taxonID	type	description	language	source
972BDD17553D986DB6E6FB52FE223185.taxon	type_taxon	Type species. Diplommatina acme Laidlaw, 1949	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553D986DB6E6FB52FE223185.taxon	diagnosis	Diagnosis. Shell small, sinistral, conical to very elongated conical; in some species, a keel emerges on the periphery and forms angular lower whorls. Shell surface usually covered with regular and dense radial ribs. In certain species, the ribs are modified into outwardly projecting triangular plates or coiled into tubular projections. Aperture oval, oblique to the shell axis. Peristome attached to the body whorl, double or triple; inner peristome protruding; outer peristome expanded but not reflected. Internal constriction (if present) ventrally situated with a thin radial palatal fold at its base. One short horizontal palatal lamella (if present) visible through translucent shell. Columellar tooth small. Umbilicus closed. Operculum corneous. Radula consists of a strong central tooth with a large central cusp and two pairs of smaller lateral cusps. Basal plate of central tooth with two attached small cusps. Lateral and marginal teeth with very short basal parts. Deferential diagnosis. All known species of Pagodapalaina gen. nov., were previously assigned to Diplommatina Benson, 1849, but recent phylogenetic studies have consistently supported the monophyly of the new genus as the sister group to Diplommatina (Webster et al., 2012; Boonmachai et al., 2023; Köhler, 2024). Morphologically, the sinistrally coiled, conical shell of Pagodapalaina gen. nov. clearly distinguishes it from Diplommatina. In terms of radular structure, Pagodapalaina gen. nov. exhibits a significantly enlarged central cusp on the central tooth compared to Diplommatina, and both the lateral and marginal teeth possess extremely short basal parts, resulting in a radula that appears notably compact (see Boonmachai et al., 2023). Additionally, the basal plate of Diplommatina typically bears a single pointed cusp, whereas Pagodapalaina gen. nov., has two relatively blunt cusps (see Boonmachai et al., 2023; Z. - Y. Chen, unpublished data). This feature closely resembles that of Luzonocoptis Páll-Gergely & Hunyadi, 2017, a genus from Luzon Island, Philippines, which also possesses a sinistral shell (Páll-Gergely & Hunyadi, 2017). However, the cusps of Pagodapalaina gen. nov., are more robust, nearly conical in shape, and the two cusps on the basal plate of the central tooth are positioned adjacent to each other rather than being widely separated. This compact radula type may potentially be associated with saxicolous habits. From a broader comparative perspective, the keeled and elongate-conical whorls of Pagodapalaina gen. nov. readily distinguish it from the sinistral taxa occurring in adjacent regions (see Vermeulen, 1993; Greķe, 2017; Nurinsiyah & Hausdorf, 2017). Although certain species of Palaina O. Semper, 1865 (e. g. Palaina attenboroughi Greķe 2017, Palaina dohertyi E. A. Smith, 1897 and Palaina parietalis Neubert & Bouchet, 2015) exhibit a similar shell outline, their weak internal lamellar structure provides a clear means of separating them from Pagodapalaina gen. nov.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553D986DB6E6FB52FE223185.taxon	etymology	Etymology. The name of the new genus is derived from “ Pagoda ”, referring to the tiered, keeled and tower-like shape of the shell, reminiscent of East Asian pagodas, and diplommatinid genus name Palaina. This genus name is feminine.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553D986DB6E6FB52FE223185.taxon	discussion	Included species. See below.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553D986DB6E6FB52FE223185.taxon	distribution	Distribution. Thailand, Malaysia, Indonesia. The known species of the new genus are distributed on the Malay Peninsula and Sumatra.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	description	(Figure 1 A)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	materials_examined	Type material. Holotype NHMUK 19930124, 2 paratypes NHMUK 19930125, Gua Kantan, Chemor, N. of Ipoh Peral, Perak, Peninsular Malaysia, 4 ° 46 ’ N, 101 ° 07 ’ E, leg. G. Davison, 20 Jan. 1991.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	etymology	Etymology. We adopted Davison’s manuscript name with minor modification. The name hortulanica, meaning “ garden-dwelling ” in Latin, may refer to the flourishing vegetation of the type locality.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	diagnosis	Diagnosis. The new species can be distinguished from most other species by its bearing relatively fewer whorls and a stouter (vs slender in other species) shell. Its overall shape resembles Pagodapalaina carinaspinosa (Maassen, 2002) comb. nov., P. gadutensis (Maassen, 2002) comb. nov., P. karoensis (Maassen, 2002) comb. nov., and P. krabiensis (Panha & Burch, 1998) comb. nov., but differs from them in its larger size and the rib structure.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	description	Description. Shell small, sinistral, pale with yellow hue, elongated conical, with 5.75 convex whorls; suture impressed; width of body whorl slightly smaller than that of penultimate whorl; a keel emerges after the penultimate whorl and formed angular whorls. Protoconch consisting of 1.5 whorls, somewhat matte; boundary between protoconch and teleoconch not clear. Teleoconch surface with fine and regularly arranged radial and prosocline ribs, approximately 8 – 9 ribs per 500 μm on the body whorl, weak and dense spiral threads situated. Aperture oval, oblique to the shell axis. Peristome attached to the body whorl, double or triple; inner peristome protruding; outer peristome expanded but not reflected. Internal constriction ventrally present with a thin radial (to the shell axis) palatal fold at base. One horizontal palatal lamella observed through translucent shell. Columellar tooth small, downward sloping. Umbilicus closed. Operculum unknown. Measurements. Shell height: 2.4 mm, shell width: 1.3 mm (holotype); shell height: 2.2 – 2.3 mm, shell width: both 1.2 mm (paratypes, n = 2)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	discussion	Remarks. Taken directly from Davison’s note on specimen label: “ In loose dry soil on the extreme north end of big limestone outcrop, about halfway up the slope to summit. In low forest of Streblus bushes, altitude 100 m. ”	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.taxon	distribution	Distribution. Malaysia: Perak. This new species is known only from the type locality.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553F986CB6E6FA06FCB034BB.taxon	description	(Figure 2 A)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553F986CB6E6FA06FCB034BB.taxon	materials_examined	Type locality. Diplommatina acme: [Malaysia] “ Kaki Bukit, Perlis ”; Diplommatina laemsonensis: Thailand, Satun Province, La-ngu District, Limestone Hill, 6 ° 54 ’ 43.7 ” N, 99 ° 41 ’ 59.0 ” E. Type material. Holotype (NHMUK 1948.11.12.5) of Diplommatina acme, Kaki Bukit, Perlis, Malaya, M. W. F. Tweedie Esq.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553F986CB6E6FA06FCB034BB.taxon	discussion	Remarks. Diplommatina laemsonensis Boonmachai, Bergey & Nantarat, 2023 does not differ morphologically from typical P. acme. Therefore, we considered it a synonym of the latter.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553F986CB6E6FA06FCB034BB.taxon	distribution	Distribution. Malaysia: Perak, Perlis; Thailand: Satun.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FF43FD073291.taxon	description	(Figure 2 B)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FF43FD073291.taxon	materials_examined	Type locality. [Malaysia] “ Gua Bama, Pahang ” Type material. Holotype (NHMUK 1948.11.12.8), Gua Bama, Pahang, Malaya, M. W. F. Tweedie Esq. Distribution. Malaysia: Pahang, Perak.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FDF8FD0430DD.taxon	description	(Figure 2 C)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FDF8FD0430DD.taxon	materials_examined	Type locality. [Indonesia] W. Sumatra: near entrance of Cave Gua Pangian, 3 km N of village Lintau, SE of Bukittinggi, 00 ° 28 ’ 19.5 ” S, 100 ° 45 ’ 11.7 ” E. Type material. 5 paratypes (NHMUK 20030224), same as type locality, vii. 1996, presented by W. J. M. Maassen. The holotype (RMNH. MOL. 92941) at NBC was not examined. Additional paratypes are housed in multiple repositories (see Maassen 2002 for details).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FDF8FD0430DD.taxon	distribution	Distribution. Indonesia: West Sumatra.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FC2FFDA6355E.taxon	description	(Figure 2 D)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FC2FFDA6355E.taxon	materials_examined	Type locality. [Malaysia] “ ad Bukit Pondong leg. cl R. Hungerford ” Type material. Holotype (SMF 105374 / 1), Malakka: Bükit Pondong (Perak), Hüngerford l., Sammlung O. v. Moellendorff. Other material examined. NHMUK 1891.7.6.5 - 6. Perak! [sic], Types P. Z. S. 1891, G. B. Sowerby Esq.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FC2FFDA6355E.taxon	distribution	Distribution. Malaysia: Perak.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6F9A8FD04348D.taxon	description	(Figure 2 E)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6F9A8FD04348D.taxon	materials_examined	Type locality. [Indonesia] W. Sumatra: Kampung Desa Gadut, limestone rocks near the village, E of Payakumbuh, 00 ° 15 ’ 36.0 ” S, 100 ° 43 ’ 58.7 ” E. Type material. 2 paratypes (NHMUK 20030225), same as type locality, vii. 1997, presented by W. J. M. Maassen. The holotype (RMNH. MOL. 92945) at NBC was not examined. Additional paratypes are housed in multiple repositories (see Maassen 2002 for details).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6F9A8FD04348D.taxon	distribution	Distribution. Indonesia: West Sumatra.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755399869B6E6F90DFDE5338C.taxon	description	(Figure 2 F)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755399869B6E6F90DFDE5338C.taxon	materials_examined	Type locality. Thailand, Khaopoo Khaoya National Park, Pattalung Province at 7 ° 39 ’ 38 ” N, 100 ° 2 ’ 30 ” E, 220 meters elevation. Type material. Holotype (CUMZ Di 040) and six paratype shells (CUMZ Di 041), same as type locality. Additional paratypes are housed in UMMZ.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755399869B6E6F90DFDE5338C.taxon	distribution	Distribution. Thailand: Phatthalung.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FE9EFD1231B3.taxon	description	(Figure 2 G)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FE9EFD1231B3.taxon	materials_examined	Type locality. [Indonesia] N. Sumatra: near the entrance of limestone cave “ Liangdehar ”, near Kuta Buluh, 40 km NW of Berastagi. Type material. 2 paratypes (NHMUK 20030226), same as type locality, vi. 1996, presented by W. J. M. Maassen. The holotype (RMNH. MOL. 92940) at NBC was not examined. Additional paratypes are housed in multiple repositories (see Maassen 2002 for details).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FE9EFD1231B3.taxon	distribution	Distribution. Indonesia: North Sumatra.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FCCCFDA33718.taxon	description	(Figure 2 H)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FCCCFDA33718.taxon	materials_examined	Type locality. [Thailand] Tam Sua, Krabi Province, at 8 ° 7 ’ 32 ” N, 98 ° 55 ’ 31 ” E, 140 meters elevation. Type material. Holotype (CUMZ Di 031) and 3 paratypes (CUMZ Di 032), same as type locality. Additional paratypes are housed in UMMZ (255345).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FCCCFDA33718.taxon	distribution	Distribution. Thailand: Krabi.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FB6AFDA634C5.taxon	description	(Figure 3 A)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FB6AFDA634C5.taxon	materials_examined	Type locality. Diplommatina (Sinica) lenggongensis: [Malaysia] “ Lenggong, Perak ”, Diplommatina lenggongensis nawawii: [Malaysia] limestone outcrop, now isolated as an island by the impounded waters of Temengor lake, at 5 ° 24 ’ N, 101 ° 18 ’ 30 ” E, at the perimeter of Temengor Forest Reserve, Hulu Perak district, Perak, Peninsular Malaysia. Type material. Syntype (NHMUK 1943.6.28.1) of Diplommatina (Sinica) lenggongensis, Lenggong, Perak, J. R. le B. Tomlin.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FB6AFDA634C5.taxon	discussion	Remarks. Contrary to the statement in the original description, the type material of Diplommatina lenggongensis nawawii could not be located in the NHM collections. The registration number ‘ 931229.1 ’ cited therein does not correspond to any recognised NHMUK catalogue number. Due to this discrepancy, and given that the original description lacks illustrations, the identity of the species cannot be confirmed based on the limited diagnostic information. We therefore treat it here as a species inquirenda, although its generic assignment appears to be unproblematic.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FB6AFDA634C5.taxon	distribution	Distribution. Malaysia: Perak.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FF43FCCD3196.taxon	description	(Figure 3 B)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FF43FCCD3196.taxon	materials_examined	Type locality. Thailand, Khaochong Wildlife Sanctuary, Trang Province at 07 ° 33 ’ 59 ” N, 099 ° 46 ’ 32 ” E, 130 meters elevation. Type material. Holotype (CUMZ Di 004) and 3 paratypes (CUMZ Di 005), same as type locality. Additional paratypes are housed in multiple repositories (see Panha 1999 b for details).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FF43FCCD3196.taxon	discussion	Remarks. In Panha (1998), the figures of Diplommatina hidagai Panha, 1998 and Diplommatina naiyanetri Panha, 1998 do not correspond with their respective descriptions. Examination of the type material by one of the present authors (CS) confirms that the descriptions are correct, and that the original figures were mistakenly interchanged. This error subsequently led to misapplied names in several later studies (e. g. Webster et al. 2012; Boonmachai et al., 2023; Chen et al., 2023; Köhler, 2023).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FF43FCCD3196.taxon	distribution	Distribution. Thailand: Phatthalung, Sutrat Thani, Trang.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FCE7FDA63716.taxon	description	(Figure 3 C)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FCE7FDA63716.taxon	materials_examined	Type locality. [Malaysia] “ Lenggong, Perak ” Type material. Holotype (NHMUK 1948.11.12.6), Lenggong, Perak, Malaya, M. W. F. Tweedie.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FCE7FDA63716.taxon	distribution	Distribution. Malaysia: Perak.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FB60FDEE357C.taxon	description	(Figure 3 D)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FB60FDEE357C.taxon	materials_examined	Type locality. Thailand, Tamkuha, Surathani Province at 8 ° 4 ’ 30 ” N, 98 ° 50 ’ 33 ” E. Type material. Holotype (CUMZ Di 043) and 4 paratypes (CUMZ Di 044), same as type locality. Additional paratypes are housed in UMMZ.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FB60FDEE357C.taxon	distribution	Distribution. Thailand: Sutrat Thani.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6F98DFDBA34DA.taxon	description	(Figure 3 E)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6F98DFDBA34DA.taxon	materials_examined	Type locality. Prakayang Cave, Ranong Province, Thailand, at 10 ° 19 ’ 37 ” N, 98 ° 48 ’ 02 ” E, 117 meters elevation. Type material. Holotype (CUMZ Di 006) and 5 paratypes (CUMZ Di 007), same as type locality. Additional paratypes are housed in multiple repositories (see Panha 1999 a for details).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6F98DFDBA34DA.taxon	distribution	Distribution. Thailand: Ranong.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755349866B6E6F88FFD67329A.taxon	description	(Figure 3 F)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755349866B6E6F88FFD67329A.taxon	materials_examined	Type locality. Diplommatina (Sinica) sinistra: [Malaysia] “ Sungei Siput, Perak ”; Diplommatina prakaiphetensis: Thailand, Satun Province, Thung Wa District, Prakaiphet Hill, 7 ° 00 ’ 00.1 ” N, 99 ° 46 ’ 08.7 ” E. Type material. Syntype (NHMUK 1938.10.25.4) of Diplommatina (Sinica) sinistra, Sungei Siput, Perak, Malaya States. J. R. le B. Tomlin Esq.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755349866B6E6F88FFD67329A.taxon	discussion	Remarks. Diplommatina prakaiphetensis Boonmachai, Bergey & Nantarat, 2023 does not differ morphologically from typical P. sinistra. Therefore, we consider it a synonym of the latter.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755349866B6E6F88FFD67329A.taxon	distribution	Distribution. Malaysia: Perak; Thailand: Satun.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FB0BFDA6350F.taxon	description	(Figure 3 G)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FB0BFDA6350F.taxon	materials_examined	Type locality. [Malaysia] “ Sungei Siput, Perak ” Type material. Holotype (NHMUK 1948.11.12.7), Sungei Siput, Perak, Malaya, M. W. F. Tweedie Esq.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FB0BFDA6350F.taxon	discussion	Remarks. Foon et al. (2017) synonymised this subspecies with the nominate one. Based on the type material, P. s. brevior exhibits a smaller shell size and less sharply developed keels compared to the nominate subspecies. Therefore, we tentatively retain the subspecies as valid.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FB0BFDA6350F.taxon	distribution	Distribution. Malaysia: Perak.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FDEBFDA637F9.taxon	description	(Figure 1 B)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FDEBFDA637F9.taxon	materials_examined	Type locality. [Malaysia] “ Perak ” Type materials. Original description states type material is in “ Indian Museum, Calcutta ” and was not examined in this study. Material examined. NHMUK 1912.4.16.237, Perak ‘ D 66 ’, Beddome collection.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FDEBFDA637F9.taxon	distribution	Distribution. Malaysia: Perak.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359865B6E6F918FDEE338C.taxon	description	(Figure 3 H)	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359865B6E6F918FDEE338C.taxon	materials_examined	Type locality. [Thailand] Klongsang Wildlife Santuary, Surathani Province at 8 ° 58 ’ 3 ” N, 98 ° 41 ’ 54 ” E, 170 meters elevation. Type material. Holotype (CUMZ Di 028) and 7 paratypes (CUMZ Di 029), same as type locality. Additional paratypes are housed in UMMZ (255344).	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359865B6E6F918FDEE338C.taxon	distribution	Distribution. Thailand: Sutrat Thani.	en	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
