identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9F37571B2220FFC7679AFB159416FD66.text	9F37571B2220FFC7679AFB159416FD66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caphodus Oman 1938	<div><p>Caphodus Oman, 1938</p><p>Figs. 1–6</p><p>Caphodus Oman, 1938: 375 .</p><p>Type species by original designation: Caphodus maculatus Oman, 1938</p><p>Length 4.2 to 5.8mm. Head as wide as or slightly narrower than pronotum. Ground color whitish, with brown and orange markings; crown with pair of large orange maculae centrally and with smaller dark brown markings anteriorly; pronotum anteriorly with 2 or 3 large orange maculae, medially with two pairs of orange maculae; forewing with two prominent brown spots- one between clavus and A1 and one basad of central anteapical cell, with numerous false brown veins, with brown coloration at wing apex, veins tan toward base and brown toward apex, reflexed costal veins brown; face and venter with extensive dark brown coloration; legs whitish with dark brown markings, setal bases dark brown. Crown produced, longer medially than next to eyes; texture shagreen on anterior 1/3, smooth on posterior 2/3. Ocelli large; close to eyes. Face with frontoclypeus long, narrow; clypellus constricted subbasally and expanded toward apex; lorae large; antennae arising near middle of eye; mesial margin of eye notched. Profemur row AV with numerous very short, stout setae along basal 2/3 of femur; intercalary row with ~15 long fine setae; AM 1 present, situated ventrally. Protibia macrosetae 1+4. Hind tarsomere I long, slender; apex with four apical platellate setae, innermost seta platellate.</p><p>Male. Pygofer longer than tall in lateral view; with several rows of long macrosetae near apex; dorsally barely incised around segment X; with two pairs of process arising posteriorly or posteroventrally. Valve triangular; width more than 2x median length. Subgenital plates long, triangular; with complete row of macrosetae near lateral margin; with numerous fine setae laterally. Connective with anterior arms closely appressed; with stem shorter than anterior arms; articulated with aedeagus. Styles with base broadly bilobed; preapical lobe present or absent; apophysis short to long. Aedeagus with one or two pairs of long processes arising from base; shaft symmetrical and tubular or asymmetrical and flattened. Segment X sclerotized dorsally and laterally; with pair of ventral processes.</p><p>Female. Sternite VII variable in shape. Ovipositor straight ventrally or slightly concave; extending beyond pygofer apex. First valvifer with or without unique posterior extension. First valvula with elongated baculiform sculpturing dorsally, baculiform elements grading together ventrally, nearly forming strigate pattern; sculpturing reaching dorsal margin, extending 2/3 to 3/4 length of valvula. Second valvula with obtusely triangular, serrated teeth dorsally on 1/2 to 2/3 length of valvula. Gonoplac with four to five stout setae apex; with several stout setae ventrally.</p><p>Notes. Given new observations of the male genitalia of C. obliquus and ovipositor of all three species, Caphodus is redescribed here.</p><p>Oman et al. (1990) listed Caphodus in Deltocephalinae but did not place the genus to a tribe, and Zahniser &amp; Dietrich (2013) placed it in Scaphoideini based on morphological similarity to some members of that group. The phylogenomic analyses of Cao et al. (2022) showed that the Caphodus is closely related to Scaphoidula Osborn, 1923 within a clade including most genera placed in Pendarini . Based on that result and a re-assessment of morphological characters defining tribes of Deltocephalinae, Zahniser et al. (in press) moved Caphodus to Pendarini .</p><p>Key to species</p><p>1. Males.............................................................................................. 2</p><p>1’. Females..............................................................................................4</p><p>2(1). Aedeagus shaft dorsoventrally flattened, wide, asymmetrical; dorsal-most pair of pygofer processes arising from dorsoapical corner of pygofer, elongate and with 4 to 5 dorsal teeth subapically................................... C. maculatus</p><p>2’. Aedeagus shaft not flattened, wide, or asymmetrical; dorsal-most pair of pygofer processes arising from posterior margin of pygofer, at mid-height or lower, not elongate and without dorsal teeth.............................................3</p><p>3(2’). Aedeagus with two pairs of processes arising from near base; dorsal-most pair of pygofer processes longer than ventral pair............................................................................................ C. obliquus</p><p>3’. Aedeagus with one pair of processes arising near base; dorsal-most pair of pygofer processes shorter than ventral pair.............................................................................................. C. panamensis</p><p>4(1’). Sternite VII strongly produced medially, apex lingulate.............................................. C. obliquus</p><p>4’. Sternite VII not produced medially, shallowly or deeply excavated; apex medially with pair of shortly produced teeth......5</p><p>5(4’). Sternite VII shallowly excavated; first valvifer without accessory process............................. C. panamensis</p><p>5’. Sternite VII deeply excavated; first valvifer with distinct club-like accessory process...................... C. maculatus</p></div>	https://treatment.plazi.org/id/9F37571B2220FFC7679AFB159416FD66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2223FFC7679AFD3492A2FAF0.text	9F37571B2223FFC7679AFD3492A2FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caphodus maculatus Oman 1938	<div><p>Caphodus maculatus Oman, 1938</p><p>Figs 1–3</p><p>Caphodus maculatus Oman, 1938: 376</p><p>Material examined. Holotype, ♂, undissected: [ARGENTINA] Loreto Missiones [sic], ARG XII-18-1931, AA Ogoblin [sic] / [red label] Type No., 51670, U.S.N.M. / Caphodus maculatus Oman, Det. P.W. Oman / USNMENT01513044 . Paratype, ♂: Loreto Missiones [sic], ARG XII-8-1931, AA Ogoblin [sic] / [red label] Paratype No., 51670, U.S.N.M. / USNMENT01513101 . Allotype, ♀, same locality data as previous, [red label] Allotype No., 51670, U.S.N.M. / USNMENT01513531 . Paratype, ♂: Loreto Missiones [sic], ARG XI-25-1931, AA Ogoblin [sic] / [red label] Paratype No., 51670, U.S.N.M. / USNMENT01513511.</p><p>Female. Sternite VII long; laterally with lobate extensions; deeply excavated medially to about 2/5 of its length, with a bipronged medial lobe. First valvifer with distinct clublike posterior extension; other characters as in genus description.</p><p>Notes. The male genitalia are unusual in possessing an aedeagal shaft that is dorsoventrally flattened and asymmetrical. The genitalia were illustrated by Linnavuori (1959) but are re-illustrated here to provide clarity. The ovipositor structures are illustrated and described here for the first time.</p></div>	https://treatment.plazi.org/id/9F37571B2223FFC7679AFD3492A2FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2223FFC1679AFA5D91F9FE3A.text	9F37571B2223FFC1679AFA5D91F9FE3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caphodus obliquus (Osborn 1923) Kramer 1963	<div><p>Caphodus obliquus (Osborn, 1923)</p><p>Figs. 4–5</p><p>Scaphoideus obliquus Osborn, 1923: 40</p><p>Caphodus obliquus (Osborn, 1923) in Oman, 1938: 376</p><p>Material examined. Holotype, ♂: [BRAZIL] Chapada, Brazil, Acc. No. 2966 / Nov. / TYPE / Scaphoideus obliquus n.sp. Osb. / Caphodus [CMNH] . 1♀: [BRAZIL] Chapada / Oct. / Collection CF Baker / Caphodus obliquus (Osb.) Kramer 1963 / USNMENT01513521.</p><p>An additional specimen at USNM has only two legs remaining on the point mount and has the same locality label data as the USNM female listed above and with determination label: Scaphoideus obliquus Osb., Compared with type by Oman [USNM].</p><p>Male.Pygofer dorsally sclerotized to apex,not incised posteriorly;with long claw like process at mid height of posterior margin; with sharp toothlike process at posteroventral corner; with long macrosetae arranged in 2–3 row posteriorly; with other shorter, fine setae on ventral half. Valve bluntly triangular; median length less than 2x width. Subgenital plates long, triangular, apex rounded and with small toothlike process; with row of macrosetae laterally along entire length; with numerous long fine setae laterally. Connective with anterior arms closely appressed; stem broad, shorter than anterior arms; articulated with aedeagus.Style relatively short; broadly bilobed at base; with blunt preapical lobe; apophysis relatively thick, short, clawlike, apex blunt. Aedeagus with two pairs of processes arising near base of shaft; shorter pair of processes rising posterodorsad along shaft, slightly shorter than shaft; longer pair extending ventrad of shaft, about 1.75x length of shaft; shaft narrow, with apical flange. Segment X dorsally, laterally, and ventrally sclerotized; ventrally with pair of long processes.</p><p>Female. Sternite VII very long; laterally sinuate and tapering to lingulate apical lobe. First valvifer with large, irregular posterior extension, broad and lobate at base, dorsally constricted to digitate apex; other characters as in genus description.</p><p>Notes. The male genitalia, female sternite VII, and ovipositor structures are illustrated and described here for the first time. “Chapada” on the labels refers to Chapada dos Guimarães in Mato Grosso State (Camisão et al., 2024).</p></div>	https://treatment.plazi.org/id/9F37571B2223FFC1679AFA5D91F9FE3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2224FFC0679AFA9793A3F80A.text	9F37571B2224FFC0679AFA9793A3F80A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caphodus panamensis Linnavuori & DeLong 1977	<div><p>Caphodus panamensis Linnavuori &amp; DeLong, 1977</p><p>Fig. 6</p><p>Caphodus panamensis Linnavuori &amp; DeLong, 1977: 564</p><p>Material examined. 1♂: PANAMA, Porterillos, Chriquí, 12 Dec 1952 / FS Blanton Collector / Caphodus n.sp. / Caphodus panamensis Lv. &amp; DeL., Det. JN Zahniser 2023 / USNMENT01513125. 1♀: PANAMA, Bat Cave, Top R of Panama, 8 X 1952 / FS Blanton Collector / USNMENT01513450. 1♀: PANAMA, La Yeguada, 21 x 52 / Coll. FS Blanton / Caphodus panamensis Lv. &amp; DeL., Det. JN Zahniser 2023 / USNMENT01513118. 1♂: PANAMA: Pan. Prov., Las Cumbres, 20 January 1976, H. Wolda / USNMENT01513039. 1♂: PANAMA, La Venta, 8 xi 52 / Coll. FS Blanton / USNMENT01513104. 1♂: [TRINIDAD, new country record] Cocoa estate about 6 mi. from LaBrea, Tr., Oct. 18, 1918. A-787, Harold Morrison / Caphodus panamensis Lv. &amp; DeL., Det. JN Zahniser 2025 / USMENT01513490.</p><p>Female. Sternite VII generally quadrate in shape; median length about 2/3 basal width; posterior margin slightly excavated, medially with pair of short teeth. First valvifer without posterior process. Second valvulae and gonoplac similar to C. maculatus .</p><p>Notes. The male genitalia were adequately described and illustrated in the original description. The first valvula and valvifer are illustrated here for the first time.</p></div>	https://treatment.plazi.org/id/9F37571B2224FFC0679AFA9793A3F80A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2229FFCD679AF9BA93BAF838.text	9F37571B2229FFCD679AF9BA93BAF838.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atanus Oman 1938	<div><p>Atanus Oman, 1938</p><p>Figs. 7–11</p><p>Atanus Oman, 1938: 381</p><p>Type species: Eutettix dentatus Osborn, 1923 by original designation by Oman (1938)</p><p>See Dmitriev et al. (2022 onward) for synonymy.</p><p>Note. In addition to the holotype specimen of Eutettix mimicus Osborn found in the Kramer loan, other types of Atanus spp. at USNM and CMNH were examined and are illustrated here.</p><p>The phylogenomic analyses of Cao et al. (2022), resolved Atanus within a large clade containing most of the New World genera previously placed in Athysanini . Zahniser et al. (in press) included these genera, including Atanus, in the resurrected tribe Platymetopiini .</p></div>	https://treatment.plazi.org/id/9F37571B2229FFCD679AF9BA93BAF838	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2228FFCC679AFF4C95D7FD2F.text	9F37571B2228FFCC679AFF4C95D7FD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atanus coronatus (Berg 1879)	<div><p>Atanus coronatus (Berg, 1879)</p><p>Fig. 7</p><p>Athysanus coronatus Berg, 1879: 261</p><p>Thamnotettix inconspicuus Jensen-Haarup, 1922: 16</p><p>Eutettix dentatus Osborn, 1923: 57</p><p>Atanus dentatus (Osborn, 1923) in Oman, 1938: 381</p><p>Atanus coronatus (Berg, 1879) in Linnavuori, 1959: 297</p><p>Eutettix dentatus Osborn, 1923 (subjective synonym of Athysanus coronatus Berg, 1879) in Linnavuori, 1959: 297</p><p>Thamnotettix inconspicuus Jensen-Haarup, 1922 (subjective synonym of Athysanus coronatus Berg, 1879) in Linnavuori, 1959: 297</p><p>Material examined. Holotype, ♀: [BOLIVIA] Prov. Del Sara, Bol. Steinbach., C M. Acc. 5064 / Feb. 1913 / TYPE / CMNH HOLOTYPE #802, Eutettix dentatus Osborn / Eutettix dentatus Osb. nsp. / Atanus cornonatus (Berg), K. Zanol det 1986.</p><p>Notes. Osborn (1923) described Eutettix dentatus Osb., and Oman (1938) listed this as the type of species Atanus . Linnavuori (1959) synonymized E. dentatus under A. coronatus (Berg) without comment. Osborn’s type female specimen of E. dentatus was borrowed from CMNH for this study and is illustrated here.</p></div>	https://treatment.plazi.org/id/9F37571B2228FFCC679AFF4C95D7FD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2228FFC8679AF956927FFE1E.text	9F37571B2228FFC8679AF956927FFE1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atanus dorsalis (Gillette 1898)	<div><p>Atanus dorsalis (Gillette, 1898)</p><p>Fig. 8</p><p>Alebra dorsalis Gillette, 1898: 713</p><p>Atanus dorsalis (Gillette, 1898) in Young, 1952: 31</p><p>Material examined. Holotype, ♂: [BRAZIL] Chapada / April / Type No. 29505, U.S.N.M. / Alebra dorsalis Gill. / USNMENT01513124</p><p>Diagnosis. A. dorsalis can be distinguished from other species in the genus by the combination of its color pattern (ground color yellow; pronotum, mesonotum, and medial half of forewing at rest dark brown), and characters of the male genitalia.</p><p>Male. Pygofer with pair of bladelike processes arising from dorsal margin and extending to near middle of pygofer height. Style apophysis long and lacking a distinct preapical lateral lobe and apex foot-like with sharp preapical medial point. Aedeagus with two pairs of apical processes; apical pair slender and directed caudolaterally and the preapical pair longer, wider, falcate, and directed anterodorsally.</p><p>Notes. In his revision of the Neotropical Deltocephalinae, Linnavuori (1959) did not observe the type specimen, and listed the species as incertae sedis, noting Young’s (1952) placement “near Atanus ”. Except for a drawing of the forewing by Gillette (1898), this species has not previously been illustrated. The male genitalia are described here for the first time, and the placement of the species in Atanus is confirmed. “Chapada” on the label refers to Chapada dos Guimarães (see Camisão et al., 2024) in Mato Grosso State.</p></div>	https://treatment.plazi.org/id/9F37571B2228FFC8679AF956927FFE1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B222CFFC8679AFDF890F6FB5E.text	9F37571B222CFFC8679AFDF890F6FB5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atanus lobatus (Osborn 1923)	<div><p>Atanus lobatus (Osborn, 1923)</p><p>Figs. 9–10</p><p>Thamnotettix lobatus Osborn, 1923: 71 .</p><p>Eutettix mimicus Osborn, 1924: 414, new synonym</p><p>Atanus lobatus (Osborn, 1923) in Oman, 1938: 381</p><p>Material examined. Holotype, ♂: Prov. del Sara, Bolivia, 450m., J. Steinbach / C.M., Acc. 4549 / Nov 1909 / TYPE / CMNH HOLOTYPE #824, Thamnotettix lobatus Osborn 1923 / Thamnotettix lobatus Osb. n.sp. H.O. Holotype, ♂: Prov. del Sara, Bol. Steinbach, C.M. Acc. 5064. / Feb, 1913 / Eutettix mimicus nsp TYPE / Tubulanus .</p><p>Notes. Among the CMNH specimens in the loan found at USNM was the type specimen of Eutettix mimicus Osborn, 1924 from Bolivia. Linnavuori (1959) did not observe the type, listed the species as incertae sedis within his broad circumscription of Euscelini, repeated the description of Osborn (1924), and made no further comment on the species. After observing the type specimen in this study and determining it belongs in Atanus, a search of the literature revealed that the genitalia structures closely resembled those of Atanus lobatus (Osborn) which were partially illustrated by Linnavuori &amp; DeLong (1977, Figs. 44–47). The type of Thamnotettix lobatus Osborn, 1923 was provided on loan from CMNH for comparison, and the two Osborn species were determined to be synonyms. Both types are illustrated here.</p></div>	https://treatment.plazi.org/id/9F37571B222CFFC8679AFDF890F6FB5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B222FFFCB679AFF4C9460FDE7.text	9F37571B222FFFCB679AFF4C9460FDE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atanus tesselatus (Osborn 1923)	<div><p>Atanus tesselatus (Osborn, 1923)</p><p>Fig. 11</p><p>Eutettix tesselatus Osborn, 1923: 56</p><p>Atanus tesselatus (Osborn, 1923) in Oman, 1938 b: 381</p><p>Material examined. Holotype, ♀: Chapada, Brazil, Acc. No. 2966 / CMNH Holotype #801, Eutettix tessellatus [sic] Osborn / Eutettix tesselatus nsp. Osb. / Atanus tessellatus [sic] (Osb.), K. Zanol det 1986.</p><p>Notes. This species is known only from female specimens from the type locality in Brazil: Mato Grosso State, Chapada dos Guimarães, and from Trinidad: Port of Spain. The habitus and sternite VII are illustrated here for the first time. Further work to identify male specimens that could be associated with females is needed to clarify the identity of this species. The female holotype specimen, borrowed from CMNH for this study, is illustrated here.</p></div>	https://treatment.plazi.org/id/9F37571B222FFFCB679AFF4C9460FDE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B222FFFCB679AFD6A92B1FC83.text	9F37571B222FFFCB679AFD6A92B1FC83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolarga Oman 1938	<div><p>Bolarga Oman, 1938</p><p>Figs. 12–13</p><p>Bolarga Oman, 1938: 366 .</p><p>type species by original designation Parabolocratus bolivianus Osborn, 1923</p></div>	https://treatment.plazi.org/id/9F37571B222FFFCB679AFD6A92B1FC83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B222FFFD5679AFC489445F821.text	9F37571B222FFFD5679AFC489445F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolarga boliviana (Osborn 1923)	<div><p>Bolarga boliviana (Osborn, 1923)</p><p>Fig. 12</p><p>Parabolocratus bolivianus Osborn, 1923: 32</p><p>Parabolocratus bolivianus var. pallida Osborn, 1923: 33</p><p>Bolarga boliviana (Osborn, 1923) in Oman, 1938: 367</p><p>Parabolocratus bolivianus var. pallida Osborn, 1923 (subjective synonym of Parabolocratus bolivianus Osborn, 1923) in Oman, 1938: 367</p><p>Material examined. Paratype, ♂: Puerto Suarez, Bolivia, 150 m ., J. Steinbach / Carn.Mus.Acc. 3844 / Parabolocratus bolivianus Osb. [CMNH] . 1♂, 1♀: same locality data, USNMENT01513134, USNMENT01513049 [USNM] . 1♂: [GUYANA, new country record] British Guiana, Victoria, July 16–20, [19]62, J. Maldonado C. / Bolarga boliviana (Osborn), Det. J.N. Zahniser 2025 / USNMENT01513494 [USNM] . 2♂, 4♀: same locality data, USNMENT01513106,USNMENT01513529, USNMENT01513537, USNMENT01513445, USNMENT01513116, USNMENT01513533 [USNM] . 1♀: [BRAZIL, new country record] Corumba, Brazil, Matto Grosso, 14–22 Dez. ’19, Cornell Univ. Exp. / Cornell U. Lot 569, Sub 140 / USNMENT01513036 [USNM] . 1♀, same locality data, Cornell U. Lot 569, Sub 128 / USNMENT01513113 [USNM] . 1♀, same label data as previous, USNMENT01513077 [USNM] . 1♂, same label data as previous, USNMENT01513073 [USNM] .</p><p>Female. Abdominal sternites mostly yellow, with diffuse brownish markings posterolaterally. Sternite VII broad; posterior margin with pair of broadly rounded lobes; roundedly excavated medially; centrally with a large dark brown marking. Pygofer yellow with pair of subapical brown spots; with tuft of light colored macrosetae before brown spot and numerous brown macrosetae apically.</p><p>Notes. Among the specimens in the recovered loan was a paratype of this species from Puerto Suarez, Bolivia. Oman (1938) described the genus Bolarga Oman, 1938, designated B. boliviana as the type species, and synonymized the variety pallida Osborn with B. boliviana . Linnavuori (1959) provided some illustrations of the male genitalia. The habitus is illustrated here for the first time, and additional images of the genitalia are provided. The apices of the forewings and hindlegs and the macrosetae of the pygofer of the paratype specimen are apparently burnt or singed.</p><p>Linnavuori (1959) described and illustrated the male genitalia of this species. However, he apparently mistakenly described the pygofer “as in the following species [ B. quadrimaculata Lv.]” which is described partly as “without a sclerified lobe”. The pygofer of B. boliviana has inner ventral margins thickened and produced anteriorly into sclerotized lobes that cross each other and extend mesad of the styles (Figs. 12D–E). Linnavuori (1959) described and illustrated similar structures in Bolarga nigriloba Linnavuori, 1959 .</p><p>The coloration on the forewings is more variable than previously described. One female specimen among the series from Corumba, Brazil lacks any fuscous spots or markings at the base and middle of the forewing, while others in the series possess markings. Within the series from Guyana, the fuscous markings at the base of the forewing vary from a roundish spot (Figs. 12A–B) to a more elongate marking along the inner cell of the clavus (Fig. 12G).</p></div>	https://treatment.plazi.org/id/9F37571B222FFFD5679AFC489445F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2230FFD4679AFF4C9494FBDF.text	9F37571B2230FFD4679AFF4C9494FBDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolarga nigriloba Linnavuori 1959	<div><p>Bolarga nigriloba Linnavuori, 1959, combination reinstated</p><p>Fig. 13</p><p>Bolarga nigriloba Linnavuori, 1956: 10 . (nomen nudum: no description)</p><p>Bolarga nigriloba Linnavuori, 1956 (unavailable or invalid, linked to Bolarga nigriloba Linnavuori, 1959) in Linnavuori, 1956: 10.</p><p>Bolarga nigriloba Linnavuori, 1959: 92 .</p><p>Daltonia nigriloba (Linnavuori, 1959) in DeLong, 1982: 616.</p><p>Material examined. 1♂: Cabañas, PARAGUAY [new country record], Dist. Caacupe, Epto. Cordillera, 9 XII 1980 / RD Cave colr / Bolarga nigriloba Linnavuori, Det. J.N. Zahniser 2025 / USNMENT01513513 [USNM]. 1♂: PARAGUAY: Central Dept., San Lorenzo, 5 December 1986, J. Kochalka / Det. J.N. Zahniser 2021 / USNMENT01513072. 2♀, same data, USNMENT01513033 and USNMENT01513469 [USNM].</p><p>Female. Abdominal sternites largely brown; with diffuse yellow markings laterally; with or without longitudinal narrow medial white band; posterior margins with narrow transverse band of white. Sternite VII mostly yellow, with some brown coloration anteriorly and with irregular brown macula medially at posterior margin. Posterior margin broadly excavated, medially with small tab-like production. Pygofer yellow with numerous brown markings; with numerous macrosetae on posterior half, some concentrated at apex but groups of setae not as well-defined as in B. boliviana .</p><p>Notes. Linnavuori (1959) described this species and originally placed it in Bolarga . DeLong (1982) transferred it to Daltonia Oman, 1949, a North American genus containing three species. However DeLong’s placement was based only on superficial similarity, stating only that B. nigriloba is “…similar in color, form, and appearance…” to Daltonia estacadus (Ball) . The more convincing placement is Bolarga, a South American genus of Deltocephalini . In contrast to Daltonia, B. nigriloba shares with the type species of Bolarga, B. boliviana, a more elongate and slender shape of the body, the unique produced ventral lobe of the pygofer (Figs. 12D–E, 13C) and aedeagus with a ventral unpaired, spinelike process (Figs. 12E, 13F). B. nigriloba therefore is here reinstated in its original genus.</p></div>	https://treatment.plazi.org/id/9F37571B2230FFD4679AFF4C9494FBDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2230FFD4679AFBBA92E4FACE.text	9F37571B2230FFD4679AFBBA92E4FACE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cruziella Linnavuori & DeLong 1979	<div><p>Cruziella Linnavuori &amp; DeLong, 1979</p><p>Fig. 14</p><p>Cruziella Linnavuori &amp; DeLong, 1979 a: 49 .</p><p>type species by original designation Cruziella trispinosa Linnavuori &amp; DeLong, 1979</p></div>	https://treatment.plazi.org/id/9F37571B2230FFD4679AFBBA92E4FACE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2230FFD6679AFA969420FF36.text	9F37571B2230FFD6679AFA969420FF36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cruziella bifasciata (Osborn 1923) Zahniser 2025	<div><p>Cruziella bifasciata (Osborn, 1923), new combination</p><p>Fig. 14</p><p>Thamnotettix bifasciatus Osborn, 1923: 69</p><p>Haldorus (Haldorus) bifasciatus (Osborn, 1923) in Linnavuori, 1959: 146</p><p>Cumora (Cumora) bifasciata (Osborn, 1923) in Dmitriev et al., 2022 onward</p><p>Material examined. Paratype, ♂: Prov. del Sara, Bolivia, 450 m., J. Steinbach / Nov 1909 / C.M. Acc. 4549 / Allotype / Bolarga not Haldorus as in Lnv. / Thamno bifasciatus Osb .</p><p>Diagnosis. C. bifasciata is very similar in external appearance and color pattern to C. trispinosa Linnavuori &amp; DeLong, currently the only other species placed in the genus, but can be distinguished by the darker coloration of the face, venter, and markings on the dorsum, the thicker style apophysis, and the shorter and less produced articulating arm of the style.</p><p>Male. Pygofer longer dorsally than ventrally; slightly bulbous anteroventrally; with numerous macrosetae on posterior half. Subgenital plates narrow, sharply triangular, with lateral margins concave; with a row of 7 to 8 macrosetae on lateral margin. Style with anterior margin deeply incised; articulating arm produced only slightly beyond lateral arm; preapical lobe broad, rounded; apophysis thick, short and blunt. Connective fused to aedeagus; anterior arms long, capitate. Aedeagus with unpaired ventral median process straight, narrow, and sharply pointed; apex strongly bifid.</p><p>Notes. The only known male specimen of this species (allotype specimen) was among those recovered in loan found at USNM. Osborn’s (1923) description was repeated by Linnavuori (1959), but the species was never illustrated.</p><p>This species clearly keys to Cruziella Linnavuori &amp; DeLong, 1979 in the key to genera of South American Deltocephalini (Duan &amp; Dietrich, 2018) based on the forewing with branches of vein R reflexed (shared with Bolarga), aedeagus with an unpaired ventral process arising preapically on the ventral side (shared with Bolarga), the pygofer lacking a sclerotized process arising from the dorsal margin and extending anteroventrad into genital capsule (present in Bolarga), and subgenital plate without a dense group of setae at apex (present in Bolarga).</p><p>Dmitriev et al. (2022 onward) noted that Cumora Oman, 1938 was described as a genus in the same paper where Haldorus Oman, 1938 was described as a subgenus. The two were later consider synonyms by Linnavuori (1959) who recognized Haldorus as the valid name, but because of the original higher rank, Cumora has priority (ICZN Art. 24.1). Thus, the combination of this species with Cumora appears on the Dmitriev et al. (2022 onward) website and in the synonymy above. The genus name Haldorus has commonly been used in the literature.</p></div>	https://treatment.plazi.org/id/9F37571B2230FFD6679AFA969420FF36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
9F37571B2232FFD6679AFE9191E0FCDF.text	9F37571B2232FFD6679AFE9191E0FCDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cruziella trispinosa Linnavuori & DeLong 1979	<div><p>Cruziella trispinosa Linnavuori &amp; DeLong, 1979</p><p>Cruziella trispinosa Linnavuori &amp; DeLong, 1979 a: 49</p><p>Material examined: 2♂, 6♀: COLOMBIA [new country record]: Meta, Puerto Lopez, 9 March 1971, S.S. Roback [USNM] [USNMENT01513505, 01513590, 01513491, 01513146, 01513011, 01513087, 01513048, 01513524]</p><p>Notes. Prior to the record in Colombia reported here, this species was known only from the type locality (Bolivia: Santa Cruz Dept., San Esteban).</p><p>Bolarga quadrimaculata Linnavuori, 1955 shares with other species of Bolarga and Cruziella an aedeagus with an unpaired ventral process but does not possess the unique inner pygofer process present in the other Bolarga, and possesses a tuft of setae at the apex of the subgenital plate which is present in the other Bolarga and is not present in Cruziella . Further study is needed to determine the proper placement of this species, but for now it is retained in Bolarga .</p></div>	https://treatment.plazi.org/id/9F37571B2232FFD6679AFE9191E0FCDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zahniser, James N.	Zahniser, James N. (2025): Notes on some Neotropical Deltocephalinae (Hemiptera: Cicadellidae) described by Herbert Osborn and a review of Caphodus. Zootaxa 5692 (2): 300-320, DOI: 10.11646/zootaxa.5692.2.5, URL: https://doi.org/10.11646/zootaxa.5692.2.5
