identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
13EA51FC53BF51B490D9365D553A5680.text	13EA51FC53BF51B490D9365D553A5680.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina affinis Theobald 1870	<div><p>9 Diplommatina affinis Theobald, 1870</p><p>Fig. 15 A</p><p>Diplommatina affinis Theobald, 1870: 398. Type locality: Shan States. Pfeiffer 1875: 70. Theobald 1876: 41. Godwin-Austen 1884: pl. 49, fig. 3. Godwin-Austen 1886: 181, 182. Gude 1921: 302.</p><p>Diplommatina (Eudiplommatina) affinis — Kobelt and Möllendorff 1898: 135.</p><p>Diplommatina (Diplommatina) affinis — Kobelt 1902: 424.</p><p>Type material examined.</p><p>Unknown or probably lost.</p><p>Diagnosis.</p><p>Shell dextral, ovate-turreted. Whorls 7, increasing regularly; ribs weak, slightly prominent on last whorl. Aperture ovate; columellar straight, columellaris small. Peristome double; outer peristome slightly expanded.</p><p>Distribution.</p><p>The species was known from Shan State (Theobald 1870) and was subsequently recorded from Modi Taung in the Arakan Hills, Rakhine State, Myanmar, by Godwin-Austen (1886).</p><p>Remarks.</p><p>This is one of four diplommatinid species described by Theobald (1870) for which the type series could not be located in either the NHM, London, nor the NMW, Cardiff (T. White, B. Rowson, and J. Gallichan, pers. comm.). As the type specimens could not be examined, the diagnosis provided here is based on the original description and the illustration by Godwin-Austen (1884: pl. 49, fig. 3).</p></div>	https://treatment.plazi.org/id/13EA51FC53BF51B490D9365D553A5680	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
5AB94CB38C3B51AFA45BC01E1EEF260F.text	5AB94CB38C3B51AFA45BC01E1EEF260F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina akron Panha & Burch 1998	<div><p>4 Diplommatina akron Panha &amp; Burch, 1998</p><p>Figs 1 D, 3 D, 7 B, 8</p><p>Diplommatina akron Panha &amp; Burch, 1998 [1996]: 52–54, fig. 3. Type locality: Doichiang Dao Wildlife Sanctuary, Chiangmai, Thailand. Panha and Burch 2005: 19, fig. 19. BEDO 2017: 75. Jirapatrasilp et al. 2023: 26.</p><p>Type material examined.</p><p>Thailand • Holotype. CUMZ 15398 (former Di- 022; Fig. 7 B) from Doichiang Dao Wildlife Sanctuary, Chiang Mai; Panha and Burch coll. • Paratypes. CUMZ 15399 (former Di- 023; 16 shells); same data as for holotype .</p><p>Other material.</p><p>Myanmar • CUMZ 15451 (1 shell; Figs 1 D, 8 A) and CUMZ 15452 (1 shell; Figs 3 D, 8 B) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.050446&amp;materialsCitation.latitude=20.791527" title="Search Plazi for locations around (long 97.050446/lat 20.791527)">Aye Say Tee Cave</a>, Taunggyi City, Shan State; 20°47'29.5"N, 97°03'01.6"E; P. Tongkerd and S. Panha leg.</p><p>Description.</p><p>Shell sinistral, fusiform to nearly conical, relatively large, and ivory white or transparent yellow. Shell height 4.9–5.8 mm; shell width 2.6–3.1 mm. Whorls ~ 7–8, rounded; spire conical; suture wide, deep. Protoconch ~ 1 ¾ whorl, nearly smooth, with weak wrinkles and coalescing round malleated pits. Teleoconch with delicate costae, later becoming stronger, regularly spaced radial ribs; ~ 8–9 ribs / mm on penultimate whorl towards aperture. Spiral striations very weak, restricted to early whorls (seen under SEM). Penultimate whorl slightly wider than last whorl. Constriction externally inconspicuous; internal constriction marked with distinct vertical ridge. Aperture circular, with four barriers: parietalis very strong, prominent, tall ridge; two horizontal palatalis situated beyond constriction – upper palatalis elongate with prominent ridge, and lower palatalis short, prominent ridge; columellaris distinctly prominent ridge, continuous with peristome, and slightly deflected downward toward palatal wall (Fig. 3 D). Peristome broadly expanded, double; outer peristome slightly broader than inner peristome. Lip thickened, multi-layered; parietal callus thickened, expanded. Columella angulate, with sinulus below. Umbilicus closed.</p><p>Distribution.</p><p>This species was originally described from northern Thailand and is also currently known from Shan State, Myanmar. In the present surveys, it was recorded from a limestone outcrop in Taunggyi, Shan State, at an elevation of 1,583 m (Fig. 1 A). However, its occurrence in other parts of Shan State cannot be ruled out.</p><p>Remarks.</p><p>Although this species shares a sinistral and fusiform shell shape with D. pupaeformis from Shan State, Myanmar, and D. inthanon Panha &amp; Burch, 2001 from Chiang Mai Province, Thailand, several diagnostic characters can be used to differentiate it from these congeners. For comparison, D. akron possesses a fusiform shell with strong, widely spaced radial ribs and a robust, prominent columellaris. In contrast, D. pupaeformis has an almost pupilliform shell, more narrowly spaced radial ribs, and a weaker columellaris. Additionally, D. inthanon (Fig. 7 C) can be distinguished by its fine, thin, and densely arranged radial ribs (Theobald 1870; Panha and Burch 2001, 2005).</p><p>Living animal pale white in colour, with white tentacles (Fig. 1 D).</p></div>	https://treatment.plazi.org/id/5AB94CB38C3B51AFA45BC01E1EEF260F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
08F56A4571E3525CA3784A852C0AE2F1.text	08F56A4571E3525CA3784A852C0AE2F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina ambigua Godwin-Austen 1892	<div><p>10 Diplommatina ambigua Godwin-Austen, 1892</p><p>Fig. 15 B</p><p>Diplommatina ambigua Godwin-Austen, 1892: 513. Type locality: South of Burrail Range, Munipur. Godwin-Austen 1897: 8, pl. 64, fig. 2. Gude 1921: 302.</p><p>Diplommatina (Eudiplommatina) ambigua — Kobelt and Möllendorff 1898: 135.</p><p>Diplommatina (Diplommatina) ambigua — Kobelt 1902: 424.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2124, (7 shells; Fig. 15 B) from South of Burrail Range (figured in Godwin-Austen 1897: pl. 64, fig. 2); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate fusiform, dull, whitish. Whorls ~ 7–8, well rounded; spire elevated conical; suture wide, shallow. Teleoconch: earlier whorls with weak, narrowly and equally spaced radial ribs; penultimate and last whorls with slightly wide-spaced ribs. Penultimate whorl convex, equal in width to last whorl. Aperture circular; columellaris strong, directed downward. Peristome expanded, double, multi-layered; inner peristome slightly broader than outer peristome; parietal callus expanded. Columella straight; umbilicus closed.</p><p>Distribution.</p><p>This species was previously known only from its type locality in the Burrail Range, Manipur State, India. The record from Myanmar lacks a precise locality (Gude 1921), and the presence there has not yet been reconfirmed.</p></div>	https://treatment.plazi.org/id/08F56A4571E3525CA3784A852C0AE2F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
502D1E16EBB855D9906A1BAC163D796D.text	502D1E16EBB855D9906A1BAC163D796D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina angulata Theobald & Stoliczka 1872	<div><p>11 Diplommatina angulata Theobald &amp; Stoliczka, 1872</p><p>Fig. 15 C – E</p><p>Diplommatina angulata Theobald &amp; Stoliczka, 1872: 331, pl. 11, fig. 3. Type locality: Prope Moulmain, provincia Martaban. Hanley and Theobald 1875: 55, pl. 140, fig. 7. Pfeiffer 1875: 78. Nevill 1878: 285. Godwin-Austen 1884: pl. 49, figs 5, 5 a. Godwin-Austen 1886: 184. Gude 1921: 302, 303.</p><p>Palaiana angulata — Theobald 1876: 43.</p><p>Diplommatina (Eudiplommatina) angulata — Kobelt and Möllendorff 1898: 135.</p><p>Diplommatina (Diplommatina) angulata — Kobelt 1902: 424, 425.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1888.12.4.232 –241 (10 shells; Fig. 15 C, D) from Chouktalon, Moulmein; Theobald coll.</p><p>Other material.</p><p>Myanmar • NHMUK 20240015 (3 shells; Fig. 15 E) from Moulmein, Burma; Sykes coll. • NHMUK 20240016 (1 shell) from Burma; Moulmein; Salisbury coll. • NHMUK 20240017 (4 shells) from India; MacAndrew coll.</p><p>Diagnosis.</p><p>Shell dextral, conical-fusiform, translucent, whitish to yellowish. Whorls ~ 5–6, angular; spire conical; suture wide, deep. Teleoconch with wide, equally spaced radial ribs throughout. Penultimate whorl well rounded, wider than last whorl. Aperture rounded; columellaris strong, distinct, directed downward. Peristome expanded, double, multi-layered. Umbilicus narrowly open.</p><p>Distribution.</p><p>The species is currently known from its type locality, corresponding to present-day Mawlamyine District in Mon State, Myanmar.</p><p>Remarks.</p><p>Diplommatina angulata Stanisic, 2010, described from Queensland, Australia, is a junior primary homonym of D. angulata Theobald &amp; Stoliczka, 1872, from Myanmar (Theobald and Stoliczka 1872; Stanisic et al. 2010; MolluscaBase 2025). The two nominal taxa are clearly distinct in shell morphology and distributional range (Stanisic et al. 2010: 74, 75 and text figure). The younger name is invalid under the ICZN (1999: Arts 57.2, 60.3). As no available junior synonym exists, a replacement name is required to maintain nomenclatural stability. We therefore propose Diplommatina stanisici Tongkerd, nom. nov., as the new substitute name for the Australian species. The holotype of D. stanisici nom. nov. is the holotype of the replaced name (QM MO 50138), and is deposited in the Queensland Museum, Australia.</p></div>	https://treatment.plazi.org/id/502D1E16EBB855D9906A1BAC163D796D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
5EE88336451E5472A7A7F72128CABD30.text	5EE88336451E5472A7A7F72128CABD30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina animula Godwin-Austen 1892	<div><p>12 Diplommatina animula Godwin-Austen, 1892</p><p>Fig. 15 F</p><p>Diplommatina animula Godwin-Austin, 1892: 516, 517. Type locality: Prowi, Lahupa Naga Hills, Munipur. Godwin-Austin 1897: 12, 13 pl. 66, fig. 2. Gude 1921: 345.</p><p>Diplommatina (Sinica) animula — Kobelt and Möllendorff 1898: 139. Kobelt 1902: 455.</p><p>Type material examined.</p><p>India • Syntype. NHMUK 1903.7.1.2164 (1 shell; Fig. 15 F) from Prowi, Lahupa, Naga Hills (figured in Godwin-Austen 1897: pl. 66, fig. 2); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, whitish. Whorls ~ 5, rounded; spire depressed conical; suture wide, deep. Teleoconch with densely placed, equally spaced, strong radial ribs throughout. Last three whorls nearly equal in width; penultimate whorl slightly wider than others. Aperture rounded; columellaris weak, indistinct. Peristome expanded, double, multi-layered; outer peristome expanded beyond inner peristome. Columella straight, with sinulus below. Umbilicus narrowly open.</p><p>Distribution.</p><p>This species was previously known only from its type locality in Prowi, Lahupa-Naga Hills, Manipur State, India. The record from Myanmar does not provide a precise locality (Gude 1921).</p></div>	https://treatment.plazi.org/id/5EE88336451E5472A7A7F72128CABD30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
61FF5028ABD9591CACAB25539F119C3E.text	61FF5028ABD9591CACAB25539F119C3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina Benson 1849	<div><p>Genus Diplommatina Benson, 1849</p><p>Diplommatina Benson, 1849: 193, 194. Godwin-Austen 1886: 166 –169. Kobelt 1902: 423. Gude 1921: 301. Thiele 1929: 110, 111. Vermeulen 1993: 5. Egorov 2013: 17.</p><p>Type species.</p><p>Bulimus folliculus Pfeiffer, 1846, by subsequent designation of Nevill (1878: 284).</p><p>Remarks.</p><p>To date, more than 420 species and subspecies have been documented within the genus (Kobelt 1902; MolluscaBase 2025). Phylogenetic evidence indicates that the genus is polyphyletic (Rundell 2008; Webster et al. 2012; Köhler 2023). Consequently, several previously recognised subgenera have been elevated to generic rank, including Moussonia Semper, 1865 and Gastroptychia Kobelt &amp; Möllendorff, 1900 (Neubert and Bouchet 2015; Chen and Ablett 2024). In addition, several new genera have been proposed, such as Pagodapalaina Chen, 2025, Sohtsuia Chen, 2023, and Sinoarinia Chen &amp; Wu, 2020 (Chen and Wu 2020; Chen et al. 2023, 2025).</p><p>To date, 26 species of Diplommatina s. l. have been reported from Myanmar. Of these, 15 species have type localities within Myanmar, whereas the remaining ten species were originally described from neighbouring countries but were listed as occurring in ‘ Burma’ or ‘ Burmah’ (Gude 1921; Theobald and Stoliczka 1872). In addition, three species, Pagodapalaina suratensis (Panha &amp; Burch, 1998), D. akron Panha &amp; Burch, 1998, and D. nimanandhi Panha, Kanchanasaka &amp; Burch, 2002, previously known only from Thailand, are recorded from Myanmar for the first time in the present study.</p></div>	https://treatment.plazi.org/id/61FF5028ABD9591CACAB25539F119C3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
E20579E0838657CAB75E5919B7781B2F.text	E20579E0838657CAB75E5919B7781B2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina butleri Godwin-Austen 1892	<div><p>13 Diplommatina butleri Godwin-Austen, 1892</p><p>Fig. 15 G</p><p>Diplommatina butleri Godwin-Austin, 1892: 512, 513. Type locality: Laisen Peak, Munipur. Godwin-Austin 1897: 6, 7, pl. 64, fig. 9. Gude 1921: 305, 306. McGhie 2008: 9.</p><p>Diplommatina (Eudiplommatina) butleri — Kobelt and Möllendorff 1898: 135.</p><p>Diplommatina (Diplommatina) butleri — Kobelt 1902: 427.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2128 (6 shells; Fig. 15 G) from Laisen Peak (Trig. Station), Munipur (figured in Godwin-Austen 1897: pl. 64, fig. 9); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, pale brownish. Whorls ~ 7–8, well rounded; spire elevated conical; suture wide, shallow. Teleoconch: earlier whorls with weak, widely, and equally spaced radial ribs; penultimate whorl and last whorl generally smooth. Penultimate whorl convex, equal in width to last whorl. Aperture circular; columellaris very strong, directed downward. Peristome expanded, double, multi-layered; outer peristome slightly broader than inner peristome. Columella angulate, with distinct sinulus below. Umbilicus closed.</p><p>Distribution.</p><p>This species was originally known from its type locality at Laiven Peak, Manipur State, India. It has also been recorded from several other localities, including the Lahupa-Naga Hills, Kezakenomih, and Klang Sing in the Naga Hills, India (Godwin-Austen 1892; Gude 1921). The record from Myanmar, however, lacks a precise locality and is cited only as ‘ Kyengdwen, Burmah’, which likely refers to the Chindwin River valley in Sagaing Region, Myanmar.</p><p>Remarks.</p><p>Three syntypes are also housed in the Manchester Museum, The University of Manchester (EE. 3511 / 109) (McGhie 2008). This species closely resembles D. ambigua in shell form but differs in having widely and regularly spaced radial ribs on the earlier whorls, and the penultimate and last whorls are generally smooth. In contrast, D. ambigua (Fig. 15 B) possesses narrowly and regularly spaced radial ribs on the earlier whorls, and the penultimate and last whorls bear widely spaced ribs.</p></div>	https://treatment.plazi.org/id/E20579E0838657CAB75E5919B7781B2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
1984A84854805010BA509F2C8319F54D.text	1984A84854805010BA509F2C8319F54D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina carneola Stoliczka 1871	<div><p>2 Diplommatina carneola Stoliczka, 1871</p><p>Figs 3 B, 4, 5</p><p>Diplommatina carneola Stoliczka, 1871: 152, 153, pl. 6, fig. 3. Type locality: Damotha, prope Moulmein. Pfeiffer 1875: 76, 77. Hanley and Theobald 1876: 55, pl. 140, fig. 4. Theobald 1876: 42. Godwin-Austen 1884: pl. 49, figs 8, 8 a. Godwin-Austen 1886: 182, 183. Gude 1921: 346.</p><p>Diplommatina carreola [sic] — Nevill 1878: 284.</p><p>Diplommatina (Sinica) carneola — Kobelt and Möllendorff 1898: 140. Kobelt 1902: 459.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1871.9.23.33 (2 shells; Fig. 4 A) from Moulmein, Tenasserim, Burma; Damon ex. Stoliczka coll. • Syntype. NHMUK 1903.7.1.2279 (1 shell; Fig. 4 C) from Moulmein, Tenasserim, Burma; Stoliczka coll.</p><p>Other material.</p><p>Myanmar • NHMUK 1888.12.4.171 –174 (4 shells; Fig. 4 B) from Damathat, Moulmein; Theobald coll. • NHMUK 1903.7.1.2199 (1 shell; Fig. 4 D); from Moulmein, Burma; Godwin-Austen coll. • NHMUK 20240011 (1 shell); from Moulmein, Burma; Sykes coll. • NHMUK 20240012 (1 shell); from Moulmein, Burma; Salisbury coll. • CUMZ 15445 (2 shells; Figs 4 E, 4 F, 5 A), CUMZ 15446 (5 shells; Figs 3 B, 5 B) and CUMZ 15447 (72 shells) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.81009&amp;materialsCitation.latitude=16.50639" title="Search Plazi for locations around (long 97.81009/lat 16.50639)">Damathat Cave</a>, Mawlamyine City, Mon State; 16°30'23.0"N, 97°48'36.3"E; P. Tongkerd and S. Panha leg.</p><p>Description.</p><p>Shell dextral, ovoid-fusiform, pale yellowish orange to yellowish. Shell height 2.0– 2.3 mm; shell width 1.3–1.4 mm. Whorls 5–6, well rounded; spire depressed conical; suture wide and deep. Protoconch ~ 1 ¾ whorls, with wrinkles and roundish malleated pits arranged in radial rows. Teleoconch with strong, regularly spaced radial ribs; ~ 5–6 ribs / 0.5 mm on both penultimate and last whorls before aperture. Last whorl slightly narrower than penultimate whorl. Spiral striations very fine and weak, and appearing throughout. Penultimate whorl wider than last whorl. Constriction externally inconspicuous; internal constriction with distinct vertical ridge. Aperture circular, with three barriers: parietalis weak, low ridge; palatalis short, slightly prominent, and situated close to internal constriction ridge; columellaris prominent, continuous with peristome, and slightly deflected downward toward palatal wall (Fig. 3 B). Peristome expanded, double; outer peristome slightly broader than inner peristome. Lip thickened, multi-layered; parietal callus thick or thin and weakly expanded. Columella short and curved. Umbilicus closed.</p><p>Distribution.</p><p>The species is currently known from its type locality ‘ Damotha, near Moulmein’, which corresponds to the modern locality name Damathat Cave (16°30'21.8"N, 97°49'11.5"E), Mawlamyine, Mon State, Myanmar (Fig. 1 A – C).</p><p>Remarks.</p><p>Among the Diplommatina species occurring in Mon and Kayin states, Myanmar, D. carneola is similar to D. polypleuris Benson, 1857 in having ovoid-fusiform shell shape (Kobelt 1902) but D. carneola presents very closely arranged transverse costae.</p></div>	https://treatment.plazi.org/id/1984A84854805010BA509F2C8319F54D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
E8ED2E64FA4C544699CEA6850A5889AF.text	E8ED2E64FA4C544699CEA6850A5889AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina commutata Godwin-Austen 1892	<div><p>14 Diplommatina commutata Godwin-Austen, 1892</p><p>Fig. 16 B</p><p>Diplommatina commutata Godwin-Austen, 1892: 513. Type locality: Prowi, Lahupa Naga Hills. Godwin-Austen 1897: 8, pl. 64, fig. 4. Gude 1921: 306, 307.</p><p>Diplommatina (Eudiplommatina) commutata — Kobelt and Möllendorff 1898: 136.</p><p>Diplommatina (Diplommatina) commutata — Kobelt 1902: 428.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2126 (7 shells; Fig. 16 B) from Prowi, Lahupa, Naga Hills (figured in Godwin-Austen 1897: pl. 64, fig. 4); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, pale brownish. Whorls ~ 7, well rounded; spire elevated-conical; suture wide, shallow. Teleoconch: earlier whorls with weak, narrowly and equally spaced radial ribs; penultimate and last whorls with very weak radial ribs to nearly smooth surface. Penultimate whorl convex, slightly wider than last whorl. Aperture circular; columellaris very strong, sharp, directed downward. Peristome expanded, double, multi-layered; outer peristome slightly broader than inner peristome. Columella curved; umbilicus closed.</p><p>Distribution.</p><p>This species was originally known from its type locality at Prowi, Lahupa-Naga Hills, Manipur State, India, and has also been recorded from Tellizo Peak, Anghami – Naga Hills, Manipur State, India. The record from Myanmar, however, lacks an exact locality (Gude 1921), and the presence there has not yet been reconfirmed.</p></div>	https://treatment.plazi.org/id/E8ED2E64FA4C544699CEA6850A5889AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
DA36122FAD245A8D85E2DD61A256335B.text	DA36122FAD245A8D85E2DD61A256335B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina compacta Godwin-Austen 1892	<div><p>15 Diplommatina compacta Godwin-Austen, 1892</p><p>Fig. 16 A</p><p>Diplommatina compacta Godwin-Austen, 1892: 515, 516. Type locality: South of Barak in Munipur. Godwin-Austen 1897: 11, 12, pl. 65, figs 7, 7 a, 7 b. Gude 1921: 340. McGhie 2008: 11.</p><p>Diplommatina (Metadiancta) compacta — Kobelt and Möllendorff 1898: 138. Kobelt 1902: 449.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2179 (28 shells; Fig. 16 A) from South of Barak, Munipur (figured in Godwin-Austen 1897: pl. 65, figs 7, 7 a); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, whitish. Whorls ~ 7, well rounded; spire elevated-conical; suture wide, shallow. Teleoconch: earlier whorls with weak, narrowly and equally spaced radial ribs; penultimate and last whorls with very weak radial ribs to nearly smooth surface. Penultimate whorl convex, wider than last whorl. Aperture circular; columellaris very strong, directed downward. Peristome expanded, double, multi-layered; outer peristome slightly broader than inner peristome; parietal callus thickened, expanded. Columella curved; umbilicus closed.</p><p>Distribution.</p><p>This species is known only from its type locality in Barak, Manipur State, India. The record from Myanmar, however, lacks a precise locality (Gude 1921), and the presence there has not yet been reconfirmed.</p><p>Remarks.</p><p>There are four additional syntype shells housed in The Manchester Museum, University of Manchester (EE. 3511 / 99) (McGhie 2008).</p></div>	https://treatment.plazi.org/id/DA36122FAD245A8D85E2DD61A256335B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
819FF82594A3508BB1B77EDEB2E86905.text	819FF82594A3508BB1B77EDEB2E86905.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina crispata Stoliczka 1871	<div><p>16 Diplommatina crispata Stoliczka, 1871</p><p>Fig. 16 D</p><p>Diplommatina (Palaina) crispata Stoliczka, 1871: 153, pl. 6, fig. 4. Type locality: Damotha, prope Moulmein. Pfeiffer 1875: 91, 92. Nevill 1878: 289. Gude 1921: 308, 309.</p><p>Palaina crispata — Theobald 1876: 43.</p><p>Diplommatina crispata — Hanley and Theobald 1876: 56, pl. 141, fig. 6. Godwin-Austen 1884: pl. 49, figs 4, 4 a, 4 b. Godwin-Austen 1886: 183.</p><p>Diplommatina (Eudiplommatina) crispata — Kobelt and Möllendorff 1898: 136.</p><p>Diplommatina (Diplommatina) crispata — Kobelt 1902: 429.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1903.7.1.2200 (2 shells; Fig. 16 D) from Moulmein, Tenasserim (figured in Godwin-Austen 1884: pl. 49, figs 4, 4 a); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, elongate-fusiform, dull, and pale brownish to pale yellowish. Whorls ~ 6–7; earlier whorls rounded-angular then becoming stronger on last two whorls; spire elongate-conical; suture wide, deep. Teleoconch with equally spaced radial ribs throughout: earlier whorls regular, strong radial ribs; last two whorls with strong, tall, sinuous radial ribs at angulation. Spiral striations fine, conspicuously present on last two whorls. Penultimate whorl much wider than last whorl; last whorl much displaced to right relative to peristome position. Aperture rounded; columellaris distinct, directed downward. Peristome expanded, double; outer peristome more broadly expanded than inner peristome, multi-layered; inner peristome slightly expanded, elevated. Umbilicus very narrow.</p><p>Distribution.</p><p>The species is currently known only from its type locality, ‘ Damotha, near Moulmein’, which corresponds to the modern Damathat Cave (16°30'21.8"N, 97°49'11.5"E) in Mawlamyine, Mon State, Myanmar.</p><p>Remarks.</p><p>Diplommatina angulifera Bavay &amp; Dautzenberg, 1912 from Vietnam and D. angulifera umpangensis Panha, 1998 from Thailand exhibit shell shapes and sculptures similar to those of D. crispata, and they are likely conspecific. Additionally, D. crispata khaochamaoensis Panha, Kanchanasaka &amp; Burch, 2002 from Chanthaburi Province, Thailand which is located approximately 600 km southeast of the type locality, differs slightly from the nominotypical subspecies. It possesses a less prominent angulation on the last whorl, whereas the nominotypical subspecies exhibits a more pronounced and sharply defined peripheral keel on the penultimate whorl that expands to nearly the same extent as that of the last whorl.</p></div>	https://treatment.plazi.org/id/819FF82594A3508BB1B77EDEB2E86905	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
54AC760F35645A7FB0889C61419ADE48.text	54AC760F35645A7FB0889C61419ADE48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina edentula Godwin-Austen 1886	<div><p>17 Diplommatina edentula Godwin-Austen, 1886</p><p>Fig. 16 C</p><p>Diplommatina edentula Godwin-Austen, 1884: pl. 49, figs 7, 7 a. [nomen nudum]</p><p>Diplommatina edentula Godwin-Austen, 1886: 185. Type locality: Moulmain [Mawlamyine, Myanmar]. Gude 1921: 312.</p><p>Diplommatina (Eudiplommatina) edentula — Kobelt and Möllendorff 1898: 136.</p><p>Diplommatina (Diplommatina) edentula — Kobelt 1902: 431.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1903.7.1.2201 (2 shells; Fig. 16 C) from Moulmein, Tenasserim (figured in Godwin-Austen 1884: pl. 49, figs 7, 7 a); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell tiny, dextral, pupate-fusiform, translucent, pale yellowish to whitish. Whorls ~ 6, slightly shouldered to rounded; spire ovate-conical; suture deep. Teleoconch with equally spaced, strong radial ribs throughout. Spiral striations thin and appearing throughout. Last three whorls nearly equal in width; penultimate whorl slightly less inflated than others. Aperture ovate; columellaris weak, not prominent. Peristome expanded; lip slightly thickened, multi-layered. Umbilicus closed.</p><p>Distribution.</p><p>This species was previously known only from its type locality in Mawlamyine, Mon State, Myanmar (Godwin-Austen 1886; Gude 1921).</p><p>Remarks.</p><p>The name edentula was first introduced by Godwin-Austen (1884) in ‘ Land and Freshwater Mollusca of India’; however, it was published without a description, definition, or indication, as only shell illustrations and the collection locality were provided. Accordingly, the name does not meet the criteria of availability under the ICZN (1999: Art. 12.2) and was therefore unavailable in Godwin-Austen (1884). Subsequently, in the same series, Godwin-Austen (1886) published a full description of the taxon under the same name, thereby rendering edentula an available name from that publication.</p></div>	https://treatment.plazi.org/id/54AC760F35645A7FB0889C61419ADE48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
ADDB7F95B9B95F4FB44DFFEB31C1059D.text	ADDB7F95B9B95F4FB44DFFEB31C1059D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina exilis Blanford 1863	<div><p>18 Diplommatina exilis Blanford, 1863</p><p>Figs 16 E, 16 F, 17 A, 17 B</p><p>Diplommatina exilis Blanford, 1863: 325. Type locality: Mya Leit Douang, Ava. Pfeiffer 1865: 10. Godwin-Austen 1868: 84, pl. 4, figs 3, 3 a. Theobald and Stoliczka 1872: 331. Hanley and Theobald 1875: 49, pl. 119, fig. 10. Pfeiffer 1875: 77. Theobald 1876: 42. Nevill 1878: 284. Godwin-Austen 1884: pl. 49, fig. 1. Godwin-Austen 1886: 180. Gude 1921: 313.</p><p>Diplommatina (Eudiplommatina) exilis — Kobelt and Möllendorff 1898: 136.</p><p>Diplommatina (Diplommatina) exilis — Kobelt 1902: 432.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1906.4.4.16 (6 shells; Fig. 16 F) from Mya Leit Doung, Ava; W. T. Blanford coll. • Syntype. NHMUK 1903.7.1.2207 (1 shell; Fig. 16 E) from Mya Leit Doung, Ava (figured in Godwin-Austen 1884: pl. 49, fig. 1); Godwin-Austen coll.</p><p>Other material.</p><p>Myanmar • NHMUK 1888.12.4.163 –166 (4 shells; Fig. 17 A) from Myaleit Doung, Ava; Theobald coll. • NHMUK 1903.7.1.2277 (1 shell; Fig. 17 B) from Ava, Burma; Godwin-Austen coll. • NHMUK 20240018 (2 shells) from Ava; Sykes coll. • NHMUK 20240019 (4 shells) from India (?); Cuming coll.</p><p>Diagnosis.</p><p>Shell dextral, elongate-conical fusiform, dull to translucent, whitish to yellowish. Whorls ~ 7–8, well rounded; spire elongate-conical; suture wide, deep. Teleoconch with equally spaced, weak to strong radial ribs throughout. Penultimate whorl wider than preceding and last whorls. Aperture rounded; columellaris strong, distinct, directed downward. Peristome expanded, double; lip thickened, multi-layered. Umbilicus closed.</p><p>Distribution.</p><p>The species is currently known only from its type locality, ‘ Mya Leit Douang, Ava’, which likely refers to the Myaleit Mountains in the Mandalay Region, Myanmar.</p></div>	https://treatment.plazi.org/id/ADDB7F95B9B95F4FB44DFFEB31C1059D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
123EA453004A5BCEB5BC9D74CEEEE193.text	123EA453004A5BCEB5BC9D74CEEEE193.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina exserta Godwin-Austen 1886	<div><p>19 Diplommatina exserta Godwin-Austen, 1886</p><p>Fig. 17 C</p><p>Diplommatina exilis var. exserta (nom. nud.) Nevill, 1878: 284. [nomen nudum]</p><p>Diplommatina exserta Godwin-Austen, 1884: pl. 49, figs 2, 2 a. [nomen nudum]</p><p>Diplommatina exserta Godwin-Austen, 1886: 184, 185. Type locality: Damotha Cave, Moulmein. Gude 1921: 314. Budha et al. 2017: 8, figs 3 b, 9 b.</p><p>Diplommatina (Eudiplommatina) exserta — Kobelt and Möllendorff 1898: 136.</p><p>Diplommatina (Diplommatina) exserta — Kobelt 1902: 432. Budha et al. 2015: 6.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1903.7.1.2208 (4 shells; Fig. 17 C) from Damathat, Moulmein (figured in Godwin-Austen 1884: pl. 49, figs 2, 2 a); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, turreted-fusiform, translucent, whitish. Whorls ~ 7–8, well rounded; spire elongate-conical; suture wide, deep. Teleoconch with slightly strong, densely and equally spaced radial ribs throughout. Spiral striations very thin and appearing throughout. Penultimate whorl slightly wider than preceding and last whorls. Aperture rounded, little tilted from columellar axis; columellaris very weak. Peristome expanded, double; basal edge slightly sinuous, protruding; outer peristome slightly broader than inner peristome, multi-layered. Umbilicus closed.</p><p>Distribution.</p><p>The species is currently known from its type locality in Mon State and was subsequently recorded from Nepal, approximately 1900 km from the type locality (Gude 1921; Budha et al. 2015, 2017).</p><p>Remarks.</p><p>This taxon name was first mentioned by Godwin-Austen (1884) solely in a figure caption, accompanied only by the collection locality. This treatment lacked a description, definition, and indication, and thus did not meet the requirements for availability under the ICZN (1999: Art. 12). Godwin-Austen (1886) subsequently published a full description of the taxon, where the name exserta became available.</p><p>The specimens from Nepal differ slightly from the syntype, having a somewhat narrower penultimate whorl at the lip, a strong columellar tooth, and a more inflated penultimate whorl compared to the preceding whorl. In contrast, the syntype shows a very weak columellar tooth, a narrower penultimate whorl at the lip, and a penultimate whorl nearly equal in swelling to the preceding whorl. These differences suggest that the specimens from Nepal may belong to a distinct species.</p></div>	https://treatment.plazi.org/id/123EA453004A5BCEB5BC9D74CEEEE193	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
0D2577298F275BCF90A3AFF42FCBA9B0.text	0D2577298F275BCF90A3AFF42FCBA9B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina henzadaensis Godwin-Austen 1886	<div><p>20 Diplommatina henzadaensis Godwin-Austen, 1886</p><p>Fig. 17 D, E</p><p>Diplommatina henzadaensis Godwin-Austen, 1884: pl. 46, figs 6, 6 a. [nomen nudum]</p><p>Diplommatina henzadaensis Godwin-Austen, 1886: 179, 180. Type locality: Kyoung Gyoung Nulla, Henzada, Pegu. Gude 1921: 318.</p><p>Diplommatina (Eudiplommatina) henzadaensis — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) henzadaensis — Kobelt 1902: 434, 435.</p><p>Type material examined.</p><p>Myanmar • Holotype. NHMUK 1906.4.4.22 (Fig. 17 D) from Khyoung Gyoung-nulla, Henzada; W. T. Blanford coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-turreted. Six whorls, rounded; antepenultimate whorl largest; sculptured with fine, widely spaced radial ribs. Spire convex; apex blunt; suture well marked. Aperture circular. Peristome double, continuous, thickened; columellaris well-developed.</p><p>Distribution.</p><p>This species is currently known only from its type locality, ‘ Kyoung Gyoung Nulla, Henzada, Pegu’, which likely corresponds to the modern Kyaunggon Township, Pathein District, Ayeyarwady Region, Myanmar.</p><p>Remarks.</p><p>This taxon was first mentioned by Godwin-Austen (1884) without a formal description, definition, or indication; only shell illustrations and the collection locality were provided. Consequently, the name does not meet the criteria for availability under the ICZN (1999: Art. 12.2) and is therefore unavailable. Godwin-Austen (1886) subsequently published a complete description of the taxon under the same name, thereby making ‘ henzadaensis ’ available. In the original description, Godwin-Austen (1886) clearly stated that only a single specimen existed in the Blanford collection. The holotype, which is affected by Byne’s disease and consequently fragile, is illustrated here for the first time.</p></div>	https://treatment.plazi.org/id/0D2577298F275BCF90A3AFF42FCBA9B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
CD057A9EAF515C69AE8AE66A78D260B5.text	CD057A9EAF515C69AE8AE66A78D260B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina lapillus Godwin-Austen 1892	<div><p>21 Diplommatina lapillus Godwin-Austen, 1892</p><p>Fig. 17 F</p><p>Diplommatina lapillus Godwin-Austen, 1892: 515. Type locality: Kopamedza Peak, Lahupa Naga Hills, 8375 ft. Godwin-Austen 1897: 11, pl. 65, figs 6, 6 a. Gude 1921: 342.</p><p>Diplommatina (Metadiancta) lapillus — Kobelt and Möllendorff 1898: 139. Kobelt 1902: 450.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2151 (2 shells; Fig. 17 F) from Kopamedza Peak, Naga Hills (figured in Godwin-Austen 1897: pl. 65, figs 6, 6 a); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, dull, whitish. Whorls ~ 7, well rounded; spire elevated-conical; suture wide, shallow. Teleoconch with very weak, narrowly and equally spaced radial ribs throughout. Penultimate whorl convex, equal in width to last whorl. Aperture circular; columellaris weak, small, directed downward. Peristome broadly expanded, double, multi-layered; outer peristome slightly broader than inner peristome; parietal callus thickened, expanded, nearly attached to suture. Columella straight; umbilicus closed.</p><p>Distribution.</p><p>This species is known from its type locality at Kopamedza Peak, Lahupa – Naga Hills, Manipur State, India, as well as from locations north of the Barail Range within the same state. Records from Myanmar lack precise locality data (Gude 1921), and the presence there has not yet been reconfirmed.</p></div>	https://treatment.plazi.org/id/CD057A9EAF515C69AE8AE66A78D260B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
A8BCFBF9A50B5AF0912631B8A62F6CCB.text	A8BCFBF9A50B5AF0912631B8A62F6CCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina munipurensis Godwin-Austen 1892	<div><p>22 Diplommatina munipurensis Godwin-Austen, 1892</p><p>Fig. 17 G</p><p>Diplommatina munipurensis Godwin-Austen, 1892: 518. Type locality: South of Barak River, between the Mao villages and Munipur. Godwin-Austen 1897: 14, pl. 66, fig. 6. Gude 1921: 323, 324. Budha et al. 2017: 13, 15, figs 5 b, 9 h. McGhie 2008: 24.</p><p>Diplommatina (Eudiplommatina) munipurensis — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) munipurensis — Kobelt 1902: 438. Budha et al. 2015: 7.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2165 (10 shells; Fig. 17 G) from South Barak, Munipur (figured in Godwin-Austen 1897: pl. 66, fig. 6); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, elongate, pupate-fusiform, translucent, pale yellowish. Whorls ~ 6–7, rounded; spire elongate-conical; suture wide, deep. Teleoconch with weak, irregularly spaced radial ribs throughout. Penultimate whorl equal to or slightly wider than last whorl. Aperture ovate; columellaris very weak, indistinct. Peristome expanded, multi-layered; lip slightly thickened; parietal callus thin, expanded. Columella thickened, straight; umbilicus closed.</p><p>Distribution.</p><p>This species was originally known only from its type locality in Manipur State, India. It was later reported from several localities in Bagmati and Sudurpashchim provinces, Nepal (Budha et al. 2015, 2017). Records from Myanmar, however, lack precise locality data (Gude 1921), and the presence there has not yet been reconfirmed.</p><p>Remarks.</p><p>There are two syntypes housed in The Manchester Museum, University of Manchester (EE. 3511 / 87) (McGhie 2008).</p></div>	https://treatment.plazi.org/id/A8BCFBF9A50B5AF0912631B8A62F6CCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
8EC55CF31F605D498D870942A97B47DB.text	8EC55CF31F605D498D870942A97B47DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina nana Blanford 1865	<div><p>23 Diplommatina nana Blanford, 1865</p><p>Fig. 18 A – E</p><p>Diplommatina nana Blanford, 1865: 85. Type locality: Akoutoung, Thondoung and Yenandoung in Henzada district, Pegu. Godwin-Austen 1868: 84, pl. 4, figs 4, 4 a. Theobald and Stoliczka 1872: 331. Hanley and Theobald 1875: 55, pl. 140, fig. 1. Pfeiffer 1875: 75. Theobald 1876: 42. Nevill, 1878: 285. Godwin-Austen 1884: pl. 49, figs 6, 6 a. Godwin-Austen 1886: 181. Gude 1921: 324.</p><p>Diplommatina (Eudiplommatina) nana — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) nana — Kobelt 1902: 439.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1888.12.4.217 –219 (3 shells; Fig. 18 D, E) from Akouktoung, Pegu; Theobald coll. • Syntypes. NHMUK 1906.4.4.18 a (6 shells; Fig. 18 B) from Akouktoung; W. T. Blanford coll. • Syntypes. NHMUK 1906.4.4.18 (6 shells; Fig. 18 C) from Thondoung, near Mya; W. T. Blanford coll.</p><p>Other material.</p><p>Myanmar • NHMUK 1903.7.1.2203 (1 shell; Fig. 18 A) from Burma (figured in Godwin-Austen 1884: pl. 49, figs 6, 6 a); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, pupate-fusiform, translucent, and pale yellowish to whitish. Whorls ~ 6, rounded; spire elongate-conical; suture wide, deep. Teleoconch with weak, irregularly spaced radial ribs. Last three whorls nearly equal in width; penultimate whorl slightly wider than others. Aperture ovate; columellaris strong, distinct, directed downward. Peristome double, expanded, multi-layered; inner peristome broadly expanded over outer peristome; lip little thickened, multi-layered. Umbilicus narrowly open.</p><p>Distribution.</p><p>The species is currently known from its type locality near the border of the present-day Bago and Ayeyarwady regions. The Akauk Taung Hill (18°30'33.7"N, 95°06'48.5"E) is located in Pyay District, Bago Region, while Thondoung and Yenangdoung likely correspond to the area of Yae Nan Taung Village, south of Akauk Taung Hill, in Hinthada District, Ayeyarwady Region.</p><p>In addition, the species has been reported from several other localities in Myanmar: Henzada (Hinthada City, Ayeyarwady Region), Thayet Myo (Thayet City, Thayet District, Magway Region), Pegu (Bago Region), Tongoop, Arakan (Taungup Township, Thandwe District, Rakhine State), and near Moulmein (Theobald and Stoliczka 1872; Nevill 1878; Gude 1921). It has also been reported from Meghalaya State, India (Ramakrishna et al. 2010).</p><p>Remarks.</p><p>In the original description, Blanford (1865: 85) did not indicate the number of type specimen lots. However, the stated collection locality suggests that the type series comprised at least three lots from distinct collecting sites: ‘ Akoutoung’ (= Akauk Taung), Pyay District, Bago Region; ‘ Thondoung’, a hill south of Akauk Taung, Hinthada District, Ayeyarwady Region; and ‘ Yenandoung’ (= Yae Nan Taung), Hinthada District, Ayeyarwady Region. The NHM collection holds three lots of this species, all originating from W. T. Blanford’s collections: NHMUK 1888.12.4.217 –219 from Akauk Taung, Pegu; NHMUK 1906.4.4.18 a from Akauk Taung; and NHMUK 1906.4.4.18 from Thondoung, near Mya. These three lots constitute the type series and are herein recognised as syntypes. Representative specimens from each syntype lot are also illustrated in this study.</p></div>	https://treatment.plazi.org/id/8EC55CF31F605D498D870942A97B47DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
599B3DC0F67E59FEAA75A0EAF8C92159.text	599B3DC0F67E59FEAA75A0EAF8C92159.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina nimanandhi Panha, Kanchanasaka & Burch 2002	<div><p>5 Diplommatina nimanandhi Panha, Kanchanasaka &amp; Burch, 2002</p><p>Figs 9, 10, 11 A</p><p>Diplommatina nimanandhi Panha, Kanchanasaka &amp; Burch, 2002 [1998]: 159–163, fig. 4. Type locality: Banpangcam Cave, Mae Hongson Province, Thailand. Panha and Burch 2005: 28, fig. 30. BEDO 2017: 81. Jirapatrasilp et al. 2023: 26.</p><p>Type material examined.</p><p>Thailand • Holotype. CUMZ 15379 (former Di- 037; Fig. 9 A) ex. from Banpangcam Cave, Mae Hongson Province; Panha coll. • Paratypes. CUMZ 15380 (former Di- 029; 14 shells); same data as for holotype .</p><p>Other material.</p><p>Myanmar • CUMZ 15453 (4 shells; Figs 9 B, 9 C, 10 A), CUMZ 15454 (5 shells; Figs 10 B, 11 A) and CUMZ 15455 (69 shells + 8 juveniles) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.81009&amp;materialsCitation.latitude=16.50639" title="Search Plazi for locations around (long 97.81009/lat 16.50639)">Damathat Cave</a>, Mawlamyine City, Mon State; 16°30'23.0"N, 97°48'36.3"E; P. Tongkerd and S. Panha leg.</p><p>Description.</p><p>Shell dextral, conical fusiform, whitish to yellowish. Shell height 2.7–2.9 mm; shell width 0.8–1.3 mm. Whorls ~ 6–7, rounded; spire conical; suture wide and deep. Protoconch ~ 1 ¾ whorl, with prominent round malleated pits throughout. Teleoconch with strong, delicate radial ribs: about first two whorls’ ribs regularly low then about last three whorls ribs gradually becoming more widely spaced, more prominent, sinuous, and very tall with trough-like shape at angulation, thinner near suture. Ribs with 2–3 ribs / 0.5 mm on penultimate whorl; 3–4 ribs / 0.5 mm on last whorl towards aperture. Spiral striations very fine, prominent, and appearing from teleoconch to aperture. Last whorl slightly narrower than penultimate whorl. Constriction externally inconspicuous; internal constriction marked with vertical ridge. Aperture circular, with two barriers: parietalis weak, low ridge to knob-like; palatalis absent; columellaris distinctly prominent ridge, continuous with peristome, slightly deflected downward toward palatal wall (Fig. 11 A). Peristome broadly expanded; outer peristome broader than inner peristome. Lip thickened, multi-layered; parietal callus thickened and expanded. Columella straight and thickened. Umbilicus closed.</p><p>Distribution.</p><p>Previously, this species was known only from its type locality in Mae Hong Son Province, Thailand (Panha et al. 2002). In the present study, despite surveys at multiple localities, the species was found exclusively in Damathat Cave, Myanmar (Fig. 1 A).</p><p>Remarks.</p><p>Among the four species D. angulata Theobald &amp; Stoliczka, 1872, D. carneola, D. crispata Stoliczka, 1871, and D. exserta Godwin-Austen, 1886, which were described from Damathat Cave, D. nimanandhi is readily distinguishable by its highly prominent, trough-shaped radial ribs. The most similar species is D. crispata, which exhibits angular whorls and dense, sinuous radial ribs (Kobelt 1902), whereas D. nimanandhi possesses more widely spaced, trough-shaped radial ribs.</p></div>	https://treatment.plazi.org/id/599B3DC0F67E59FEAA75A0EAF8C92159	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
29C0E79C52905241A607ECD8408BA111.text	29C0E79C52905241A607ECD8408BA111.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina oligopleuris Blanford 1868	<div><p>24 Diplommatina oligopleuris Blanford, 1868</p><p>Fig. 18 F, G</p><p>Diplommatina oligopleuris Blanford, 1868: 82, pl. 3, fig. 4. Type locality: Teria Ghat ad latus meridionale montium Khasi. Hanley and Theobald 1875: 49, pl. 119, figs 2, 3. Pfeiffer 1875: 74. Theobald 1876: 42. Nevill 1878: 285. Godwin-Austen 1884: pl. 50, fig. 1. Gude 1921: 325, 326.</p><p>Diplommatina (Eudiplommatinu) oligopleuris — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina olygopleuris [sic] — Theobald and Stoliczka 1872: 330, 331.</p><p>Diplommatina (Diplommatina) oligopleuris — Kobelt 1902: 440.</p><p>Type material examined.</p><p>India • Syntype. NHMUK 1903.7.1.2273 (1 shell; Fig. 18 F) from Teria Ghat, Khasi; Godwin-Austen coll.</p><p>Other material.</p><p>• NHMUK 1903.7.1.2197 (1 shell; Fig. 18 G) from Dikrang Valley, Dafla Hills; Godwin-Austen coll. NHMUK 1903.7.1.2274 (7 shells) from Moyong, N. W. Khasi; Godwin-Austen coll. NHMUK 1903.7.1.2275 (9 shells) from Rywuk, Garo Hills; Godwin-Austen coll. NHMUK 1903.7.1.273 (37 shells) from Khasi Hills; Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, pale yellowish to whitish. Whorls ~ 5–6, well rounded; spire depressed-conical; suture wide, deep. Teleoconch with weak, widely and equally spaced radial ribs throughout. Penultimate whorl well rounded, much wider than preceding and last whorls. Aperture circular; columellaris weak or strong, directed downward. Peristome expanded, double, multi-layered; outer peristome slightly broader than inner peristome. Umbilicus closed.</p><p>Distribution.</p><p>This species was originally discovered from Teria Ghat, Khasi Hills and later reported from Kumah Hill and Baom in Rakhine State, Myanmar (Theobald and Stoliczka 1872: 330).</p></div>	https://treatment.plazi.org/id/29C0E79C52905241A607ECD8408BA111	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
64984CA9EF1D50D7AE57A9D5249DDC56.text	64984CA9EF1D50D7AE57A9D5249DDC56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina polypleuris Benson 1857	<div><p>25 Diplommatina polypleuris Benson, 1857</p><p>Fig. 19 A, B</p><p>Diplommatina polypleuris Benson, 1857: 203. Type locality: ad Nancla. Theobald 1858: 318; Pfeiffer, 1858: 11. Godwin-Austen 1868: 83, pl. 3, fig. 1. Theobald and Stoliczka 1872: 330. Hanley and Theobald 1875: 56, pl. 140, fig. 10. Pfeiffer 1875: 73. Theobald 1876: 42. Nevill 1878: 285. Godwin-Austen 1884: pl. 45, figs 1, 1 a. Godwin-Austen 1886: 176. Gude 1921: 328.</p><p>Diplommatina polypleuris var. Godwin-Austen, 1870: 4, pl. 1, figs 4, 4 a. Type locality: North Cachar and north Jaintia hills [mountainous region in Assam and Meghalaya state, India].</p><p>Diplommatina polypleuris var. minuta Godwin-Austen, 1876: 178 (nomen nudum).</p><p>Diplommatina polypleuris var. minuta Godwin-Austen, 1886: 176 [in the name usage list].</p><p>Diplommatina (Eudiplommatina) polypleuris — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) polypleuris — Kobelt 1902: 441. Preece et al. 2022: 86, fig. 34 e.</p><p>Type material examined.</p><p>• Syntype. UMZC I.102640 (1 shell), no locality data; R. McAndrew coll.</p><p>Other material.</p><p>• NHMUK 1903.7.1.2314 (2 shells) from Jawai, Jaintia; Godwin-Austen coll. NHMUK 1903.7.1.2177 (1 shell) from Khasi (figured in Godwin-Austen 1884: pl. 45, figs 1, 1 a); Godwin-Austen coll. NHMUK 1903.7.1.2269 (24 shells) from Shillong, Khasi Hills; Godwin-Austen coll. NHMUK 1905.5.5.49 (12 shells) from North Scarp, Khasi Hills; W. T. Blanford coll. UMZC I.102645 (1 shell), no locality data; R. McAndrew coll. NHMUK 1891.3.17.728 –730 (3 shells; Fig. 19 B) labelled as D. polypleuris var. from Nattoung; Sowerby coll. NHMUK 1903.7.1.2252 (5 shells; Fig. 19 A) labelled as D. polypleuris var. minuta from Nongtung, Khasi; Godwin-Austen coll. NHMUK 1903.7.1.2262 (1 shell) labelled as D. polypleuris var. minuta from Japvo Peak, Naga; Godwin-Austen coll. NHMUK 1903.7.1.263 (1 shell) labelled as D. polypleuris var. minuta from Munken Valley, Jaintia; Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, whitish to yellowish. Whorls ~ 5, rounded; spire depressed conical; suture wide, deep. Teleoconch with strong, densely and equally spaced radial ribs throughout. Last three whorls nearly equal in width; penultimate whorl slightly wider than others. Aperture rounded; columellaris weak, indistinct. Peristome expanded, double, multi-layered; outer peristome expanded beyond inner peristome. Umbilicus narrowly open.</p><p>Distribution.</p><p>This species was originally known from its type locality in Nanclai Poonjee, Assam, India, and was later recorded from the Dafla Hills, North Khasi Hills, and Darjeeling (Hanley and Theobald 1870; Gude 1921; Preece et al. 2022). In Myanmar, it has been reported from Nattoung, Sandoway, which likely corresponds to the Thandwe District area in Rakhine State (Theobald and Stoliczka 1872; Preece et al. 2022).</p><p>Remarks.</p><p>Godwin-Austen (1870: 4) recognised specimens from ‘ North Cachar and North Jaintia Hills’ as a distinct variety, but did not provide a name for it. Subsequently, he introduced the name var. minuta for specimens from ‘ Shengorh and Torúpútú Peaks’ (Godwin-Austen 1876: 178). However, this nomination lacks a definition, indication, description, and any reference to earlier work; therefore, this varietal name was not made available (ICZN 1999: Art. 12). Later, in the list of name usages under D. polypleuris, Godwin-Austen (1886: 176) stated ‘ from N. Jaintia is distinct; I have named it minuta ’, with an explicit indication referring to the published description in Godwin-Austen (1870). It seems that the name var. minuta was made available from Godwin-Austen (1886). Regarding the type series, the NHM holds one lot, NHMUK 1903.7.1.2252 (4 shells), registered as syntypes of D. polypleuris minuta . However, the recorded locality is from ‘ Nongtung, Khasi’, which does not match the locality ‘ North Jaintia Hills’ associated with the original description. Therefore, this specimen lot is unlikely to be the syntype, since it neither originates from the type locality nor appears to be part of the type series examined by Godwin-Austen.</p><p>The specimen figured in Preece et al. (2022: fig. 34 e), labelled as D. polypleuris, appears to have prominent radial ribs and a distinct horizontal palatalis, whereas the specimens from Nattoung (Fig. 19 B), referred to as D. polypleuris var., tend to have weaker radial ribs and a conspicuous vertical plica. These differences suggest that the two are unlikely to represent the same taxon.</p></div>	https://treatment.plazi.org/id/64984CA9EF1D50D7AE57A9D5249DDC56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
918F5E998F6C5051AD3CD8F3AB0156C8.text	918F5E998F6C5051AD3CD8F3AB0156C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina prolixa Tongkerd 2026	<div><p>6 Diplommatina prolixa Tongkerd sp. nov.</p><p>Figs 11 B, 12 A</p><p>Type material.</p><p>Myanmar • Holotype. CUMZ 15458 (height 2.2 mm, width 1.2 mm; Figs 11 B, 12 A) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.570694&amp;materialsCitation.latitude=16.835056" title="Search Plazi for locations around (long 97.570694/lat 16.835056)">Rathye Pyan Cave</a>, Hpa An City, Kayin State; 16°50'6.2"N, 97°34'14.5"E, 15 m a. s. l.; P. Tongkerd and S. Panha leg. • Paratype. CUMZ 25459 (1 damaged shell); same data as for holotype .</p><p>Diagnosis.</p><p>Shell small, dextral, elongate-pyramidal, fusiform, ~ 6 ¼ convex whorls. Penultimate whorl widest. Shell with broadly spaced radial ribs, fine spiral striations throughout. Peristome double, expanded. Aperture rounded with three barriers: parietalis, palatalis and columellaris.</p><p>Description.</p><p>Shell dextral, elongate-pyramidal fusiform, whisper white or transparent yellow, apex orange. Shell height 2.0 and 2.2 mm; shell width 1.1 and 1.2 mm. Whorls 6 ¼, and broadly convex; spire elongate conical; suture wide and deep. Protoconch ~ 1 ½ whorls, with round malleated pits. Teleoconch with wide, regularly spaced radial ribs; ribs slightly sinuous, prominent near suture. Ribs with 3–4 ribs / 0.5 mm on penultimate whorl, 4–5 ribs / 0.5 mm on last whorl towards aperture. Spiral striations very weak, fine on only earlier whorls. Last whorl narrower than penultimate whorl. Constriction externally weakly conspicuous; internal constriction marked with distinct vertical ridge. Aperture circular with three barriers: parietalis very strong, prominent ridge; palatalis moderately long, strong, horizontal ridge; columellaris distinctly prominent, tall ridge, continuous with peristome, and slightly deflected downward toward palatal wall (Fig. 11 B). Peristome broadly expanded, double; outer peristome broader than inner peristome. Lip thickened, multi-layered; parietal callus thick, expanded. Columella straight. Umbilicus closed.</p><p>Differential diagnosis.</p><p>Diplommatina prolixa sp. nov. is most similar in shell sculpture to D. oligopleuris Blanford, 1868 from India and D. sperata Blanford, 1862 from Rakhine State, Myanmar. However, it differs from D. oligopleuris in having an elongate fusiform shell, whereas D. oligopleuris has an ovate-fusiform shell. In addition, the new species can be distinguished from D. sperata by its more widely spaced radial ribs and a peristome edge that lacks protrusion, while D. sperata has more narrowly spaced radial ribs and a peristome with a protruding columellar edge.</p><p>For further comparison, D. prolixa sp. nov. can be distinguished from D. naiyanetri Panha, 1998 from southern Thailand by its wider spacing between the radial ribs, a rounded aperture, and without a pointed columellar edge. In contrast, D. naiyanetri possesses more narrowly spaced radial ribs, somewhat squarish aperture, and a clearly defined columellar notch. In addition, D. naiyanetri (Fig. 12 B) was recorded from a locality approximately 1,040 km south of the type locality of this new species.</p><p>Distribution.</p><p>Diplommatina prolixa sp. nov. is currently known only from its type locality in Kayin State, Myanmar. This new species inhabits limestone cliff faces and rock crevices, with surrounding vegetation (Fig. 1 A).</p><p>Etymology.</p><p>The specific name prolixus is a Latin word meaning wide or broad and alludes to broadly spaced radial ribs, which characterises this species.</p><p>Remarks.</p><p>The shell of D. prolixa sp. nov. is comparable to that of D. exilis from Mandalay Region, Myanmar (Kobelt 1902). However, the radial ribs of D. exilis are narrower than those of D. prolixa sp. nov.</p></div>	https://treatment.plazi.org/id/918F5E998F6C5051AD3CD8F3AB0156C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
BBD32EDF393E5876ACC2F2EE970D5C3D.text	BBD32EDF393E5876ACC2F2EE970D5C3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina pupaeformis Theobald 1870	<div><p>26 Diplommatina pupaeformis Theobald, 1870</p><p>Fig. 19 C – F</p><p>Diplommatina pupaeformis Theobald, 1870: 398. Type locality: Shan States. Pfeiffer 1875: 84, 85. Theobald 1876: 42. Godwin-Austen 1884: pl. 46, figs 4, 4 a. Godwin-Austen 1886: 182. Gude 1921: 329.</p><p>Diplommatina salwiniana var. pupaeformis — Nevill 1878: 285.</p><p>Diplommatina (Eudiplommatina) pupaeformis — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) pupaeformis — Kobelt 1902: 442.</p><p>Type material examined.</p><p>Myanmar • Possible syntypes. NMW 1894.015. 00102 (3 shells; Fig. 19 D – F) from Shan State; Fedden coll.</p><p>Other material.</p><p>Myanmar • NMW 1955.158. 27985 (1 shell; Fig. 19 C) from Upper Salwin, Shan State Godwin-Austen ex. Melvill-Tomlin coll.</p><p>Diagnosis.</p><p>Shell sinistral, fusiform to almost pupilliform, whitish or transparent yellow. Whorls ~ 7 ½ and slightly convex; spire conical; suture wide, deep. Protoconch ~ 1 ¾ whorl, nearly smooth, without radial ribs or with weak wrinkles and coalescing round malleated pits. Teleoconch with delicate costae then becoming strong, regularly spaced radial ribs towards aperture. No spiral striation. Penultimate whorl slightly wider than last whorl. Aperture rounded; columellaris distinct, slightly directed downward to palatal wall. Peristome broadly expanded; lip thickened, multi-layered. Umbilicus sealed.</p><p>Distribution.</p><p>This species is currently known only from Shan State and is likely endemic to the region.</p><p>Remarks.</p><p>This is the only one of the four diplommatinid species described by Theobald (1870) for which the type series could be located (T. White, B. Rowson, and J. Gallichan, pers. comm.). In the introduction, Theobald (1870: 398) stated that the specimens were obtained from Mr. Fedden, who had collected them after returning from the Upper Salwin. The NMW, Cardiff, holds a lot from the Fedden collection containing three shells, and with the collection locality labelled ‘ Shan States, Burmah’, which are considered syntypes.</p></div>	https://treatment.plazi.org/id/BBD32EDF393E5876ACC2F2EE970D5C3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
00AFCD96C0BE55B9A047AA85D1EC7DC8.text	00AFCD96C0BE55B9A047AA85D1EC7DC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina puppensis Blanford 1863	<div><p>27 Diplommatina puppensis Blanford, 1863</p><p>Fig. 20 A, B</p><p>Diplommatina puppensis Blanford, 1863: 324, 325, Type locality: Puppa Hill in Upper Burma. Godwin-Austen 1868: 84, pl. 4, figs 2, 2 a. Hanley and Theobald 1875: 55, pl. 139, figs 8, 9. Pfeiffer 1875: 75, 76. Theobald 1876: 42. Nevill 1878: 284. Godwin-Austen 1884: pl. 49, fig. 9. Godwin-Austen 1886: 180, 181. Gude 1921: 329, 330.</p><p>Diplommatina pappensis [sic.] — Blanford 1864: 443.</p><p>Diplommatina (Eudiplommatina) puppensis — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) puppensis — Kobelt 1902: 442.</p><p>Type material examined.</p><p>Myanmar • Syntypes. NHMUK 1906.4.4.15 (4 shells; Fig. 20 A) from Puppa Hill, extinct volcano, Upper Burma; W. T. Blanford coll. • Syntype. NHMUK 1903.7.1.2278 (1 shell; Fig. 20 B) from Puppa Hill, extinct volcano, Upper Burma (figured in Godwin-Austen 1884: pl. 49, fig. 9); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, conical-fusiform, translucent, whitish to pale yellowish. Whorls ~ 6–7, rounded; spire elongate-conical; suture wide, deep. Teleoconch slightly thin, with closely and equally spaced radial ribs throughout. Penultimate whorl slightly wider than last whorl. Aperture rounded; columellaris distinct, directed downward. Peristome expanded, double, multi-layered; basal edge slightly sinuous, protruded; inner peristome broadly expanded on parietal side. Umbilicus closed.</p><p>Distribution.</p><p>This species was originally known only from its type locality, ‘ Puppa Hill, Ava’, which likely corresponds to present-day Mount Popa (20°55'N, 95°14'E) in central Mandalay Region.</p></div>	https://treatment.plazi.org/id/00AFCD96C0BE55B9A047AA85D1EC7DC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
73C70A59278F5FCDAAE33578A255CEBF.text	73C70A59278F5FCDAAE33578A255CEBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina richthofeni Theobald & Stoliczka 1872	<div><p>3 Diplommatina richthofeni Theobald &amp; Stoliczka, 1872</p><p>Figs 3 C, 6, 7 A</p><p>Diplommatina richthofeni Theobald &amp; Stoliczka, 1872: 331, 332, pl. 11, fig. 4. Type locality: Prope Moulmain [Mawlamyine, Myanmar]. Pfeiffer 1875: 78, 79. Hanley and Theobald 1876: 56, pl. 141, figs 7, 8. Theobald 1876: 42. Nevill 1878: 285. Gude 1921: 331.</p><p>Diplommatina (Eudiplommatina) richthofeni — Kobelt and Möllendorff 1898: 137.</p><p>Diplommatina (Diplommatina) richthofeni — Kobelt 1902: 443.</p><p>Type material examined.</p><p>Myanmar • Holotype. NHMUK 1888.12.4.162 (Fig. 7 A) from Moulmein; Theobald coll.</p><p>Other material.</p><p>Myanmar • CUMZ 15448 (1 shell; Fig. 6 A), CUMZ 15449 (1 shell; Figs 3 C, 6 B) and CUMZ 15450 (85 shells) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.70638&amp;materialsCitation.latitude=16.316221" title="Search Plazi for locations around (long 97.70638/lat 16.316221)">Naga Mauk Cave</a>, Mawlamyine City, Mon State; 16°18'58.4"N, 97°42'23.0"E; P. Tongkerd and S. Panha leg.</p><p>Description.</p><p>Shell small, dextral, spindle-shaped, pale whitish to pale yellowish. Shell height 2.1–2.5 mm; shell width 0.8–1.1 mm. Whorls ~ 6, slightly angular-rounded; spire conical; suture wide and deep. Protoconch ~ 1 ¾ whorl, with prominent round malleated pits throughout. Teleoconch with strong, irregularly spaced, and sinuous radial ribs; ~ 10–11 ribs / 0.5 mm on penultimate whorl, ~ 8–9 ribs / 0.5 mm on last whorl towards aperture. Spiral striations very fine, appearing throughout from teleoconch to last whorl near aperture. Penultimate whorl slightly wider than last whorl. Constriction externally inconspicuous; internal constriction marked with low vertical ridge. Aperture circular, with two barriers: parietalis weak, low ridge; palatalis absent; columellaris prominent ridge, continuous with peristome, and slightly deflected downward toward palatal wall (Fig. 3 C). Peristome expanded; outer peristome broader than inner peristome. Lip multi-layered; parietal callus thin, expanded. Columella short and curved. Umbilicus sealed.</p><p>Distribution.</p><p>This species is currently known only from Naga Mauk Cave, Mawlamyine City, Mon State, Myanmar (Fig. 1 A). The original locality was recorded as ‘ Moulmain’.</p><p>Remarks.</p><p>Theobald and Stoliczka (1872) clearly stated in the original description that only a single specimen was obtained from Farm Caves.</p></div>	https://treatment.plazi.org/id/73C70A59278F5FCDAAE33578A255CEBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
E9B2814194A35A949E2871E380F936F4.text	E9B2814194A35A949E2871E380F936F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina salwiniana Theobald 1870	<div><p>28 Diplommatina salwiniana Theobald, 1870</p><p>Diplommatina salwiniana Theobald, 1870: 398. Type locality: Shan States. Pfeiffer 1875: 84. Nevill 1878: 285. Gude 1921: 332.</p><p>Diplommatina salewiniana [sic.] — Theobald 1876: 42.</p><p>Diplommatina (Eudiplommatina) salwiniana — Kobelt and Möllendorff 1898: 138.</p><p>Diplommatina (Diplommatina) salwiniana — Kobelt 1902: 444.</p><p>Type material examined.</p><p>Unknown.</p><p>Diagnosis.</p><p>Shell sinistral, ovate-turreted, pale yellowish; suture impressed. Whorls 7 ½, convex, increasing regularly; radial ribs distinct, more widely spaced on last whorl. Aperture rounded-oval; columella straight; columellaris distinct.</p><p>Distribution.</p><p>The species is currently known only from its type locality in Shan State, Myanmar.</p><p>Remarks.</p><p>Among the four diplommatinid species described by Theobald (1870), the status of this species remains uncertain. The original description lacked an illustration, the type specimens could not be located (T. White, B. Rowson, and J. Gallichan, pers. comm.), and no subsequent illustrations have been published. The diagnosis provided here is based on the original description and the account by Gude (1921).</p></div>	https://treatment.plazi.org/id/E9B2814194A35A949E2871E380F936F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
7188197ACAB55B3ABAEBBE8D1940B550.text	7188197ACAB55B3ABAEBBE8D1940B550.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina scalaroidea Theobald 1870	<div><p>1 Diplommatina scalaroidea Theobald, 1870</p><p>Figs 2, 3 A</p><p>Diplommatina scalaroidea Theobald, 1870: 399, pl. 18, fig. 5. Type locality: Mandalay, regno Burmanico. Pfeiffer 1875: 80. Hanley and Theobald 1876: 56, pl. 141, fig. 10. Theobald 1876: 42. Nevill 1878: 285. Gude 1921: 333.</p><p>Diplommatina (Eudiplommatina) scalaroidea — Kobelt and Möllendorff 1898: 138.</p><p>Diplommatina (Diplommatina) scalaroidea — Kobelt 1902: 445.</p><p>Type material examined.</p><p>The type series could not be located in the Natural History Museum, London, nor the National Museum of Wales, Cardiff (T. White, B. Rowson, and J. Gallichan, pers. comm.).</p><p>Other material.</p><p>Myanmar • CUMZ 15442 (2 shells; Fig. 2 A), CUMZ 15443 (1 shell; Figs 2 B, 3 A) and CUMZ 15444 (12 shells + 3 juveniles) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.40344&amp;materialsCitation.latitude=20.846529" title="Search Plazi for locations around (long 96.40344/lat 20.846529)">Pyiyaung city</a>, Mandalay Region; 20°50'47.5"N, 96°24'12.4"E; P. Tongkerd and S. Panha leg.</p><p>Description.</p><p>Shell sinistral, ovoid-fusiform, pale yellowish to whitish colour. Shell height 4.6–5.2 mm; shell width 2.3–2.6 mm. Whorls ~ 7–8, convex; spire conical; suture wide and deep. Protoconch ~ 1 ¾ whorl, with fine growth lines, wrinkles, and roundish malleated pits. Teleoconch with strong, nearly regularly spaced radial ribs; penultimate whorl with widest spacing ribs, ~ 3–4 ribs / mm; last whorl before aperture with ~ 5–6 ribs / mm. Spiral striations very thin, fine and appear from teleoconch to about fourth whorl. Penultimate whorl wider than last whorl. Constriction externally inconspicuous; internal constriction marked by distinct vertical ridge. Aperture circular, with four barriers: parietalis very strong and prominent ridge; two horizontal palatalis situated beyond constriction – upper palatalis elongate with prominent ridge, lower palatalis short and strong; columellaris distinctly prominent ridge, continuous with peristome, and slightly deflected downward toward palatal wall (Fig. 3 A). Peristome broadly expanded, double; outer peristome broader than inner peristome. Lip thickened, multi-layered; parietal callus thin, expanded. Columellar angulate, with distinct sinulus below. Umbilicus closed.</p><p>Distribution.</p><p>This species was originally described from the Mandalay Region. During our surveys across multiple localities in the Mandalay Region and Shan State, the species was found exclusively on limestone karsts near Pyiyaung City, suggesting that its distribution is likely restricted to the Mandalay Region (Fig. 1 A).</p><p>Remarks.</p><p>Theobald (1870: 398) described four Diplommatina species, namely D. affinis Theobald, 1870, D. pupaeformis Theobald, 1870, D. salwiniana Theobald, 1870 and D. scalaroidea, consecutively in the same publication but without clearly indicating the number of specimens examined, illustrations, or specifying the type localities. However, in the introduction, Theobald (1870) mentioned that the specimens were obtained from Mr. Fedden, who had collected them after returning from the Upper Salwin. The type series of these four species could not be located in the NHM, London, nor the NMW, Cardiff collections, except for D. pupaeformis (see under D. pupaeformis; T. White, B. Rowson, and J. Gallichan, pers. comm.).</p><p>Although the type specimen of D. scalaroidea is not available, the diagnostic characters given in the original description and the type locality correspond well with the specimens examined in this study. This species is readily distinguished from all other congeners by its sinistral shell coiling, whereas all other species possess dextral shells. The most closely related species in Myanmar is D. exilis Blanford, 1863 from Ava, which differs in having a dextral shell and more narrowly spaced radial ribs (Hanley and Theobald 1876).</p></div>	https://treatment.plazi.org/id/7188197ACAB55B3ABAEBBE8D1940B550	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
7FD3F5F37ADF5BCA8B3041AD9596C857.text	7FD3F5F37ADF5BCA8B3041AD9596C857.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina somsakpanhai Tongkerd 2026	<div><p>7 Diplommatina somsakpanhai Tongkerd sp. nov.</p><p>Figs 11 C, 13</p><p>Type material.</p><p>Myanmar • Holotype. CUMZ 15460 (height 1.4 mm, width 2.2 mm; Fig. 13 A) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.17619&amp;materialsCitation.latitude=11.2295" title="Search Plazi for locations around (long 99.17619/lat 11.2295)">Buddha Cave</a>, Lenya City, Tanintharyi Region; 11°13'46.2"N, 99°10'34.3"E, 65 m a. s. l.; C. Sutcharit, R. Chanabun and R. Srisonchai leg. • Paratypes. CUMZ 15461 (6 shells; Figs 11 C, 13 B) and CUMZ 15462 (30 shells + 20 juveniles); same data as for holotype . • NHMUK 20260057 (5 shells); same data as for holotype . • SMF (5 shells); same data as for holotype .</p><p>Diagnosis.</p><p>Shell small, sinistral, ovoid fusiform, ~ 5 ½ rounded whorls. Last whorl nearly same diameter as penultimate whorl. Shell with narrowly spaced radial ribs, fine spiral striations throughout. Peristome thickened, double, broadly expanded. Aperture rounded, with three barriers: parietalis, palatalis and columellaris.</p><p>Description.</p><p>Shell sinistral, ovoid-fusiform, pale orange or light yellowish. Shell height 1.3–1.5 mm; shell width 2.1–2.2 mm. Whorls ~ 5 ½, rounded; spire depressed conic; suture wide, deep. Protoconch ~ 1 ¾ whorls, nearly smooth to very weakly wrinkled or with roundish pits. Teleoconch with regular and narrowly spaced radial ribs; ribs spaced slightly wider near aperture; 10–11 ribs / 0.5 mm on penultimate whorl, 7–8 ribs / 0.5 mm on last whorl towards aperture. Spiral striations very fine, weakly appearing throughout. Last whorl slightly narrower than penultimate whorl. Constriction externally inconspicuous; internal constriction marked with indistinct vertical ridge. Aperture circular with three barriers: parietalis and palatalis weak, low ridges; columellaris distinctly prominent ridge, continuous with peristome, slightly deflected downward toward palatal wall (Fig. 11 C). Peristome double, broadly expanded, and nearly or completely reaches suture; inner peristome broader than outer peristome. Lip thickened, multi-layered; parietal callus thickened, broadly expanded. Columella thickened and curved. Umbilicus sealed.</p><p>Differential diagnosis.</p><p>Diplommatina somsakpanhai sp. nov. can be distinguished from D. siriphumi Panha &amp; Burch, 2001 from Chiang Mai Province, Thailand, by several diagnostic characters. The new species possesses more ovate whorls, a narrow space between radial ribs, a broadly expanded peristomal lip, and a large, distinct columellaris. In contrast, D. siriphumi (Fig. 7 D) has a less ovate whorl, a wide space between radial ribs, a narrow peristomal lip, and a small, less prominent columellaris. In addition, D. siriphumi was recorded from a non-limestone habitat located approximately 700 km north of the type locality of this new species (Panha and Burch 2001, 2005).</p><p>Furthermore, the new species differs from Pagodapalaina krabiensis (Panha &amp; Burch, 1998) from southern Thailand by its ovate-fusiform shell, ovate whorls, narrowly spaced radial ribs, and broadly expanded peristomal lip. In comparison, P. krabiensis possesses a fusiform shell with shouldered whorls, widely spaced radial ribs, and a narrowly expanded peristomal lip (Panha and Burch 1998, 2005).</p><p>Distribution.</p><p>Diplommatina somsakpanhai sp. nov. is currently known only from its type locality in the Tanintharyi Region, Myanmar. It inhabits limestone cliff faces and rock crevices, with surrounding vegetation (Fig. 1 A).</p><p>Etymology.</p><p>The specific name somsakpanhai is derived from the full name of Professor Dr. Somsak Panha of Chulalongkorn University, in recognition of his remarkable contributions to the study of Southeast Asian land molluscs and biodiversity research in Thailand. This new species is named in his honour to celebrate the anniversary of his sixty-fifth birthday.</p><p>Remarks.</p><p>This new species is superficially similar to the type species of Gastroptychia, G. adversa (Adams &amp; Adams, 1851), in having a sinistral, ovate-fusiform shell and a strongly expanded lip that is nearly attached to the suture (Adams and Adams 1851; Vermeulen 1993: 36, fig. 42). Although Gastroptychia has recently been resurrected as a valid genus, its diagnostic characters and distribution remain intermixed with the highly diverse genus Diplommatina, and the characters supporting its separation are still provisional (Egorov 2013; Chen and Ablett 2024). Current recognition of Gastroptychia is mainly based on molecular phylogenetic evidence (Webster et al. 2012), but this analysis did not include the type species. Therefore, in the absence of clear generic diagnostic characters, we provisionally place this new species in the widespread genus Diplommatina, pending a clearer definition of the diagnostic characteristics of Gastroptychia .</p><p>There are very few sinistral Diplommatina species that have been recorded from Myanmar, only five of 31 known species that occur in Myanmar, namely D. salwiniana, D. scalaroidea, D. akron, P. suratensis, and the new species described herein.</p></div>	https://treatment.plazi.org/id/7FD3F5F37ADF5BCA8B3041AD9596C857	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
DC7DF65F026E5A029F5D3B1DF6C8D9F7.text	DC7DF65F026E5A029F5D3B1DF6C8D9F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina sperata Blanford 1862	<div><p>29 Diplommatina sperata Blanford, 1862</p><p>Fig. 20 C</p><p>Diplommatina sperata Blanford, 1862: 143, 144. Type locality: in montibus Arakan a Pegu secernentibus. Pfeiffer 1865: 10. Godwin-Austen 1868: 84, pl. 4, figs 1, 1 a. Theobald and Stoliczka 1872: 330. Theobald 1876: 42. Nevill 1878: 284. Godwin-Austen 1884: pl. 46, figs 5, 5 a. Godwin-Austen 1886: 179. Gude 1921: 335, 336. Budha et al. 2017: 22, figs 8 b, 9 n.</p><p>Diplommatina (Eudiplommatina) sperata — Kobelt and Möllendorff 1898: 138.</p><p>Diplommatina (Diplommatina) sperata — Kobelt 1902: 446. Budha et al. 2015: 7.</p><p>Type material examined.</p><p>Myanmar • Possible syntypes. NHMUK 1888.12.4.134 –137 (4 shells; Fig. 20 C) from Maii, Arakan Coast; W. Theobald coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, pale yellowish or whitish. Whorls ~ 5–6, well rounded; spire depressed-conical; suture wide, deep. Teleoconch with wide, equally spaced radial ribs throughout. Penultimate whorl well rounded, wider than last whorl. Aperture ovate; columellaris prominent, directed downward. Peristome expanded, double, multi-layered; basal edge slightly sinuous and protruded; inner peristome broadly expanded on parietal side. Umbilicus closed.</p><p>Distribution.</p><p>This species was originally known from its type locality in the Arakan Hills, Bago Region, Myanmar. It was later recorded from ‘ Moditoung’, likely corresponding to the Modi Taung Hills in Padauang Township, Pyay District, Bago Region, and from Mai-i, likely in the area of Ma-ei Town, Thandwe District, Rakhine State (Theobald and Stoliczka 1872). More recently, it has been reported from Nepal (Kuznetsov and Schileyko 1997; Budha et al. 2015, 2017).</p><p>Remarks.</p><p>This species has also been reported from several localities in southwestern to eastern Myanmar, including Ayeyarwady, Bago, and Tanintharyi regions (Blanford 1865).</p><p>It appears to have a broad distribution across Bangladesh, India, and Nepal (Budha et al. 2015; Preece et al. 2022). However, Budha et al. (2017) noted that specimens from Nepal tend to have thinner shells and much thinner peristomal lips than the syntypes, and are therefore provisionally assigned to this species.</p></div>	https://treatment.plazi.org/id/DC7DF65F026E5A029F5D3B1DF6C8D9F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
6A512F5AD75258F8B68B44D7CC890105.text	6A512F5AD75258F8B68B44D7CC890105.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina thomsoni Godwin-Austen 1892	<div><p>30 Diplommatina thomsoni Godwin-Austen, 1892</p><p>Fig. 20 D</p><p>Diplommatina thomsoni Godwin-Austen, 1892: 514. Type locality: South Burrail. Godwin-Austen 1897: 9, 10, pl. 65, figs 2, 2 a. Gude 1921: 344. McGhie 2008: 32.</p><p>Diplommatina (Metadiancta) thomsoni — Kobelt and Möllendorff 1898: 139. Kobelt 1902: 451.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2191 (14 shells; Fig. 20 D) from South Burrail; Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, translucent, whitish. Whorls ~ 7, well rounded; spire elevated conical; suture wide, shallow. Teleoconch: earlier whorls with strong, narrowly and equally spaced radial ribs; penultimate and last whorls with slightly wider-spaced radial ribs. Penultimate whorl convex, slightly wider than last whorl. Aperture circular; columellaris weak, directed downward. Peristome expanded, double, multi-layered; inner peristome slightly broader than outer peristome; parietal callus thickened. Columella straight; umbilicus closed.</p><p>Distribution.</p><p>This species is known from its type locality in the Barail Range, Manipur State, India. Records from Myanmar, however, lack precise locality data (Gude 1921), and the presence there has not yet been reconfirmed.</p><p>Remarks.</p><p>There are three syntypes housed in The Manchester Museum, University of Manchester (EE. 3511 / 130) (McGhie 2008).</p></div>	https://treatment.plazi.org/id/6A512F5AD75258F8B68B44D7CC890105	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
6B552319D7BF5B00BFDA8E82EA44EFC3.text	6B552319D7BF5B00BFDA8E82EA44EFC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina tumida subsp. laisenensis Gude 1921	<div><p>31 Diplommatina tumida laisenensis Gude, 1921</p><p>Fig. 20 E</p><p>Diplommatina tumida var. Godwin-Austen, 1892: 512. Type locality: Laisen Peak and Trigonometrical Station, Munipur. Godwin-Austen 1897: 7, 8, pl. 64, fig. 7.</p><p>Diplommatina tumida var. laisenensis Gude, 1921: 338 . Type locality: Burma: Laisen Peak and Nongmaiching Trigonometrical Station, Munipur.</p><p>Type material examined.</p><p>India • Syntypes. NHMUK 1903.7.1.2139 (5 shells) labelled as D. tumida var. from Laisen H. S. Munipur; Godwin-Austen coll. • Syntypes. NHMUK 1903.7.1.2136 (2 shells; Fig. 20 E) labelled as D. tumida small var. from Nongmaiching, Munipur (figured in Godwin-Austen 1897: pl. 64, fig. 7); Godwin-Austen coll.</p><p>Diagnosis.</p><p>Shell dextral, ovate-fusiform, dull, whitish. Whorls ~ 7, well rounded; spire elevated conical; suture wide, shallow. Teleoconch: earlier whorls with weak, narrowly and equally spaced radial ribs; penultimate and last whorls with slightly wider-spaced radial ribs to nearly smooth. Penultimate whorl convex, wider than last whorl. Aperture circular; columellaris strong, directed downward. Peristome expanded, double, multi-layered; outer peristome slightly broader than inner peristome; parietal callus expanded. Columella straight; umbilicus closed.</p><p>Distribution.</p><p>This species is known from its type localities at Laisen Peak and the Trigonometrical Station in Manipur State, India. Records from Myanmar, however, lack precise locality data (Gude 1921), and the presence there has not yet been reconfirmed.</p><p>Remarks.</p><p>This subspecies was originally described in detail by Godwin-Austen (1892) under the designation ‘ var. ’ and later illustrated by Godwin-Austen (1897), although no formal name was proposed at that time. Gude (1921) subsequently republished the original description and introduced ‘ laisenensis ’ as a nomen novum, which has been made available. The type series of this subspecies corresponds to the specimens referred to as ‘ var. ’ and the ‘ illustrated specimen’ by Godwin-Austen (1892, 1897) from Lisein Peak and the Nongmaiching Trigonometrical Station, Munipur. The NHM, London, holds two specimen lots considered syntypes: one containing five shells labelled ‘ Diplommatina tumida var. from Laisen H. S., Munipur’, and another containing two shells labelled ‘ Diplommatina tumida small var. from Nongmaiching, Manipur’.</p></div>	https://treatment.plazi.org/id/6B552319D7BF5B00BFDA8E82EA44EFC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
A73011C5B78A5B75B418659D728C96C9.text	A73011C5B78A5B75B418659D728C96C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina Chen 2025	<div><p>Genus Pagodapalaina Chen, 2025</p><p>Pagodapalaina Chen in Chen et al. 2025: 390, 391.</p><p>Type species.</p><p>Pagodapalaina hortulanica Chen, 2025, by original designation.</p><p>Remarks.</p><p>The genus was recently separated from the more diverse Diplommatina s. l. based on differences in radular and shell morphology, as well as published molecular data (Webster et al. 2012; Boonmachai et al. 2023; Chen et al. 2025). It is characterised by a small, sinistral, conical to elongate-conical shell with angular whorls and prominent radial ribs that often form trough-like projections along the shell periphery. The genus currently comprises 17 species, which are mainly distributed in the Malay Peninsula and Sumatra, Indonesia. In Myanmar, the genus is recorded for the first time, as P. suratensis from the southern part of the Tanintharyi Region.</p></div>	https://treatment.plazi.org/id/A73011C5B78A5B75B418659D728C96C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
FE717513F89D56CF8963226871607C17.text	FE717513F89D56CF8963226871607C17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagodapalaina suratensis (Panha & Burch 1998)	<div><p>8 Pagodapalaina suratensis (Panha &amp; Burch, 1998)</p><p>Figs 7 E, 11 D, 14</p><p>Diplommatina suratensis Panha &amp; Burch, 1998: 57, fig. 5. Type locality: Klongsang Wildlife Sanctuary, Surathani Province, Thailand. Panha and Burch 2005: 33, fig. 35. BEDO 2017: 83. Jirapatrasilp et al. 2023: 26.</p><p>Pagodapalaina suratensis — Chen et al. 2025: 398, 399, fig. 3 h.</p><p>Type material examined.</p><p>Thailand • Holotype. CUMZ 15370 (former Di 028; Fig. 7 E) from Klongsang Wildlife Sanctuary, Surathani Province; Panha and Burch coll. • Paratypes. CUMZ 15371 (former Di 029; 7 shells); same data as for holotype .</p><p>Other material.</p><p>Myanmar • CUMZ 15463 (1 shell; Fig. 14 A), CUMZ 15464 (1 shell; Figs 11 D, 14 B), CUMZ 15465 (327 shells + 6 juveniles) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.17619&amp;materialsCitation.latitude=11.2295" title="Search Plazi for locations around (long 99.17619/lat 11.2295)">Buddha Cave</a>, Lenya City, Tanintharyi Region; 11°13'46.2"N, 99°10'34.3"E; C. Sutcharit, R. Chanabun and R. Srisonchai leg.</p><p>Description.</p><p>Shell sinistral, turreted-fusiform, reddish to yellowish. Shell height 1.9–2.2 mm; shell width 0.9–1.2 mm. Whorls ~ 7, earlier whorls rounded, last three whorls angular; spire elongate conical; suture wide and very deep. Protoconch ~ 1 ½ whorls, with roundish malleated pits throughout. Teleoconch with strong, regularly spaced radial ribs; earlier whorls ribs regularly strong, last three whorls’ ribs slightly elevated, weakly sinuous at angulation, and then thinner and lower near suture. Ribs with 6–7 ribs / 0.5 mm on penultimate whorl; 10–11 ribs / 0.5 mm on last whorl towards aperture. Spiral striations very thin and fine, appearing from teleoconch to aperture. Last whorl narrower than penultimate whorl and displaced rightward relative to peristome position. Constriction inconspicuous externally and internally. Aperture circular bearing, with three barriers: parietalis short, very weak, low knob-like; palatalis elongate horizontally and form tall ridge; columellaris prominent, continuous with peristome, slightly deflected downward toward palatal wall (Fig. 11 D). Peristome broadly expanded, double; outer peristome broader than inner peristome. Lip thickened, multi-layered; parietal callus thickened and expanded. Columella straight, thickened. Umbilicus closed.</p><p>Distribution.</p><p>Pagodapalaina suratensis occurs in several areas of southern Thailand (Panha and Burch 2005). It has recently been recorded from limestone outcrops in the Tanintharyi Region, southern Myanmar (Fig. 1 A).</p><p>Remarks.</p><p>This is the first record of this species in Myanmar. Diplommatina somsakpanhai sp. nov. was also found syntopic with this species. The new species differs in having opposite shell coiling, an ovate shell shape, and a very large, thickened shell aperture.</p></div>	https://treatment.plazi.org/id/FE717513F89D56CF8963226871607C17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tongkerd, Piyoros;Lwin, Ngwe;Ablett, Jonathan D.;Pholyotha, Arthit;Annate, Supunya;Sutcharit, Chirasak	Tongkerd, Piyoros, Lwin, Ngwe, Ablett, Jonathan D., Pholyotha, Arthit, Annate, Supunya, Sutcharit, Chirasak (2026): Contributions of a small collection of the terrestrial microsnails genus Diplommatina s. l. (Caenogastropoda, Cyclophoroidea, Diplommatinidae) from Myanmar, with a catalogue of recorded species and description of two new species. ZooKeys 1285: 71-124, DOI: 10.3897/zookeys.1285.186884
