identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
74A47139EBC15D288C68EC5AEB0CD09E.text	74A47139EBC15D288C68EC5AEB0CD09E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Unihamus concavus Luo & Che 2026	<div><p>Unihamus concavus Luo &amp; Che sp. nov.</p><p>Figs 2 E – H, 5, 7 E, 7 F, 8 C, 8 G, 8 K, 9 C</p><p>Unihamus sp. 2: Luo et al. 2025.</p><p>Type materials.</p><p>Holotype: China • ♂; Hainan, Mt Wuzhishan; 18–20. V. 2014; Shunhua Gui &amp; Xinran Li leg.; SWU -B-BL-083101 . Paratypes: China • 1 ♀; Hainan, Jianfengling National Forest Park; 24. IV. 2020; Yong Li &amp; Jing Zhu leg.; SWU -B-BL-083102 • 1 ♂; Hainan, Jianfengling National Forest Park; 11–22. IV. 2011; Wenxuan Bi leg.; 07002189 (Shanghai Entomological Museum, Chinese Academy of Sciences) .</p><p>Diagnosis.</p><p>Vertex slightly exposed. Pronotum subelliptical, widest posterior to midpoint. Posterior margin of metanotum mesally produced. Tegmina and wings well developed. Posterior branch of R reaching end of tegmina. Hind metatarsus longer than remaining tarsomeres combined. First abdominal tergite with visible gland. Hind margin of supra-anal plate deeply concave inwards, W-shaped. Posterior lateral angles of subgenital plate extended outward; hind margin slightly concave. L 1 weakly sclerotized. L 2 with two long ventral spines. L 3 not bifurcate. L 4 C elongate and distal part sharp and curved, and with spine-like process near basal right margin. Basal part of R 1 H with an elongate spine; distal margin with two spines. Distal margin of ventral surface of R 1 G extended into a strong spine and another slender spine.</p><p>This species can be distinguished from its congeners by the following characters: 1) supra-anal plate of male W-shaped (Fig. 5 F); 2) hind margin of subgenital plate convex and middle part concave (Fig. 5 G); 3) end of sclerite L 2 with two long spines (Figs 5 H, 5 I, 7 E); 4) R 1 H somewhat broad and basal part with an elongate spine (Figs 5 H, 7 F). In addition, U. concavus can be distinguished from U. elegans and U. longispinus by the curved distal part of L 4 C (Figs 5 H, 7 E).</p><p>Description.</p><p>Colouration. Body reddish brown. Ocelli white. Abdominal ventral surface reddish brown to blackish brown.</p><p>Male. Total length: 29.8–31.5 mm; body length: 23.71–24.0 mm; pronotum length × width: 5.64–6.0 × 7.89–10.1 mm; tegmina length × width: 24.6–25.5 × 8.1–9.2 mm. Head and thorax. Vertex slightly exposed; interocular space wider than interocellar space and shorter than interantennal space (Fig. 5 B). Antennae longer than body. Pronotum subelliptical, with anterior margin straight and hind margin slightly convex, surface smooth, median slightly elevated, widest posterior to midpoint (Fig. 5 A). Posterior margin of metanotum mesally produced; posterior-lateral angles with symmetrical projections (Fig. 8 C). Tegmina and wings well developed, surpassing tip of abdomen (Figs 2 E, 2 F, 5 D, 5 E). Tegmina ScP slightly thickened; posterior branch of R approaching end of tegmina (Fig. 5 D). Front femur of type A 2 (Fig. 5 C). Hind metatarsus longer than remaining tarsomeres combined; metatarsi with two rows of spinules ventrally and with one spinule on each side of pulvillus. Pulvilli present; claws symmetrical and unspecialized; arolium medium-sized. Abdomen. First abdominal tergite gland with downward-pointing setae (Fig. 8 C). Lateral margin of supra-anal plate contracted inward; middle of hind margin deeply concave inwards, W-shaped; margin of hind margin with a row of long setae (Fig. 5 F). Paraprocts (pp.) elongate, distally protruding downward. Cerci strong and long. Subgenital plate overall nearly square; posterior lateral angles extended outward; hind margin of interstylar slightly concave; styli slender (Fig. 5 G). Genitalia (Figs 5 H, 5 I, 7 E, 7 F, 8 G). L 1 small and weakly sclerotized (Figs 5 H, 7 E). L 2 irregular, with one long dorsal spine and another long ventral spine (Figs 5 H, 7 E). L 3 unciform and well sclerotized, not bifurcate, with apex sharp (Fig. 8 G). L 4 C elongate, with spine-like process near basal right margin and distal part sharp and curved (Figs 5 H, 7 E). R 1 H rather broad and irregular; basal part with an elongate spine; distal margin of broad sclerite with a large spine and a small spine (Figs 5 H, 7 F). R 1 G well sclerotized; distal margin of ventral surface extending a strong spine and another slender spine (Figs 5 I, 7 F).</p><p>Female (Figs 2 G, 2 H, 8 K, 9 C). Body slightly broader than male. Total length: 26.38 mm; body length: 23.71 mm; pronotum length × width: 6.01 × 8.44 mm; tegmina length × width: 19.4 × 6.85 mm. Head and thorax. As in male. Abdomen. Hind margin of tergum X with median invagination and with a membrane line inside (Fig. 9 C). Genitalia (Fig. 9 C). First valve with small, sparse punctures. First valvifer slightly sclerotized. Second valve strip-like. Third valve large and wrapped around second valve. Posterior lobes of valvifer II less sclerotized; outer margin connected to laterosternite IX. Laterosternite IX broad and irregular. Paratergites long, strip-like, and slender. Anterior arch broad, with sparse microtrichia. Spermathecal plate integrated into basivalvula. Spermathecal opening located at middle of basivalvula. Spermatheca branched, with duct rather short and end capsule rod-shaped (Fig. 8 K). Basivalvulae nearly fan-shaped and symmetrical; surface with sparse microtrichia. Laterosternal shelf symmetrical, with sparse punctures.</p><p>Etymology.</p><p>The Latin word concavus, meaning “ concave ”, refers to the deeply concave hind margin of the supra-anal plate.</p><p>Distribution.</p><p>China (Hainan).</p><p>Remarks.</p><p>Pairwise COI distances between this species and the other three congeners range from 6.54 % to 8.69 %, supporting the establishment of this species based on DNA barcoding.</p></div>	https://treatment.plazi.org/id/74A47139EBC15D288C68EC5AEB0CD09E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Luo, Xin-Xing;Qiu, Meng-Yu;Cai, Ting-Ting;Wang, Zong-Qing;Che, Yan-Li	Luo, Xin-Xing, Qiu, Meng-Yu, Cai, Ting-Ting, Wang, Zong-Qing, Che, Yan-Li (2026): Taxonomic update on the genus Unihamus Luo & Wang, 2025 (Blattidae, Blattinae), with three new species described. ZooKeys 1277: 175-197, DOI: 10.3897/zookeys.1277.163465
9AD71B2CA63E5182B8C95A91EDA52B09.text	9AD71B2CA63E5182B8C95A91EDA52B09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Unihamus elegans (Hanitsch 1927)	<div><p>Unihamus elegans (Hanitsch, 1927)</p><p>Figs 1, 3, 7 A, B, 8 A, E, I, 9 A</p><p>Periplaneta elegans Hanitsch, 1927: 25 (holotype: ♂, Vietnam; OUMNH; England); Bey-Bienko 1957: 896; Princis 1966: 445.</p><p>Unihamus elegans: Luo et al. 2025: 844.</p><p>Material examined.</p><p>Holotype • ♂ (OUM, TYPE ENT-ORTH 318) Vietnam: “ ♂ TYPE. Periplaneta elegans, Hanitsch. J. Siam Soc., Vol. vii, 1927, p. 25, fig. ”; “ Dran, Langbian, S. Annam, 3,000 '. Apl. May 1918. CB. Kloss. Coll. ”, “ Periplaneta elegans n. sp. ”; “ TYPE ORTH: 318 Periplaneta elegans Hanitsch / HOPE DEPT. OXFORD ”.</p><p>China • 2 ♂♂, 1 ♀; Yunnan, Butterfly Valley, Maandi village, Jinping County; 14–16. V. 2015; Jianyue Qiu leg; SWU -B-BL-080301 to 080303 • 2 ♀♀; Yunnan, Daweishan Mountain, Pingbian County; 16–17. V. 2015; Chenglong Ren &amp; Jianyue Qiu leg.; SWU -B-BL-080304 to 080305 .</p><p>Diagnosis.</p><p>Vertex exposed. Pronotum subelliptical, widest point posterior to midpoint. Posterior margin of metanotum mesally produced. Tegmina and wings well developed. Posterior branch of R not reaching end of tegmina. Hind metatarsus longer than remaining tarsomeres combined. First abdominal tergite with visible gland. Surface of supra-anal plate with small, sparse spines; median of hind margin incised; middle part of ventral surface with transverse projection, covered with small spines. Hind margin of subgenital plate slightly concave. L 1 weakly sclerotized. L 2 with long ventral spine pointing inward. L 3 not bifurcate. L 4 C lanceolate, with blade-like process. Left margin of R 1 H extending into two spines; ventral surface with a spine. Right ventral surface of R 1 G with a finger-like spine and another long spine.</p><p>This species can be easily distinguished from congeners by the following characters: 1) ventral surface of supra-anal plate with transverse, striate projection covered with small spines (Fig. 3 F); 2) acute blade-like process present near basal right margin of L 4 C (Figs 3 H, 7 A); 3) L 4 C lanceolate and not curved (differs from U. flavus and U. concavus) (Figs 4 H, 7 A); 4) R 1 H broad (differs from U. longispinus) (Figs 3 H, 7 B).</p><p>Redescription.</p><p>Colouration. Body black, except: ocelli white, antennae brown, tegmina reddish brown, tarsi blackish brown.</p><p>Male. Total length: 27.2–27.5 mm; body length: 18.5 mm; pronotum length × width: 4.8–5.2 × 5.7–6.1 mm; tegmina length × width: 22.0–22.3 × 6.6–6.7 mm. Head and thorax. Vertex exposed; interocular space almost equal to interocellar space, shorter than interantennal space (Fig. 3 B). Pronotum subelliptical, with anterior margin nearly straight and hind margin convex, surface smooth, widest point posterior to midpoint (Fig. 3 A). Posterior margin of metanotum mesally produced; posterior-lateral angles with symmetrical projections (Fig. 8 A). Tegmina and wings well developed, surpassing tip of abdomen (Figs 1 D, 1 E, 3 D, 3 E). Tegmina ScP slightly thickened; posterior branch of R not reaching end of tegmina (Fig. 3 D). Front femur of type A 2 (Fig. 3 C). Hind metatarsus longer than remaining tarsomeres combined; metatarsi with two rows of spinules ventrally and one spinule on each side of pulvillus. Pulvilli present; claws symmetrical and unspecialized; arolium medium-sized. Abdomen. First abdominal tergite gland with downward-pointing setae (Fig. 8 A). Lateral margin of supra-anal plate contracted inward near distal part and less sclerotized at margin; surface with small, sparse spines; middle of hind margin incised and divided into two subobtusely triangular lobules; middle part of ventral surface with transverse projection, covered with small spines (Fig. 3 F). Paraprocts long and strip-like. Cerci strong and symmetrical. Subgenital plate overall nearly square; hind margin of interstylar slightly concave (Fig. 3 G). Genitalia (Figs 3 H, 3 I, 7 A, 7 B, 8 E). L 1 weakly sclerotized, bearing pubescence (Fig. 7 A). L 2 irregular, with long, ventral, inward-pointing spine (Fig. 7 A). L 3 unciform and well sclerotized, not bifurcate (Figs 7 A, 8 E). L 4 C lanceolate, with distally acute, blade-like process near inner basal margin (Fig. 7 A). R 1 H broad and irregular; left margin extending into two spines; ventral surface with a spine (Fig. 7 B). R 1 G well sclerotized; right side of ventral surface with a finger-like spine and another long spine (Figs 3 I, 7 B).</p><p>Female (Figs 1 F, 1 G, 8 I, 9 A). Body broader than male. Total length: 23.1–28.1 mm; body length: 20.5–25.0 mm; pronotum length × width: 6.0–6.4 × 7.4–8.0 mm; tegmina length × width: 17.7–23.2 × 6.3–8.1 mm. Head and thorax. Same as in male. Abdomen. Hind margin of tergum X with median invagination and a membrane line inside (Fig. 9 A). Genitalia (Fig. 9 A). First valve sclerotized, with small, dense punctures, and basal part with microtrichia; left and right valve asymmetrical; right margin of left valve with a rhombic projection. First valvifer broad and slightly curved. Second valve strip-like. Third valve large and wrapping around second valve. Posterior lobes of valvifer II less sclerotized and outer margin connected to irregular laterosternite IX. Paratergites long, strip-like, and slender. Anterior arch nearly rectangular and outer and hind margin with small, dense spines. Spermathecal plate somewhat fan-shaped, connected to basivalvula by membrane. Spermathecal opening located at middle of basivalvula. Spermatheca branched into a leading duct and branching duct, with duct short and end capsule rod-shaped (Fig. 8 I). Basivalvulae (bsv.) nearly rectangular, asymmetrical, with microtrichia. Laterosternal shelf (ltst. sh.) symmetrical, with sparse punctures.</p><p>Distribution.</p><p>Vietnam (Lam Dong), China (Yunnan).</p><p>Remarks.</p><p>Unihamus elegans (Hanitsch, 1927) was first described from Da lat, Lam Dong Province, Vietnam, and was subsequently recorded from Baoshan, Yunnan Province, China (Bey-Bienko 1957). Our specimens were collected in Jinping and Pingbian counties, Yunnan Province, which borders Vietnam; these specimens correspond to Hanitsch’s (1927) original description and to photographs of the holotype.</p></div>	https://treatment.plazi.org/id/9AD71B2CA63E5182B8C95A91EDA52B09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Luo, Xin-Xing;Qiu, Meng-Yu;Cai, Ting-Ting;Wang, Zong-Qing;Che, Yan-Li	Luo, Xin-Xing, Qiu, Meng-Yu, Cai, Ting-Ting, Wang, Zong-Qing, Che, Yan-Li (2026): Taxonomic update on the genus Unihamus Luo & Wang, 2025 (Blattidae, Blattinae), with three new species described. ZooKeys 1277: 175-197, DOI: 10.3897/zookeys.1277.163465
63334C338BBA550399BBF39D1740E9C0.text	63334C338BBA550399BBF39D1740E9C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Unihamus flavus Luo & Che 2026	<div><p>Unihamus flavus Luo &amp; Che sp. nov.</p><p>Figs 2 A – D, 4, 7 C, 7 D, 8 B, 8 F, 8 J, 9 B</p><p>Unihamus sp. 1: Luo et al. 2025.</p><p>Type materials.</p><p>Holotype: China • ♂; Yunnan, Meizihu Reservoir, Pu’er City; 30. IV. 2014; Xinran Li &amp; Jianyue Qiu leg.; SWU -B-BL-082001 . Paratypes: China • 1 ♀; Yunnan, Meizihu Reservoir, Pu’er City; 30. IV. 2014; Xinran Li &amp; Jianyue Qiu leg.; SWU -B-BL-082006 • 2 ♂♂; Yunnan, Dadugang Tea Garden and Secondary Plants, Xishuangbanna Prefecture; 1296–1299 m alt.; 28. IV. 2014; Xinran Li leg.; SWU -B-BL-082002 to 082003 • 1 ♀; Yunnan, Meizihu Reservoir Highway, Pu’er City; 20–21. V. 2016; Lu Qiu &amp; Zhiwei Qiu leg.; SWU -B-BL-082004 • 1 ♀; Yunnan, Meizihu Reservoir, Pu’er City; 1384–1418 m alt.; 2014; Hongguang Liu &amp; Xinran Li leg.; SWU -B-BL-082005 • 1 ♀; Guangxi, Yinshan Park, Jinxiu County; 16–17. VII. 2015; Lu Qiu &amp; Qikun Bai leg.; SWU -B-BL-082007 .</p><p>Diagnosis.</p><p>Vertex slightly exposed. Pronotum subelliptical, widest posterior to midpoint. Posterior margin of metanotum mesally produced. Tegmina and wings well developed. Posterior branch of R not reaching end of tegmina. Hind metatarsus longer than or nearly equal to remaining tarsomeres combined. First abdominal tergite with visible gland. Hind margin of supra-anal plate mesally incised. Hind margin of supra-anal plate incised and divided into two subobtusely triangular lobules. Hind margin of subgenital plate slightly concave. L 1 weakly sclerotized. L 2 with a long ventral spine. L 3 not bifurcate. L 4 C lanceolate, with distal part sharp and curved, with strip-like process near basal right margin. Left margin of R 1 H extended into a small spine; middle part of distal margin with a small projection. Distal margin of ventral surface of R 1 G extended into a long spine and another short spine.</p><p>Although closely resembling U. elegans, this species can be distinguished by the following combination of characters: 1) hind margin of supra-anal plate without small, sparse spines (Fig. 4 F); 2) ventral surface of supra-anal plate without projections (Fig. 4 F); 3) L 4 C with strip-like process near basal right margin and distal part curved (Figs 4 H, 7 C). In addition, U. flavus can be distinguished from U. concavus and U. longispinus by the following characteristics: 1) hind margin of supra-anal plate slightly concave (vs deeply concave) (Fig. 4 F); 2) L 2 with ventral spine shorter than U. concavus and U. longispinus (Figs 4 H, 4 I, 7 C); 3) R 1 H broader than U. concavus and U. longispinus (Figs 4 H, 7 D).</p><p>Description.</p><p>Colouration. Body blackish brown to brown. Ocelli white, antennae yellowish brown. Tegmina reddish brown. Legs brown to reddish brown. Abdominal ventral surface unevenly coloured: margin black, middle reddish brown to yellowish brown.</p><p>Male. Body medium-sized. Total length: 26.1–29.5 mm; body length: 20.8–21.4 mm; pronotum length × width: 4.7–5.0 × 6.4–7.7 mm; tegmina length × width: 23.5–25.3 × 6.4–7.7 mm. Head and thorax. Vertex slightly exposed; interocular space wider than interocellar space and shorter than interantennal space (Fig. 4 B). Pronotum subelliptical, with anterior margin slightly concave and hind margin convex, surface smooth, widest posterior to midpoint (Fig. 4 A). Posterior margin of metanotum mesally produced; posterior-lateral angles with symmetrical projections (Fig. 8 B). Tegmina and wings well developed, surpassing tip of abdomen (Figs 2 A, 2 B, 4 D, 4 E). Tegmina ScP slightly thickened; posterior branch of R not reaching end of tegmina (Fig. 4 D). Front femur of type A 2 (Fig. 4 C). Hind metatarsus longer than or nearly equal to remaining tarsomeres combined; metatarsi with two rows of spinules ventrally; each side of pulvillus with one spinule. Pulvilli present; claws symmetrical and unspecialized; arolium medium-sized. Abdomen. First abdominal tergite gland with downward-pointing setae (Fig. 8 B). Lateral margin of supra-anal plate contracted inward near distal part; median of hind margin incised and divided into two subobtusely triangular lobules (Fig. 4 F). Paraprocts (pp.) elongate. Cerci strong and symmetrical. Subgenital plate overall nearly square; posterior lateral angles extended outward; hind margin of interstylar slightly concave (Fig. 4 G). Genitalia (Figs 4 H, 4 I, 7 C, 7 D, 8 F). L 1 small and weakly sclerotized (Figs 4 H, 7 C). L 2 irregular, with a ventral spine (Figs 4 H, 7 C). L 3 unciform and well sclerotized, not bifurcate, with apex sharp (Fig. 8 F). L 4 C lanceolate and broad, with strip-like process near basal right margin; distal part sharp and curved (Figs 4 H, 7 C). R 1 H broad and irregular; left margin extending into a small spine and middle part of distal margin with a small projection (Figs 4 H, 7 D). R 1 G well sclerotized; distal margin of ventral surface extending into a long spine and another short spine (Figs 4 I, 7 D).</p><p>Female (Figs 2 C, 2 D, 8 J, 9 B). Body slightly broader than male. Total length: 17.9–27.2 mm; body length: 17.9–27.2 mm; pronotum length × width: 4.7–6.2 × 6.3–8.0 mm; tegmina length × width: 15.3–21.2 × 5.3–8.4 mm. Head and thorax. As in male. Abdomen. Hind margin of tergum X with median invagination and a membrane line inside (Fig. 9 B). Genitalia (Fig. 9 B). First valve with small punctures. First valvifer slender and slightly curved. Second valve strip-like. Third valve large and wrapped around second valve. Posterior lobes of valvifer II less sclerotized; outer margin connected to laterosternite IX. Laterosternite IX broad and membranous. Paratergites long, strip-like, and slender. Anterior arch membranous, with dense microtrichia. Spermathecal plate hyaline, connected to basivalvula by membrane. Spermathecal opening at middle of basivalvula. Spermatheca branched, with duct long and curved and end capsule reniform (Fig. 8 J). Basivalvulae nearly fan-shaped and symmetrical, membranous; surface with microtrichia. Laterosternal shelf symmetrical and membranous; surface with sparse punctures.</p><p>Etymology.</p><p>The species epithet is derived from the Latin word flavus, “ yellow ”, referring to the yellowish-brown ventral surface of the species’ abdomen.</p><p>Distribution.</p><p>China (Guangxi, Yunnan).</p><p>Remarks.</p><p>In their phylogenetic analysis, Luo et al. (2025) placed this species as the sister lineage of U. elegans . DNA barcoding supported the separation of these two species, revealing a genetic divergence of 5.92–6.26 %.</p></div>	https://treatment.plazi.org/id/63334C338BBA550399BBF39D1740E9C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Luo, Xin-Xing;Qiu, Meng-Yu;Cai, Ting-Ting;Wang, Zong-Qing;Che, Yan-Li	Luo, Xin-Xing, Qiu, Meng-Yu, Cai, Ting-Ting, Wang, Zong-Qing, Che, Yan-Li (2026): Taxonomic update on the genus Unihamus Luo & Wang, 2025 (Blattidae, Blattinae), with three new species described. ZooKeys 1277: 175-197, DOI: 10.3897/zookeys.1277.163465
128C5E34F4135B40966FCDAFDDDF5545.text	128C5E34F4135B40966FCDAFDDDF5545.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Unihamus longispinus Luo & Che 2026	<div><p>Unihamus longispinus Luo &amp; Che sp. nov.</p><p>Figs 2 I – L, 6, 7 G, 7 H, 8 D, 8 H, 8 L, 9 D</p><p>Type materials.</p><p>Holotype: China • ♂; Hainan, Ganshiling, Baoting County; 9–10. IV. 2015; Lu Qiu &amp; Qikun Bai leg.; SWU -B-BL-083601 . Paratype: China • 1 ♀; Hainan, Diaoluoshan National Forest Park; 2. V. 2013; Shunhua Gui &amp; Yan Shi leg.; SWU -B-BL-083602 .</p><p>Diagnosis.</p><p>Vertex exposed. Pronotum subelliptical, widest posterior to midpoint. Posterior margin of metanotum mesally produced. Tegmina and wings well developed. Posterior branch of R reaching end of tegmina. First abdominal tergite with visible gland. Hind margin of supra-anal plate deeply concave inwards, with a row of long setae. Hind margin of subgenital plate slightly concave. L 1 weakly sclerotized. L 2 with one long, strong ventral spine. L 3 not bifurcate. L 4 C lanceolate, with spine-like process near basal right margin. Basal part of R 1 H with a sharp spine. Distal margin of ventral surface of R 1 G extends into a strong spine and another slender spine.</p><p>This species can be distinguished from U. concavus by the shape of the supra-anal and subgenital plates (Fig. 6 F, G). In U. concavus, the supra-anal plate is W-shaped and the subgenital plate is convex towards the back with the middle concave. It differs from U. elegans and U. flavus in having L 2 longer than L 3 and in having R 1 H spine-like (Figs 6 H, 6 I, 7 G, 7 H).</p><p>Description.</p><p>Colouration. Body brown to blackish brown. Ocelli white. Tegmina of male reddish brown and of female dark blackish brown to black.</p><p>Male. Total length: 24.0 mm; body length: 19.3 mm; pronotum length × width: 4.98 × 6.99 mm; tegmina length × width: 21.3 × 7.26 mm. Head and thorax. Vertex exposed; interocular space wider than interocellar space, shorter than interantennal space (Fig. 6 B). Antennae longer than body. Pronotum subelliptical, anterior margin straight and hind margin slightly convex, smooth, and widest posterior to midpoint (Fig. 6 A). Posterior margin of metanotum mesally produced; posterior-lateral angles with symmetrical projections (Fig. 8 D). Tegmina and wings well developed, surpassing tip of abdomen (Figs 2 I, 2 J, 6 D, 6 E). Tegmina ScP slightly thickened; posterior branch of R approaching end of tegmina (Fig. 6 D). Front femur type of A 2 (Fig. 6 C). Hind legs unknown. Pulvilli present; claws symmetrical and unspecialized; arolium medium-sized. Abdomen. First abdominal tergite gland with downward-pointing setae (Fig. 8 D). Lateral margin of supra-anal plate contracted inward, with median of hind margin deeply concave inwards, and with a row of long setae (Fig. 6 F). Paraprocts (pp.) elongate. Cerci strong and long. Subgenital plate overall nearly square; hind margin of interstylar slightly concave; styli slender (Fig. 6 G). Genitalia (Figs 6 H, 6 I, 7 G, 7 H, 8 H). L 1 weakly sclerotized (Figs 6 H, 7 G). L 2 irregular; end with a long, strong spine (Figs 6 H, 7 G). L 3 unciform, well sclerotized, and not bifurcate (Fig. 8 H). L 4 C lanceolate, with spine-like process near basal right margin; distal part sharp (Figs 6 H, 7 G). R 1 H irregular; basal part with a sharp spine (Figs 6 H, 7 H). R 1 G well sclerotized; distal margin of ventral surface extended into a strong spine and another slender spine (Figs 6 I, 7 H).</p><p>Female (Figs 2 K, 2 L, 8 L, 9 D). Body slightly broader than male. Total length: 23.55 mm; body length: 21.29 mm; pronotum length × width: 5.9 × 8.3 mm; tegmina length × width: 18.8 × 8.0 mm. Head and thorax. As in male. Abdomen. Hind margin of tergum X with median invagination, and with a membrane line inside (Fig. 9 D). Genitalia (Fig. 9 D). First valve with small punctures. First valvifer rather broad. Second valve sclerotized. Third valve large and wrap second valve. Posterior lobes of valvifer II connected to laterosternite IX. Laterosternite IX broad and irregular. Paratergites strip-like and slender. Anterior arch broad; surface with sparse microtrichia. Spermathecal plate fan-shaped and integrated into basivalvula. Spermathecal opening at middle of basivalvula. Spermatheca branched, with duct short and end capsule rod-shaped (Fig. 8 L). Basivalvulae symmetrical; basal part of lateral margin contracted inward; surface with sparse microtrichia. Laterosternal shelf symmetrical and with sparse punctures.</p><p>Etymology.</p><p>The species epithet is derived from the Latin word longispinus, meaning “ long spined ”, in reference to L 2 ventral spine in males, which is longer than in congeneric species.</p><p>Distribution.</p><p>China (Hainan).</p><p>Remarks.</p><p>Pairwise COI distances between this species and the other three congeners range from 5.73 % to 7.65 %. The genetic distance between U. longispinus and U. flavus is 5.73 %, which is less than the genetic distance between most species. However, they strongly differ in their morphology.</p><p>Molecular analysis based on COI</p><p>Two COI sequences were obtained from U. elegans, U. longispinus Luo &amp; Che, sp. nov., U. concavus Luo &amp; Che, sp. nov., and seven COI sequences were generated for U. flavus sp. nov. All new sequences were deposited in GenBank with accession numbers PZ 151070 to PZ 151078. In our maximum-likelihood (ML) analysis, the Unihamus species each formed a well-supported monophyletic group (Fig. 10), including female and male specimens. Intraspecific COI genetic divergence (K 2 P) was 0.00 % in U. elegans, 0.00–0.30 % in U. flavus sp. nov., 0.46 % in U. longispinus sp. nov., and 1.85 % in U. concavus sp. nov. Interspecific K 2 P distances between the four species ranged from 5.73 % ( U. flavus sp. nov. and U. longispinus sp. nov.) to 8.69 % ( U. elegans and U. concavus sp. nov.) (Suppl. material 2).</p></div>	https://treatment.plazi.org/id/128C5E34F4135B40966FCDAFDDDF5545	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Luo, Xin-Xing;Qiu, Meng-Yu;Cai, Ting-Ting;Wang, Zong-Qing;Che, Yan-Li	Luo, Xin-Xing, Qiu, Meng-Yu, Cai, Ting-Ting, Wang, Zong-Qing, Che, Yan-Li (2026): Taxonomic update on the genus Unihamus Luo & Wang, 2025 (Blattidae, Blattinae), with three new species described. ZooKeys 1277: 175-197, DOI: 10.3897/zookeys.1277.163465
9254BDCF31E6537AA6E87EF4AD16F9A1.text	9254BDCF31E6537AA6E87EF4AD16F9A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Unihamus Luo & Wang 2025	<div><p>Genus Unihamus Luo &amp; Wang, 2025</p><p>Unihamus Luo &amp; Wang in Luo et al. 2025. Type species: Periplaneta elegans Hanitsch, 1927, by original designation.</p><p>Generic diagnosis.</p><p>Body medium-sized. Sexual dimorphism weak: female broader than male; tegmina and wings of female shorter than male, but all surpassing tip of abdomen. Male. Antennae longer than body. Pronotum subelliptical, widest point posterior to midpoint. Posterior margin of metanotum mesally produced. Tegmina ScP slightly thickened. Front femur of type A 2. Pulvilli present; claws symmetrical and unspecialized; arolium medium-sized. Hind metatarsus longer than or equal to remaining tarsomeres combined; metatarsi with two rows of spinules ventrally; each side of pulvillus with one spinule. First abdominal tergite with visible, setose gland. Middle part of hind margin of supra-anal plate and subgenital plate concave inwards. L 3 unciform and not bifurcated. L 4 C with process near basal right margin. Distal part of R 1 G with two spines. R 2 large and complex. Anterior margin of R 3 triangular. Female. Outer margin of posterior lobes of valvifer II merged with laterosternite IX.</p><p>The simple antenna of Unihamus differ from that of Thyrsocera Burmeister, 1838, which has the antenna plumose or with plumose portions in males and moniliform in females.</p><p>This macropterous genus can be readily distinguished from the apterous and micropterous blattid genera (i. e. Afrostylopyga Anisyutkin, 2014, Apterisca Princis, 1963, Brinckella Princis, 1963, Henicotyle Rehn &amp; Hebard, 1927, Macrostylopyga Anisyutkin et al., 2013, Miostylopyga Princis, 1966, and Neostylopyga Shelford, 1911). The weak sexual dimorphism (macroptery of males and females) in Unihamus differs from distinct sexual dimorphism in other genera:</p><p>Macropterous male and apterous female; i. e. Deropeltis Burmeister, 1838, Archiblatta Snellen van Vollenhoven, 1862, and Catara Walker, 1868);</p><p>Macropterous male and micropterous female; i. e. Blatta Linnaeus, 1758, Planiblatta Luo &amp; Wang, 2023, Pseudoderopeltis Krauss, 1890);</p><p>Macropterous male and brachypterous female; i. e. Arcicaulis Luo &amp; Wang, 2025, Bundoksia Lucañas, 2021, Cartoblatta Shelford, 1910, Tenumembrana Luo &amp; Wang, 2025, some Periplaneta Burmeister, 1838, Protagonista Shelford, 1908, Scabinopsis Bey-Bienko, 1969, and Vittiblatta Luo &amp; Wang, 2023).</p><p>The hind metatarsus of Unihamus is longer than or equal to the remaining tarsal segments combined, which differs from Atemeleta Brunner von Wattenwyl, 1893, Eumethana Princis, 1951, and Scabinopsis .</p><p>The first abdominal tergite has a visible, setose gland in Unihamus . Setose glands are absent in Blatta, Cartoblatta, Homalosilpha Stål, 1874, Periplaneta s. s., and Valaraukar Lucañas, 2025.</p><p>Species of Unihamus in which L 3 is not bifurcated can be distinguished from other genera with a bifurcated L 3. These genera are Archiblatta, Arcicaulis, Blatta, Bundoksia, Catara, Crescispina Luo &amp; Wang, 2025, Deropeltis, Dorylaea Stål, 1877, Eroblatta Shelford, 1910, Hobbitoblatta Lucañas, 2023, Homalosilpha, Nazgultaure Lucañas, 2023, Tenumembrana, Valaraukar, Validiblatta Luo &amp; Wang, 2025, and Vittiblatta.</p><p>Key to males of known species of Unihamus</p></div>	https://treatment.plazi.org/id/9254BDCF31E6537AA6E87EF4AD16F9A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Luo, Xin-Xing;Qiu, Meng-Yu;Cai, Ting-Ting;Wang, Zong-Qing;Che, Yan-Li	Luo, Xin-Xing, Qiu, Meng-Yu, Cai, Ting-Ting, Wang, Zong-Qing, Che, Yan-Li (2026): Taxonomic update on the genus Unihamus Luo & Wang, 2025 (Blattidae, Blattinae), with three new species described. ZooKeys 1277: 175-197, DOI: 10.3897/zookeys.1277.163465
