taxonID	type	description	language	source
9C1AF9689C01FFFD74EF68CCFE82FC02.taxon	type_taxon	Type and only species. Densipsychops pectinatus sp. nov.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF68CCFE82FC02.taxon	diagnosis	Diagnosis. Sc fused with RA distally; RP densely branches (about 30 branches), and most RP branches shallowly forked; MA simply branched with a few distal branches; MP and CuA pectinately branched near its middle part, sharing similar branching pattern; anterior branch of CuP pectinately branched, posterior branch of CuP deeply forked.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF68CCFE82FC02.taxon	etymology	Etymology. From the Latin prefix Dens - (meaning “ dense ”), and - psychops, a traditional generic ending of Osmylopsychopidae, in reference to the dense veins of forewing. Gender is masculine.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF68CCFE82FC02.taxon	discussion	Remarks. Densipsychops gen. nov. belongs to Osmylopsychopidae for sharing the distinct characters of this family, i. e. same broad wing shape, costal space distinctly narrowed towards end, and RP 1 deeply branched. Moreover, this new genus can be easily distinguished from other osmylopsychopid genera by its shallowly forked RP branches (vs. most RP branches deeply forked in Archepsychops, Brongniartiella, Daopsychops, Eupypsychops, Osmylopsychoides, Petropsychops, Triassopsychops and Undulopsychopsis) (Peng et al., 2011; Lambkin, 2014; Peng et al., 2015; Khramov et Makarkin, 2015), simple MA with a few distal branches (vs. MA forked near wing base or middle part in Gayndahpsychops, Minipsychops and Nipponopsychops) (Lambkin, 2014; Khramov et al., 2024; Liu et al., 2023), more than ten pectinate MP near middle part (vs. MP forked with less than five branches distally in Nematopsychops, Osmylopsychostoechus and Psychostoechotes) (Peng et al., 2015; Khramov et Makarkin, 2015) and CuP deeply forked several times near wing base (vs. CuP deeply forked only 1 – 2 times in Osmylopsychops, Ochropsychops and Puripsychops; CuP forked once time distally in Stenopteropsychops) (Ellenberger et al., 1952; Peng et al., 2015).	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF6B93FA7FF90F.taxon	description	Figure 1	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF6B93FA7FF90F.taxon	materials_examined	Material. Holotype: CNU-NEU-NN- 2024004, two incomplete forewings and a poorly preserved hind wing.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF6B93FA7FF90F.taxon	etymology	Etymology. From the Latin pectinatus (meaning “ pectinate ”), referring to the pectinately branched MP of this new species. Locality and horizon. latest Middle Jurassic (Upper Callovian), Jiulongshan Formation; Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF6B93FA7FF90F.taxon	diagnosis	Diagnosis. As for the genus.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFD74EF6B93FA7FF90F.taxon	description	Description. Forewing relatively elongated, ca. 17 – 18 mm long and 9 – 10 mm wide. Trichosors indistinct. Basal costal space broad, distinctly narrowed towards the distal. Costal gradate series present. Subcostal veinlets dense, most forked 2 – 3 times. Humeral veinlets strongly recurrent with several sub-branches. Sc fused with RA distally. At least one crossvein present at subcostal space. RA simple with few branches distally. RP pectinate with about 30 primary branches, most shallowly forked. RP 1 deeply branched twice. M forked near wing base, MA forked with few distal branches, MP pectinately branched near the middle. Cu forked at wing base; CuA pectinately branched, similar to MP branching; CuP forked near wing base, anterior branch of CuP pectinately branched, posterior branch of CuP deeply forked. A 1, A 2 and A 3 forked with complicated sub-branches. Large dark irregularly transverse markings present on forewing. Hind wing poorly preserved, ca. 15.5 mm long and 8.5 mm wide. All preserved venations similar to forewing.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFB74EF6E8DFEB7FD99.taxon	type_taxon	Type and only species. Puripsychops eurypterus sp. nov.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFB74EF6E8DFEB7FD99.taxon	diagnosis	Diagnosis. Outer wing margin smooth; most RP branches shallowly forked; M forked near the middle, MA branched with several sub-branches, MP deeply dichotomously branched near the middle part of wing, two primary branches forming several sub-branches; CuP deeply dichotomously forked once.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFB74EF6E8DFEB7FD99.taxon	etymology	Etymology. Derived from the Latin prefix puri - (meaning “ pure ”), and - psychops, a traditional generic ending of Osmylopsychopidae, referring to the absence of wing markings. Gender is masculine.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C01FFFB74EF6E8DFEB7FD99.taxon	discussion	Remarks. Puripsychops gen. nov. is most characterized by its configuration of M. M forked at its middle part in this new genus while M forked near the wing base in Archepsychops, Brongniartiella, Gayndahpsychops, Osmylopsychops, Petropsychops, Triassopsychops and Nipponopsychops (Lambkin, 2014; Peng et al., 2015; Khramov et al., 2024); Additionally, MA forked distally in Puripsychops while MA forked near the middle or near the base of wing in Minipsychops (Liu et al., 2023); MP forked near the middle part of wing in Puripsychops while MP forked distally in Daopsychops, Eupypsychops, Nematopsychops, Osmylopsychoides, Osmylopsychostoechus, Psychostoechotes and Stenopteropsychops (Peng et al., 2015; Khramov et Makarkin, 2015). Moreover, this new genus can be easily distinguished from other genera by its smooth wing margin (vs. outer wing margin undulate in Undulopsychopsis) (Peng et al., 2011), the deeply forked RP 1 (vs. RP 1 shallowly forked in Ochropsychops) (Peng et al., 2015) and CuP deeply forked once time (vs. CuP deeply forked several times with numerous branches in Densipsychops).	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C07FFF874EF6A31FC66FD99.taxon	description	Figure 2	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C07FFF874EF6A31FC66FD99.taxon	materials_examined	Material. Holotype: CNU-NEU-NN- 2024008, a nearly completed specimen including two relatively completed forewings and two relatively completed hind wings.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C07FFF874EF6A31FC66FD99.taxon	etymology	Etymology. From the Latin Eurypterus (meaning “ wide wing ”), referring that the forewing of this new species is significantly wider compared to other species of Osmylopsychopidae. Locality and horizon. latest Middle Jurassic (Upper Callovian), Jiulongshan Formation; Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C07FFF874EF6A31FC66FD99.taxon	diagnosis	Diagnosis. As for the genus.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C07FFF874EF6A31FC66FD99.taxon	description	Description. Medium body size. Forewing wide and nearly triangular, ca. 16 - 17 mm long and 8.5 - 10 mm wide. Basal costal space broad and significantly narrowed towards the distal. Subcostal veinlets dense, most forked distally. Humeral veinlets strongly recurrent with several branches. Sc fused with RA distally. At least one crossvein present at subcostal space. RA with several pectinate branches distally. RP pectinate with about 20 primary branches, most shallowly forked. RP 1 deeply forked with several branches. M forked near its middle part, MA forked medially with a few distal branches, MP deeply dichotomously forked with several branches. Cu forked at wing base, CuA pectinately branched with several branches. CuP dichotomously forked once, two primary branches forming a few sub-branches distally. A 1 and A 2 complicatedly forked with numerous branches, A 3 forked near the wing margin. Hind wing smaller than forewing, relatively elongated, ca. 15 – 17 mm long and 7.5 – 8.5 mm wide. Costal space nearly equal width. Other preserved venations similar to forewing.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6B3DFA82F8CD.taxon	description	Figure. 3	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6B3DFA82F8CD.taxon	materials_examined	Material. Holotype: CNU-NEU-NN- 2024009, an incomplete forewing and an incomplete hind wing. Paratype: CNU-NEU-NN- 2024010, an incomplete and compressed forewing.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6B3DFA82F8CD.taxon	etymology	Etymology. From the Latin oligophlebius (meaning “ a few branches ”), referring that the number of its CuP branches is relatively small compared to other species of Eupypsychops. Locality and horizon. latest Middle Jurassic (Upper Callovian), Jiulongshan Formation; Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6B3DFA82F8CD.taxon	diagnosis	Diagnosis. The outer radial gradates series of forewing absent, CuP dichotomously branched with a few branches near its middle part, no distinct wing patterns.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6B3DFA82F8CD.taxon	description	Description. Forewing broad, ca. 32 – 36 mm long and 20 – 22 mm wide. Trichosors indistinct. Basal costal space broad and significantly narrowed towards end. Subcostal veinlets dense and abundant, most forked 1 – 2 times. A costal gradate series of crossveins present. Humeral veinlets strongly recurrent with several branches. Sc not fused with RA distally. RA with few distal pectinate branches. RP pectinately branched with about 15 primary branches, most deeply forked more than once. RP 1 deeply forked 3 – 4 times with several branches. M forked near wing base, MA and MP simple with similar configuration. Cu forked at wing base, CuA pectinately branched with numerous branches medially, CuP dichotomously branched with a few branches near its middle part. Anal space broad, A 1, A 2 and A 3 complicated dichotomously branched. Wing patterns indistinct. Hind wing broad and slightly smaller than forewing, ca. 33 mm long and 19 mm wide. Numerous radial crossveins forming an outer gradate series. Other venations similar to forewing.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6B3DFA82F8CD.taxon	discussion	Remark. This new species is assigned to Eupypsychops Peng et al., 2015 for sharing the similar venation with this genus, i. e. Sc not fused with RA distally, most RP branches deeply forked, RP 1 deeply forked at least three times, M forked slightly earlier than the first branched point of CuA and MP dichotomously forked distally. Additionally, it can be clearly differentiated from other species of Eupypsychops by the absence of radial gradate series in forewing (vs. presence of a radial gradate series in others), simply dichotomously branched CuP (vs. CuP deeply complicated branched in others) and the absence of distinct wing patterns (vs. presence of distinct wing patterns in others).	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
9C1AF9689C04FFF874EF6A31FD33FC95.taxon	type_taxon	Type species. Eupypsychops confinis Peng, Makarkin et Ren, 2015 Included Species. Eupypsychops confinis Peng, Makarkin et Ren, 2015; Eupypsychops ferox Peng, Makarkin et Ren, 2015; Eupypsychops oligophlebius sp. nov.	en	Liu, Xiaotian, Ren, Dong, Wang, Yongjie (2025): New Middle Jurassic Osmylopsychopidae (Neuroptera) from Northeastern China. Zootaxa 5723 (2): 292-300, DOI: 10.11646/zootaxa.5723.2.8, URL: https://doi.org/10.11646/zootaxa.5723.2.8
