taxonID	type	description	language	source
99080D3C3416FFE1EF194C51FBA37FAA.taxon	description	urn: lsid: zoobank. org: pub: 9 CEB 1804 - D 1 FB- 4467 - 9272 - 82 AA 78 FFD 63 A Figs 4 – 9 A; Table 3	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	diagnosis	Diagnosis Cyrtodactylus jayadityai sp. nov. (n = 5) is a small to moderately sized bent-toed gecko ranging in SVL from 47.8 mm to 54.9 mm (up to 70.7 mm including BNHS 2230), original TL ranging from 55.7 mm to 57.0 mm, having 9 to 12 supralabials and 9 to 11 infralabials. The dorsal tubercles are usually feebly keeled, bluntly conical and irregularly arranged in 19 to 21 rows (DTR) between the indistinct ventrolateral folds at the midbody; dorsal tubercles are usually four to five times the size of dorsal granular scales; 44 to 50 paravertebral tubercles (PVT 1), originating from the most anterior tubercle on the occiput to the mid-sacrum; 30 to 31 paravertebral tubercles (PVT 2) between the level of the axilla and the level of the groin and 37 to 43 mid-ventral scale rows (MVSR) between the indistinct ventrolateral folds. Males with a total of 18 to 28 pre-cloacal femoral pores (PcFP) (see ‘ Variation’ section for further details) in two different arrangements: (1) a continuous series of 18 pre-cloacal pores or PcP (9 on each side) with no visible pitted scales (2) a discontinuous series of 22 to 28 pre-cloacal femoral pores or PcFP (11 to 15 PcFP on either side of which 7 to 13 occur continuously separated by 1 to 5 poreless scales from the remaining 2 to 4 pores) with no visible pitted scales; females with 0 to 14 continuous series of pre-cloacal pits (up to 19 continuous series of precloacal-femoral pits including BNHS 2230); 13 to 15 subdigital lamellae on finger IV (excluding non-lamellar scales between the proximal and apical lamellae series); 15 to 18 subdigital lamellae on toe IV (excluding non-lamellar scales between the proximal and apical lamellae series). The dorsum has a total of 9 – 10 paired dark brown blotches posteriorly edged with whitish punctuations, occurring on either side of the mid-vertebral region on the dorsum, somewhat parallel to each other, thereby forming a thin mid-dorsal line like appearance (the dark blotches close to the vertebral region are surrounded by another irregular row of somewhat lateral and similarly bi-colored, i. e., black and white blotches on either side). A continuous series of alternating dark and light transverse bands on the dorsum of the tail; not a single row of transversely enlarged, paired subcaudals on the ventral aspect of the tail; subcaudal scales are instead arranged in small, heterogeneous and irregular transverse series.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	etymology	Etymology The specific epithet is a patronym honoring Dr. Jayaditya Purkayastha (Help Earth, Guwahati, Assam, India), an Indian herpetologist known for his remarkable contributions in the field of herpetology, particularly regarding the systematics of natricid snakes (mainly Fowlea, Hebius and Xenochrophis) and the herpetofauna of the Indo-Burmese region.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	materials_examined	Type material Holotype INDIA • ♂ adult; Tripura State, North Tripura District, Chamtilla village roadside mud bunds, surrounded by degraded mixed moist deciduous to semi-evergreen lowland forests; 24 ° 17 ′ 8.55 ″ N, 92 ° 10 ′ 32.90 ″ E, WGS 84; ca 56 to 60 m a. s. l.; 16 Aug. 2024, 11: 30 h; Arnab Deb leg.; MZMU 3795 (Figs 4 – 5, 8 A). Paratypes INDIA • 2 ♂♂ adult; same data as for holotype; MZMU 3791, MZMU 3792 • 2 ♀♀ adult; same data as for holotype MZMU 3793, MZMU 3794 (Figs 7, 8 B – D). Referred material (based on genetic data only) INDIA • 1 ♀ adult; Tripura State, North Tripura District, Rowa Wildlife Sanctuary; 24.29062 ° N 92.16496 ° E; ca 56 m a. s. l., 2 Oct. 2010; A. Datta-Roy, T. Khichi, NPI Das and I. Agarwal leg.; BNHS 2230 (Agarwal et al. 2018 a).	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	description	Description (holotype MZMU 3795) Specimen is generally in good preservation condition (Fig. 4); the abdomen has a small incision made for tissue extraction and slightly shows signs of discoloration due to preservation. Adult specimen (Figs 4 – 5) with a SVL of 50.9 mm. HEAD. Length less than one-third of SVL (HL / SVL 0.3), longer than wide (HW / HL 0.56), distinct from neck and somewhat depressed (HD / HW 0.59); interorbital area flat, canthus rostralis broadly rounded; loreal region inflated with granular scales; snout less than half of the head length (SO / HL 0.38); dorsal scales on the forehead are almost homogeneous with the exception of few rows of scales close to the bony interorbital region (between the bony orbital sockets in the frontal region) and canthus rostralis that are comparatively smaller and slightly larger in size respectively; dorsal scales above the bony interorbital region (those occurring between the bony orbital sockets in the frontal region) are sub-equal / sub-homogeneous in size with the exception of very few slightly enlarged tubercle like scales scattered irregularly; dorsal scales just above the orbit are sub-equal in size and visibly larger than the ones occurring between the bony orbital sockets in the frontal region; dorsal scales beginning slightly anterior to the bony interorbital region of the head and extending to the nape are smaller than those on the forehead and canthus rostralis, somewhat blunt and juxtaposed; scales on the bony interorbital and occipital regions are almost homogeneous in size interspersed with small but distinct tubercles (the tubercles are usually two to three times the size of the dorsal scales on the head); eye slightly more than a quarter of the head length (OD / HL 0.26); vertical pupil with crenulate margins; supraciliaries small to moderate in size, blunt, somewhat oval and slightly circular, those present in the mid-portion (above the orbit) are the largest and are more prominent; ear opening oval, obliquely orientated, small (EL / HL 0.11); orbit to ear distance is slightly greater than the eye diameter (OE / OD 1.1); rostral wider than long (RL / RW 0.57), partially and dorsally divided (nearly invisible) by an indistinct rostral groove; a single enlarged supranasal on either side, separated by three small granular internasals which are in contact with the canthus rostralis and are slightly larger than the scales on the forehead, particularly frontal region; rostral in contact with the first supralabials, nasals, two supranasals and three internasals; nostrils semicircular, openings laterally orientated, posterior half covered by the nasal pad; each nasal is in broad contact with the rostral and surrounded by the supranasal, first supralabial, and five to six small post-nasal scales on either side (left and right) of the head; a single row of small granular scales separates the orbit from the supralabials. Mental scale is wider than long (ML / MW 0.7) and somewhat triangular; two well-developed postmentals on either side; the inner pair of postmentals (PMIL) are longer than the outer pair (PMIIL) (PMIIL / PMIL 0.55); the inner pair of postmentals are bordered by the mental, infralabial I, the outer postmental and eight gular scales whereas the outer pair is bordered by inner postmental, infralabials I and II, two to three gular scales and post mental III on either side; ten supralabials on either side of the head (the last one being the smallest and is somewhat sub-equal to the bordering granular scales), bordered by a row of small to medium sized lateral scales, somewhat flat and elongated; 10 infralabials on either side of the head (the first infralabial on the left side of the head is partially divided on its posterior half by a weak suture); a row of medium to slightly large sized gular scales (largest anteriorly) ventrally bordering infralabials II to VII; gular region with small and roughly homogeneous granular scales throughout except for a few scale rows bordering the postmentals and infralabials which are larger, flat and juxtaposed. BODY. Moderately slender, trunk length less than half of the snout to vent length (TRL / SVL 0.45); dorsal scales are mostly homogeneous, i. e., small irregularly rounded granule-like scales intermixed with irregularly arranged, bluntly conical and feebly keeled enlarged tubercles (four to five times the size of granular scales) becoming more conical towards the flanks, those tubercles on the dorsolateral region of the mid-dorsum appears to be the largest; large and distinct tubercles extend posteriorly from the occipital region of the head to the second dark segment of the tail and are smaller on the occiput as well as nape than those on the dorsum; 50 paravertebral tubercles, originating from the most anterior tubercle on the occiput to the mid-sacrum; 31 paravertebral tubercles between the level of the axilla and the level of the groin; ventrolateral folds are poorly developed, not denticulate but with a single row of continuous, projected smooth tubercle like scales; ventral scales smooth, cycloid, imbricate to sub-imbricate, much larger than dorsals; 41 mid-ventral scale rows between the ventrolateral folds; 22 distinct precloacal-femoral pores in a discontinuous series (11 pore-bearing scales on the right side of which nine occur in a continuous row separated by five non pore-bearing scales from the remaining two pore bearing ones; 11 continuous series of pore-bearing scales on the left side); a single row of ten enlarged sub-equal scales (nearly identical to the size of the pore-bearing scales) posteriorly bordering the pore-bearing scales in the pre-cloacal region; three post-cloacal spurs or tubercles on both the left and right sides of the tail base ventrally, particularly on the hemipenile bulge. LEGS. Forearm (FL / SVL 0.16) and tibia (CL / SVL 0.19) roughly moderate; digits narrow, without an adhesive pad, strongly inflected at each joint, all bearing robust, recurved claws; sub-digital lamellae transversely widened beneath the basal phalanx; basal lamellae 4 - 4 - 5 - 6 - 4 on the right manus, 4 - 5 - 5 - 8 - 5 on the right pes; distal lamellae (intervening rows of broken / non-lamellar granular scales between the basal and distal lamellae series in parentheses): 8 (1) - 8 (0) - 9 (1) - 9 (2) - 8 (2) on the right manus, 9 (1) - 9 (2) - 10 (3) - 10 (3) - 11 (2) on the right pes; interdigital webbing absent from both the manus and pes; relative length of digits I <II <V <III <IV on the right manus, I <II <V <III <IV on the right pes; scales on the palms and soles are smooth, weakly raised and somewhat subimbricate. Forelimbs usually comprising of flat, imbricate to subimbricate small granular scales; the dorsal granular scales on the forearms (of the forelimbs) are usually heterogeneous whereas those on the hind-arms (of the forelimbs) are somewhat homogeneous except for a few slightly enlarged granular scales; ventral portion of the forelimbs covered with smooth, somewhat homogeneous and sub-imbricate scales except for a few slightly enlarged ones close to the wrist and elbow; dorsal scales on the hind limbs are heterogeneous in size, dorsal surfaces of the thighs and shanks have slightly enlarged scales, intermixed with scattered, enlarged, conical, feebly keeled tubercles; a quarter of the dorsal portion of thighs anteriorly (not transversely but longitudinally) and ventral aspect of hind limbs has homogeneous, slightly enlarged, smooth, imbricate to sub-imbricate scales. TAIL. Original and complete, oval in cross section, dorsoventrally depressed; dorsal tubercles on the tail base (tail portion containing the hemipenis and the first dark segment of the tail) are enlarged, flat, strongly keeled and pointed; a few enlarged and keeled tubercles randomly scattered in and around the second dark segment of the tail, remaining dorsal caudal scales smooth, flat and somewhat homogeneous in size and shape; dorsal tubercles reaches up to second dark tail blotch; subcaudal scales on the ventral aspect of the tail smooth, sub-imbricate, small, visibly heterogeneous in size and arranged in irregular transverse series, although superficially granule-like in appearance; not a single row of transversely enlarged, paired subcaudals on the ventral aspect of the tail (Fig. 5 D). COLORATION IN LIFE (Fig. 8 A). Head is primarily dark (tan) brown in colour with a combination of dark and creamish patches scattered irregularly; a thin, narrow but distinct dark streak begins at the end of the posterior orbital margin and continues to the retroarticular process of the jaw dorso-laterally on either side of the head. Posterior region of the head (back of the occiput) has a pair of distinct darkcolored blotch bordered by white punctuations posteriorly and are slightly in contact with each other, thereby giving a transverse band like appearance close to the neck. Dorsal aspect of the body has shades of brown, somewhat slightly tan brown to beige brown intermixed with paired dark blotches and each dark blotch is posteriorly bordered by a small whitish spot, thereby forming a single bi-colored blotch; collectively rows of these bi-colored blotches run parallel to each other on either side of the vertebral region, thereby forming a total of ten distinct, somewhat paired bi-colored (white edged black) blotches on the dorsum; the paired, parallel blotches on the dorsum are barely in contact with each other, thereby giving a typical mid-vertebral or mid-dorsal line like appearance. Furthermore, the dark blotches close to the vertebral region are densely surrounded by another row of somewhat lateral and similar bi-colored (black and white) blotches on either side. Dorsal aspect of all the limbs with shades of brownish-grey, somewhat tan to beige brown intermixed with irregular dark and whitish blotches. Tail is complete, having 12 distinct alternate dark and whitish transverse bands (the dark bands are much larger than the whitish ones which are reduced to narrow transverse bands) dorsally which continue to the tail tip. Ventral region (excluding the underside of the tail) is mostly light, somewhat faded or beige in coloration, along with small, irregular creamish spots scattered irregularly close to the ventro-lateral edges of the abdomen, forelimbs as well as head and ventral aspect of hindlimbs as well. Ventral aspect of the tail with 12 distinct alternate dark and whitish transverse bands (the dark bands are much larger than the whitish ones which are reduced to narrow transverse bands; some of the whitish bands in the central region of the tail are broken into spots) which continue to the tail tip. COLORATION IN PRESERVATIVE (Fig. 4). The specimen has become dull in preservative comparatively to its living condition. Variation Please refer to Table 3 and Fig. 6 for variations within the new species. Most of the morphological characters in all the male and female paratypes are within the range of the holotype except for the precloacal regions. Detail descriptions of the pre-cloacal regions and their variations: MZMU 3792 (an adult ♂) has a continuous series of 18 PcP (9 continuous pores on eich side with no poreless scales); MZMU 3791 (an adult ♂) has a total of 28 PcFP occurring in a discontinuous series (right side with 15 pores of which 13 are present in a continuous series followed by four poreless scales and then two additional pores whereas left side with 13 of which seven pores are present in a continuous row followed by a single poreless scale, then three pores, then five poreless scales and then a final series of three pores); MZMU 3794 (an adult ♀) and MZMU 3793 (an adult ♀) have 0 and 14 precloacal pits, respectively; BNHS 2230 (an adult ♀) has 19 pitted scales in the precloacal-femoral region as mentioned in Agarwal et al. (2018 a) versus the holotype (MZMU 3795, an adult ♂) with a discontinuous series of 22 PcFP (right side with 11 pores of which nine are present in a continuous series followed by five poreless scales and then two additional pores whereas left side with a continuous series of 11 pores with no intermediate poreless scales). Sexual dimorphism Please refer to Table 3 for sexual dimorphism within the new species. Apart from the strong differences in the precloacal and femoral region, i. e., 18 to 28 PcFP in males versus 0 to 14 pitted (0 to 19 pitted scales including BNHS 2230) pre-cloacal scales in females, no other prominent differences were found between the two sexes (Fig. 6). Comparisons Genetically, Cyrtodactylus jayadityai sp. nov. is a sister to C. tripuraensis, both of them members of the‘ khasiensis ’ group and can be differentiated from each other by a pairwise genetic distance of at least 4.7 to 5.2 % in the mitochondrial ND 2 gene (Table 2) and certain non-overlapping morphological characters (Table 3). Morphologically, Cyrtodactylus jayadityai sp. nov. differs from the closest members of the ‘ khasiensis ’ group, including C. tripuraensis, for the combination of the following characters: (1) males with 18 PcP to 28 PcFP in a continuous to discontinuous series, respectively (versus 29 to 37 continuous series of PcFP in males of C. tripuraensis; 27 PcFP in a continuous series in males of C. ngengpuiensis Boruah, Narayanan, Lalronunga, Deepak & Das 2024); (2) females with 0 to 14 pitted scales (up to 19 pits including BNHS 2230) in the pre-cloacal region (versus 20 to 29 pitted scales in females of C. tripuraensis); (3) 19 to 21 rows of DTR at midbody (versus 22 to 24 rows in C. ayeyarwadyensis Bauer, 2002); (4) 37 to 43 MVSR between the indistinct ventro-lateral folds (versus 32 to 37 MVSR in C. ayeyarwadyensis); (5) dorsum with a total of 9 – 10 small-sized, paired dark brown blotches posteriorly edged with whitish punctuations, occurring on either side of the mid-vertebral region on the dorsum, somewhat parallel to each other, thereby forming a thin mid-dorsal line like appearance (versus dorsum mostly with paired, enlarged irregular blotches on either side of the vertebral region that are parallel and strongly connected to each other alongside the presence of a distinct, thin dark dorso-lateral stripe on either side in C. ngengpuiensis; the paired dark blotches on either side of the vertebral region are at least 2 × longer in C. tripuraensis). For further morphological differences between C. jayadityai and its congeners in the ‘ khasiensis ’ group, refer to the comparative data provided in Table 4. Sequence divergence (Table 2) Cyrtodactylus jayadityai sp. nov. is supported as a distinct species based on the mitochondrial ND 2 gene, exhibiting 0 % intraspecific divergence among the sequenced samples, with no current evidence of intraspecific phylogenetic splitting within the lineage. The new species shows an interspecific genetic distance of 4.7 – 5.2 % from its closest congener C. tripuraensis, 6.2 – 7.6 % from C. ngengpuiensis, and 9.7 – 10 % from C. ayeyarwadyensis. Furthermore, it is more than 10 % genetically divergent from all other congeners within the‘ khasiensis ’ group.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	biology_ecology	Natural history and biology The individuals of Cyrtodactylus jayadityai sp. nov. predominantly inhabit small burrows and crevices along roadside mud bunds, often covered with moss and large tree roots connected to these bunds in association with mixed moist deciduous to semi-evergreen lowland forests (Fig. 10 A – B) (56 to 100 m a. s. l.). These geckos exhibit strictly nocturnal behavior, with individuals regularly observed emerging from their retreats to forage between 20: 00 h and 03: 30 h. Most individuals were recorded at heights ranging from 1.5 to 1.8 m above ground level; however, some were observed as high as approximately 10 meters. Uncollected gravid females ranging in SVL from 49 mm to 60 mm were observed during the months of March to August, suggesting that breeding occurs during pre and peak monsoon. Notably, several individuals were also recorded during the peak winter month of January, despite ambient temperatures as low as 14 ° C. The species is sympatric with other gekkonid species, including Hemidactylus frenatus Duméril & Bibron 1836, H. aquilonius Mcmahan & Zug, 2007, H. platyurus Schneider, 1797 and H. cf. brooki Gray, 1845. Dietary observations suggest a preference for small to medium-sized arthropods like forest cockroaches and huntsman spiders. Behaviorally, individuals are generally shy and avoidant when approached, typically retreating into their burrows. However, when handled, they exhibit a strong biting behavior, indicating a possible defense mechanism against predators or threats.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	distribution	Distribution Based on the current data, Cyrtodactylus jayadityai sp. nov. is found only in a small patch of forests in the lowlands of North Tripura District covering an area of 8 to 12 km 2 ranging from the western (including Chamtilla Village) and southern boundaries of Rowa WLS towards its eastern boundary, further stretching into Indurail reserve forests close to Assam-Tripura border (Fig. 1).	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	conservation	Conservation status Based on our current data of distribution, we consider this species Data Deficient (DD) as per the IUCN. However, the species restricted distribution (8 to 12 km 2) in fragmented and anthropogenically pressured habitats suggests that it may qualify for a threatened category (Endangered or Critically Endangered) if assessed formally, given its vulnerability to habitat alteration.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	vernacular_names	Suggested common name Jayaditya’s forest bent-toed gecko.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3416FFE1EF194C51FBA37FAA.taxon	discussion	Remarks Agarwal et al. (2018 a) described Cyrtodactylus tripuraensis based on 11 specimens from Tripura, northeast India of which 10 were collected from Gomati and Sephaijala Districts (including type locality of C. tripuraensis) whereas the remaining one paratype (BNHS 2230) was collected from Rowa WLS, North Tripura District (very close to the type locality of the currently described new species). Our genetic data excised from MZMU 3795 (holotype of Cyrtodactylus jayadityai sp. nov.) collected from the North Tripura District has an intraspecific genetic divergence of 0 % from BNHS 2230 (KM 255201). As a result, we refer BNHS 2230 as additional material of C. jayadityai based on the existing genetic data. Together, these two samples (BNHS 2230 and MZMU 3795) of C. jayadityai are 4.7 to 5.2 % divergent from all other samples of C. tripuraensis in the mitochondrial ND 2 gene. To date, this specimen (BNHS 2230) has not been morphologically examined by us, so we decide not to include it amongst our type series (holotype and paratypes) of C. jayadityai. Apart from the size and its related parameters (e. g., SVL, HL, etc., as BNHS 2230 is a bit larger in size (Agarwal et al. 2018 a )), the majority of the diagnostic morphological characters of BNHS 2230 as mentioned in Agarwal et al. (2018 a) falls well within the range of the type series of the newly described species (Table 3). Furthermore, those diagnostic parameters show the slightest of variations between the type series (holotype and paratypes) and the referred material (BNHS 2230) of the new species is mentioned separately within brackets in the Diagnosis section of C. jayadityai to avoid any sort of confusion in the future.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3407FFE6EDFC4850FA847FA5.taxon	discussion	Cyrtodactylus khasiensis was described based on three specimens from the Khasi Hills of Meghalaya, northeast India (precise locality unknown) by Jerdon (1870) without the designation of a type specimen for the species. As a result, Agarwal et al. (2018 a) designated BMNH 1906.8.10.4 (formerly ZSIK 6198) as a lectotype of C. khasiensis for the taxonomic stability of the species and also provided a detailed redescription of the species based on two paralectotypes and seven freshly collected specimens. Agarwal et al. (2018 a) also provided the first mitochondrial genetic data of one of the topotypical specimens of Cyrtodactylus khasiensis sensu stricto. However, Agarwal et al. (2018 a) mentioned that C. khasiensis had ‘ subcaudal scalation of original tail with median series of paired enlarged plates, covering underside of tail entirely toward tail-tip’. Upon examination of three freshly collected adult specimens of Cyrtodactylus khasiensis sensu stricto from Sohra, East Khasi Hills (MZMU 3798, ADBU / GT / 1 and ADBU / GT / 2) of Meghalaya, we found that all of these specimens never possessed a distinct median series of paired enlarged plate-like subcaudals across the underside of tail as mentioned in Agarwal et al. (2018 a). Rather, our collected topotypical specimens of C. khasiensis exhibit a small to medium, visibly heterogeneous and irregular transverse series of sub-imbricate subcaudal scales that are superficially granule-like in appearance across the ventral aspect of the tail, similar to all other described members of the ‘ khasiensis ’ group. However, our C. khasiensis specimens also bear fewer rows (6 to 12 continuous or discontinuous rows) of subcaudal scales possessing a somewhat paired, median series of minutely to slightly enlarged scales when compared to its flanking / bordering smaller subcaudal scales. This suggests that Agarwal et al. (2018 a) most likely used the terminology ‘ paired enlarged plates’ for C. khasiensis to describe these very few rows of weakly enlarged, median series of somewhat paired subcaudal scales. Many species within the ‘ khasiensis ’ group (e. g., Cyrtodactylus urbanus Purkayastha, Das, Bohra, Bauer & Agarwal, 2020; C. lungleiensis Lalremsanga, Chinliansiama, Bohra, Biakzuala, Vabeiryureilai, Muansanga, Malsawmdawngliana, Hmar, Decemson, Siammawii, Das & Purkayastha, 2022; C. guwahatiensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 and C. bapme Kamei & Mahony, 2021) also exhibit a subcaudal scalation completely similar to C. khasiensis sensu stricto whereas in some species (e. g., C. tripuraensis, C. jayadityai sp. nov. and C. siahaensis Purkayastha, Lalremsanga, Litho, Rathee, Bohra, Mathipi, Biakzuala & Muansanga, 2022), all the subcaudal scale rows exhibit a small to medium, irregularly sized, heterogeneous scalation throughout the underside of the tail. In terms of morphology, the case of the ‘ subcaudal scalation’ within all the members of the Cyrtodactylus khasiensis group is completely overlapping across species. This is why we recommend not using ‘ subcaudal scalation’ as a valid morphological character for differentiating new species with the congeners within the ‘ khasiensis ’ group, since previously this character was used as a strong diagnostic feature by several workers in their ‘ comparisons and tables’ section to differentiate their new species with the congeners, particularly C. khasiensis (Agarwal et al. 2018 a, 2018 b; Purkayastha et al. 2020, 2021, 2022; Mirza et al. 2021, 2022; Bohra et al. 2022; Lalremsanga et al. 2022, 2023; Boruah et al. 2024; Bharali et al. 2025). Prior to this study, Cyrtodactylus khasiensis was known precisely only from the hills of East Khasi Hills District, Sohra, Meghalaya. Another collected adult male specimen (MZMU 3797) from Umdohlun, South West Khasi Hills District, Meghalaya, is only ~ 1.9 % genetically divergent from the available mitochondrial ND 2 data of C. khasiensis (KM 255188). As a result, we confirm the occurrence of C. khasiensis sensu stricto further westwards from its currently known locality (Sohra, East Khasi Hills District) towards South-West Khasi Hills District close to India-Bangladesh border based on the molecular data.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C3407FFE6EDFC4850FA847FA5.taxon	conservation	Conservation status Based on our current data of distribution, we consider this species Data Deficient (DD) as per the IUCN.	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343EFFDCED984824FD477F3E.taxon	materials_examined	Holotype INDIA • ♂ adult; Durtlang, Aizawl District, Mizoram State; MZMU 2015 Paratypes INDIA • 3 ♂♂ adult; Mizoram University Campus, Aizawl District, Mizoram State; MZMU 2012, MZMU 2021, MZMU 2032 • 1 ♀ adult; Tamdil National Wetland Saitual District, Mizoram State; MZMU 2067 • ♂ adult; same collection data as for preceding; MZMU 2020 • ♂ adult; Zawngtahlipui stream, Sihphir, Aizawl District, Mizoram State; MZMU 2014 • 2 ♀♀ adult; Durtlang, Aizawl District, Mizoram State; MZMU 2018, MZMU 2056	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343EFFDCEDAE4A15FAE97856.taxon	materials_examined	Holotype INDIA • ♂ adult; Outskirts of Siju village, South Garo Hills District, Meghalaya State; MZMU 2160. Paratypes INDIA • 3 ♂♂ adult; Outskirts of Siju village, South Garo Hills District, Meghalaya State; MZMU 2157, MZMU 2159, MZMU 2161 • 1 ♂ sub-adult; same collection data as for preceding; MZMU 2158	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343EFFDCED994DEDFD6F7A08.taxon	materials_examined	Holotype INDIA • ♂ adult; Sailam Community Reserved Forest outskirts of Sailam Village, Aizawl District, Mizoram State; MZMU 1989 Paratypes INDIA • 2 ♂♂ adult; Sailam Community Reserved Forest outskirts of Sailam village, Aizawl District, Mizoram State; MZMU 1987; MZMU 1990 • 3 ♀♀ adult; same collection data as for preceding; MZMU 1985, MZMU 1986, MZMU 1988	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343EFFDCEC054F3BFDCA7B07.taxon	materials_examined	Referred materials INDIA • 5 ♂♂ adult; Guwahati, Assam State, India; VR / ERS / ZSI / 690, VR / ERS / ZSI / 691, VR / ERS / ZSI / 693, VR / ERS / ZSI / 694, VR / ERS / ZSI / 696 • 2 ♀♀ adult; same collection data as for preceding VR / ERS / ZSI / 692, VR / ERS / ZSI / 695	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343FFFDDEDA04ABAFB78793D.taxon	materials_examined	Holotype INDIA • adult ♂; Umroi military cantonment area, Ri-Bhoi District, Meghalaya State; MZMU 2542. Paratypes INDIA • 1 ♀ adult; Umroi military cantonment area, Ri-Bhoi District, Meghalaya State; MZMU 2543 • 2 ♂♂ adult; same collection data as for preceding; MZMU 2544 and MZMU 2545	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343FFFDDEDA94850FBDF7E19.taxon	materials_examined	Holotype INDIA • ♂ adult; Outskirts of Siju village, South Garo Hills District, Meghalaya State; MZMU 2155 Paratypes INDIA • 2 ♂♂ adult; Outskirts of Siju village, South Garo Hills District, Meghalaya State; MZMU 2154, MZMU 2156 • ♀ adult; same collection data as for preceding; MZMU 2153	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343FFFDDED984C08FC9A7A8F.taxon	materials_examined	Holotype INDIA • ♂ adult; New Colony-I, Siaha District, Siaha Town, Mizoram State; MZMU 2445 Paratypes INDIA • 2 ♂♂ adult; New Colony-I, Siaha District, Siaha Town, Mizoram State; MZMU 2444, MZMU 2446 • 1 ♀ adult; same collection data as for preceding; MZMU 2443 • 2 ♀♀ juvenile; same collection data as for preceding; MZMU 244, MZMU 2449	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343FFFDDEC294B2BFEA87F8A.taxon	materials_examined	Holotype INDIA • ♂ adult; Basistha, Guwahati, Assam State; VR / ERS / ZSI / 683 Paratypes INDIA • 3 ♂♂ adult; Basistha, Guwahati, Assam State; VR / ERS / ZSI / 686, VR / ERS / ZSI / 685, VR / ERS / ZSI / 687 • 3 ♀♀ adult; same collection data as for preceding; VR / ERS / ZSI / 684, VR / ERS / ZSI / 688, VR / ERS / ZSI / 689	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343FFFDEED9F4F86FD6E7DA3.taxon	materials_examined	Holotype INDIA • ♂ adult; Tuikhur veng, Chawngbawla section, Ngopa Town, Champhai District, Mizoram State; MZMU 2553 Paratypes INDIA • 1 ♂ adult; Tuikhur veng, Chawngbawla section, Ngopa Town, Champhai District, Mizoram State; MZMU 2555 • 5 ♀♀ adult; same collection data as for preceding; MZMU 2552, MZMU 2554, MZMU 2556, MZMU 2557, MZMU 2558	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343CFFDEEF3B4A69FBEF7817.taxon	materials_examined	Holotype INDIA • ♂ adult; Electric Veng, Zobawk, Lunglei District, Mizoram State; MZMU 2432 Paratypes INDIA • 2 ♂♂ adult; Lunglei District, Mizoram State; MZMU 2431, MZMU 2426 • 3 ♀♀ adult; same collection data as for preceding; MZMU 2430, MZMU 2427, MZMU 2429	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343CFFDEEC104D2EFB5C78DB.taxon	materials_examined	Additional material INDIA • ♂ adult; Roadside loose soils in Gomati District, Tripura State; MZMU 3796	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
99080D3C343CFFDEEDA44892FAF77EDA.taxon	materials_examined	Holotype INDIA • ♂ adult; Hall Veng, Vairengte, Kolasib District, Mizoram State; MZMU 2903 Paratypes INDIA • 3 ♂♂ adult; Vairengte, Kolasib District, Mizoram State; MZMU 2904, MZMU 2901, MZMU 2902 • 3 ♀♀ adult; same collection data as for preceding; MZMU 2905, MZMU 2626, MZMU 2742	en	Bohra, Sanath Chandra, Deb, Arnab, Thongni, Goldenstar, Bhattacharjee, Rupankar, Biakzuala, Lal, Lalremsanga, Hmar Tlawmte, Swargiary, Pranjal, Roy, Rita (2026): A new micro-endemic species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from the lowlands of Northeast India, with additional morphological notes on Cyrtodactylus khasiensis Jerdon, 1870 based on topotypical specimens from Meghalaya, India. European Journal of Taxonomy 1048: 256-303, DOI: 10.5852/ejt.2026.1048.3255, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3255/14405
