (Romania). B1–B2. Paratype, NHMW-ZOO-MO-75000-E-11797, Bad Vöslau (Austria). C1–C3. Paratype, NHMW-ZOO-MO- 75000-E-11797, Bad Vöslau (Austria). D–E. Vellicolla pannonica (Csepreghy-Meznerics, 1952). D1–D2. Holotype, M.61.4301, Szob (Hungary). E1–E3. NHMW-ZOO-MO-75000-E-3330, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC6670BF2BD04C3FC80F827.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC6670BF2BD04C3FC80F827.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC6670BF2BD04C3FC80F827.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC76705F2BD06CAFF37FECB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166287/files/figure.png	https://doi.org/10.5281/zenodo.19166287	FIGURE 50. A1–A2. Melanochlamys radwanskii (Bałuk, 2018), holotype, MWG UW ZI/93/1252, Badenian, Korytnica (Poland). B1–B2. Chelidonura piserai sp. nov., holotype, MWG UW ZI/93/1251, Badenian, Korytnica (Poland).	FIGURE 50. A1–A2. Melanochlamys radwanskii (Bałuk, 2018), holotype, MWG UW ZI/93/1252, Badenian, Korytnica (Poland). B1–B2. Chelidonura piserai sp. nov., holotype, MWG UW ZI/93/1251, Badenian, Korytnica (Poland).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC76705F2BD06CAFF37FECB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC76705F2BD06CAFF37FECB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC86705F2BD057BFF37F9BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166287/files/figure.png	https://doi.org/10.5281/zenodo.19166287	FIGURE 50. A1–A2. Melanochlamys radwanskii (Bałuk, 2018), holotype, MWG UW ZI/93/1252, Badenian, Korytnica (Poland). B1–B2. Chelidonura piserai sp. nov., holotype, MWG UW ZI/93/1251, Badenian, Korytnica (Poland).	FIGURE 50. A1–A2. Melanochlamys radwanskii (Bałuk, 2018), holotype, MWG UW ZI/93/1252, Badenian, Korytnica (Poland). B1–B2. Chelidonura piserai sp. nov., holotype, MWG UW ZI/93/1251, Badenian, Korytnica (Poland).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC86705F2BD057BFF37F9BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC86705F2BD057BFF37F9BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC96707F2BD0786FD5BFC1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166289/files/figure.png	https://doi.org/10.5281/zenodo.19166289	FIGURE 51. A–B. Roxania pila sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24799A, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-24799, Lăpugiu de Sus (Romania). C1–C3. Roxania oviformis sp. nov., holotype, NHMW 2025/0152/0003, Bad Vöslau (Austria). D1–D3. Roxania bomba nov. nom., Holotype, NHMW-ZOO-MO-75000-E- 18181A, Guntersdorf (Austria).	FIGURE 51. A–B. Roxania pila sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24799A, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-24799, Lăpugiu de Sus (Romania). C1–C3. Roxania oviformis sp. nov., holotype, NHMW 2025/0152/0003, Bad Vöslau (Austria). D1–D3. Roxania bomba nov. nom., Holotype, NHMW-ZOO-MO-75000-E- 18181A, Guntersdorf (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC96707F2BD0786FD5BFC1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFC96707F2BD0786FD5BFC1F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCA6706F2BD030AFAA1F80D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166291/files/figure.png	https://doi.org/10.5281/zenodo.19166291	FIGURE 52. A–B. Roxania callifera (Boettger, 1907).A1–A2. NHMW 1847/0037/0131, Baden (Austria). B1–B2. NHMW-ZOO- MO-75000-E-47090, Bad Vöslau (Austria). C–D. Sabatia quasimodoi sp. nov. C1–C2. Holotype, NHMW 1863/0015/1154, Rudice (Czech Republic). D1–D2. Paratype, NHMW 1863/0015/1154a, Rudice (Czech Republic).	FIGURE 52. A–B. Roxania callifera (Boettger, 1907).A1–A2. NHMW 1847/0037/0131, Baden (Austria). B1–B2. NHMW-ZOO- MO-75000-E-47090, Bad Vöslau (Austria). C–D. Sabatia quasimodoi sp. nov. C1–C2. Holotype, NHMW 1863/0015/1154, Rudice (Czech Republic). D1–D2. Paratype, NHMW 1863/0015/1154a, Rudice (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCA6706F2BD030AFAA1F80D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCA6706F2BD030AFAA1F80D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCC6701F2BD0786FAE6FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166289/files/figure.png	https://doi.org/10.5281/zenodo.19166289	FIGURE 51. A–B. Roxania pila sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24799A, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-24799, Lăpugiu de Sus (Romania). C1–C3. Roxania oviformis sp. nov., holotype, NHMW 2025/0152/0003, Bad Vöslau (Austria). D1–D3. Roxania bomba nov. nom., Holotype, NHMW-ZOO-MO-75000-E- 18181A, Guntersdorf (Austria).	FIGURE 51. A–B. Roxania pila sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24799A, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-24799, Lăpugiu de Sus (Romania). C1–C3. Roxania oviformis sp. nov., holotype, NHMW 2025/0152/0003, Bad Vöslau (Austria). D1–D3. Roxania bomba nov. nom., Holotype, NHMW-ZOO-MO-75000-E- 18181A, Guntersdorf (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCC6701F2BD0786FAE6FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCC6701F2BD0786FAE6FB13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCC6700F2BD0206FEC2FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166289/files/figure.png	https://doi.org/10.5281/zenodo.19166289	FIGURE 51. A–B. Roxania pila sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24799A, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-24799, Lăpugiu de Sus (Romania). C1–C3. Roxania oviformis sp. nov., holotype, NHMW 2025/0152/0003, Bad Vöslau (Austria). D1–D3. Roxania bomba nov. nom., Holotype, NHMW-ZOO-MO-75000-E- 18181A, Guntersdorf (Austria).	FIGURE 51. A–B. Roxania pila sp. nov. A1–A3. Holotype, NHMW-ZOO-MO-75000-E-24799A, Lăpugiu de Sus (Romania). B1–B3. Paratype, NHMW-ZOO-MO-75000-E-24799, Lăpugiu de Sus (Romania). C1–C3. Roxania oviformis sp. nov., holotype, NHMW 2025/0152/0003, Bad Vöslau (Austria). D1–D3. Roxania bomba nov. nom., Holotype, NHMW-ZOO-MO-75000-E- 18181A, Guntersdorf (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCC6700F2BD0206FEC2FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCC6700F2BD0206FEC2FE5F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCD6702F2BD06CAFDC9FADB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166295/files/figure.png	https://doi.org/10.5281/zenodo.19166295	FIGURE 53. Roxania utriculus (Brocchi, 1814). A1–A3. NHMW 2013/0078/0555, Bad Vöslau (Austria). B1–B3. NHMW 2013/0078/0555a, Bad Vöslau (Austria). C1–C2. NHMW 1867/0019/0324, Coşteiu de Sus (Romania). D1–D3. NHMW 2013/0078/0555b,BadVöslau (Austria).E1–E2. NHMW1867/0019/0321,Coşteiu de Sus(Romania).F. NHMW1997z01781211, Bad Vöslau (Austria).	FIGURE 53. Roxania utriculus (Brocchi, 1814). A1–A3. NHMW 2013/0078/0555, Bad Vöslau (Austria). B1–B3. NHMW 2013/0078/0555a, Bad Vöslau (Austria). C1–C2. NHMW 1867/0019/0324, Coşteiu de Sus (Romania). D1–D3. NHMW 2013/0078/0555b,BadVöslau (Austria).E1–E2. NHMW1867/0019/0321,Coşteiu de Sus(Romania).F. NHMW1997z01781211, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCD6702F2BD06CAFDC9FADB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCD6702F2BD06CAFDC9FADB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFCD6702F2BD06CAFDC9FADB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD0671DF2BD04E7FF37F97B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166297/files/figure.png	https://doi.org/10.5281/zenodo.19166297	FIGURE 54. A–C. Hermania scabra (Müller, 1784). A1–A4. NHMW 2010/0004/1648, Bad Vöslau (Austria). B1–B2. NHMW 2010/0004/1648, Vöslau (Austria). C1–C2. NHMW-ZOO-MO-75000-E-47001, Lăpugiu de Sus (Romania). D1–D3. Philine catena (Montagu, 1803), NHMW-ZOO-MO-75000-E-9340, Enzesfeld (Austria). E1–E3. Cosmophiline marinellii Berger, 1949, holotype, NHMW-ZOO-MO-75000-E-4172, Bad Vöslau (Austria).	FIGURE 54. A–C. Hermania scabra (Müller, 1784). A1–A4. NHMW 2010/0004/1648, Bad Vöslau (Austria). B1–B2. NHMW 2010/0004/1648, Vöslau (Austria). C1–C2. NHMW-ZOO-MO-75000-E-47001, Lăpugiu de Sus (Romania). D1–D3. Philine catena (Montagu, 1803), NHMW-ZOO-MO-75000-E-9340, Enzesfeld (Austria). E1–E3. Cosmophiline marinellii Berger, 1949, holotype, NHMW-ZOO-MO-75000-E-4172, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD0671DF2BD04E7FF37F97B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD0671DF2BD04E7FF37F97B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD2671FF2BD0786FECEF976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166297/files/figure.png	https://doi.org/10.5281/zenodo.19166297	FIGURE 54. A–C. Hermania scabra (Müller, 1784). A1–A4. NHMW 2010/0004/1648, Bad Vöslau (Austria). B1–B2. NHMW 2010/0004/1648, Vöslau (Austria). C1–C2. NHMW-ZOO-MO-75000-E-47001, Lăpugiu de Sus (Romania). D1–D3. Philine catena (Montagu, 1803), NHMW-ZOO-MO-75000-E-9340, Enzesfeld (Austria). E1–E3. Cosmophiline marinellii Berger, 1949, holotype, NHMW-ZOO-MO-75000-E-4172, Bad Vöslau (Austria).	FIGURE 54. A–C. Hermania scabra (Müller, 1784). A1–A4. NHMW 2010/0004/1648, Bad Vöslau (Austria). B1–B2. NHMW 2010/0004/1648, Vöslau (Austria). C1–C2. NHMW-ZOO-MO-75000-E-47001, Lăpugiu de Sus (Romania). D1–D3. Philine catena (Montagu, 1803), NHMW-ZOO-MO-75000-E-9340, Enzesfeld (Austria). E1–E3. Cosmophiline marinellii Berger, 1949, holotype, NHMW-ZOO-MO-75000-E-4172, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD2671FF2BD0786FECEF976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD2671FF2BD0786FECEF976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD2671FF2BD0786FECEF976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD3671EF2BD0786FF08F9BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166297/files/figure.png	https://doi.org/10.5281/zenodo.19166297	FIGURE 54. A–C. Hermania scabra (Müller, 1784). A1–A4. NHMW 2010/0004/1648, Bad Vöslau (Austria). B1–B2. NHMW 2010/0004/1648, Vöslau (Austria). C1–C2. NHMW-ZOO-MO-75000-E-47001, Lăpugiu de Sus (Romania). D1–D3. Philine catena (Montagu, 1803), NHMW-ZOO-MO-75000-E-9340, Enzesfeld (Austria). E1–E3. Cosmophiline marinellii Berger, 1949, holotype, NHMW-ZOO-MO-75000-E-4172, Bad Vöslau (Austria).	FIGURE 54. A–C. Hermania scabra (Müller, 1784). A1–A4. NHMW 2010/0004/1648, Bad Vöslau (Austria). B1–B2. NHMW 2010/0004/1648, Vöslau (Austria). C1–C2. NHMW-ZOO-MO-75000-E-47001, Lăpugiu de Sus (Romania). D1–D3. Philine catena (Montagu, 1803), NHMW-ZOO-MO-75000-E-9340, Enzesfeld (Austria). E1–E3. Cosmophiline marinellii Berger, 1949, holotype, NHMW-ZOO-MO-75000-E-4172, Bad Vöslau (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD3671EF2BD0786FF08F9BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD3671EF2BD0786FF08F9BE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD46719F2BD0786FDE0FAAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166291/files/figure.png	https://doi.org/10.5281/zenodo.19166291	FIGURE 52. A–B. Roxania callifera (Boettger, 1907).A1–A2. NHMW 1847/0037/0131, Baden (Austria). B1–B2. NHMW-ZOO- MO-75000-E-47090, Bad Vöslau (Austria). C–D. Sabatia quasimodoi sp. nov. C1–C2. Holotype, NHMW 1863/0015/1154, Rudice (Czech Republic). D1–D2. Paratype, NHMW 1863/0015/1154a, Rudice (Czech Republic).	FIGURE 52. A–B. Roxania callifera (Boettger, 1907).A1–A2. NHMW 1847/0037/0131, Baden (Austria). B1–B2. NHMW-ZOO- MO-75000-E-47090, Bad Vöslau (Austria). C–D. Sabatia quasimodoi sp. nov. C1–C2. Holotype, NHMW 1863/0015/1154, Rudice (Czech Republic). D1–D2. Paratype, NHMW 1863/0015/1154a, Rudice (Czech Republic).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD46719F2BD0786FDE0FAAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD46719F2BD0786FDE0FAAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD4671BF2BD01B2FD28FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166299/files/figure.png	https://doi.org/10.5281/zenodo.19166299	FIGURE 55. Scaphander alatissimus sp. nov. A1–A3. Holotype, NHMW 2007z0086/0000, Grund (Austria). B1–B3. Paratype, NHMW 2007z0086/0000a, Grund (Austria). C1–C3. Paratype, NHMW 2007z0086/0000b, Grund (Austria). D1–D3. Paratype, NHMW 2007z0086/0000c, Grund (Austria).	FIGURE 55. Scaphander alatissimus sp. nov. A1–A3. Holotype, NHMW 2007z0086/0000, Grund (Austria). B1–B3. Paratype, NHMW 2007z0086/0000a, Grund (Austria). C1–C3. Paratype, NHMW 2007z0086/0000b, Grund (Austria). D1–D3. Paratype, NHMW 2007z0086/0000c, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD4671BF2BD01B2FD28FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD4671BF2BD01B2FD28FDB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD66715F2BD05A3FA87FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166302/files/figure.png	https://doi.org/10.5281/zenodo.19166302	FIGURE 56. A–C. Scaphander grundincola (De Gregorio, 1885).A1–A2. Holotype, NHMW 1851/0002/0080, Grund (Austria). B1–B3. NHMW 1855/0045/0795, Grund (Austria). C1–C3. NHMW 1997z0178/1216, Bad Vöslau (Austria). D1–D2. Scaphander helveticus Peyrot, 1932, NHMW 2000z0001/0053, Niederkreuzstetten (Austria).	FIGURE 56. A–C. Scaphander grundincola (De Gregorio, 1885).A1–A2. Holotype, NHMW 1851/0002/0080, Grund (Austria). B1–B3. NHMW 1855/0045/0795, Grund (Austria). C1–C3. NHMW 1997z0178/1216, Bad Vöslau (Austria). D1–D2. Scaphander helveticus Peyrot, 1932, NHMW 2000z0001/0053, Niederkreuzstetten (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD66715F2BD05A3FA87FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD66715F2BD05A3FA87FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD66715F2BD05A3FA87FC57.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD86714F2BD04C3FEEEFE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166302/files/figure.png	https://doi.org/10.5281/zenodo.19166302	FIGURE 56. A–C. Scaphander grundincola (De Gregorio, 1885).A1–A2. Holotype, NHMW 1851/0002/0080, Grund (Austria). B1–B3. NHMW 1855/0045/0795, Grund (Austria). C1–C3. NHMW 1997z0178/1216, Bad Vöslau (Austria). D1–D2. Scaphander helveticus Peyrot, 1932, NHMW 2000z0001/0053, Niederkreuzstetten (Austria).	FIGURE 56. A–C. Scaphander grundincola (De Gregorio, 1885).A1–A2. Holotype, NHMW 1851/0002/0080, Grund (Austria). B1–B3. NHMW 1855/0045/0795, Grund (Austria). C1–C3. NHMW 1997z0178/1216, Bad Vöslau (Austria). D1–D2. Scaphander helveticus Peyrot, 1932, NHMW 2000z0001/0053, Niederkreuzstetten (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD86714F2BD04C3FEEEFE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD86714F2BD04C3FEEEFE97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD96714F2BD0683FB2CF91D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166304/files/figure.png	https://doi.org/10.5281/zenodo.19166304	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD96714F2BD0683FB2CF91D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD96714F2BD0683FB2CF91D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD96716F2BD000CFEC2FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166304/files/figure.png	https://doi.org/10.5281/zenodo.19166304	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD96716F2BD000CFEC2FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFD96716F2BD000CFEC2FDCF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDB6716F2BD057BFE8BF820.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166306/files/figure.png	https://doi.org/10.5281/zenodo.19166306	FIGURE 58. A1–A2. Scaphander schollnbergeri sp. nov., Holotype, NHMW 1836/0012/6576a, Hrušovany nad Jevišovkou (Czech Republic). B–D. Pseudavena bella sp. nov. B1–B3. Holotype, NHMW 1863/0015/1155, Rudice (Czech Republic). C1– C3. Paratype, NHMW 1863/0015/1155a, Rudice (Czech Republic). D1–D2. Paratype, NHMW 2013/0300/0556, Baden-Sooss (Austria).	FIGURE 58. A1–A2. Scaphander schollnbergeri sp. nov., Holotype, NHMW 1836/0012/6576a, Hrušovany nad Jevišovkou (Czech Republic). B–D. Pseudavena bella sp. nov. B1–B3. Holotype, NHMW 1863/0015/1155, Rudice (Czech Republic). C1– C3. Paratype, NHMW 1863/0015/1155a, Rudice (Czech Republic). D1–D2. Paratype, NHMW 2013/0300/0556, Baden-Sooss (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDB6716F2BD057BFE8BF820.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDB6716F2BD057BFE8BF820.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDB6716F2BD057BFE8BF820.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166304/files/figure.png	https://doi.org/10.5281/zenodo.19166304	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDC6710F2BD011AFAEAFC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166304/files/figure.png	https://doi.org/10.5281/zenodo.19166304	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	FIGURE 57. A1–A3. Scaphander perforatus sp. nov., holotype, NHMW 1866/0050/0227, Baden Sooss (Austria). B1–B2. Scaphander perforatus sp. nov., paratype, NHMW 1836/0012/6576, Hrušovany nad Jevišovkou (Czech Republic). C1–C3. Scaphander krijgsmani sp. nov., holotype, NHMW 1860/0004/0375, Lăpugiu de Sus (Romania). D1–D3. Scaphander catena sp. nov., holotype, NHMW 1997z0178/1217, Lăpugiu de Sus (Romania).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDC6710F2BD011AFAEAFC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDC6710F2BD011AFAEAFC73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDD6713F2BD01FBFAC3FAE7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166306/files/figure.png	https://doi.org/10.5281/zenodo.19166306	FIGURE 58. A1–A2. Scaphander schollnbergeri sp. nov., Holotype, NHMW 1836/0012/6576a, Hrušovany nad Jevišovkou (Czech Republic). B–D. Pseudavena bella sp. nov. B1–B3. Holotype, NHMW 1863/0015/1155, Rudice (Czech Republic). C1– C3. Paratype, NHMW 1863/0015/1155a, Rudice (Czech Republic). D1–D2. Paratype, NHMW 2013/0300/0556, Baden-Sooss (Austria).	FIGURE 58. A1–A2. Scaphander schollnbergeri sp. nov., Holotype, NHMW 1836/0012/6576a, Hrušovany nad Jevišovkou (Czech Republic). B–D. Pseudavena bella sp. nov. B1–B3. Holotype, NHMW 1863/0015/1155, Rudice (Czech Republic). C1– C3. Paratype, NHMW 1863/0015/1155a, Rudice (Czech Republic). D1–D2. Paratype, NHMW 2013/0300/0556, Baden-Sooss (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDD6713F2BD01FBFAC3FAE7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166175/files/figure.png	https://doi.org/10.5281/zenodo.19166175	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	FIGURE 1. Terminology for shell morphology; SL: shell height, MD: maximum diameter, LWH: last whorl height, AH: aperture height, PH: height of periphery, ApD: diameter of apex, AA: apical angle (°), AP: apical plane (°), SA: angle of shell sides, MAW: maximum aperture width, AW: width of adapical aperture.	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDD6713F2BD01FBFAC3FAE7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166179/files/figure.png	https://doi.org/10.5281/zenodo.19166179	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	FIGURE 3. A–B. Acteon gutta sp. nov. A1–A2. Holotype, NHMW 2018/0248/0485, Coșteiu de Sus (Romania). B1–B2. Paratype, NHMW 2018/0248/0485a, Coșteiu de Sus (Romania). C1–C2. Paratype, NHMW 2018/0248/0485b, Coșteiu de Sus (Romania). D–F. Acteon laevigatus (Grateloup, 1828). D. SMF 363155, Coșteiu de Sus (Romania). E1–E2. NHMW 2020/0004/1666b, Bad Vöslau (Austria). F. RGM 776 928/1, Seyithasan (Turkey).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
A67987E6FFDD6713F2BD01FBFAC3FAE7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19166177/files/figure.png	https://doi.org/10.5281/zenodo.19166177	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	FIGURE 2. Acteon antipinguis sp. nov. A1–A2. Paratype, NHMW 1855/0045/0782, Grund (Austria). B1–B2. Paratype, NHMW 1855/0045/0782b, Grund (Austria). C1–C2. Paratype, NHMW 1855/0045/0782c, Grund (Austria). D1–D2. Holotype, NHMW 1855/0045/0782a, Grund (Austria). E1–E2. Paratype, NHMW 1855/0045/0782b, Grund (Austria).	2026-01-08	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.		Zenodo	biologists	Harzhauser, Mathias;Landau, Bernard M.;Malaquias, Manuel António E.			
