identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
972BDD17553D986DB6E6FB52FE223185.text	972BDD17553D986DB6E6FB52FE223185.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina Z. - Y. Chen 2025	<div><p>Pagodapalaina Z.-Y. Chen gen. nov.</p><p>Diplommatina in part (Group superba)—Laidlaw, 1949: 209.</p><p>Diplommatina in part (Group diminuta)—Laidlaw, 1949: 212.</p><p>Diplommatina in part (Clade D)— Webster et al., 2012: 634.</p><p>Diplommatina in part (Clade C)— Boonmachai, Bergey &amp; Nantarat, 2023: 203.</p><p>Type species. Diplommatina acme Laidlaw, 1949</p><p>Diagnosis. Shell small, sinistral, conical to very elongated conical; in some species, a keel emerges on the periphery and forms angular lower whorls. Shell surface usually covered with regular and dense radial ribs. In certain species, the ribs are modified into outwardly projecting triangular plates or coiled into tubular projections. Aperture oval, oblique to the shell axis. Peristome attached to the body whorl, double or triple; inner peristome protruding; outer peristome expanded but not reflected. Internal constriction (if present) ventrally situated with a thin radial palatal fold at its base. One short horizontal palatal lamella (if present) visible through translucent shell. Columellar tooth small. Umbilicus closed. Operculum corneous. Radula consists of a strong central tooth with a large central cusp and two pairs of smaller lateral cusps. Basal plate of central tooth with two attached small cusps. Lateral and marginal teeth with very short basal parts.</p><p>Deferential diagnosis. All known species of Pagodapalaina gen. nov., were previously assigned to Diplommatina Benson, 1849, but recent phylogenetic studies have consistently supported the monophyly of the new genus as the sister group to Diplommatina (Webster et al., 2012; Boonmachai et al., 2023; Köhler, 2024). Morphologically, the sinistrally coiled, conical shell of Pagodapalaina gen. nov. clearly distinguishes it from Diplommatina . In terms of radular structure, Pagodapalaina gen. nov. exhibits a significantly enlarged central cusp on the central tooth compared to Diplommatina, and both the lateral and marginal teeth possess extremely short basal parts, resulting in a radula that appears notably compact (see Boonmachai et al., 2023). Additionally, the basal plate of Diplommatina typically bears a single pointed cusp, whereas Pagodapalaina gen. nov., has two relatively blunt cusps (see Boonmachai et al., 2023; Z.-Y. Chen, unpublished data). This feature closely resembles that of Luzonocoptis Páll-Gergely &amp; Hunyadi, 2017, a genus from Luzon Island, Philippines, which also possesses a sinistral shell (Páll-Gergely &amp; Hunyadi, 2017). However, the cusps of Pagodapalaina gen. nov., are more robust, nearly conical in shape, and the two cusps on the basal plate of the central tooth are positioned adjacent to each other rather than being widely separated. This compact radula type may potentially be associated with saxicolous habits. From a broader comparative perspective, the keeled and elongate-conical whorls of Pagodapalaina gen. nov. readily distinguish it from the sinistral taxa occurring in adjacent regions (see Vermeulen, 1993; Greķe, 2017; Nurinsiyah &amp; Hausdorf, 2017). Although certain species of Palaina O. Semper, 1865 (e.g. Palaina attenboroughi Greķe 2017, Palaina dohertyi E. A. Smith, 1897 and Palaina parietalis Neubert &amp; Bouchet, 2015) exhibit a similar shell outline, their weak internal lamellar structure provides a clear means of separating them from Pagodapalaina gen. nov.</p><p>Etymology. The name of the new genus is derived from “Pagoda”, referring to the tiered, keeled and tower-like shape of the shell, reminiscent of East Asian pagodas, and diplommatinid genus name Palaina . This genus name is feminine.</p><p>Included species. See below.</p><p>Distribution. Thailand, Malaysia, Indonesia. The known species of the new genus are distributed on the Malay Peninsula and Sumatra.</p></div>	https://treatment.plazi.org/id/972BDD17553D986DB6E6FB52FE223185	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553E986DB6E6FC96FB0C358B.text	972BDD17553E986DB6E6FC96FB0C358B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina hortulanica Z. - Y. Chen 2025	<div><p>Pagodapalaina hortulanica Z.-Y. Chen sp. nov.</p><p>(Figure 1A)</p><p>Diplommatina hortulana Davison MS (non Diplommatina (Sinica) hortulana Leschke, 1914: 215, figs 10–11)</p><p>Type material. Holotype NHMUK 19930124, 2 paratypes NHMUK 19930125, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.11667&amp;materialsCitation.latitude=4.7666664" title="Search Plazi for locations around (long 101.11667/lat 4.7666664)">Gua Kantan</a>, Chemor, N. of Ipoh Peral, Perak, Peninsular Malaysia, 4°46’N, 101°07’E, leg. G. Davison, 20 Jan. 1991.</p><p>Etymology. We adopted Davison’s manuscript name with minor modification. The name hortulanica, meaning “garden-dwelling” in Latin, may refer to the flourishing vegetation of the type locality.</p><p>Diagnosis. The new species can be distinguished from most other species by its bearing relatively fewer whorls and a stouter (vs slender in other species) shell. Its overall shape resembles Pagodapalaina carinaspinosa (Maassen, 2002) comb. nov., P. gadutensis (Maassen, 2002) comb. nov., P. karoensis (Maassen, 2002) comb. nov., and P. krabiensis (Panha &amp; Burch, 1998) comb. nov., but differs from them in its larger size and the rib structure.</p><p>Description. Shell small, sinistral, pale with yellow hue, elongated conical, with 5.75 convex whorls; suture impressed; width of body whorl slightly smaller than that of penultimate whorl; a keel emerges after the penultimate whorl and formed angular whorls. Protoconch consisting of 1.5 whorls, somewhat matte; boundary between protoconch and teleoconch not clear. Teleoconch surface with fine and regularly arranged radial and prosocline ribs, approximately 8–9 ribs per 500 μm on the body whorl, weak and dense spiral threads situated. Aperture oval, oblique to the shell axis. Peristome attached to the body whorl, double or triple; inner peristome protruding; outer peristome expanded but not reflected. Internal constriction ventrally present with a thin radial (to the shell axis) palatal fold at base. One horizontal palatal lamella observed through translucent shell. Columellar tooth small, downward sloping. Umbilicus closed. Operculum unknown.</p><p>Measurements. Shell height: 2.4 mm, shell width: 1.3 mm (holotype); shell height: 2.2–2.3 mm, shell width: both 1.2 mm (paratypes, n = 2)</p><p>Remarks. Taken directly from Davison’s note on specimen label: “In loose dry soil on the extreme north end of big limestone outcrop, about halfway up the slope to summit. In low forest of Streblus bushes, altitude 100 m. ”</p><p>Distribution. Malaysia: Perak. This new species is known only from the type locality.</p></div>	https://treatment.plazi.org/id/972BDD17553E986DB6E6FC96FB0C358B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553F986CB6E6FA06FCB034BB.text	972BDD17553F986CB6E6FA06FCB034BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina acme (Laidlaw 1949) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina acme (Laidlaw, 1949) comb. nov.</p><p>(Figure 2A)</p><p>Diplommatina acme Laidlaw, 1949: 212, fig. 4d.</p><p>Diplommatina acme — Maassen, 2001: 28, 145.</p><p>Diplommatina cf. diminuta — Foon et al., 2017: 25, fig. 9C. misidentification.</p><p>Diplommatina laemsonensis Boonmachai, Bergey &amp; Nantarat, 2023: 198, figs 2B, 3D–F. new synonym</p><p>Type locality. Diplommatina acme: [Malaysia] “Kaki Bukit, Perlis ”; Diplommatina laemsonensis: Thailand, Satun Province, La-ngu District, Limestone Hill, 6°54’43.7”N, 99°41’59.0”E.</p><p>Type material. Holotype (NHMUK 1948.11.12.5) of Diplommatina acme, Kaki Bukit, Perlis, Malaya, M.W.F. Tweedie Esq.</p><p>Remarks. Diplommatina laemsonensis Boonmachai, Bergey &amp; Nantarat, 2023 does not differ morphologically from typical P. acme . Therefore, we considered it a synonym of the latter.</p><p>Distribution. Malaysia: Perak, Perlis; Thailand: Satun.</p></div>	https://treatment.plazi.org/id/972BDD17553F986CB6E6FA06FCB034BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FF43FD073291.text	972BDD175538986BB6E6FF43FD073291.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina attenuata (Laidlaw 1949) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina attenuata (Laidlaw, 1949) comb. nov.</p><p>(Figure 2B)</p><p>Diplommatina attenuata Laidlaw, 1949: 210, fig. 4c.</p><p>Diplommatina attenuata — Maassen, 2001: 29, 145, fig. 17.</p><p>Diplommatina lenggongensis — Foon et al., 2017: 27, fig. 10A. misidentification.</p><p>Type locality. [Malaysia] “ Gua Bama, Pahang ”</p><p>Type material. Holotype (NHMUK 1948.11.12.8), Gua Bama, Pahang, Malaya, M.W.F. Tweedie Esq.</p><p>Distribution. Malaysia: Pahang, Perak.</p></div>	https://treatment.plazi.org/id/972BDD175538986BB6E6FF43FD073291	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FDF8FD0430DD.text	972BDD175538986BB6E6FDF8FD0430DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina carinaspinosa (Maassen 2002) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina carinaspinosa (Maassen, 2002) comb. nov.</p><p>(Figure 2C)</p><p>Diplommatina carinaspinosa Maassen, 2002: 167, figs 5–7.</p><p>Type locality. [Indonesia] W. Sumatra: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.75325&amp;materialsCitation.latitude=-0.47208333" title="Search Plazi for locations around (long 100.75325/lat -0.47208333)">near entrance of Cave Gua Pangian</a>, 3 km N of village Lintau, SE of Bukittinggi, 00°28’19.5”S, 100°45’11.7”E.</p><p>Type material. 5 paratypes (NHMUK 20030224), same as type locality, vii.1996, presented by W.J.M. Maassen. The holotype (RMNH.MOL.92941) at NBC was not examined. Additional paratypes are housed in multiple repositories (see Maassen 2002 for details) .</p><p>Distribution. Indonesia: West Sumatra.</p></div>	https://treatment.plazi.org/id/972BDD175538986BB6E6FDF8FD0430DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6FC2FFDA6355E.text	972BDD175538986BB6E6FC2FFDA6355E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina diminuta (Mollendorff 1891) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina diminuta (Möllendorff, 1891) comb. nov.</p><p>(Figure 2D)</p><p>Diplommatina (Sinica) diminuta Möllendorff, 1891: 344, pl. 30, fig. 15.</p><p>Diplommatina (Sinica) diminuta — Kobelt, 1902: 462.</p><p>Diplommatina (Sinica) diminuta — Laidlaw, 1928: 37.</p><p>Diplommatina diminuta — Laidlaw, 1949: 212.</p><p>Diplommatina (Sinica) diminuta — Zilch, 1953: 29, pl. 11, fig. 170.</p><p>Diplommatina diminuta — Maassen, 2001: 30, 145.</p><p>Diplommatina parabates — Foon et al., 2017: 29, fig. 10C. misidentification.</p><p>Diplommatina sinistra — Foon et al., 2017: 29, fig. 10D. misidentification?</p><p>Type locality. [Malaysia] “ad Bukit Pondong leg. cl R. Hungerford ”</p><p>Type material. Holotype (SMF 105374/1), Malakka: Bükit Pondong (Perak), Hüngerford l., Sammlung O. v. Moellendorff.</p><p>Other material examined. NHMUK 1891.7.6.5-6. Perak! [sic], Types P.Z.S. 1891, G. B. Sowerby Esq .</p><p>Distribution. Malaysia: Perak.</p></div>	https://treatment.plazi.org/id/972BDD175538986BB6E6FC2FFDA6355E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD175538986BB6E6F9A8FD04348D.text	972BDD175538986BB6E6F9A8FD04348D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina gadutensis (Maassen 2002) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina gadutensis (Maassen, 2002) comb. nov.</p><p>(Figure 2E)</p><p>Diplommatina gadutensis Maassen, 2002: 168, figs 8–10.</p><p>Type locality. [Indonesia] W. Sumatra: Kampung Desa Gadut, limestone rocks near the village, E of Payakumbuh, 00°15’36.0”S, 100°43’58.7”E.</p><p>Type material. 2 paratypes (NHMUK 20030225), same as type locality, vii. 1997, presented by W.J.M. Maassen. The holotype (RMNH.MOL.92945) at NBC was not examined. Additional paratypes are housed in multiple repositories (see Maassen 2002 for details).</p><p>Distribution. Indonesia: West Sumatra.</p></div>	https://treatment.plazi.org/id/972BDD175538986BB6E6F9A8FD04348D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755399869B6E6F90DFDE5338C.text	972BDD1755399869B6E6F90DFDE5338C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina jirasaki (Panha, Kanchanasaka & Burch 2002) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina jirasaki (Panha, Kanchanasaka &amp; Burch, 2002) comb. nov.</p><p>(Figure 2F)</p><p>Diplommatina jirasaki Panha, Kanchanasaka &amp; Burch, 2002: 163, fig. 5.</p><p>Diplommatina jirasaki — Panha &amp; Burch, 2008: 25, fig. 26.</p><p>Type locality. Thailand, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.041664&amp;materialsCitation.latitude=7.660556" title="Search Plazi for locations around (long 100.041664/lat 7.660556)">Khaopoo Khaoya National Park</a>, Pattalung Province at 7°39’38”N, 100°2’30”E, 220 meters elevation.</p><p>Type material. Holotype (CUMZ Di 040) and six paratype shells (CUMZ Di 041), same as type locality. Additional paratypes are housed in UMMZ.</p><p>Distribution. Thailand: Phatthalung.</p></div>	https://treatment.plazi.org/id/972BDD1755399869B6E6F90DFDE5338C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FE9EFD1231B3.text	972BDD17553A9869B6E6FE9EFD1231B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina karoensis (Maassen 2002) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina karoensis (Maassen, 2002) comb. nov.</p><p>(Figure 2G)</p><p>Diplommatina karoensis Maassen, 2002: 169, figs 11–12.</p><p>Type locality. [Indonesia] N. Sumatra: near the entrance of limestone cave “Liangdehar”, near Kuta Buluh, 40 km NW of Berastagi.</p><p>Type material. 2 paratypes (NHMUK 20030226), same as type locality, vi. 1996, presented by W.J.M. Maassen. The holotype (RMNH.MOL.92940) at NBC was not examined. Additional paratypes are housed in multiple repositories (see Maassen 2002 for details).</p><p>Distribution. Indonesia: North Sumatra.</p></div>	https://treatment.plazi.org/id/972BDD17553A9869B6E6FE9EFD1231B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FCCCFDA33718.text	972BDD17553A9869B6E6FCCCFDA33718.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina krabiensis (Panha & Burch 1998) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina krabiensis (Panha &amp; Burch, 1998) comb. nov.</p><p>(Figure 2H)</p><p>Diplommatina krabiensis Panha &amp; Burch, 1998: 61, fig. 6.</p><p>Diplommatina krabiensis — Panha &amp; Burch, 2008: 26, fig. 28.</p><p>Type locality. [Thailand] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.92528&amp;materialsCitation.latitude=8.125556" title="Search Plazi for locations around (long 98.92528/lat 8.125556)">Tam Sua</a>, Krabi Province, at 8°7’32”N, 98°55’31”E, 140 meters elevation.</p><p>Type material. Holotype (CUMZ Di 031) and 3 paratypes (CUMZ Di 032), same as type locality. Additional paratypes are housed in UMMZ (255345).</p><p>Distribution. Thailand: Krabi.</p></div>	https://treatment.plazi.org/id/972BDD17553A9869B6E6FCCCFDA33718	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553A9869B6E6FB6AFDA634C5.text	972BDD17553A9869B6E6FB6AFDA634C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina lenggongensis (Tomlin 1941) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina lenggongensis (Tomlin, 1941) comb. nov.</p><p>(Figure 3A)</p><p>Diplommatina (Sinica) lenggongensis Tomlin, 1941: 319, pl.13, fig. 1.</p><p>Diplommatina lenggongensis — Laidlaw, 1949: 210, fig. 4a.</p><p>Diplommatina lenggongensis nawawii Davison, 1995: 237 . species inquirenda</p><p>Diplommatina lenggongensis lenggongensis — Maassen, 2001: 30, 145, fig. 15.</p><p>Diplommatina lenggongensis nawawii — Maassen, 2001: 30, 145.</p><p>Type locality. Diplommatina (Sinica) lenggongensis: [Malaysia] “ Lenggong, Perak ”, Diplommatina lenggongensis nawawii: [ Malaysia] limestone outcrop, now isolated as an island by the impounded waters of Temengor lake, at 5°24’N, 101°18’30”E, at the perimeter of Temengor Forest Reserve, Hulu Perak district, Perak, Peninsular Malaysia.</p><p>Type material. Syntype (NHMUK 1943.6.28.1) of Diplommatina (Sinica) lenggongensis, Lenggong, Perak, J.R. le B. Tomlin .</p><p>Remarks. Contrary to the statement in the original description, the type material of Diplommatina lenggongensis nawawii could not be located in the NHM collections. The registration number ‘931229.1’ cited therein does not correspond to any recognised NHMUK catalogue number. Due to this discrepancy, and given that the original description lacks illustrations, the identity of the species cannot be confirmed based on the limited diagnostic information. We therefore treat it here as a species inquirenda, although its generic assignment appears to be unproblematic.</p><p>Distribution. Malaysia: Perak.</p></div>	https://treatment.plazi.org/id/972BDD17553A9869B6E6FB6AFDA634C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FF43FCCD3196.text	972BDD17553B9868B6E6FF43FCCD3196.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina hidagai (Panha 1998) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina hidagai (Panha, 1998) comb. nov.</p><p>(Figure 3B)</p><p>Diplommatina hidagai Panha, 1998b: 56, fig. 4 (erroneously state as figure 3 in original publication).</p><p>Diplommatina hidagai — Panha &amp; Burch, 2008: 23, 24, fig. 29 (erroneously state as figure 24).</p><p>Diplommatina naiyanetri — Boonmachai, Bergey &amp; Nantarat, 2023: 198, fig. 2D, misidentification).</p><p>Type locality. Thailand, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.77556&amp;materialsCitation.latitude=7.566389" title="Search Plazi for locations around (long 99.77556/lat 7.566389)">Khaochong Wildlife Sanctuary</a>, Trang Province at 07°33’59”N, 099°46’32”E, 130 meters elevation.</p><p>Type material. Holotype (CUMZ Di 004) and 3 paratypes (CUMZ Di 005), same as type locality. Additional paratypes are housed in multiple repositories (see Panha 1999b for details).</p><p>Remarks. In Panha (1998), the figures of Diplommatina hidagai Panha, 1998 and Diplommatina naiyanetri Panha, 1998 do not correspond with their respective descriptions. Examination of the type material by one of the present authors (CS) confirms that the descriptions are correct, and that the original figures were mistakenly interchanged. This error subsequently led to misapplied names in several later studies (e.g. Webster et al. 2012; Boonmachai et al., 2023; Chen et al., 2023; Köhler, 2023).</p><p>Distribution. Thailand: Phatthalung, Sutrat Thani, Trang.</p></div>	https://treatment.plazi.org/id/972BDD17553B9868B6E6FF43FCCD3196	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FCE7FDA63716.text	972BDD17553B9868B6E6FCE7FDA63716.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina parabates (Laidlaw 1949) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina parabates (Laidlaw, 1949) comb. nov.</p><p>(Figure 3C)</p><p>Diplommatina parabates Laidlaw, 1949: 210, fig. 4b.</p><p>Diplommatina parabates — Maassen, 2001: 31, 145, fig. 16.</p><p>Type locality. [Malaysia] “ Lenggong, Perak ”</p><p>Type material. Holotype (NHMUK 1948.11.12.6), Lenggong, Perak, Malaya, M.W.F. Tweedie.</p><p>Distribution. Malaysia: Perak.</p></div>	https://treatment.plazi.org/id/972BDD17553B9868B6E6FCE7FDA63716	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6FB60FDEE357C.text	972BDD17553B9868B6E6FB60FDEE357C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina pongrati (Panha, Kanchanasaka & Burch 2002) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina pongrati (Panha, Kanchanasaka &amp; Burch, 2002) comb. nov.</p><p>(Figure 3D)</p><p>Diplommatina pongrati Panha, Kanchanasaka &amp; Burch, 2002: 167, fig. 6.</p><p>Diplommatina pongrati — Panha &amp; Burch, 2008: 29, fig. 31.</p><p>Type locality. Thailand, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.8425&amp;materialsCitation.latitude=8.075" title="Search Plazi for locations around (long 98.8425/lat 8.075)">Tamkuha</a>, Surathani Province at 8°4’30”N, 98°50’33”E.</p><p>Type material. Holotype (CUMZ Di 043) and 4 paratypes (CUMZ Di 044), same as type locality. Additional paratypes are housed in UMMZ.</p><p>Distribution. Thailand: Sutrat Thani.</p></div>	https://treatment.plazi.org/id/972BDD17553B9868B6E6FB60FDEE357C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD17553B9868B6E6F98DFDBA34DA.text	972BDD17553B9868B6E6F98DFDBA34DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina prakayangensis (Panha 1998) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina prakayangensis (Panha, 1998) comb. nov.</p><p>(Figure 3E)</p><p>Diplommatina prakayangensis Panha, 1998a: 41, figs 1–3a.</p><p>Diplommatina prakayangensis — Panha &amp; Burch, 2008: 30, fig. 32.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.80056&amp;materialsCitation.latitude=10.326944" title="Search Plazi for locations around (long 98.80056/lat 10.326944)">Prakayang Cave</a>, Ranong Province, Thailand, at 10°19’37”N, 98°48’02”E, 117 meters elevation.</p><p>Type material. Holotype (CUMZ Di 006) and 5 paratypes (CUMZ Di 007), same as type locality. Additional paratypes are housed in multiple repositories (see Panha 1999a for details).</p><p>Distribution. Thailand: Ranong.</p></div>	https://treatment.plazi.org/id/972BDD17553B9868B6E6F98DFDBA34DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755349866B6E6F88FFD67329A.text	972BDD1755349866B6E6F88FFD67329A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina sinistra (Tomlin 1938) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina sinistra (Tomlin, 1938) comb. nov.</p><p>(Figure 3F)</p><p>Diplommatina (Sinica) sinistra Tomlin, 1938: 74, pl. 11, fig. 4.</p><p>Diplommatina sinistra —Laidlaw, 1949: 212.</p><p>Diplommatina sinistra — Maassen, 2001: 31, 145.</p><p>Diplommatina prakaiphetensis Boonmachai, Bergey &amp; Nantarat, 2023: 199, figs 2C, 3G–I. new synonym</p><p>Type locality. Diplommatina (Sinica) sinistra: [Malaysia] “ Sungei Siput, Perak ”; Diplommatina prakaiphetensis: Thailand, Satun Province, Thung Wa District, Prakaiphet Hill, 7°00’00.1”N, 99°46’08.7”E.</p><p>Type material. Syntype (NHMUK 1938.10.25.4) of Diplommatina (Sinica) sinistra, Sungei Siput, Perak, Malaya States. J. R. le B. Tomlin Esq .</p><p>Remarks. Diplommatina prakaiphetensis Boonmachai, Bergey&amp; Nantarat,2023 does not differ morphologically from typical P. sinistra . Therefore, we consider it a synonym of the latter.</p><p>Distribution. Malaysia: Perak; Thailand: Satun.</p></div>	https://treatment.plazi.org/id/972BDD1755349866B6E6F88FFD67329A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FB0BFDA6350F.text	972BDD1755359866B6E6FB0BFDA6350F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina superba subsp. brevior (Laidlaw 1949)	<div><p>Pagodapalaina superba brevior (Laidlaw, 1949) comb. nov.</p><p>(Figure 3G)</p><p>Diplommatina superba brevior Laidlaw, 1949: 210, fig. 3e.</p><p>Diplommatina superba brevior — Maassen, 2001: 32, 145.</p><p>Type locality. [Malaysia] “ Sungei Siput, Perak ”</p><p>Type material. Holotype (NHMUK 1948.11.12.7), Sungei Siput, Perak, Malaya, M.W.F. Tweedie Esq.</p><p>Remarks. Foon et al. (2017) synonymised this subspecies with the nominate one. Based on the type material, P. s. brevior exhibits a smaller shell size and less sharply developed keels compared to the nominate subspecies. Therefore, we tentatively retain the subspecies as valid.</p><p>Distribution. Malaysia: Perak.</p></div>	https://treatment.plazi.org/id/972BDD1755359866B6E6FB0BFDA6350F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359866B6E6FDEBFDA637F9.text	972BDD1755359866B6E6FDEBFDA637F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina superba subsp. superba (Godwin-Austen & Nevill 1880) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina superba superba (Godwin-Austen &amp; Nevill, 1880) comb. nov.</p><p>(Figure 1B)</p><p>Diplommatina (Palaina) superba Godwin-Austen &amp; Nevill, 1880: 739, pl. 60, figs 5, 5a.</p><p>Diplommatina superba —Möllendorff, 1886: 313.</p><p>Diplommatina superba — Tenison-Woods, 1888: 1070.</p><p>Diplommatina (Sinica) superba — Möllendorff, 1891: 344.</p><p>Diplommatina (Sinica) superba — Kobelt, 1902: 474.</p><p>Diplommatina (Sinica) superba — Laidlaw, 1928: 37.</p><p>Diplommatina superba superba — Laidlaw, 1949: 209.</p><p>Diplommatina superba superba — Maassen, 2001: 32, 145.</p><p>Diplommatina superba superba — Foon et al., 2017: 31, fig. 11B.</p><p>Type locality. [Malaysia] “ Perak ”</p><p>Type materials. Original description states type material is in “Indian Museum, Calcutta” and was not examined in this study.</p><p>Material examined. NHMUK 1912.4.16.237, Perak ‘D66’, Beddome collection .</p><p>Distribution. Malaysia: Perak.</p></div>	https://treatment.plazi.org/id/972BDD1755359866B6E6FDEBFDA637F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
972BDD1755359865B6E6F918FDEE338C.text	972BDD1755359865B6E6F918FDEE338C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina suratensis (Panha & Burch 1998) Chen & Sutcharit & Ablett 2025	<div><p>Pagodapalaina suratensis (Panha &amp; Burch, 1998) comb. nov.</p><p>(Figure 3H)</p><p>Diplommatina suratensis Panha &amp; Burch, 1998: 57, fig. 5.</p><p>Diplommatina suratensis — Panha &amp; Burch, 2008: 33, fig. 35.</p><p>Type locality. [Thailand] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.69833&amp;materialsCitation.latitude=8.9675" title="Search Plazi for locations around (long 98.69833/lat 8.9675)">Klongsang Wildlife Santuary</a>, Surathani Province at 8°58’3”N, 98°41’54”E, 170 meters elevation .</p><p>Type material. Holotype (CUMZ Di 028) and 7 paratypes (CUMZ Di 029), same as type locality. Additional paratypes are housed in UMMZ (255344).</p><p>Distribution. Thailand: Sutrat Thani.</p></div>	https://treatment.plazi.org/id/972BDD1755359865B6E6F918FDEE338C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Zhe-Yu;Sutcharit, Chirasak;Ablett, Jonathan D.	Chen, Zhe-Yu, Sutcharit, Chirasak, Ablett, Jonathan D. (2025): Pagodapalaina, a new sinistral genus of Diplommatinidae from the Malay Peninsula and Sumatra, with description of a new species from Perak, Peninsular Malaysia (Gastropoda: Caenogastropoda: Cyclophoroidea). Zootaxa 5737 (3): 389-400, DOI: 10.11646/zootaxa.5737.3.5, URL: https://doi.org/10.11646/zootaxa.5737.3.5
