identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6ED1EB58CB1F597F85FB672DF59DAA42.text	6ED1EB58CB1F597F85FB672DF59DAA42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus altus Gojsina, Hunyadi & Pall-Gergely 2026	<div><p>Acinolaemus altus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 7 A, 8, 9, 17</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell; Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.30041&amp;materialsCitation.latitude=20.133368" title="Search Plazi for locations around (long 97.30041/lat 20.133368)">ca 6 km east from centre of Hsihseng, right side of the road, 500 m on a side road</a>, limestone rock; 20°8.0021'N, 97°18.0243'E: 1000 m a. s. l.; 07 Oct 2018, A. Hunyadi, K. Okubo, J. U. Otani leg; CUMZ 15424 . Paratypes. Myanmar • 8 shells; same data as for holotype; coll. HA • 2 shells; Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.29805&amp;materialsCitation.latitude=20.15598" title="Search Plazi for locations around (long 97.29805/lat 20.15598)">ca 4.5 km east from centre of Hsihseng, left side of the road, ca 1.7 km on a side road</a>, limestone rock; 20°9.3587'N, 97°17.8831'E; 1140 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; coll. HA • 11 shells; Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.268555&amp;materialsCitation.latitude=20.187475" title="Search Plazi for locations around (long 97.268555/lat 20.187475)">Hsihseng centre NNE ca 3.6 km</a>, limestone hill; 20°11.2485'N, 97°16.1135'E; 1040 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; coll. HA .</p><p>Additional material examined.</p><p>Myanmar • 3 shells (upper whorls broken = not paratypes); same data as for holotype; coll. HA • 3 shells (juveniles = not paratypes); Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.268555&amp;materialsCitation.latitude=20.187475" title="Search Plazi for locations around (long 97.268555/lat 20.187475)">Hsihseng centre NNE ca 3.6 km</a>, limestone hill; 20°11.2485'N, 97°16.1135'E; 1040 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg; coll. HA .</p><p>Diagnosis.</p><p>Shell high concave-conical, colorless to light-yellowish. Surface sculpture honeycomb-like. Palatal plicae, basal plica, as well as the inner part of the angular lamella, are hooked.</p><p>Description.</p><p>Shell high, slightly concave-conical, colorless to light-yellowish, consisting of 4.75–5.25 regularly increasing, convex whorls separated by deep suture; protoconch consisting of ~ 1.5 whorls, very weakly spirally striated (~ 12 spiral striae), striae clearer terminally; boundary between protoconch and teleoconch visible due to rougher surface sculpture of teleoconch whorls; last whorl slightly shouldered, adnate to penultimate (in some specimens only peristome weakly detached) and slightly descending near aperture; surface of shell honeycomb-like, surface sculpture most prominent on last two whorls; peristome finely granulated, expanded (except around sinulus), not reflected; constriction or cervical swelling behind peristome both absent; aperture equipped with numerous (~ 12), long barriers originating deep inside aperture; parietal lamella long, curved; deeper in aperture (between angular and parietal lamellae) there can be a low additional lamella with reduced outer part closer to peristome and distant from inner part; angular lamella almost twice as long as parietal, consisting of inner, hooked part (damaged in holotype) pointing outside aperture and outer, slender part projecting towards peristome edge and reaching it; sometimes, hooked inner part is underdeveloped, seemingly blunt (although in some specimens could also be damaged); inner and outer parts of angular lamella connected by much thinner, blade-like interval; sinulus with two or three low and slender barriers; three palatal plicae present (upper, inter and lower), all with inner parts in form of elevated, sharp hooks pointing outside; outer parts of palatal plicae in form of low, slender, blade-like projections, getting thicker as getting closer to peristome; palatal plicae approximately equally developed; there can be a distinct furrow visible on last whorl behind aperture, corresponding to upper palatal plica; palatal tubercle (which can sometimes appear as double, as in holotype, see Fig. 8) distinct and located in front of upper palatal plica (and visible as slightly below it in standard view); basal plica of same morphology as palatals but lower and shorter, positioned closer to columellar lamella than to lower palatal plica; subcolumellar lamella low and blunt; columellar lamella high and blunt, not reaching expanding peristome; one or two supracolumellar lamellae present, first low (or absent), second almost as high as columellar; sinulus well-separated from rest of aperture; umbilicus not much expanded at last whorl and measuring ~ 1 / 4 of shell width, sometimes with shallow groove appearing at last whorl and terminating almost immediately behind the peristome.</p><p>Differential diagnosis.</p><p>This species can be separated from other congeners by the combination of slender shell and hooked apertural barriers (angular, basal and the palatal plicae). See under A. cryptidentatus .</p><p>Measurements (in mm, n = 5).</p><p>SH = 1.77–1.99; SW = 1.56–1.72; AH = 0.58–0.66; AW = 0.63–0.69.</p><p>Distribution.</p><p>This species is known from three sampling sites in southern Shan State, Myanmar.</p><p>Etymology.</p><p>This species is named after for altus = high in Latin, in reference to its high and narrow shell.</p><p>Remarks.</p><p>Last whorl can be more or less high (e. g., lower in the holotype, Fig. 8, than in the paratype, Fig. 9), which is regarded as intraspecific variability.</p></div>	https://treatment.plazi.org/id/6ED1EB58CB1F597F85FB672DF59DAA42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
1ECEA283794B5B23BCB7EFA2F72E0EF8.text	1ECEA283794B5B23BCB7EFA2F72E0EF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus asper Gojsina, Tongkerd & Pall-Gergely 2026	<div><p>Acinolaemus asper Gojšina, Tongkerd &amp; Páll-Gergely sp. nov.</p><p>Figs 7 G, 18, 19, 30</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Tak province, 5 km ENE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.66389&amp;materialsCitation.latitude=15.776945" title="Search Plazi for locations around (long 98.66389/lat 15.776945)">Ban Huei Hin Fong</a>; 15°46.6167'N, 98°39.8333'E; 570 m a. s. l.; 26 Apr 1988; F. G. Thompson leg.; locality code FGT- 4390; UF 347324 . Paratypes. Thailand • 2 shells; same data as for holotype; UF 593502 • 6 shells; Tak Province, 9.3 km ENE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.69833&amp;materialsCitation.latitude=15.802221" title="Search Plazi for locations around (long 98.69833/lat 15.802221)">Ban Huei Hin Fong</a>; 15°48.1333'N, 98°41.9'E; 660 m a. s. l.; 26 Apr 1988; F. G. Thompson leg.; locality code FGT- 4387; UF 347311 .</p><p>Additional material examined.</p><p>Thailand • 3 shells (damaged / juveniles = not paratypes); Tak Province, 9.3 km ENE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.69833&amp;materialsCitation.latitude=15.802221" title="Search Plazi for locations around (long 98.69833/lat 15.802221)">Ban Huei Hin Fong</a>; 15°48.1333'N, 98°41.9'E; 660 m a. s. l.; 26 Apr 1988; F. G. Thompson leg.; locality code FGT- 4387; UF 593503 .</p><p>Diagnosis.</p><p>Shell depressed, concave-conical, with honeycomb-like sculpture. Numerous apertural barriers terminating in small peristomal pliculae. All apertural barriers strongly tuberculate. Umbilicus wide, measuring ~ 1 / 3 of the shell width.</p><p>Description.</p><p>Shell depressed, concave-conical, light yellowish, consisting of 3.5–4 convex whorls separated by deep suture; protoconch consisting of ~ 1.5 whorls, spirally striated (11 distinct, equidistant spiral striae terminally); boundary between protoconch and teleoconch clearly visible due to change of surface sculpture (becoming honeycomb-like on teleoconch); last whorl weakly shouldered, adnate to penultimate and slightly descending near aperture; shell surface honeycomb-like, equally strong on whole shell; peristome expanded (except around sinulus), not reflected; cervical swelling and constriction immediately behind peristome both present but weak; aperture equipped with numerous barriers, all terminating in small peristomal pliculae; parietal lamella long, moderately high, reaching peristome; angular lamella slender, higher and blunter at its inner portion, reaching peristome; blunt topped inner and outer portions of angular lamella separated by short, blade-like and less elevated interval; all palatal plicae blunt and elevated at their inner portions, sloping and getting blade-like at outer portions; upper palatal plica higher than interpalatal; lower palatal plica higher and broader than both inter and upper palatals; distinct palatal tubercle sits in front of upper palatal, visible as slightly below it in standard view; infrapalatal plica two times weaker (lower and narrower) than lower palatal; basal plica approximately of same appearance as infrapalatal; one or two subcolumellar lamellae present, nearly as strong as infrapalatal plica; columellar lamella not blade-like, oblique, reaching peristome where it becomes quite low and slender; supracolumellar lamella present but very low, equally developed infraparietal lamella present above it, both without blade-like intervals; surface of all barriers very rough, with strong, raised tubercles; sinulus small, distinctly separated from the rest of aperture; umbilicus wide, measuring ~ 1 / 3 of shell width.</p><p>Differential diagnosis.</p><p>This species can be separated from all other congeners most clearly by the strongly tuberculate apertural barriers.</p><p>Measurements (n = 5, in mm).</p><p>SH = 1.03–1.29; SW = 1.34–1.46; AH = 0.48–0.56; AW = 0.42–0.54.</p><p>Distribution.</p><p>This species is known only from the Tak Province.</p><p>Etymology.</p><p>The specific epithet if from the Latin asper, meaning rough, referring to the peculiar surface of apertural barriers.</p></div>	https://treatment.plazi.org/id/1ECEA283794B5B23BCB7EFA2F72E0EF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
F11A7C15D5BE5CAF9691D07D18BFC219.text	F11A7C15D5BE5CAF9691D07D18BFC219.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus atypicus Gojsina, Tongkerd & Pall-Gergely 2026	<div><p>Acinolaemus atypicus Gojšina, Tongkerd &amp; Páll-Gergely sp. nov.</p><p>Figs 7 B, 10, 11, 17</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Lampang Province, limestone ridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.842224&amp;materialsCitation.latitude=18.585278" title="Search Plazi for locations around (long 99.842224/lat 18.585278)">Ban Pang La</a>; 18°35.1167'N, 99°50.5333'E; 400 m a. s. l., 14 May 1988; F. G. Thompson leg.; locality code FGT- 4438; UF 591353 . Paratypes. Thailand • 2 shells; Lampang Province, limestone dome at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.80639&amp;materialsCitation.latitude=18.466112" title="Search Plazi for locations around (long 99.80639/lat 18.466112)">pass 11 km SSW of Ban Pang La</a>; 18°27.9667'N, 99°48.3833'E; 610 m a. s. l.; 14 May 1988; F. G. Thompson leg.; sample code FGT- 4441; UF 591358 • 3 shells; Lampang Province, limestone knoll 1 km NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.86972&amp;materialsCitation.latitude=18.554167" title="Search Plazi for locations around (long 99.86972/lat 18.554167)">Ban Pang La</a>; 18°33.25'N, 99°52.1833'E; 400 m a. s. l.; 14 May 1988; F. G. Thompson leg.; locality code FGT- 4436; UF 591352 • 1 shell; Lampang Province, 6 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.851944&amp;materialsCitation.latitude=17.964722" title="Search Plazi for locations around (long 98.851944/lat 17.964722)">Mae Pak</a>, limestone hill; 17°57.8833'N, 98°51.1167'E (incorrect coordinates, provided on the original label); 600 m a. s. l.; 13 Jun 1987; F. G. Thompson leg.; locality code FGT- 4328; UF 593495 .</p><p>Additional material examined.</p><p>• 1 shell (damaged, not paratype); same data as for holotype; UF 593494 .</p><p>Diagnosis.</p><p>Shell concave-conical, colorless, with honeycomb-like surface pattern mixed with more regular rectangular fields. Two palatal plicae present, with a distinct palatal tubercle. Columellar lamella wide. Umbilicus measuring ~ 1 / 4 of the shell width.</p><p>Description.</p><p>Shell concave-conical, colorless and consisting of 4–4.25 regularly increasing, convex whorls; protoconch initially weakly spirally striated, terminally with ten stronger spiral striae; boundary between protoconch and teleoconch clearly visible due to abrupt change of surface sculpture (appearance of strong radial growth lines); last whorl adnate to penultimate, rounded to slightly shouldered, weakly descending immediately near aperture; surface sculpture of teleoconch honeycomb-like, also with some clearly rectangular fields; peristome finely granulated, expanded (especially in direction below from palatal tubercle, towards parietal wall), not reflected; cervical swelling and constriction behind peristome both absent; aperture equipped with 5–7 barriers (parietal, angular, upper palatal, lower palatal, basal, columellar, and infraparietal); parietal lamella moderately thick, not very high, situated deep and directed towards palatal side of aperture; angular lamella longer, higher and thinner than parietal, reaching expanding peristome; angular lamella consists of two parts (higher inner and lower outer) separated by thinner and less elevated, blunt interval; two palatal plicae (upper and lower) not very slender and of similar appearance (upper palatal being thinner than lower); upper palatal plica situated deeper in aperture than lower; relatively weak palatal tubercle sits in front of upper palatal plica and visible as slightly below it in standard view; basal plica barely visible or even absent, situated halfway between lower palatal plica and columellar lamella; columellar lamella usually strongest (widest) barrier in aperture, high and not very long; infraparietal lamella very weak (low and short), in form of a small tubercle-like swelling, sometimes absent; sinulus small, distinctly separated from rest of aperture; umbilicus not much expanded at last whorl and measuring between 1 / 3 and 1 / 4 of shell width.</p><p>Differential diagnosis.</p><p>This species has a combination of a honeycomb-like surface sculpture and a small number of barriers which makes it distinguishable from all Acinolaemus and Clostophis species. See under A. simplex sp. nov.</p><p>Measurements (n = 3, in mm).</p><p>SH = 1.16–1.18; SW = 1.17–1.2; AH = 0.49–0.5; AW = 0.45–0.47.</p><p>Distribution.</p><p>This species is known from Lampang Province, Thailand.</p><p>Etymology.</p><p>This species is named for its apertural barrier arrangement, which is generally atypical for Acinolaemus .</p><p>Remarks.</p><p>In some specimens (e. g., UF 591358, see Fig. 11 B), the columellar lamella is weak so that the lower palatal plica is the strongest apertural barrier. In the same specimen, the infraparietal lamella was absent and the basal was exceptionally weak.</p></div>	https://treatment.plazi.org/id/F11A7C15D5BE5CAF9691D07D18BFC219	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
0E4D7EAED5CD51D3A759593A36383CE9.text	0E4D7EAED5CD51D3A759593A36383CE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus corusticorus Tongkerd & Panha 2025	<div><p>Acinolaemus corusticorus Tongkerd &amp; Panha, 2025</p><p>Figs 7 H, 20, 30</p><p>Acinolaemus corusticorus Tongkerd &amp; Panha in Tongkerd et al., 2025: 59–62, figs 13 D, E, 14, 15, table 4.</p><p>Type locality.</p><p>“ Phra Phutthabat Doi Khao Nam, Ban Na subdistrict, Sam Ngao District, Tak Province; 17°14'56.4"N, 98°56'16.3"E ” (Thailand).</p><p>Material examined.</p><p>Thailand • holotype; CUMZ 15363.1 .</p><p>Differential diagnosis.</p><p>Acinolaemus cryptidentatus normally has no peristomal pliculae and has more apertural barriers (including several small barriers in the sinulus). Also, the last whorl is not so strongly shouldered in A. cryptidentatus as it is in A. corusticorus . Acinolaemus solitus sp. nov. is smaller, has blade-like barriers and has no peristomal pliculae. See also under A. mueangonensis .</p><p>Distribution.</p><p>This species is only known from the type locality (Tongkerd et al. 2025).</p></div>	https://treatment.plazi.org/id/0E4D7EAED5CD51D3A759593A36383CE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
32596C4AF33A50E69B5E7601DEB06686.text	32596C4AF33A50E69B5E7601DEB06686.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus cryptidentatus Changlom, Chan-ard & Dumrongrojwattana 2019	<div><p>Acinolaemus cryptidentatus Changlom, Chan-ard &amp; Dumrongrojwattana, 2019</p><p>Figs 7 C, 12, 17</p><p>Acinolaemus cryptidentatus Changlom, Chan-ard &amp; Dumrongrojwattana in Changlom et al., 2019: 158–159, fig. 2.</p><p>Acinolaemus cryptidentatus — Dumrongrojwattana and Wongkamhaeng 2024: 321, 323, 330, fig. 7, 13, table 3; Tongkerd et al. 2024: 164, fig. 2 B; Tongkerd et al. 2025: 52 –53, table 4.</p><p>Type locality.</p><p>“ Tham Wua (Wua Cave), Mueang District, Mae Hong Son Province, 19°31.77'N, 98°04.92'E ” (Thailand).</p><p>Material examined.</p><p>Myanmar • 12 shells; Shan State, 13.5 km from centre of Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.61323&amp;materialsCitation.latitude=20.59044" title="Search Plazi for locations around (long 96.61323/lat 20.59044)">Myinmati Taung</a>; 20°35.4264'N, 96°36.7938'E; 1350 m a. s. l.; 03 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 7 shells; Shan State, WSW Taunggyi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.01762&amp;materialsCitation.latitude=20.7547" title="Search Plazi for locations around (long 97.01762/lat 20.7547)">Montawa Cave</a>; 20°45.282'N, 97°01.0572'E; 1260 m a. s. l.; 05 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 22 shells; Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.30041&amp;materialsCitation.latitude=20.133368" title="Search Plazi for locations around (long 97.30041/lat 20.133368)">ca 6 km from center of Hsihseng, right side of the road, 500 m on a side road</a>, limestone rock; 20°08.0021'N, 97°18.0243'E; 1000 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 19 shells; Mandalay, Kyaukse, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.33715&amp;materialsCitation.latitude=21.735884" title="Search Plazi for locations around (long 96.33715/lat 21.735884)">Kho Nan Shin Cave Pagoda</a>; 21°44.153'N, 96°20.229'E; 110 m a. s. l.; 17 Oct 2018; A. Hunyadi leg.; coll. HA • 12 shells; Mandalay, Patheingyi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.21515&amp;materialsCitation.latitude=21.988016" title="Search Plazi for locations around (long 96.21515/lat 21.988016)">vicinity of Shin Bodh Mell Waterfall</a>; 21°59.281'N, 96°12.909'E; 225 m a. s. l.; 16 Oct 2018; A. Hunyadi leg.; coll. HA • 15 shells; Mandalay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.61812&amp;materialsCitation.latitude=22.096033" title="Search Plazi for locations around (long 96.61812/lat 22.096033)">vicinity of Maha Nandamu Peik Chin Myaung Cave and waterfall</a>; 22°05.762'N, 96°37.087'E; 740 m a. s. l.; 19 Oct 2018; A. Hunyadi leg.; coll. HA • 3 shells; Shan State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.302414&amp;materialsCitation.latitude=20.13305" title="Search Plazi for locations around (long 97.302414/lat 20.13305)">ca 6 km from center of Hsihseng, right side of the road, 400 m on a side road</a>, limestone rock; 20°07.983'N, 97°18.1448'E; 1010 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 15 shells; Shan State, 7.4 km from center of Hopong towards Namsang, 5 km north on road # 4, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.23779&amp;materialsCitation.latitude=20.84938" title="Search Plazi for locations around (long 97.23779/lat 20.84938)">Parpant Cave</a>; 20°50.9628'N, 97°14.2674'E; 1170 m a. s. l.; 06 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 40 shells; Shan State, ca 16 km from center of Taunggyi towards Hopong, 1.5 km north on road # 4, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.13732&amp;materialsCitation.latitude=20.787724" title="Search Plazi for locations around (long 97.13732/lat 20.787724)">Shwe Pyi Aunchonda Monastery</a>; 20°47.2634'N, 97°08.2394'E; 1110 m a. s. l.; 08 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 3 shells; Shan State, ca 3.6 km NNE from center of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.268555&amp;materialsCitation.latitude=20.187475" title="Search Plazi for locations around (long 97.268555/lat 20.187475)">Hsihseng</a>, limestone rock; 20°11.2485'N, 97°16.1135'E; 1040 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 4 shells; Shan State, Hopong, Sam Phu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.269966&amp;materialsCitation.latitude=20.685051" title="Search Plazi for locations around (long 97.269966/lat 20.685051)">cave Ae- 5 at ridge above village Ho Hwe</a>; 20°41.103'N, 97°16.198'E; 12 Feb 2019; J. Grego leg.; coll. JG • 45 shells; Shan State, 5.8 km from the center of Hopong towards Namsang, left side of the road no. 4, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.22448&amp;materialsCitation.latitude=20.81713" title="Search Plazi for locations around (long 97.22448/lat 20.81713)">Hopong Spring Cave</a>; 20.81713°N, 97.22448°E; 1110 m a. s. l.; 06 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; coll. HA .</p><p>Thailand • 14 shells; Mae Hong Son Province, 34.0 km NW of Pai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.333336&amp;materialsCitation.latitude=19.45" title="Search Plazi for locations around (long 98.333336/lat 19.45)">Ban Nam Rin village</a>; 19°27'N, 98°20'E; 940 m a. s. l.; 18 Mar 1988; K. Auffenberg leg.; locality code KA- 0573; UF 593496 • 5 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.083334" title="Search Plazi for locations around (long 97.96667/lat 19.083334)">35.1 km S of Mae Hong Son, 1.7 km E of Route 108</a>; 19°05'N, 97°58'E; 850 m a. s. l.; 23 Mar 1988; K. Auffenberg leg.; locality code KA- 0597; UF 591346 • 130 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.38333&amp;materialsCitation.latitude=19.383333" title="Search Plazi for locations around (long 98.38333/lat 19.383333)">13.9 km NW of Pai, along Road 1095</a>; 19°23'N, 98°23'E; 780 m a. s. l.; 18 Mar 1988; K. Auffenberg leg.; sample code KA- 0571; UF 591351 • 10 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.15&amp;materialsCitation.latitude=19.566668" title="Search Plazi for locations around (long 98.15/lat 19.566668)">Ban Tham Lod village, 7 km N of Soppong on Road 1095</a>; 19°34'N, 98°09'E; 790 m a. s. l.; 18 Mar 1988; K. Auffenberg leg.; locality code KA- 0574; UF 345500 • 6 shells; Chiang Mai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.048836&amp;materialsCitation.latitude=19.4075" title="Search Plazi for locations around (long 99.048836/lat 19.4075)">NW side of Doi Pha Sam Sao</a>, ravine with dipterocarp forest, leaf litter; 19°24.45'N, 99°02.93'E; 800 m a. s. l.; 20 May 1988; F. G. Thompson leg.; locality code FGT- 4453; UF 593497 • 33 shells; Chiang Mai Province, 5 km NW of Chiang Dao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.92852&amp;materialsCitation.latitude=19.3938" title="Search Plazi for locations around (long 98.92852/lat 19.3938)">vicinity of Wat Tham Chiang Dao</a>; 19°23.628'N, 98°55.711'E; 450 m a. s. l.; 07 Feb 2015; A. Hunyadi leg.; coll. HA .</p><p>Differential diagnosis.</p><p>This species differs from A. altus sp. nov. by the clearly less slender shell and absence of hooked apertural barriers. See also under A. solitus sp. nov., A. corusticorus and A. mueangonensis .</p><p>Distribution.</p><p>This species has a wider distribution from Mae Hong Son and Chiang Mai provinces in Thailand westwards to the Shan State in Myanmar.</p><p>Remarks.</p><p>Tongkerd et al. (2024) figured a specimen of A. cryptidentatus with a bit more slender apertural barriers that reach the peristome edge in the form of small pliculae. Even though this is not typical for A. cryptidentatus, no other differences could be observed which is why we agree that it most probably represents an unusual form of this species.</p></div>	https://treatment.plazi.org/id/32596C4AF33A50E69B5E7601DEB06686	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
1F872EEFBDEB59178F68787217883512.text	1F872EEFBDEB59178F68787217883512.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus dayanus (Stoliczka 1871)	<div><p>Acinolaemus dayanus (Stoliczka, 1871)</p><p>Figs 7 I, 21, 30</p><p>Hypselostoma dayanum Stoliczka, 1871: 172, 173, pl. 7, fig. 2.</p><p>Hypselostoma dayanum — Hanley and Theobald 1876: 59, pl. 147, fig. 10; Pfeiffer 1876: 488; Pfeiffer 1880: 344.</p><p>Boysidia (?) dayana — Pilsbry 1917: 205, 206, pl. 34, figs 5, 6.</p><p>Pupa (Hypselostoma) dayana — Nevill 1878: 193.</p><p>Pupa (Hypselostoma) dayanum — Gude 1914: 300, 301.</p><p>Hypselostoma dayana — Thompson and Upatham 1997: 225.</p><p>Acinolaemus dayanum — Tongkerd et al. 2024: 162, figs 2 A, 13 A; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Damotha, prope Moulmein ” [Kayon Hill, Dhammasa Village, Mawlamyine District, Mon State, Myanmar].</p><p>Material examined.</p><p>Myanmar • 11 shells; Mon State, Mawlamyine center NEE ca 26 km, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.81076&amp;materialsCitation.latitude=16.506716" title="Search Plazi for locations around (long 97.81076/lat 16.506716)">Dhammasa Cave</a>; 16°30.4029'N, 97°48.6458'E; 8 m a. s. l.; 11 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; coll. HA .</p><p>Differential diagnosis.</p><p>Acinolaemus asper sp. nov. has a wider umbilicus and more peristomal pliculae than A. dayanus as well as the very rough surface of the barriers. Acinolaemus solitus sp. nov. has a wider umbilicus, less slender apertural barriers and lacks the peristomal pliculae. This species differs from A. humilis sp. nov. by the more slender palatal plicae which are situated deeper in the aperture and reach the peristome edge in the form of small pliculae. Furthermore, the shell of A. humilis sp. nov. is lower and apertural barriers are stronger than those of A. dayanus . See also under A. mueangonensis, A. microcubus sp. nov., and A. zed sp. nov.</p><p>Distribution.</p><p>This species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/1F872EEFBDEB59178F68787217883512	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
5A79D2E0FCA95AEEA6F8F89B667038A7.text	5A79D2E0FCA95AEEA6F8F89B667038A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus F. G. Thompson & Upatham 1997	<div><p>Genus Acinolaemus F. G. Thompson &amp; Upatham, 1997</p><p>Acinolaemus F. G. Thompson &amp; Upatham, 1997: 223–225 .</p><p>Type species.</p><p>Acinolaemus ptychochilus F. G. Thompson &amp; Upatham, 1997, by original designation.</p><p>Remarks.</p><p>We have subdivided Acinolaemus into six species groups based on similarities in surface sculpture and apertural barrier morphology as follows:</p></div>	https://treatment.plazi.org/id/5A79D2E0FCA95AEEA6F8F89B667038A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
EB6C4B378F565C90A00E50F6CD185AF1.text	EB6C4B378F565C90A00E50F6CD185AF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus ferox Gojsina, Tongkerd & Pall-Gergely 2026	<div><p>Acinolaemus ferox Gojšina, Tongkerd &amp; Páll-Gergely sp. nov.</p><p>Figs 7 O, 31, 33</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Chiang Rai Province, limestone knoll, 4 km NE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.98722&amp;materialsCitation.latitude=19.571945" title="Search Plazi for locations around (long 99.98722/lat 19.571945)">Ban Pa Ngae</a> limestone knoll, base of limestone ledge; 19°34.3167'N, 99°59.2333'E; 410 m a. s. l.; 12 May 1988; F. G. Thompson leg.; locality code FGT- 4429; UF 591354.</p><p>Diagnosis.</p><p>Shell concave-conical, colorless. Last whorl weakly shouldered, teleoconch strongly spirally straited. All palatal plicae and angular lamella with hooked inner portions and slender outer portions which reach the peristome edge as small pliculae. Umbilicus wide, ~ 1 / 2 of the shell width with a blunt periumbilical keel.</p><p>Description.</p><p>Shell concave-conical, probably light yellowish (but not well visible due to slightly corroded specimen), consisting of almost 5 convex whorls separated by deep suture; protoconch consisting of ~ 1.5 whorls and ~ 15 dense, equidistant spiral striae, clearer terminally; boundary between protoconch and teleoconch distinct due to change of surface sculpture (becoming more prominent on teleoconch and with stronger radial growth lines); last whorl weakly shouldered, adnate to penultimate and slightly descending near aperture; surface sculpture of shell consisting of raised spiral striae crossed by weaker and much sparser radial growth lines; peristome finely granulated, expanded (except around sinulus), but not reflected; no distinct cervical swelling or constriction present; aperture equipped with nine barriers; parietal lamella strongly oblique, high and very thick, directed towards palatal wall, almost fully reaching peristome where it becomes quite low; angular lamella bipartite, consisting of inner, hooked part directed outside aperture and outer, slender portion which reaches peristome edge; two portions are blunt-topped, separated by blade-like, less elevated, short interval; all palatal plicae and basal plica with inner, hooked portions pointing outside, and much lower, slender and blade-like outer portions that reach expanding peristome in form of blunt pliculae; upper palatal plica slightly thinner than interpalatal and lower palatal; interpalatal and lower palatal plicae equally strong; infrapalatal and basal plicae approximately as same as upper palatal; columellar lamella broad, blunt and approximately as high as parietal, oblique and almost reaching peristome; columellar lamella getting lower towards peristome; infraparietal lamella blunt, half as high as columellar and parietal, situated halfway between them; sinulus small, distinctly separated from remainder of aperture; umbilicus wide, ~ 1 / 3 of shell width; blunt periumbilical keel present behind peristome and terminating at ~ 1 / 2 of last whorl.</p><p>Differential diagnosis.</p><p>See under A. ptychochilus .</p><p>Measurements (n = 1, in mm).</p><p>SH = 1.5; SW = 1.5; AH = 0.62; AW = 0.65.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Etymology.</p><p>The specific epithet is the Latin ferox meaning fierce or harsh, referring to numerous hooked apertural barriers.</p></div>	https://treatment.plazi.org/id/EB6C4B378F565C90A00E50F6CD185AF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
418A31CFEE17599A8C3A5EECDDD92C02.text	418A31CFEE17599A8C3A5EECDDD92C02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus humilis Gojsina, Hunyadi & Pall-Gergely 2026	<div><p>Acinolaemus humilis Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 7 D, 13, 17</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell; Kayin State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.71161&amp;materialsCitation.latitude=16.780243" title="Search Plazi for locations around (long 97.71161/lat 16.780243)">2.8 km southwest from Naung Lon towards – Zwegabin junction</a>, right side of the road; 16°46.8146'N, 97°42.6965'E; 30 m a. s. l.; 12 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; CUMZ 15425 . Paratypes. Myanmar • 6 shells; same data as for holotype; coll. HA .</p><p>Diagnosis.</p><p>Shell depressed concave-conical, low, strongly spirally striated. Spiral striae crossed by equally strong radial growth lines. Nine strong apertural barriers. Palatal plicae and columellar lamella are especially strong. Umbilicus wide, measuring ~ 1 / 3 of the shell width.</p><p>Description.</p><p>Shell depressed concave-conical, slightly yellowish, 3–3.25 whorls separated by deep suture; protoconch of ~ 1.25 whorls, with ~ 15 spiral striae; boundary between protoconch and teleoconch discernible due to strong radial growth lines; last whorl strongly enlarged, shouldered, distinctly spirally striated, around 20 spiral striae visible in standard view; spiral striae crossed by rather randomly positioned radial growth lines which are equally strong as spiral striae and form rectangular to polygonal fields which produce honeycomb surface pattern; honeycomb-like fields appear in combination with distinct spiral striae making surface sculpture intermediary between two types; last whorl slightly descending near aperture; peristome finely granulated, strongly expanded, not reflected; slight indications of cervical swelling and constriction present behind peristome; aperture equipped with nine barriers (parietal, angular, suprapalatal, upper palatal, interpalatal, lower palatal, basal, columellar, and infraparietal); angular and parietal lamellae developed to same extent, both high, curved and long; angular lamella separated into shorter (outer) and longer (inner) parts by clear blade-like, thin interval; angular lamella reaching peristome edge, parietal almost reaching; main palatal plicae (upper, inter and lower) all long and high; inner portion of upper palatal plica high and lowering towards peristome; interpalatal plica stronger (broader and higher) than both upper and lower palatal, highest in middle portion; lower palatal plica of similar morphology as interpalatal but approximately as high and broad as upper palatal; one lower suprapalatal plica situated immediately above upper palatal; none of palatal plicae reach peristome edge; palatal tubercle strong, situated in front of upper palatal plica and appears slightly below it in standard view; basal plica low and short, situated approximately halfway between lower palatal plica and columellar lamella; latter is widest in aperture, oblique and almost reaching peristome edge; infraparietal lamella low and short, positioned halfway between columellar and parietal lamellae; sinulus small and well-separated from remainder of aperture; umbilicus wide, significantly wider towards last whorl and measuring ~ 1 / 3 of shell width; very shallow umbilical groove present, visible approximately at last quarter of last whorl.</p><p>Differential diagnosis.</p><p>See under A. dayanus and A. zed sp. nov.</p><p>Measurements (in mm, n = 5).</p><p>SH = 0.84–0.99; SW = 1.25–1.33; AH = 0.49–0.52; AW = 0.52–0.61.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Etymology.</p><p>This species’ epithet refers to its low, modest shell.</p></div>	https://treatment.plazi.org/id/418A31CFEE17599A8C3A5EECDDD92C02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
A9E42EABCFB352A1BE3D8AEAFD6E56C4.text	A9E42EABCFB352A1BE3D8AEAFD6E56C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus microcubus Gojsina, Hunyadi & Pall-Gergely 2026	<div><p>Acinolaemus microcubus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 7 J, 22, 30</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell; Kayin State, ca 51 km east-southeast from centre of Mudon, northeast from Htimahto, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.97&amp;materialsCitation.latitude=16.039932" title="Search Plazi for locations around (long 97.97/lat 16.039932)">northeastern end of Kwooprai Hill</a>; 16°2.3959'N, 97°58.2001'E; 25 m a. s. l.; 07 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; CUMZ 15427 . Paratypes. Myanmar • 7 shells; same data as for holotype; coll. HA .</p><p>Diagnosis.</p><p>Shell depressed conical, yellowish, with honeycomb-like surface sculpture. Last whorl strongly shouldered, umbilicus very wide. Aperture equipped with eight low, elongated barriers and a distinct palatal tubercle.</p><p>Description.</p><p>Shell depressed conical, light yellowish to brownish, consisting of 3.5–4 whorls separated by deep suture; protoconch spirally striated (approximately 15 equidistant spiral striae), consisting of ~ 1.5–1.75 whorls; boundary between protoconch and teleoconch distinct due to abrupt change of surface sculpture (distinct spiral striae on protoconch vs honeycomb-like on teleoconch); protoconch and all teleoconch whorls, except last one, are rounded, weakly convex; last whorl enlarged, distinctly shouldered; surface sculpture equally prominent on whole teleoconch; last whorl very weakly descending near aperture but apertural profile almost parallel to shell axis; peristome finely granulated, expanded (except around sinulus), not reflected: no constriction or cervical swelling behind peristome; aperture equipped with eight barriers (parietal, angular, upper palatal, interpalatal, lower palatal, basal, columellar and infraparietal); angular and parietal lamellae developed approximately to same extent, both long, parietal lamella bit higher, both almost reaching expanding peristome; parietal lamella not blade-like; angular lamella consists of two parts (inner (higher) and outer (lower)) separated by blade-like, less elevated interval; three palatal plicae (upper, inter and lower) all equally wide and long, higher and blunt at their inner portions and abruptly sloping as blade-like towards peristome; palatal tubercle distinct, situated in front of upper palatal plica and appears slightly below it in standard view; interpalatal and lower palatal plicae projecting all the way to expanding peristome and terminating in moderate, blunt pliculae below strong palatal tubercle; basal plica low, positioned closer to columellar than to lower palatal, reaching peristome similarly as described for interpalatal and lower palatal plicae, and of same morphology; columellar lamella long, blunt along entire length, oblique, almost fully reaching peristome edge; infraparietal lamella low but long, blunt, nearly reaching peristome edge; sinulus small and distinctly separated from remainder of aperture; umbilicus wide, between 1 / 2 and 1 / 3 of shell width, widening near last whorl; blunt periumbilical keel present.</p><p>Differential diagnosis.</p><p>This species is similar to A. dayanus in general shell shape and apertural barrier arrangement. However, the last whorl is much more strongly shouldered in the new species and there is neither a cervical swelling nor the constriction behind the peristome (present in A. dayanus). The umbilicus is also narrower in A. dayanus and there is no periumbilical keel. See also under A. corusticorus and A. zed sp. nov.</p><p>Measurements (in mm, n = 5).</p><p>SH = 1.16–1.28; SW = 1.41–1.42; AH = 0.61–0.71; AW = 0.59–0.65.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Etymology.</p><p>The specific epithet refers to the combination of a small and cube-shaped shell (due to the strongly shouldered last whorl).</p></div>	https://treatment.plazi.org/id/A9E42EABCFB352A1BE3D8AEAFD6E56C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
984272E237DF5CC1B8CCC05EC1BE929C.text	984272E237DF5CC1B8CCC05EC1BE929C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus mueangonensis Changlom, Chan-ard & Dumrongrojwattana 2019	<div><p>Acinolaemus mueangonensis Changlom, Chan-ard &amp; Dumrongrojwattana, 2019</p><p>Figs 7 K, 23 K – 26, 30</p><p>Acinolaemus mueangonensis Changlom, Chan-ard &amp; Dumrongrojwattana, 2019: 159–161, fig. 3 A – G.</p><p>Acinolaemus mueangon [sic] — Dumrongrojwattana and Wongkamhaeng 2024: 321, 323, 330, fig. 7, 13, table 3.</p><p>Acinolaemus mueangonensis — Tongkerd et al. 2025: 55 –59, table 4.</p><p>Type locality.</p><p>“ Tham Mueang on [Mueang On Cave], Mae On District, Chiang Mai Province, 18°47.39'N, 99°14.43'E ” (Thailand).</p><p>Material examined.</p><p>Thailand • 3 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Mae Hong Son, road 1095</a>; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar 1988; K. Auffenberg leg.; locality code KA- 0595 B; UF 345719 • 31 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.98333&amp;materialsCitation.latitude=19.45" title="Search Plazi for locations around (long 97.98333/lat 19.45)">40 km N of Mae Hong Son</a>; 19°27'N, 97°59'E; 550 m a. s. l.; 21 Mar 1988; K. Auffenberg leg.; sample code KA- 0588; UF 345640 • 24 shells; Loei Province, Mueang Loei district, rock wall <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.85963&amp;materialsCitation.latitude=17.46493" title="Search Plazi for locations around (long 101.85963/lat 17.46493)">above Wat Tham Piya Thammarangsi</a>; 17°27.8958'N, 101°51.5778'E; 390 m a. s. l.; 27 Feb 2023; A. Hunyadi, J. U. Otani leg.; coll. HA • 1 shell; Loei Province, Nong Hin district, Puan Phu subdistrict, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.72758&amp;materialsCitation.latitude=17.04118" title="Search Plazi for locations around (long 101.72758/lat 17.04118)">20.3 km southwest from the centre of Nong Hin, Pha Wai, left side of the road no. 3029</a>; 17°2.4708'N, 101°43.6548'E; 705 m a. s. l.; 28 Feb 2023; A. Hunyadi, J. U. Otani leg.; coll. HA .</p><p>Differential diagnosis.</p><p>This species differs from A. cryptidentatus by the much more slender, blade-like apertural barriers which reach the peristome edge in small pliculae as well as the brownish coloration of the shell. This species differs from A. corusticorus by the brown shell as well as longer and blade-like apertural barriers.</p><p>Distribution.</p><p>This species is known from Chiang Mai, Mae Hong Son, Loei, and Tak provinces in Thailand.</p><p>Remarks.</p><p>Specimens from Tak Province reported by Tongkerd et al. (2025) share the same apertural barrier arrangement, surface sculpture, and coloration with the typical A. mueangonensis but have more slender shells. Since the slenderness of the shell is found to be variable within several species, the population from Tak Province is identified as A. mueangonensis . Some specimens from Mae Hong Son Province (UF 345640) (Fig. 25) shared the same shell shape, apertural barrier arrangement and umbilicus width with typical A. mueangonensis but had a less pronounced honeycomb-like surface, which is distinct only on isolated parts of the shell. Since other specimens from the same sample had a surface more similar to typical A. mueangonensis, we also treat the population from Mae Hong Son as A. mueangonensis . Specimens from Loei Province had a more depressed shell with a wider umbilicus (Fig. 26) and are therefore identified as Acinolaemus cf. mueangonensis .</p></div>	https://treatment.plazi.org/id/984272E237DF5CC1B8CCC05EC1BE929C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
9C6ECD1C0DD25DD5A42805F9A00327AE.text	9C6ECD1C0DD25DD5A42805F9A00327AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus paucidentatus Gojsina, Tongkerd & Pall-Gergely 2026	<div><p>Acinolaemus paucidentatus Gojšina, Tongkerd &amp; Páll-Gergely sp. nov.</p><p>Figs 7 Q, 34, 35, 39</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Chiang Mai Province, 20 k SSW of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.081&amp;materialsCitation.latitude=19.578" title="Search Plazi for locations around (long 99.081/lat 19.578)">Ban Mae Khi</a> 700 m limestone leaf litter; 19°34.68'N, 99°4.86'E; 20 Jun 1987; F. G. Thompson leg.; locality code FGT- 4334; UF 529566.</p><p>Additional material examined.</p><p>Thailand • 10 shells (not paratypes); Chiang Rai Province, 7 + 2 km south-southwest from Mae Sai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.86362&amp;materialsCitation.latitude=20.328716" title="Search Plazi for locations around (long 99.86362/lat 20.328716)">vicinity of Wat Tham Pla</a>; 20°19.723'N, 99°51.817'E; 400 m a. s. l.; 12 Feb 2015; A. Hunyadi leg.; coll. HA . • 1 shell (damaged = not paratype); same data as for holotype; UF 593510 .</p><p>Diagnosis.</p><p>A colorless Acinolaemus with six main blunt and short apertural barriers. Shell sculpture reticulated (i. e., raised spiral striae crossed by equally strong radial growth lines). Umbilicus wide, ~ 1 / 3 of the shell width.</p><p>Description.</p><p>Shell concave-conical, colorless, consisting of 4–4.25 convex, rounded whorls; protoconch very finely, coarsely spirally striated and consisting of ~ 1.5 whorls; boundary between protoconch and teleoconch distinct because sculpture becomes stronger on initial teleoconch whorls (including presence of radial growth lines); last whorl very weakly shouldered, almost parallel to shell axis near aperture; surface sculpture of teleoconch densely spirally striated; spiral striae crossed by equally strong, dense radial growth lines forming reticulated pattern; peristome finely granulated, expanded (except around sinulus), not reflected; cervical swelling and constriction behind peristome present, both vary from weak to conspicuous; aperture equipped with six barriers (parietal, angular, upper palatal, lower palatal, basal, and columellar); parietal lamella moderately high, not reaching peristome; angular lamella longer and more slender than parietal, reaching peristome edge, bipartite with inner (higher) and outer (lower) portions separated by less elevated but also blunt interval; two palatal plicae (upper and lower) relatively high, not very slender and of similar appearance (upper palatal being slightly lower than lower palatal); weak tubercle present in front of lower palatal plica, probably representing a reduced peristomal plicula; distinct palatal tubercle situated in front of upper palatal plica; basal plica low and short (more than two times lower than lower palatal), situated halfway between lower palatal plica and columellar lamella; columellar lamella broad, oblique, not reaching peristome; two tubercle-like swellings present above columellar lamella, one probably corresponding to supracolumellar lamella and other to infraparietal; sinulus small, distinctly separated from remainder of aperture; umbilicus wide, ~ 1 / 3 of shell width.</p><p>Differential diagnosis.</p><p>Acinolaemus paucidentatus sp. nov. has a reticulated shell surface sculpture (not honeycomb-like) and a wider umbilicus than A. simplex sp. nov.</p><p>Measurements (n = 1, in mm).</p><p>SH = 1.64; SW = 1.54; AH = 0.67; AW = 0.66.</p><p>Distribution.</p><p>This species is known from Chiang Mai and Chiang Rai Provinces in Thailand.</p><p>Etymology.</p><p>The specific epithet is a combination of the Latin pauci (few) and dentatus (teeth) referring to a relatively few apertural barriers when compared to the majority of congeners.</p><p>Remarks.</p><p>In specimens from Chiang Rai Province, we have noticed that they have slightly more reduced apertural barriers. All other characters (surface sculpture type, general barrier morphology and arrangement and umbilicus width) are shared with specimens from Chiang Mai Province (type locality) which is why we treat them conspecific. However, specimens from Chiang Rai Province are not designated as paratypes.</p></div>	https://treatment.plazi.org/id/9C6ECD1C0DD25DD5A42805F9A00327AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
6608C6853FD95A4A8BCDD63F93201B16.text	6608C6853FD95A4A8BCDD63F93201B16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus profundus Gojsina, Auffenberg & Pall-Gergely 2026	<div><p>Acinolaemus profundus Gojšina, Auffenberg &amp; Páll-Gergely sp. nov.</p><p>Figs 7 P, 32, 33</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Mae Hong Son Province, 34.0 km NW Pai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.333336&amp;materialsCitation.latitude=19.45" title="Search Plazi for locations around (long 98.333336/lat 19.45)">Ban Nam Rin village</a>; 19°27'N, 98°20'E; 940 m a. s. l.; 18 Mar 1988; K. Auffenberg leg.; locality code KA- 0573; UF 345958 . Paratypes. Thailand • 130 shells; same data as for holotype; UF 593505 • 2 shells; same data as for holotype; CUMZ 15428 • 7 shells; same data as for holotype; locality code KA- 0572; UF 345951 • 1 shell; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.083334" title="Search Plazi for locations around (long 97.96667/lat 19.083334)">35.1 km S Mae Hong Son, 1.7 km E route 108</a>; 19°5'N, 97°58'E; 850 m a. s. l.; 23 Mar 1988; K. Auffenberg leg.; locality code KA- 0597; UF 591367 • 6 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Mae Hong Son, road 1095</a>; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar 1988; K. Auffenberg leg.; locality code KA- 0595 B; UF 593507 • 151 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.21667&amp;materialsCitation.latitude=19.533333" title="Search Plazi for locations around (long 98.21667/lat 19.533333)">42 km NW of Pai, road 1095</a>; 19°32'N, 98°13'E; 750 m a. s. l.; 20 Mar 1988; K. Auffenberg leg.; locality code KA- 0584; UF 345583 • 6 shells; Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.2799&amp;materialsCitation.latitude=19.565466" title="Search Plazi for locations around (long 98.2799/lat 19.565466)">Tham Nam Lod 800 m – Buddhist temple</a>; 19°33.928'N, 98°16.794'E; 650 m a. s. l.; 09 Feb 2015; A. Hunyadi leg.; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 10 shells (damaged / juveniles = not paratypes); same data as for holotype; UF 593506 • 8 shells (damaged / juveniles = not paratypes); same data as for holotype; locality code KA- 0572; UF 593508 • 1 shell (juvenile = not paratype); Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.96667&amp;materialsCitation.latitude=19.416666" title="Search Plazi for locations around (long 97.96667/lat 19.416666)">Mae Hong Son, road 1095</a>; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar 1988; K. Auffenberg leg.; locality code KA- 0595 B; UF 593509 • 12 shells (damaged / juveniles = not paratypes); Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.21667&amp;materialsCitation.latitude=19.533333" title="Search Plazi for locations around (long 98.21667/lat 19.533333)">42 km NW of Pai, road 1095</a>; 19°32'N, 98°13'E; 750 m a. s. l.; 20 Mar 1988; K. Auffenberg leg.; locality code KA- 0584; UF 591348 • 2 shells (damaged and juvenile = not paratypes); Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.2799&amp;materialsCitation.latitude=19.565466" title="Search Plazi for locations around (long 98.2799/lat 19.565466)">Tham Nam Lod 800 m – Buddhist temple</a>; 19°33.928'N, 98°16.794'E; 650 m a. s. l.; 09 Feb 2015; A. Hunyadi leg.; coll. HA .</p><p>Diagnosis.</p><p>Shell concave-conical, whorls very convex, suture very deep. Aperture equipped with nine or ten elongated barriers and a palatal tubercle. Umbilicus wide, ~ 1 / 4 of the shell width.</p><p>Description.</p><p>Shell concave-conical, colorless, consisting of 3.75–4 convex whorls separated by very deep suture; protoconch spirally striated, with ~ 13 terminally clearer spiral striae, consisting of ~ 1.25 whorls; boundary between protoconch and teleoconch distinct due to change in surface sculpture (much stronger radial and spiral growth elements on teleoconch); last whorl convex, rounded, descending near aperture; teleoconch sculpture consisting of raised spiral striae crossed by nearly equally strong, but irregular radial growth lines; peristome finely granulated, weakly expanded (except around sinulus), not reflected; no cervical swelling or constriction visible behind peristome; aperture equipped with nine or ten barriers (parietal, angular, suprapalatal (sometimes absent), upper palatal, interpalatal, lower palatal, basal, subcolumellar (sometimes absent), columellar, infraparietal); parietal lamella high, blunt, long, higher in middle portions and sloping towards peristome and almost reaching its edge; angular lamella of similar appearance as parietal, slightly lower, thinner, more curved, but longer, and reaching peristome; all palatal plicae and basal plica blunt and higher at their inner portions, sloping as less elevated, blade-like; upper palatal plica low and slender, thinnest among main palatals; suprapalatal plica very short, low and indistinct, occasionally absent; interpalatal plica slightly higher than upper palatal but slightly lower than lower palatal (both of similar morphology as upper palatal); distinct palatal tubercle (sometimes very weak, rarely absent) situated in front of upper palatal plica and as appears slightly below it in standard view; basal plica low and slender, situated approximately halfway between lower palatal and columellar; subcolumellar lamella, if present, low and shorts, columellar lamella oblique, nearly as broad as parietal; infraparietal lamella moderate, situated halfway between columellar and parietal lamellae; two weak peristomal pliculae present, corresponding to lower palatal plica and basal plica; sinulus small and distinctly separated from remainder of aperture; umbilicus wide, ~ 1 / 3 – 1 / 4 of shell width.</p><p>Differential diagnosis.</p><p>Acinolaemus cryptidentatus has a honeycomb-like surface, less elongated barriers, less rounded whorls, and a more slender shell. Acinolaemus mueangonensis has a honeycomb-like surface and slenderer shell.</p><p>Measurements (n = 5, in mm).</p><p>SH = 1.19–1.37; SW = 1.39–1.47; AH = 0.52–0.61; AW = 0.55–0.58.</p><p>Distribution.</p><p>This species is known from several localities in Mae Hong Son Province, Thailand.</p><p>Etymology.</p><p>The species name is from the Latin profundus, meaning deep, in reference to its deep suture.</p></div>	https://treatment.plazi.org/id/6608C6853FD95A4A8BCDD63F93201B16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
D226A057C184573485E9781EC412F0D2.text	D226A057C184573485E9781EC412F0D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus ptychochilus F. G. Thompson & Upatham 1997	<div><p>Acinolaemus ptychochilus F. G. Thompson &amp; Upatham, 1997</p><p>Figs 7 R, 36 –39</p><p>Acinolaemus ptychochilus F. G. Thompson &amp; Upatham, 1997: 225–226, figs 7–11.</p><p>Acinolaemus ptychochilus — Changlom et al. 2019: 156; Panha and Burch 2008: 41 –43, fig. 40; Schileyko 2011: 4; Dumrongrowjattana and Wongkamhaeng 2024: 321, 323, 330, fig. 7, 13, table 3; Tongkerd et al. 2025: 50 –52, figs 8, 9, table 4.</p><p>Type locality.</p><p>“ Thailand, Chiang Mae Province, Doi Pha San Sao (Mountain), 3 km west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.058334&amp;materialsCitation.latitude=19.433332" title="Search Plazi for locations around (long 99.058334/lat 19.433332)">Ban Prang Ma-O</a>; 500 m altitude (19°26.0'N, 99°03.5'E) ”.</p><p>Type material examined.</p><p>Thailand • holotype; UF 113502 • 2 paratypes from the type locality; UF 113504 • 4 paratypes from the type locality; UF 113503 .</p><p>Additional material examined.</p><p>Thailand • 4 shells; Nan Province, Ban Pha Tun, 11 km N of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.73583&amp;materialsCitation.latitude=18.849167" title="Search Plazi for locations around (long 100.73583/lat 18.849167)">Nan</a>; 18°50.95'N, 100°44.15'E; 260 m a. s. l.; 28 Apr 1988; F. G. Thompson leg.; sample code FGT- 4392; UF 347336 • 5 shells; Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.55167&amp;materialsCitation.latitude=20.193611" title="Search Plazi for locations around (long 99.55167/lat 20.193611)">limestone knoll, Ban Mae Song Nai, 4 km N, 6 km NW of Mae Chan</a>, limestone domes, shaded ledges; 20°11.6167'N, 99°33.1'E; 520 m a. s. l.; 10 May 1988; F. G. Thompson leg.; locality code FGT- 4425; UF 593511 • 1 shell; Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.98722&amp;materialsCitation.latitude=19.571945" title="Search Plazi for locations around (long 99.98722/lat 19.571945)">limestone knoll, 4 km NE of Ban Pa Ngae</a>, base of limestone ledge; 19°34.3167'N, 99°59.2333'E; 410 m a. s. l.; 12 May 1988; F. G. Thompson leg.; locality code FGT- 4429; UF 593504 • 3 shells; Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.444725&amp;materialsCitation.latitude=19.648611" title="Search Plazi for locations around (long 99.444725/lat 19.648611)">limestone mountain, 6 km by road W of Ban Mae Suai</a>; 19°38.9167'N, 99°26.6833'E; 650 m a. s. l.; 7 May 1988; F. G. Thompson leg.; locality code FGT- 4415; UF 347100 .</p><p>Differential diagnosis.</p><p>This species is similar to A. ferox sp. nov. which however has all palatal plicae hooked, as well as the inner portions of the angular lamella ( A. ptychochilus has only the lower palatal hooked). The apertural barrier arrangement in A. mueangonensis is similar to A. ptychochilus but the latter has a different shell surface sculpture (distinctly spirally striated, not honeycomb-like). Also, no apertural barriers are hooked in A. mueangonensis . See also under A. singularis sp. nov.</p><p>Distribution.</p><p>This species is known from Chiang Mai, Chiang Rai and Nan provinces, Thailand.</p><p>Remarks.</p><p>One of the three paratypes of A. ptychochilus in UF 113504 represents A. cryptidentatus . Specimens from Nan Province (UF 347336) showed the same shell morphology as that of the type series, only with a slightly denser surface sculpture which we regard as intraspecific variability. Specimens from Chiang Rai Province (UF 593511) have non-hooked lower palatal plica but since this is the only difference observed, we treat them as A. ptychochilus .</p></div>	https://treatment.plazi.org/id/D226A057C184573485E9781EC412F0D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
68E100D0DC4A5B11A90CA4156EB8846B.text	68E100D0DC4A5B11A90CA4156EB8846B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus rhamphodontis Tongkerd & Panha 2025	<div><p>Acinolaemus rhamphodontis Tongkerd &amp; Panha, 2025</p><p>Figs 7 L, 27, 30</p><p>Acinolaemus rhamphodontis Tongkerd &amp; Panha in Tongkerd et al., 2025: 63–66, figs 16, 17, table 4.</p><p>Type locality.</p><p>“ Phra Phutthabat Doi Khao Nam, Ban Na subdistrict, Sam Ngao District, Tak Province; 17°14'56.4"N, 98°56'16.3"E ” (Thailand).</p><p>Material examined.</p><p>Thailand • holotype; CUMZ 14449 .</p><p>Differential diagnosis.</p><p>This species differs from all other congeners by the strongly detached last whorl, which is also descending near the aperture. From Clostophis species with similar shell morphology ( C. sankeyi W. H. Benson, 1860, C. yoga Páll-Gergely &amp; Hunyadi, 2022, C. udayaditinus Sutcharit &amp; Panha, 2025, C. rhynchotes Tongkerd &amp; Panha, 2025), this species differs by the presence of numerous, elongated apertural barriers reaching the peristome edge in small pliculae.</p><p>Distribution.</p><p>This species is only known from the type locality (Tongkerd et al. 2025).</p></div>	https://treatment.plazi.org/id/68E100D0DC4A5B11A90CA4156EB8846B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
D02523F5845C5C54A9F004882478ECC6.text	D02523F5845C5C54A9F004882478ECC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus rugolabialis Gojsina, Hunyadi & Pall-Gergely 2026	<div><p>Acinolaemus rugolabialis Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 7 T, 41, 42</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell; Shan State, Hopong ca 22 km from the center of Hopong towards Namsang, Hkoche, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.33532&amp;materialsCitation.latitude=20.81806" title="Search Plazi for locations around (long 97.33532/lat 20.81806)">Htem Sann Cave</a>; 20°49.0836'N, 97°20.1192'E; 1240 m a. s. l.; 06 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; CUMZ 15457 . Paratypes. Myanmar • 42 shells; same data as for holotype; coll. HA • 40 shells; Shan State, Kalaw, north-northeast, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.582115&amp;materialsCitation.latitude=20.655333" title="Search Plazi for locations around (long 96.582115/lat 20.655333)">Osei Mountain Pagoda, northeast 150 m</a>; 20°39.320'N, 96°34.927'E; 1565 m a. s. l.; 03 Oct 2018; A. Hunyadi leg.; coll. HA • 17 shells; Shan State, Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.55789&amp;materialsCitation.latitude=20.621027" title="Search Plazi for locations around (long 96.55789/lat 20.621027)">Shwe Oo Min Paya</a>; 20°37.2616'N, 96°33.4735'E; 1340 m a. s. l.; 02 Oct 2018; A. Hunyadi leg.; coll. HA • 96 shells; Shan State, 13.5 km east-southeast from the center of Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.61323&amp;materialsCitation.latitude=20.59044" title="Search Plazi for locations around (long 96.61323/lat 20.59044)">Myinmati Taung</a>; 20°35.4264'N, 96°36.7938'E; 1350 m a. s. l.; 03 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 63 shells; Shan State, Taunggyi, west-southwest, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.01762&amp;materialsCitation.latitude=20.7547" title="Search Plazi for locations around (long 97.01762/lat 20.7547)">Montawa Cave</a>; 20°45.282'N, 97°01.0572'E; 1260 m a. s. l.; 05 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 9 shells; Shan State, Taunggyi, hillside above Aye Say Tee, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.050606&amp;materialsCitation.latitude=20.791477" title="Search Plazi for locations around (long 97.050606/lat 20.791477)">Dragon Cave</a>; 20°47.4886'N, 97°03.0363'E; 1380 m a. s. l.; 08 Oct 2018; A. Hunyadi, K. Okubo &amp; J. U. Otani leg.; coll. HA • 2 paratypes; Shan State, Hopong, Sam Phu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.269966&amp;materialsCitation.latitude=20.685051" title="Search Plazi for locations around (long 97.269966/lat 20.685051)">Cave Ae- 5 at ridge above village Ho Hwe</a>; 20°41.103'N, 97°16.198'E; 12 Feb 2019; J. Grego leg.; coll. JG .</p><p>Additional material examined.</p><p>Myanmar • 15 shells (juveniles = not paratypes); same data as for holotype; coll. HA • 13 shells (juvenile = not paratypes); Shan State, 13.5 km east-southeast from the center of Kalaw, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.61323&amp;materialsCitation.latitude=20.59044" title="Search Plazi for locations around (long 96.61323/lat 20.59044)">Myinmati Taung</a>; 20°35.4264'N, 96°36.7938'E; 1350 m a. s. l.; 03 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; coll. HA .</p><p>Diagnosis.</p><p>Shell conical, weakly spirally striated. Apertural barriers numerous, all main barriers except the parietal are hooked. Peristome wrinkled on the columellar side. Umbilicus wide, ~ 1 / 3 of the shell width.</p><p>Description.</p><p>Shell conical, colorless, consisting of 3.75–4 regularly increasing whorls separated by deep suture; protoconch consisting of ~ 1.25 whorls, initially almost smooth, terminally finely spirally striated (with ~ 12 spiral striae); boundary between protoconch and teleoconch distinct due to rougher surface sculpture of teleoconch; last whorl rounded, adnate to penultimate and almost parallel to shell axis near aperture; surface sculpture of shell consisting of fine spiral striae crossed by even finer, irregularly spaced radial growth lines; surface sculpture most prominent on last whorl, particularly around umbilicus; peristome strong, thick, finely granulated, expanded (except around sinulus) but not reflected; weak cervical swelling present behind peristome (slightly stronger than one found in e. g., A. dayanus, see Tongkerd et al. 2024: fig. 2 A), followed by slight constriction immediately behind it; aperture equipped with eight main barriers (parietal, angular, upper palatal, interpalatal, lower palatal, basal, columellar, and infraparietal) and occasionally additional swelling-like barriers; parietal lamella highest and broadest in aperture, heart shaped in lateral view due to medial, blade-like interval separating blunter, higher and thicker inner and outer portions; outer portion of parietal lamella almost reaches peristome edge as very low, slender projection; angular lamella bipartite: inner portion high, in form of sharp hook pointing outside aperture and outer portion low, not hooked and reaching, or almost reaching peristome edge; two portions of angular lamella connected by less elevated, blade-like interval; palatal plicae (upper, inter, and lower) basal and those on columellar side are all of same morphology: with high inner, hooked portions and low, slender, long, and blade-like portions which do not reach peristome edge; upper palatal plica smallest among palatals, and lower palatal largest; two rounded swellings present on palatal wall: one between upper palatal and interpalatal plica and one between interpalatal and lower palatal plica; these swellings do not develop into proper barriers; palatal tubercle distinct, situated in front of upper palatal plica appears slightly below it in standard view; basal plica small, hooked, situated closer to lower palatal than to columellar; there may be small, blunt swelling between basal plica and lower palatal plica which most probably represents infrapalatal plica; subcolumellar lamella may be present with same form as infrapalatal plica; columellar lamella hooked and not very broad; peristome always distinctly wrinkled in front of columellar lamella; infraparietal lamella low, hooked, and positioned between columellar and parietal lamellae (but closer to parietal); sinulus small, well-separated from remainder of aperture and usually with two or three low plicae inside; umbilicus wide, ~ 1 / 3 of shell width.</p><p>Differential diagnosis.</p><p>This species is unique with the combination of a low, depressed shell, presence of numerous apertural barriers, all of which except the parietal are hooked (including the inner portion of the angular lamella) and a wide umbilicus. It is also readily recognized by the wrinkled peristome on the columellar side.</p><p>Measurements (in mm, n = 5).</p><p>SH = 1.19–1.31; SW = 1.28–1.41; AH = 0.49–0.57; AW = 0.5–0.58.</p><p>Distribution.</p><p>This species is known from seven localities in Shan State, Myanmar.</p><p>Etymology.</p><p>The specific epithet is derived from the Latin rugosa (wrinkled) and labia (lip), referring to the wrinkled columellar side of the peristome.</p><p>Remarks.</p><p>Shells may appear more or less depressed. Specimens from Dragon Cave showed a much stronger lower palatal plica.</p></div>	https://treatment.plazi.org/id/D02523F5845C5C54A9F004882478ECC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
3032F2049E315C738F2474294AEC2DC2.text	3032F2049E315C738F2474294AEC2DC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus simplex Gojsina, Tongkerd & Pall-Gergely 2026	<div><p>Acinolaemus simplex Gojšina, Tongkerd &amp; Páll-Gergely sp. nov.</p><p>Figs 7 E, 14, 17</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Phrae Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.07056&amp;materialsCitation.latitude=18.178055" title="Search Plazi for locations around (long 100.07056/lat 18.178055)">11 km W of Phrae, on Road 1023</a>, limestone knoll, leaf litter; 18°10.6833'N, 100°4.2333'E; 290 m a. s. l.; 16 May 1988; F. G. Thompson leg.; locality code FGT- 4447; UF 591359 . Paratypes. Thailand • 13 shells; same data as for holotype; UF 593498 • 1 shell; same data as for holotype; CUMZ 15426 .</p><p>Additional material examined.</p><p>Thailand • 1 shell (juvenile = not paratype); same data as for holotype; UF 593499 .</p><p>Diagnosis.</p><p>Shell concave-conical, colorless, last whorl weakly shouldered. Shell surface sculpture honeycomb-like. Six apertural barriers (angular, parietal, upper palatal, lower palatal, basal and columellar).</p><p>Description.</p><p>Shell concave-conical, colorless and consisting of 5–5.25 regularly increasing, convex whorls; protoconch spirally striated (approximately 11 spiral striae, clearer terminally) and consisting of 1.5 whorls; boundary between protoconch and teleoconch clearly discernible due to abrupt change of surface sculpture in form of stronger radial growth lines; last whorl weakly shouldered, adnate to penultimate whorl, slightly descending near aperture; surface sculpture honeycomb-like; peristome finely granulated, expanded (except around sinulus), not reflected; cervical swelling distinct, situated immediately behind peristome, and with clear constriction behind it; aperture equipped with six barriers (parietal, angular, upper palatal, lower palatal, basal and columellar); parietal lamella not very high, not reaching peristome; angular lamella longer and thinner than parietal, almost reaching expanding peristome; angular lamella consists of two parts, inner (higher) and outer (lower); two palatal plicae (upper and lower) relatively high, not very slender and of similar appearance (upper palatal being slightly lower and thinner than lower palatal); weak to distinct palatal tubercle sits in front of upper palatal plica and appears slightly below it in standard view; basal plica slender, more than two times lower than lower palatal, situated halfway between lower palatal plica and columellar lamella; columellar lamella usually widest barrier in aperture, oblique, high and not very long, not reaching peristome; sinulus small, distinctly separated from remainder of aperture; umbilicus relatively narrow initially, expanding at last whorl and measuring between 1 / 3 and 1 / 4 of shell width and with shallow umbilical groove; indistinct periumbilical keel visible only near cervical constriction.</p><p>Differential diagnosis.</p><p>In comparison to A. atypicus sp. nov., Acinolaemus simplex sp. nov. has a slenderer shell with more whorls, strong basal plica, narrower umbilicus with a periumbilical keel as well as cervical swelling and a constriction. See also under A. paucidentatus sp. nov.</p><p>Measurements (n = 5, in mm).</p><p>SH = 1.86–1.99; SW = 1.63–1.77; AH = 0.71–0.82; AW = 0.68–0.75.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Etymology.</p><p>The specific epithet refers to the simple appearance of apertural barriers.</p></div>	https://treatment.plazi.org/id/3032F2049E315C738F2474294AEC2DC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
AF27F75C325A589F8B9E30D5D0EF5E81.text	AF27F75C325A589F8B9E30D5D0EF5E81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus singularis Gojsina, Tongkerd & Pall-Gergely 2026	<div><p>Acinolaemus singularis Gojšina, Tongkerd &amp; Páll-Gergely sp. nov.</p><p>Figs 7 S, 40, 42</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Chiang Rai Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.55167&amp;materialsCitation.latitude=20.193611" title="Search Plazi for locations around (long 99.55167/lat 20.193611)">limestone knoll, Ban Mae Song Nai, 4 km N, 6 km NW Mae Chan</a>, limestone domes, shaded ledges; 20°11.6167'N, 99°33.1'E; 520 m a. s. l.; 10 May 1988; F. G. Thompson leg.; locality code FGT- 4425; UF 583728 . Paratypes. Thailand • 3 shells; same data as for holotype; UF 591349 .</p><p>Diagnosis.</p><p>Acinolaemus species with reticulated surface sculpture and no peristomal pliculae. There are 11 apertural barriers. Umbilicus wide, ~ 1 / 3 of the shell width.</p><p>Description.</p><p>Shell weakly concave-conical, light brownish, consisting of 4.5–5 convex whorls separated by deep suture; protoconch spirally striated and consisting of ~ 1.5 whorls with ~ 11 spiral striae; boundary between protoconch and teleoconch distinct because sculpture becomes stronger on initial teleoconch whorls (including presence of radial growth lines); last whorl shouldered, almost parallel to shell axis near aperture; surface sculpture of teleoconch spirally striated; spiral striae crossed by equally strong, radial growth lines forming reticulated pattern; peristome finely granulated, expanded, not reflected; moderate swelling present behind peristome, followed by distinct constriction; aperture equipped with 11 barriers (parietal, angular, upper palatal, two interpalatals, lower palatal, infrapalatal, basal, columellar, supracolumellar, and infraparietal); no peristomal pliculae present; parietal lamella high, not reaching peristome and sometimes with small additional portion anterior to it; angular lamella more slender than parietal, almost reaching peristome edge, bipartite with inner (higher) and outer (lower) portions separated by less elevated, short and blunt interval; upper palatal plica weaker than two interpalatal plicae below it; lower palatal plica widest and highest among all palatals; infrapalatal plica approximately same as interpalatals; palatal tubercle distinct; basal plica slightly weaker than infrapalatal, situated approximately halfway between lower palatal and columellar; columellar lamella strong, thick, slightly oblique, almost reaching peristome; supracolumellar lamella weak, tubercle-like; infraparietal lamella situated closer to parietal than to columellar, approximately as strong as interpalatal plicae; sinulus small, distinctly separated from remainder of aperture; umbilicus wide, ~ 1 / 3 of shell width, expanding at last whorl.</p><p>Differential diagnosis.</p><p>This is the only Acinolaemus species that has a combination of reticulated surface sculpture and no peristomal pliculae. Otherwise, it is most similar to A. ptychochilus . Apart from the absence of peristomal pliculae, the new species lacks the hooked apertural barriers which can separate it from A. ptychochilus .</p><p>Measurements (in mm, n = 4).</p><p>SH = 1.48–1.64; SW = 1.46–1.51; AH = 0.69–0.74; AW = 0.62–0.65.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Etymology.</p><p>The specific epithet is derived from the Latin singularis, meaning singular, the only one, which is a reference to the fact that this species is the only currently known representative of Acinolaemus with reticulated surface pattern and no peristomal pliculae.</p></div>	https://treatment.plazi.org/id/AF27F75C325A589F8B9E30D5D0EF5E81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
BC74D0108CC65D8BBD8C49C1EAF1A4CD.text	BC74D0108CC65D8BBD8C49C1EAF1A4CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus solitus Gojsina, Hunyadi & Pall-Gergely 2026	<div><p>Acinolaemus solitus Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 7 F, 15 –17</p><p>Type material.</p><p>Holotype. Thailand • 1 shell; Tak Province, knoll 5.5 km NNW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.20417&amp;materialsCitation.latitude=17.270279" title="Search Plazi for locations around (long 98.20417/lat 17.270279)">Tha Song Yang</a>; 17°16.2167'N, 98°12.25'E; 180 m a. s. l.; 03 May 1988; F. G. Thompson leg.; locality code FGT- 4407; UF 347039 . Paratypes. Thailand • 2 shells; same data as for holotype; UF 593500 • 90 shells; Tak Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.2109&amp;materialsCitation.latitude=17.256016" title="Search Plazi for locations around (long 98.2109/lat 17.256016)">6 km NNW from Tha Song Yang, left side of road no. 105</a>; 17°15.361'N, 98°12.654'E; 140 m a. s. l.; 15 Feb 2015; A. Hunyadi leg.; coll. HA .</p><p>Additional material examined.</p><p>Thailand • 1 shell (broken = not paratype); same data as for holotype; UF 593501 • 20 shells (juveniles / broken = not paratypes); Tak Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.2109&amp;materialsCitation.latitude=17.256016" title="Search Plazi for locations around (long 98.2109/lat 17.256016)">6 km NNW from Tha Song Yang, left side of road no. 105</a>; 17°15.361'N, 98°12.654'E; 140 m a. s. l.; 15 Feb 2015; A. Hunyadi leg.; coll. HA .</p><p>Diagnosis.</p><p>Shell depressed concave-conical. Last whorl shouldered, surface sculpture honeycomb-like. Aperture equipped with numerous elongated barriers. Umbilicus wide, measuring ~ 1 / 3 of the shell width.</p><p>Description.</p><p>Shell depressed concave-conical, light brownish, consisting of 3.5–3.75 convex whorls separated by deep suture; protoconch consisting of ~ 1.5 whorls and with ~ 10 clear spiral striae; boundary between protoconch and teleoconch clearly visible due to change of surface sculpture (honeycomb-like on teleoconch); last whorl shouldered, flattened below shoulder, adnate to penultimate whorl and almost parallel to shell axis in lateral view; surface sculpture honeycomb-like; peristome finely granulated, expanded (except around sinulus) but not reflected; cervical swelling and constriction both absent; aperture equipped with numerous barriers; parietal lamella long, high, slowly elevating towards its middle, not reaching peristome edge; angular lamella lower, more slender, surpassing profile of parietal lamella and reaching peristome edge; angular lamella consists of inner and outer, blunt-topped portions separated by blade-like (sharp), less elevated interval; three palatal plicae (upper, inter and lower), equally strong, higher and blunt at their inner portions and sloping towards peristome in form of slender, blade-like projections; palatal tubercle distinct, sitting in front of upper palatal plica, appearing slightly below it in standard view; basal plica of same morphology but slightly lower than palatals; columellar lamella strong (high and long), oblique (curved towards basal lip) and almost reaching peristome; infraparietal lamella moderate, two times lower than columellar, situated halfway between columellar and parietal lamella; sinulus small, distinctly separated from remainder of aperture; umbilicus wide, ~ 1 / 3 of shell width.</p><p>Differential diagnosis.</p><p>Acinolaemus cryptidentatus is larger, less depressed, has a narrower umbilicus and more apertural barriers (including additional small barriers inside the sinulus). See under A. dayanus and A. corusticorus .</p><p>Measurements (n = 3, in mm).</p><p>SH = 1.03–1.12; SW = 1.18–1.26; AH = 0.49–0.53; AW = 0.49–0.51.</p><p>Distribution.</p><p>This species is known from two closely situated localities in Tak Province.</p><p>Etymology.</p><p>The Latin-derived specific epithet solitus, meaning usual, refers to the fact that there are no peculiar, striking, or unique characters that separate this species from its similar congeners (i. e., it is unremarkable).</p></div>	https://treatment.plazi.org/id/BC74D0108CC65D8BBD8C49C1EAF1A4CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
AFD997A92C3150928450E79AF73095B7.text	AFD997A92C3150928450E79AF73095B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus undefined-1	<div><p>Acinolaemus sp. 1</p><p>Figs 7 N, 29, 30</p><p>Material examined.</p><p>Thailand • 1 whole shell and 3 fragments; Tak Province, ravine 16 km NNW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.444725&amp;materialsCitation.latitude=17.058332" title="Search Plazi for locations around (long 98.444725/lat 17.058332)">Mae Ramat</a>; 17°3.5'N, 98°26.6833'E; 220 m a. s. l.; 03 May 1988; F. G. Thompson leg.; locality code FGT- 4405; UF 347020 .</p><p>Remarks.</p><p>This is most likely an undescribed species. The single complete specimen is by the apertural barrier arrangement and shell shape most similar to A. corusticorus . However, the plicae in the undescribed species are much thinner (slender, blade-like) and there are also barriers inside the sinulus (absent in A. corusticorus). However, as we have just a single complete specimen, we choose not to describe it here due to the striking similarity with A. corusticorus .</p></div>	https://treatment.plazi.org/id/AFD997A92C3150928450E79AF73095B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
3D7FC59397DE5471B2FA74C7D7583D97.text	3D7FC59397DE5471B2FA74C7D7583D97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acinolaemus zed Gojsina, Hunyadi & Pall-Gergely 2026	<div><p>Acinolaemus zed Gojšina, Hunyadi &amp; Páll-Gergely sp. nov.</p><p>Figs 7 M, 28, 30</p><p>Type material.</p><p>Holotype. Myanmar • 1 shell; Mon State, ca 14 km northeast from the center of Mawlamyine, Kalagon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.71517&amp;materialsCitation.latitude=16.533092" title="Search Plazi for locations around (long 97.71517/lat 16.533092)">Kha Yone cave</a>; 16°31.9856'N, 97°42.9102'E; 11 Oct 2018; A. Hunyadi, K. Okubo, J. U. Otani leg.; CUMZ 15456 . Paratypes. Myanmar • 37 shells; same data as for holotype; coll. HA .</p><p>Additional material examined.</p><p>Myanmar • 3 shells (damaged / juveniles = not paratypes); same data as for holotype; coll. HA .</p><p>Diagnosis.</p><p>Shell colorless, depressed concave-conical. Last whorl shouldered, surface sculpture honeycomb-like. Apertural barriers numerous, slender. Umbilicus wide, with both a strong periumbilical keel and a deep groove.</p><p>Description.</p><p>Shell depressed, concave-conical, light yellowish, consisting of 3.75–4 whorls separated by deep suture; protoconch with ~ 10 spiral striae (clearer terminally), consisting of ~ 1.25 whorls; boundary between protoconch and teleoconch distinct due to change of surface sculpture (rougher, with both radial and spiral growth elements); teleoconch surface sculpture honeycomb-like; lower portion of the body whorl sometimes with more distinct spiral striae while upper portion has sculpture of honeycomb-like pattern; in some specimens, last whorl is entirely sculptured with honeycomb-like pattern; last whorl shouldered and slightly descending near aperture; peristome finely granulated, expanded (especially at palatal and columellar sides, not around sinulus), not reflected; no distinct cervical swelling or constriction present; aperture equipped with numerous slender apertural barriers; parietal lamella high, curved, blunt, almost fully reaching peristome edge; angular lamella of similar appearance, reaching peristome edge, consisting of inner (higher and broader) and outer (lower and narrower) blunt-topped portions separated by short, less elevated, blade-like interval; outer portion of angular lamella sigmoid; three main palatal plicae present; upper palatal plica higher and thicker at inner portion, while lower and more slender (blade-like) towards peristome; low and slender suprapalatal plica present above upper palatal; interpalatal plica widest among all palatals but becomes significantly narrower and blade-like towards peristome, reaching it in form of blunt plicula; palatal tubercle strong, sitting in front of upper palatal plica and appearing slightly below it in standard view; lower palatal plica of same morphology as upper palatal, slightly thinner and lower, reaching the peristome in form of blunt plicula; infrapalatal plica slightly lower than lower palatal; inner portion of infrapalatal plica blunt, sloping as blade-like, not reaching peristome; columellar lamella wide, oblique, blunt and almost fully reaching peristome edge; infraparietal lamella lower than both parietal and columellar, situated halfway between them, blunt; umbilicus wide, ~ 1 / 3 of shell width; strong periumbilical keel running along umbilicus; deep groove present inside the umbilicus.</p><p>Differential diagnosis.</p><p>This species differs from A. microcubus sp. nov. by the stronger periumbilical keel and the presence of a deep groove inside the umbilicus as well as the much less strongly shouldered last whorl. Acinolaemus microcubus sp. nov. is also a bit smaller than A. zed sp. nov. Acinolaemus dayanus has a narrower umbilicus, lacks both the periumbilical keel and the umbilical groove, has a less distinctly shouldered last whorl and has overall weaker (lower) apertural barriers. Acinolaemus humilis sp. nov. is similar in apertural barrier arrangement and strength but the shell is smaller, has a less shouldered last whorl, and lacks the periumbilical keel as well as the umbilical groove. Also, peristomal pliculae are lacking in A. humilis sp. nov.</p><p>Measurements (in mm, n = 5).</p><p>SH = 1.21–1.34; SW = 1.56–1.68, AH = 0.57–0.67; AW = 0.65–0.71.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Etymology.</p><p>The specific epithet zed is derived from the Latin zeta referring to the fact that this is the last species to be described herein in this species group of Acinolaemus .</p><p>Remarks.</p><p>The difference between the sculpture of the upper portion of the last whorl (typically honeycomb-like) and the lower portion (with distinct spiral striae) was observed in 14 paratypes. The pronounced periumbilical keel and a deep umbilical groove seem to be characteristic for this species as these characters were consistent in all paratypes.</p></div>	https://treatment.plazi.org/id/3D7FC59397DE5471B2FA74C7D7583D97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
2EFF43A18C0B5AE4A56CE0F3126DC122.text	2EFF43A18C0B5AE4A56CE0F3126DC122.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis carcharodon (Vermeulen, Phung & Truong 2007) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis carcharodon (Vermeulen, Phung &amp; Truong, 2007) comb. nov.</p><p>Acinolaemus carcharodon Vermeulen, Phung &amp; Truong, 2007: 87, fig. 4.</p><p>Acinolaemus carcharodon — Changlom et al. 2019: 156; Schileyko 2011: 4; Sutcharit et al. 2023: 1289 –1290, fig. 1, table 1; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Vietnam, Kien Giang Prov., Kien Luong, Hon Chong Hill ”.</p><p>Differential diagnosis.</p><p>When compared to C. carcharodon, C. udayaditinus Sutcharit &amp; Panha, 2025 has a less depressed shell, more strongly detached last whorl from the penultimate, and has a columellar lamella consisting of two hooked portions directed towards each other. See under C. pyramidalis .</p><p>Distribution.</p><p>This species is known from the Kien Giang Province, Vietnam and the Kampot Province, Cambodia.</p><p>Remarks.</p><p>This species is transferred to Clostophis due to the similar apertural barrier arrangement and morphology as in C. lacrima (Páll-Gergely &amp; Hunyadi, 2015), C. obliquus Páll-Gergely &amp; Hunyadi, 2022 and C. socialis (Páll-Gergely &amp; Hunyadi, 2015) (i. e., strong and moderately long barriers, including two strong palatal plicae). Furthermore, C. carcharodon lacks the spiral sculpture on the protoconch, the peculiar honeycomb-like surface sculpture of the shell (present in many Acinolaemus species) and peristomal pliculae (present in A. ptychochilus, the type species of Acinolaemus).</p></div>	https://treatment.plazi.org/id/2EFF43A18C0B5AE4A56CE0F3126DC122	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
D27D840500305DB7A73F210ED4B7B8EF.text	D27D840500305DB7A73F210ED4B7B8EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis colpodon (F. G. Thompson & Upatham 1997) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis colpodon (F. G. Thompson &amp; Upatham, 1997) comb. nov.</p><p>Fig. 43</p><p>Acinolaemus colpodon F. G. Thompson &amp; Upatham, 1997: 229–230, figs 3, 26–31.</p><p>Acinolaemus colphodon [sic] — Changlom et al. 2019: 156.</p><p>Acinolaemus colpodon — Boonngam et al. 2008: 258, table 1; Panha and Burch 2008: 40 –41, fig. 29; Dumrongrowjattana and Wongkamhaeng 2024: 321, 323, 330, fig. 7, 13, table 3; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Thailand, Rayong Province, Khao Bot, ca 6 km north and 2 km west of Ban Syaek Batan; 150 m altitude (13°02.8'N, 101°38.5'E) ”.</p><p>Type material examined.</p><p>Thailand • holotype; UF 113528 • 5 paratypes (UF 113529) .</p><p>Additional material examined.</p><p>Thailand • 386 shells; Rayong Province, Khao Chamao district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.5975&amp;materialsCitation.latitude=13.073983" title="Search Plazi for locations around (long 101.5975/lat 13.073983)">Wat Tham Khao Pratun</a>, gorge above the temple; 13°04.439'N, 101°35.850'E; 115 m a. s. l.; 09 Mar 2023; A. Hunyadi leg.; coll. HA . Cambodia • Banteay Meanchey Province, Sisophon N, Pongro W 1500 m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.94427&amp;materialsCitation.latitude=13.63241" title="Search Plazi for locations around (long 102.94427/lat 13.63241)">Prasat Phnom Korngva</a>; 13.63241°N, 102.94427°E; 50 m a. s. l.; 28 Oct 2023; A. Hunyadi leg.; coll. HA .</p><p>Differential diagnosis.</p><p>This species is readily distinguished from other Clostophis and Acinolaemus species by the strikingly upwardly directed aperture (making it virtually parallel to the shell axis). This shape makes it superficially more similar to Hypselostoma erawan Panha &amp; J. B. Burch, 2003 and H. populare Gojšina, Hunyadi &amp; Páll-Gergely, 2025 . Hypselostoma erawan has more apertural barriers (especially on the palatal side), and its angular and columellar lamellae are not hooked. Hypselostoma populare has none of its barriers hooked. Furthermore, both H. erawan and H. populare are much larger than C. colpodon, both are brownish ( C. colpodon is colorless) and both have more elevated shells.</p><p>Distribution.</p><p>This species is known from Rayong and Chanthaburi Provinces, Thailand, as well as from Banteay Meanchey Province, Cambodia.</p><p>Remarks.</p><p>This species is transferred to Clostophis because it has only a single lamella on the parietal side (two in all Acinolaemus species) and it lacks the typical reticulated surface sculpture (as found in the type species of Acinolaemus) or characteristic honeycomb-like surface (as found in a number of Acinolaemus species). Furthermore, the apertural barrier arrangement is of the same type as that of C. udayaditinus, i. e., there is a strong palatal plica, hooked angular lamella and bipartite columellar lamella consisting of hooked inner and outer parts. See Dumrongrojwattana and Wongkamhaeng (2024) for additional sampling sites in Chanthaburi and Rayong provinces.</p></div>	https://treatment.plazi.org/id/D27D840500305DB7A73F210ED4B7B8EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
B5608DC2829E5843AB3107713F3584B7.text	B5608DC2829E5843AB3107713F3584B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis pyramidalis (Vermeulen, Phung & Truong 2007) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis pyramidalis (Vermeulen, Phung &amp; Truong, 2007) comb. nov.</p><p>Montapiculus pyramidalis Vermeulen, Phung &amp; Truong, 2007: 91, fig. 5.</p><p>Montapiculus pyramidalis — Schileyko 2011: 4.</p><p>Acinolaemus pyramidalis — Vermeulen et al. 2019: 32 –34, fig. 52; Sutcharit et al. 2023: 1289 –1290, fig. 1, table 1; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Vietnam, Kien Giang Prov., Kien Luong, Hon Chong Hill ”.</p><p>Differential diagnosis.</p><p>This species, in comparison to C. carcharodon, has no hooked barriers and it possesses a less depressed shell. See also under C. rectus .</p><p>Distribution.</p><p>This species is known from Kien Giang Province in Vietnam and Kampot Province in Cambodia.</p><p>Remarks.</p><p>This species is transferred to Clostophis due to the similar apertural barrier arrangement and morphology as in C. lacrima, C. obliquus and C. socialis (i. e., strong and moderately long barriers, including two strong palatal plicae). Furthermore, C. pyramidalis lacks the peculiar honeycomb-like surface sculpture of the shell (present in many Acinolaemus species) and peristomal pliculae (present in A. ptychochilus, the type species of Acinolaemus).</p></div>	https://treatment.plazi.org/id/B5608DC2829E5843AB3107713F3584B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
E48872860B3F5D749EC22EA502111CC8.text	E48872860B3F5D749EC22EA502111CC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis rectus (Vermeulen, Luu, Theary & Anker 2019) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis rectus (Vermeulen, Luu, Theary &amp; Anker, 2019) comb. nov.</p><p>Fig. 44</p><p>Acinolaemus rectus Vermeulen, Luu, Theary &amp; Anker, 2019: 34–35, 52, 54–56.</p><p>Acinolaemus rectus — Sutcharit et al. 2023: 1289 –1290, fig. 1, table 1; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Cambodia: Kampot Province, Kampong Trach area: Phnom Kampong Trach ”.</p><p>Type material examined.</p><p>Cambodia • holotype; NHMUK 20180572 .</p><p>Differential diagnosis.</p><p>This species differs from A. pyramidalis by the slightly upturning aperture near the last whorl (down-turning in A. pyramidalis). See also under C. rhamphodon and C. sphinctinion .</p><p>Distribution.</p><p>This species is known only from Kampot Province in Cambodia.</p><p>Remarks.</p><p>This species is transferred to Clostophis due to the similar apertural barrier arrangement and morphology as in C. lacrima, C. obliquus and C. socialis (i. e., strong and moderately long barriers, including two strong palatal plicae). Furthermore, C. pyramidalis lacks the peculiar honeycomb-like shell surface sculpture (present in many Acinolaemus species) and peristomal pliculae (present in A. ptychochilus, the type species of Acinolaemus).</p></div>	https://treatment.plazi.org/id/E48872860B3F5D749EC22EA502111CC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
CF163693EEDE50A08A3C32F980B1CD25.text	CF163693EEDE50A08A3C32F980B1CD25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis rhamphodon (F. G. Thompson & Upatham 1997) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis rhamphodon (F. G. Thompson &amp; Upatham, 1997) comb. nov.</p><p>Fig. 45</p><p>Acinolaemus rhamphodon F. G. Thompson &amp; Upatham, 1997: 227–228, figs 4, 20–25.</p><p>Acinolaemus ramphodon [sic] — Changlom et al. 2019: 156.</p><p>Acinolaemus rhamphodon — Panha and Burch 2008: 43 –44, fig. 41; Dumrongrowjattana and Wongkamhaeng 2024: 321, 323, 330, 331, fig. 7, 13, table 3, 5; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Thailand, Chachaengsao Province, Khao Tam Raet, 5 km ENE Ban Nan Khok, 100 m altitude (13°26.1'N, 101°44.2'E) ”.</p><p>Type material examined.</p><p>Thailand • holotype; UF 113505 • 1 paratype; UF 113506 .</p><p>Additional material examined.</p><p>Thailand • 4 shells; Rayong Province, Khao Chamao District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.63533&amp;materialsCitation.latitude=13.037884" title="Search Plazi for locations around (long 101.63533/lat 13.037884)">rock wall behind Wat Tham Khao Bot</a>; 13°02.273'N, 101°38.120'E; 70 m a. s. l.; 09 Mar 2023; A. Hunyadi leg.; coll. HA • 21 shells; Chonburi Province, Bo Thong District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.59763&amp;materialsCitation.latitude=13.162916" title="Search Plazi for locations around (long 101.59763/lat 13.162916)">rock wall behind Wat Khao Ha Yot</a>; 13°09.775'N, 101°35.858'E; 150 m a. s. l.; 09 Mar 2023; A. Hunyadi leg.; coll. HA • 52 shells; Rayong Province, Khao Chamao District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.5975&amp;materialsCitation.latitude=13.073983" title="Search Plazi for locations around (long 101.5975/lat 13.073983)">Wat Tham Khao Pratun</a>, gorge above the temple; 13°04.439'N, 101°35.850'E; 115 m a. s. l.; 09 Mar 2023; A. Hunyadi leg.; coll. HA .</p><p>Differential diagnosis.</p><p>Clostophis rectus has a wider umbilicus, narrower (lower) last whorl and lacks the hooked columellar lamella. See under C. sphinctinion and C. stenopus .</p><p>Distribution.</p><p>This species is known from Chachoengsao, Rayong, Chanthaburi and Chonburi provinces, Thailand.</p><p>Remarks.</p><p>This species is transferred to Clostophis due to the similar apertural barrier arrangement and morphology as in C. lacrima, C. obliquus, and C. socialis (i. e., strong and moderately long barriers, including two strong palatal plicae). Furthermore, C. rhamphodon lacks both the peculiar honeycomb-like shell surface sculpture (present in many Acinolaemus species) and peristomal pliculae (present in A. ptychochilus, the type species of Acinolaemus). Specimens from Wat Tham Khao Bot (Rayong Province) have a blunt (not hooked) columellar lamella. Since this was the only significant difference from typical A. rhamphodon we could find, we treat this as intraspecific variability. See Dumrongrojwattana and Wongkamhaeng (2024) for additional sampling sites.</p></div>	https://treatment.plazi.org/id/CF163693EEDE50A08A3C32F980B1CD25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
F9B1E4B1CD235DC78C0CBB755E351CEE.text	F9B1E4B1CD235DC78C0CBB755E351CEE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis sphinctinion (F. G. Thompson & Upatham 1997) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis sphinctinion (F. G. Thompson &amp; Upatham, 1997) comb. nov.</p><p>Fig. 46</p><p>Acinolaemus sphinctinion F. G. Thompson &amp; Upatham, 1997: 226–227, figs 16–19.</p><p>Acinolaemus sphinctinion — Panha and Burch 2008: 44 –45, fig. 42; Changlom et al. 2019: 156; Dumrongrowjattana and Wongkamhaeng 2024: 321, 323, 330, fig. 7, 13, table 3; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Thailand, Prachuap Khiri Khan Province, Khao Sam Roi Yoi National Park, limestone knoll on east side of brackish marsh 3 km south of park entrance; (12°12.5'N, 99°59.6'E) ”.</p><p>Type material examined.</p><p>Thailand • holotype; UF 113507 .</p><p>Differential diagnosis.</p><p>This species differs from C. rectus and C. rhamphodon by the merged angular and parietal lamellae into the angulo-parietal. It also has more angled whorls and a narrower (lower) last whorl with a distinct constriction behind the aperture.</p><p>Distribution.</p><p>This species is known only from the type locality.</p><p>Remarks.</p><p>This species is provisionally assigned to Clostophis due to similar apertural barrier arrangement as in C. lacrima, C. obliquus and C. socialis . It does not have a honeycomb-like shell surface, or peristomal pliculae, and its protoconch is not spirally striated which is why it is excluded from Acinolaemus . Lastly, it has a merged angulo-parietal lamella (separated in all Acinolaemus species). It is possible that this species might warrant its own genus, introducing a new name herein would be premature due to the unclear generic boundaries in the entire family.</p></div>	https://treatment.plazi.org/id/F9B1E4B1CD235DC78C0CBB755E351CEE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
BF1879DF10615F9E8FDA2FE4255B0BF2.text	BF1879DF10615F9E8FDA2FE4255B0BF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis stenopus (F. G. Thompson & Upatham 1997) Gojšina & Hunyadi & Sutcharit & Tongkerd & Auffenberg & Grego & Páll-Gergely 2026	<div><p>Clostophis stenopus (F. G. Thompson &amp; Upatham, 1997) comb. nov.</p><p>Fig. 47</p><p>Acinolaemus stenopus F. G. Thompson &amp; Upatham, 1997: 228–229, figs 2, 12–15.</p><p>Acinolaemus stenopus — Changlom et al. 2019: 156; Panha and Burch 2008: 45 –46, fig. 43; Dumrongrowjattana and Wongkamhaeng 2024: 321, 323, 330, 331, fig. 7, 13, table 3, 5; Tongkerd et al. 2025: 53, table 4.</p><p>Type locality.</p><p>“ Thailand, Chanthaburi Province, limestone ridge 3 km w N Yai Ham; 25 m altitude (12°44.3'N, 101°52.9'E) ”.</p><p>Type material examined.</p><p>Thailand • holotype; UF 113530 • 5 paratypes; UF 113531 .</p><p>Differential diagnosis.</p><p>This species differs from C. rhamphodon most clearly by the presence of a strong and curved angular lamella which is directed towards the upper palatal plica and almost merging with it (leaving only a very narrow passage) and enclosing the sinulus.</p><p>Distribution.</p><p>This species is known from Chanthaburi and Rayong provinces, Thailand (Dumrongrojwattana and Wongkamhaeng 2024).</p><p>Remarks.</p><p>This species is excluded from Acinolaemus due to the apertural barrier arrangement which is strikingly different from A. ptychochilus and more similar to some Clostophis species, namely C. lacrima, C. obliquus and C. socialis . The trend of the curved angular lamella almost merging with the upper palatal plica in C. stenopus is also apparent in cases of the angulo-parietal lamella in mentioned Clostophis species.</p></div>	https://treatment.plazi.org/id/BF1879DF10615F9E8FDA2FE4255B0BF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
F7EBC9B2BDA956B48BB28321304992DB.text	F7EBC9B2BDA956B48BB28321304992DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clostophis W. H. Benson 1860	<div><p>Genus Clostophis W. H. Benson, 1860</p><p>Clostophis Benson, 1860: 95 .</p><p>Montapiculus Panha &amp; Burch, 1999: 148 .</p><p>Type species.</p><p>Clostophis sankeyi Benson, 1860, by monotypy.</p><p>Remarks.</p><p>The following species are transferred from Acinolaemus to Clostophis:</p></div>	https://treatment.plazi.org/id/F7EBC9B2BDA956B48BB28321304992DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Gojšina, Vukašin;Hunyadi, András;Sutcharit, Chirasak;Tongkerd, Piyoros;Auffenberg, Kurt;Grego, Jozef;Páll-Gergely, Barna	Gojšina, Vukašin, Hunyadi, András, Sutcharit, Chirasak, Tongkerd, Piyoros, Auffenberg, Kurt, Grego, Jozef, Páll-Gergely, Barna (2026): A taxonomic revision of the land snail genus Acinolaemus F. G. Thompson & Upatham, 1997 (Gastropoda, Eupulmonata, Hypselostomatidae), with notes on Clostophis W. H. Benson, 1860 species from Cambodia, Thailand, and Vietnam. ZooKeys 1279: 101-165, DOI: 10.3897/zookeys.1279.186771
