taxonID	type	description	language	source
90278E13FFE9FF9657C5FD9F494E9273.taxon	distribution	Distribution type: Peregrine Distribution in Brazil: Present in 15 Brazilian states (Brown & Fragoso, 2007, Sousa et al. 2020 b)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE9FF9657C5FD9F494E9273.taxon	materials_examined	MARANHÃO: Centro Novo do Maranhão: LB _ ANL 000271, 57 adults, in plinthosol in a pasture, (3 ° 40 ' 25.49 " S 46 ° 47 ' 20.48 " W), April, 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000274, 5 adults, in plinthosol in a pasture, (3 ° 55 ' 3.55 " S 46 ° 45 ' 49.55 " W), April, 2017, L. M. Hernández-García, G. X. Rousseau, colls.; Itinga do Maranhão: LB _ ANL 000272, 129 adults, in argisol in a pasture, (4 ° 5 ' 43.00 " S 46 ° 52 ' 20.00 " W), April, 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000273, 14 juveniles, in plinthosol in a logged forest, (3 ° 55 ' 23.40 " S 46 ° 45 ' 57.04 " W), April, 2017, L. M. Hernández-García, G. X. Rousseau, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE9FF9657C5FB4C4EDE902B.taxon	distribution	Distribution type: Exotic Distribution in Brazil: Present in 14 Brazilian states (Brown & Fragoso, 2007) MARANHÃO: Caxias (Righi, 1972). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE9FF9757C5F8D44FE49770.taxon	distribution	Distribution type: Exotic (Brown & Fragoso, 2007) Distribution in Brazil: Present in 10 Brazilian states (Brown & Fragoso, 2007; This study)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE9FF9757C5F8D44FE49770.taxon	materials_examined	MARANHÃO: Caxias: LB _ ANL 000375, 5 adults, in Vermicomposting, (4 ° 51 ' 32 " S 43 ° 21 ' 10 " W), February, 2019, L. M. Hernández-García.; Raposa: LB _ ANL 000376, 4 adults, in soil next to vermicomposting, (2 ° 25 ' 44 " S 44 ° 06 ' 34 " W), May, 2022, L. M. Hernández-García.; São Luís: LB _ ANL 000376, 10 adults, in Vermicomposting, (2 ° 31 ' 47 " S 44 ° 18 ' 10 " W), May, 2025, L. M. Hernández-García. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9757C5FC7F4E2D92D0.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Hernandez et al. 2018 a, This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9757C5FC7F4E2D92D0.taxon	materials_examined	MARANHÃO: Centro Novo do Maranhão: LB _ ANL 000275, 1 adult, in plinthosol in a logged forest, (3 ° 40 ’ 50.06 ” S 46 ° 46 ’ 12.95 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000276, 2 adults, in plinthosol in an old-growth forest, (3 ° 41 ’ 27.68 ” S 46 ° 46 ’ 2.88 ” W), April, 2016, L. M. Hernández-García, G. X. Rousseau, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9057C5FAE34D64936D.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve, Itinga (Hernandez et al. 2018 a, This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9057C5FAE34D64936D.taxon	materials_examined	MARANHÃO: Centro Novo do Maranhão: LB _ ANL 000277, 9 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 11.69 " S 46 ° 45 ' 34.87 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000278 — 12 juveniles and LB _ ANL 000279, 8 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 16.35 " S 46 ° 45 ' 34.73 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000280 — 6 juveniles and LB _ ANL 000281, 8 adults, in plinthosol in a logged forest, (3 ° 40 ' 50.95 " S 46 ° 46 ' 13.06 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000282 — 9 juveniles and LB _ ANL 000283, 12 adults, in plinthosol in a logged forest, (3 ° 40 ' 59.74 " S 46 ° 46 ' 1.11 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000284 — 6 adults and LB _ ANL 000285, 2 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ' 27.21 " S 46 ° 45 ' 6.19 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000286 — 7 juveniles and LB _ ANL 000287, 6 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 24.87 " S 46 ° 45 ' 13.87 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000288 — 1 juvenile and LB _ ANL 000289, 7 adults, in plinthosol in a logged forest, (3 ° 41 ' 10.36 " S 46 ° 45 ' 56.36 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000290 — 12 adults and LB _ ANL 000291, 23 juveniles, in plinthosol in a logged forest, (3 ° 41 ' 20.33 " S 46 ° 46 ' 1.16 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000292 — 1 adult and LB _ ANL 000293, 1 juvenile, in plinthosol in a pasture, (3 ° 40 ' 25.49 " S 46 ° 47 ' 20.47 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000294 — 4 adults and LB _ ANL 000295, 1 juvenile, in plinthosol in a pasture, (3 ° 37 ' 59.90 " S 46 ° 48 ' 57.10 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000296, 4 adults, in plinthosol in an intermediate secondary forest, (3 ° 41 ' 56.61 " S 46 ° 45 ' 57.81 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000297, 7 adults, in plinthosol in an intermediate secondary forest, (3 ° 41 ' 57.30 " S 46 ° 46 ' 4.80 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000298, 9 adults, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 7.90 " S 46 ° 45 ' 42.63 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000299 — 10 juveniles and LB _ ANL 000300, 12 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 26.11 " S 46 ° 46 ' 14.11 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000301 — 27 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ' 18.50 " S 46 ° 45 ' 56.20 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000302 — 2 juveniles and LB _ ANL 000303, 11 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 27.72 " S 46 ° 46 ' 2.26 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000304 — 29 adults and LB _ ANL 000305, 61 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ' 21.00 " S 46 ° 46 ' 24.20 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000306 — 5 adults and LB _ ANL 000307, 5 juveniles, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 18.22 " S 46 ° 45 ' 47.40 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000308 — 12 juveniles and LB _ ANL 000309, 10 adults, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 7.3 " S 46 ° 45 ' 36.1 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls.; Itinga do Maranhão: LB _ ANL 000310, 7 adults, in latosol in a pasture, (4 ° 2 ' 20.00 " S 46 ° 56 ' 57.01 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000311, 1 adult, in latosol in an old-growth forest, (4 ° 2 ' 28.80 " S 46 ° 54 ' 30.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000312, 2 adults, in latosol in an old-growth forest, (4 ° 2 ' 3.64 " S 46 ° 5 ' 3.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000313, 3 adults, in latosol in an old-growth forest, (4 ° 2 ' 12.66 " S 46 ° 2 ' 12.20 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000314 — 9 juveniles and LB _ ANL 000315, 1 adult, in argisol in an old-growth forest, (4 ° 5 ' 57.60 " S 46 ° 5 ' 57.60 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000316 — 5 adults and LB _ ANL 000317, 3 juveniles, in argisol in an old-growth forest, (4 ° 6 ' 4.00 " S 46 ° 54 ' 58.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000318, 2 juveniles, in argisol in a pasture, (4 ° 4 ' 20.98 " S 46 ° 52 ' 59.78 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000319, 1 adult, in argisol in an intermediate secondary forest, (4 ° 4 ' 50.30 " S 46 ° 52 ' 51.30 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000320, 2 juveniles, in argisol in an intermediate secondary forest, (4 ° 4 ' 31.00 " S 46 ° 4 ' 31.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000321 — 3 juveniles and LB _ ANL 000322, 7 adults, in plinthosol in a logged forest, (3 ° 55 ' 23.60 " S 46 ° 45 ' 57.04 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000323 — 3 adults and LB _ ANL 000324, 4 juveniles, in plinthosol in a logged forest, (3 ° 58 ' 59.00 " S 46 ° 47 ' 56.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000325 — 5 adults and LB _ ANL 000326, 1 juvenile, in plinthosol in a logged forest, (3 ° 55 ' 3.55 " S 46 ° 45 ' 49.55 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE8FF9757C5FE0B4D2794BC.taxon	distribution	Distribution type: Exotic (Brown & Fragoso, 2007) Distribution in Brazil: Maranhão: São Luís (Sousa et al. 2020 b). Pernambuco: Ponta de Pedras (Righi, 1972). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEFFF9157C5FA3F4F4D97C5.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve, Itinga (Hernandez et al. 2018 a, This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEFFF9157C5FA3F4F4D97C5.taxon	materials_examined	MARANHÃO: Centro Novo do Maranhão: LB _ ANL 000327, 3 adults, in plinthosol in a logged forest, (3 ° 40 ’ 50.95 ” S 46 ° 46 ’ 13.06 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000328, 1 adult, in plinthosol in a logged forest, (3 ° 41 ’ 20.35 ” S 46 ° 46 ’ 1.16 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000329 — 1 adult and LB _ ANL 000330, 10 juveniles, in plinthosol in a logged forest, (3 ° 40 ’ 25.50 ” S 46 ° 47 ’ 20.46 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000331 — 15 adults and LB _ ANL 000332, 7 juveniles, in plinthosol in a logged forest, (3 ° 40 ’ 22.00 ” S 46 ° 47 ’ 46.00 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000333 — 13 adults and LB _ ANL 000334, 12 juveniles, in plinthosol in a logged forest, (3 ° 40 ’ 17.20 ” S 46 ° 47 ’ 54.90 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000335, 4 adults, in plinthosol in a logged forest, (3 ° 40 ’ 8.20 ” S 46 ° 47 ’ 54.20 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000336 — 4 juveniles and LB _ ANL 000337, 7 adults, in plinthosol in a logged forest, (3 ° 40 ’ 7.20 ” S 46 ° 47 ’ 42.50 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000338, 2 juveniles, in plinthosol in a logged forest, (3 ° 36 ’ 59.90 ” S 46 ° 48 ’ 57.50 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000339, 2 juveniles, in plinthosol in an intermediate secondary forest, (3 ° 41 ’ 57.30 ” S 46 ° 46 ’ 04.80 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000340, 2 adults, in plinthosol in an intermediate secondary forest, (3 ° 42 ’ 18.20 ” S 46 ° 45 ’ 47.40 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls.; Itinga do Maranhão: LB _ ANL 000341, 6 juveniles, in argisol in a pasture, (4 ° 5 ' 43.00 " S 46 ° 52 ' 21.10 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000342, 3 juveniles, in argisol in an intermediate secondary forest, (4 ° 4 ' 32.00 " S 46 ° 52 ' 49.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEEFF9157C5FD9349279505.taxon	distribution	Distribution type: Native. Distribution in Brazil: Amapá: Serra do Navio (Righi, 1971); Maranhão: Caxias (Righi, 1972); Roraima: Ilha de Maracá, Estrada Boa Vista — Bonfim, km 100 (Hamoui & Donatelli, 1983; Righi, 1990). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEEFF9157C5FC534D8491ED.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Dos Santos et al. 2017), Alcântara (This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEEFF9157C5FC534D8491ED.taxon	materials_examined	MARANHÃO: Centro Novo do Maranhão: LB _ ANL 000351 — 3 juveniles and LB _ ANL 000352, 4 adults, in plinthosol in an old-growth forest, (3 ° 41 ’ 11.71 ” S 46 ° 45 ’ 34.91 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000353 — 2 adults and LB _ ANL 000354, 2 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ’ 16.35 ” S 46 ° 45 ’ 34.74 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000355, 1 juvenile, in plinthosol in a logged forest, (3 ° 40 ’ 59.74 ” S 46 ° 46 ’ 1.49 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000356, 1 adult, in plinthosol in an old-growth forest, (3 ° 41 ’ 24.87 ” S 46 ° 45 ’ 13.00 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000357, 3 adults, in plinthosol in a logged forest, (3 ° 41 ’ 10.76 ” S 46 ° 45 ’ 56.54 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000358, 1 adult, in plinthosol in a logged forest, (3 ° 41 ’ 20.33 ” S 46 ° 46 ’ 1.16 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000362, 2 adults, in plinthosol in an intermediate secondary forest, (3 ° 41 ’ 57.30 ” S 46 ° 46 ’ 4.80 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000363, 1 adult, in plinthosol in an old-growth forest, (3 ° 41 ’ 26.10 ” S 46 ° 46 ’ 14.20 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000364, 8 adults, in plinthosol in an old-growth forest, (3 ° 41 ’ 18.50 ” S 46 ° 45 ’ 57.20 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000365, 7 adults, in plinthosol in an old-growth forest, (3 ° 41 ’ 27.7 ” S 46 ° 46 ’ 2.7 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000370, 6 adults, in plinthosol in a logged forest, (3 ° 55 ’ 3.55 ” S 46 ° 45 ’ 49.97 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls.; Alcântara: LB _ ANL 000359 — 4 adults and LB _ ANL 000360, 2 juveniles, in latosol in a young secondary forest, (2 ° 22 ' 12.99 " S 44 ° 26 ' 40.30 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000361, 5 adults, in latosol in an intermediate secondary forest, (2 ° 21 ' 56.30 " S 44 ° 27 ' 10.50 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000366 — 2 juveniles and LB _ ANL 000367, 1 adult, in latosol in a young secondary forest, Alcântara (2 ° 21 ' 51.10 " S 44 ° 27 ' 16.40 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000368, 1 adult, in latosol in a logged forest, (2 ° 21 ' 57.00 " S 44 ° 27 ' 15.98 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000369, 1 juvenile, in latosol in a young secondary forest, (2 ° 22 ' 3.10 " S 44 ° 26 ' 42.10 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000371, 1 juvenile, in latosol in a young secondary riparian forest, (2 ° 20 ' 55.50 " S 44 ° 27 ' 56.40 " W), June 2013, A. R. Piedade. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FD0E490995F9.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Rosário (Hernández-García et al. 2018 b, This study)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FD0E490995F9.taxon	materials_examined	MARANHÃO: Rosário: LB _ ANL 000252, 14 juveniles, in argisol in an advanced AFS, (2 ° 51 ' 4.04 " S 44 ° 8 ' 51.42 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000253, 1 juvenile, in argisol in a young AFS, (2 ° 51 ' 14.37 " S 44 ° 9 ' 13.37 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FBCA4F989181.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Alcântara (Hernández-García et al. 2018 b, This study), São José de Ribamar (This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FBCA4F989181.taxon	materials_examined	MARANHÃO: Alcântara: LB _ ANL 000254, 5 adults, in gleysol in a logged forest, (2 ° 25 ' 3.40 " S 44 ° 25 ' 20.00 " W), June 2010, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000255 — 3 juveniles andLB _ ANL 000256, 9 adults, in gleysol in a young secondary forest, (2 ° 25 ' 25.50 " S 44 ° 25 ' 31.60 " W), June 2010, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000257, 7 adults, in gleysol in a grassland, (2 ° 25 ' 26.9 " S 44 ° 25 ' 30.7 " W), June 2010, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000258, 18 juveniles, in gleysol in a logged forest, (2 ° 24 ' 59.15 " S 44 ° 25 ' 30.36 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000259 — 2 juveniles an LB _ ANL 000260, 3 adults, in gleysol in an intermediate secondary forest, (2 ° 24 ' 58.18 " S 44 ° 25 ' 28.40 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000261 — 12 adults and LB _ ANL 000262, 2 juveniles, in gleysol in an intermediate secondary forest, (2 ° 25 ' 33.19 " S 44 ° 25 ' 26.59 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000264, 1 juvenile, in latosol in a slash-burn, (2 ° 21 ' 26.1 " S 44 ° 28 ' 41.1 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000265, 3 juveniles, in latosol in a logged forest, (2 ° 21 ' 58.60 " S 44 ° 27 ' 16.50 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000266, 16 juveniles, in latosol in a young secondary forest, (2 ° 22 ' 13.00 " S 44 ° 26 ' 40.3 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000267, 1 juvenile, in latosol in an intermediate secondary forest, (2 ° 21 ' 56.30 " S 44 ° 27 ' 10.51 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000268, 4 juveniles, in latosol in a young secondary forest, (2 ° 21 ' 51.10 " S 44 ° 27 ' 16.40 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000269 — 3 adults and LB _ ANL 000270, 13 juveniles, in latosol in a young secondary forest, (2 ° 22 ' 3.1 " S 44 ° 26 ' 42.1 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls.; São José de Ribamar: LB _ ANL 000263, 1 juvenile, in argisol in an old-growth forest, (2 ° 38 ' 59.24 " S 44 ° 8 ' 4.1 " W), January 2014, J. A. M. Gutiérrez. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FF624F86940D.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Caxias (This study), São Luís (Sousa et al. 2020 b). Pará: Cametá, Mocajuba (Righi, 1972), Itupiranga, Marabá, Tucuruí (Righi, 1975). Paraíba: João Pessoa (Righi, 1975). Mato Grosso: Pontes e Lacerda (Righi, 1975).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFEDFF9257C5FF624F86940D.taxon	materials_examined	MARANHÃO: Caxias: LB _ ANL 000372, 15 juveniles, in neosol in a logged riparian forest, (4 ° 53 ’ 33.60 ” S 43 ° 25 ’ 36.37 ” W), February 2019, A. M. Sousa, L. M. Hernández-García, colls. LB _ ANL 000373, 2 juveniles, in neosol in a logged riparian forest, (4 ° 53 ’ 27.92 ” S 43 ° 25 ’ 32.98 ” W), February 2019, A. M. Sousa, L. M. Hernández-García, colls. LB _ ANL 000374, 8 juveniles, in neosol in an old-growth forest, (4 ° 54 ’ 43.91 ” S 43 ° 27 ’ 54.28 ” W), February 2019, A. M. Sousa, L. M. Hernández-García, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5F8FA4D5991AD.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Alcântara, Centro Novo do Maranhão, Itinga, Rosário, São Luís (Hernández-García et al. 2018 c)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5FD9E4F8B90A9.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão (This study), Itinga do Maranhão (Hernández-García et al. 2018 c, This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5FD9E4F8B90A9.taxon	materials_examined	MARANHÃO: Centro Novo do Maranhão: LB _ ANL 000230, 9 juveniles, in latosol in a logged forest, (3 ° 59 ’ 52.12 ” S 46 ° 50 ’ 8.77 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000232, 2 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ’ 27.19 ” S 46 ° 45 ’ 6.91 ” W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000233, 3 juveniles, in plinthosol in an intermediate secondary forest, (3 ° 41 ’ 56.70 ” S 46 ° 45 ’ 57.80 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000234, 4 juveniles, in plinthosol in an intermediate secondary forest, (3 ° 41 ’ 57.30 ” S 46 ° 46 ’ 4.80 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000235, 2 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ’ 26.10 ” S 46 ° 46 ’ 14.20 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000236, 2 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ’ 27.70 ” S 46 ° 46 ’ 2.70 ” W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls.; Itinga do Maranhão: LB _ ANL 000231, 6 juveniles, in latosol in an intermediate secondary forest, (4 ° 2 ' 3.80 " S 46 ° 54 ' 19.80 " W), Jule, 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000238, 3 juveniles, in latosol in a pasture, (4 ° 2 ' 35.30 " S 46 ° 55 ' 56.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000239, 6 juveniles, in latosol in an intermediate secondary forest, (4 ° 1 ' 53.70 " S 46 ° 52 ' 58.30 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000240, 2 juveniles, in latosol in a pasture, (4 ° 2 ' 17.99 " S 46 ° 54 ' 45.50 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000241, 1 juvenile, in latosol in an old-growth forest, (4 ° 2 ' 28.80 " S 46 ° 54 ' 30.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000242, 8 juveniles, in latosol in an old-growth forest, (4 ° 2 ' 3.80 " S 46 ° 54 ' 19.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000243, 3 juveniles, in argisol in an old-growth forest, (4 ° 6 ' 4.00 " S 46 ° 6 ' 4.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000244, 1 juvenile, in argisol in an intermediate secondary forest, (4 ° 4 ' 50.4 " S 46 ° 4 ' 50.3 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000245, 6 juveniles, in latosol in a logged forest, (3 ° 59 ' 57.69 " S 46 ° 49 ' 43.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000246, 8 juveniles, in plinthosol in a logged forest, (3 ° 55 ' 23.60 " S 46 ° 45 ' 57.04 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000247, 1 juvenile, in plinthosol in a logged forest, (3 ° 58 ' 50.00 " S 46 ° 47 ' 9.99 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000248, 4 juveniles, in plinthosol in a logged forest, (3 ° 59 ' 59.00 " S 46 ° 47 ' 56.80 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFECFF9357C5FF2A4D27979D.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Caxias, São Luís (Sousa, 2020 b). Paraíba: João Pessoa (Righi, 1994). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FE664DB89429.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Hernández-García et al. 2024). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FD7A4DB8952D.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve (Hernández-García et al. 2024). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FF624FD39755.taxon	discussion	To date only two species of Atatina were recorded to Maranhão: Atatina arimikuriaka and Atatina amirikuritea (Hernández-García et al. 2024). The accession codes LB 000001 – LB 000010 were recodified as presented in the current model (LB _ ANL) of the Maranhão Invertebrate Collection (CISMA).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FC264E2D9389.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Bom Jardim * (Sousa et al. 2020 b), Itinga do Maranhão (This study).	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9C57C5FC264E2D9389.taxon	materials_examined	MARANHÃO: Itinga do Maranhão: LB _ ANL 000343, 2 adults, in latosol in an old-growth forest, (4 ° 02 ’ 34.50 ” S 46 ° 54 ’ 36.10 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000344, 1 adult, in argisol in an old-growth forest, (4 ° 06 ’ 04.00 ” S 46 ° 54 ’ 59.00 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000345 — 3 adults and LB _ ANL 000346, 4 juveniles, in argisol in a pasture, (4 ° 4 ’ 20.79 ” S 46 ° 52 ’ 59.80 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000347 — 8 adults and LB _ ANL 000348, 6 juveniles, in argisol in an intermediate secondary forest, (4 ° 4 ’ 50.30 ” S 46 ° 52 ’ 37.40 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000349 — 2 juveniles and LB _ ANL 000350, 1 adult, in argisol in an intermediate secondary forest, (4 ° 4 ’ 32.00 ” S 46 ° 52 ’ 49.00 ” W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9857C5FA1A49439165.taxon	distribution	Distribution type: Peregrine. Distribution in Brazil: Present in 17 Brazilian states (Brown & Fragoso, 2007; This study)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE3FF9857C5FA1A49439165.taxon	materials_examined	MARANHÃO: Alcântara: LB _ ANL 000039 — 8 adults and LB _ ANL 000040, 4 juveniles, in gleysol in an intermediate secondary forest, (2 ° 25 ' 33.19 " S 44 ° 25 ' 26.59 " W), July 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000077 — 8 adults and LB _ ANL 000078, 11 juveniles, in latosol in an advanced secondary riparian forest, (2 ° 21 ' 32.60 " S 44 ° 27 ' 43.30 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000079 — 10 adults and LB _ ANL 000080, 72 juveniles, in latosol in an advanced secondary riparian forest, (2 ° 21 ' 32.60 " S 44 ° 27 ' 43.30 " W), June 2013, A. R. Piedade. LB _ ANL 000081, 8 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 21 ' 5.50 " S 44 ° 27 ' 51.40 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000082 — 4 adults and LB _ ANL 000083, 3 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 21 ' 5.50 " S 44 ° 27 ' 51.40 " W), June 2013, A. R. Piedade. LB _ ANL 000084, 3 juveniles, in latosol in a young secondary riparian forest, (2 ° 21 ' 2.10 " S 44 ° 27 ' 56.50 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000085, 35 juveniles, in latosol in an advanced secondary riparian forest, (2 ° 21 ' 37.80 " S 44 ° 27 ' 16.50 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000086 — 9 adults and LB _ ANL 000087, 46 juveniles, in latosol in an advanced secondary riparian forest, (2 ° 21 ' 37.80 " S 44 ° 27 ' 16.50 " W), June 2013, A. R. Piedade. LB _ ANL 000088, 22 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 21 ' 37.10 " S 44 ° 27 ' 36.10 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000089 — 7 adults and LB _ ANL 000090, 43 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 21 ' 37.10 " S 44 ° 27 ' 36.10 " W), June 2013, A. R. Piedade. LB _ ANL 000091 — 1 adult and LB _ ANL 000092, 4 juveniles, in latosol in a young secondary riparian forest, (2 ° 21 ' 35.70 " S 44 ° 27 ' 26.70 " W), June 2013, A. R. Piedade. LB _ ANL 000093, 35 juveniles, in latosol in an advanced secondary riparian forest, (2 ° 21 ' 39.20 " S 44 ° 26 ' 30.10 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000094 — 2 adults and LB _ ANL 000095, 45 juveniles, in latosol in an advanced secondary riparian forest, (2 ° 21 ' 39.20 " S 44 ° 26 ' 30.10 " W), June 2013, A. R. Piedade. LB _ ANL 000096 — 6 adults and LB _ ANL 000097, 31 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 21 ' 43.90 " S 44 ° 26 ' 36.00 " W), June 2013, A. R. Piedade. LB _ ANL 000098 — 11 juveniles and LB _ ANL 000099, 2 juveniles, in latosol in a young secondary riparian forest, (2 ° 21 ' 42.00 " S 44 ° 26 ' 31.90 " W), June 2013, A. R. Piedade. LB _ ANL 000100, 1 adult, in latosol in a manihot crop, (2 ° 21 ' 37.90 " S 44 ° 26 ' 54.60 " W), June 2013, A. R. Piedade. LB _ ANL 000101, 65 juveniles, in latosol in a logged advanced secondary riparian forest, (2 ° 20 ' 54.00 " S 44 ° 28 ' 46.10 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000102 — 10 adults and LB _ ANL 000103, 67 juveniles, in latosol in a logged advanced secondary riparian forest, (2 ° 20 ' 54.00 " S 44 ° 28 ' 46.10 " W), June 2013, A. R. Piedade. LB _ ANL 000104 — 2 juveniles and LB _ ANL 000105, 7 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 20 ' 47.20 " S 44 ° 28 ' 29.90 " W), June 2013, A. R. Piedade. LB _ ANL 000106 — 4 juveniles and LB _ ANL 000107, 4 juveniles, in latosol in a young secondary riparian forest, (2 ° 20 ' 52.60 " S 44 ° 28 ' 39.60 " W), June 2013, A. R. Piedade. LB _ ANL 000108, 44 juveniles, in latosol in a logged advanced secondary riparian forest, (2 ° 20 ' 53.4 " S 44 ° 28 ' 50.7 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000109 — 3 adults and LB _ ANL 000110, 42 juveniles, in latosol in a logged advanced secondary riparian forest, (2 ° 20 ' 53.4 " S 44 ° 28 ' 50.7 " W), June 2013, A. R. Piedade. LB _ ANL 000111, 62 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 20 ' 55.50 " S 44 ° 28 ' 54.30 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000112 — 2 adults and LB _ ANL 000113, 44 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 20 ' 55.50 " S 44 ° 28 ' 54.30 " W), June 2013, A. R. Piedade. LB _ ANL 000114 — 1 juvenile and LB _ ANL 000115, 1 juvenile, in latosol in a young secondary riparian forest, (2 ° 20 ' 55.70 " S 44 ° 28 ' 56.40 " W), June 2013, A. R. Piedade. LB _ ANL 000116, 3 juveniles, in latosol in a manihot crop, (2 ° 20 ' 52.50 " S 44 ° 28 ' 50.00 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000117 — 1 adult and LB _ ANL 000118, 46 juveniles, in latosol in a logged advanced secondary riparian forest, (2 ° 20 ' 56.0 " S 44 ° 29 ' 0.96 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000119 — 1 adult and LB _ ANL 000120, 56 juveniles, in latosol in a logged advanced secondary riparian forest, (2 ° 20 ' 56.0 " S 44 ° 29 ' 0.96 " W), June 2013, A. R. Piedade. LB _ ANL 000121, 25 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 20 ' 52.71 " S 44 ° 29 ' 1.33 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000122 — 1 adult and LB _ ANL 000123, 15 juveniles, in latosol in an intermediate secondary riparian forest, (2 ° 20 ' 52.71 " S 44 ° 29 ' 1.33 " W), June 2013, A. R. Piedade. LB _ ANL 000124, 4 juveniles, in latosol in a young secondary riparian forest, (2 ° 20 ' 38.64 " S 44 ° 29 ' 10.6 " W), November 2012, S. P. Triana, M. L. C. Zelarayán, colls. LB _ ANL 000125 — 1 adult and LB _ ANL 000126, 39 juveniles, in latosol in a young secondary riparian forest, (2 ° 20 ' 38.64 " S 44 ° 29 ' 10.6 " W), June 2013, A. R. Piedade. LB _ ANL 000127, 2 adults, in latosol in a manihot crop Crop, (2 ° 20 ' 36.02 " S 44 ° 29 ' 8.14 " W), June 2013, A. R. Piedade. LB _ ANL 000179 — 16 juveniles and LB _ ANL 000180, 1 adult, in latosol in a fallow, (2 ° 21 ' 27.90 " S 44 ° 28 ' 40.70 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000181, 2 juveniles, in latosol in a young secondary forest, (2 ° 21 ' 52.60 " S 44 ° 27 ' 14.10 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000182, 5 juveniles, in latosol in an intermediate secondary forest, (2 ° 21 ' 56.30 " S 44 ° 27 ' 10.51 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000183 — 13 adults and LB _ ANL 000184, 12 juveniles, in latosol in a young secondary forest, (2 ° 21 ' 29.90 " S 44 ° 28 ' 39.90 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000185 — 22 juveniles and LB _ ANL 000186, 16 adults, in latosol in an intermediate secondary forest, (2 ° 21 ' 30.00 " S 44 ° 28 ' 32.23 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000187, 5 juveniles, in latosol in a young secondary forest, (2 ° 22 ' 26.70 " S 44 ° 27 ' 43.00 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000188 — 3 adults and LB _ ANL 000189, 5 juveniles, in latosol in a logged forest, (2 ° 24 ' 2.70 " S 44 ° 27 ' 40.80 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000190 — 3 adults and LB _ ANL 000191, 7 juveniles, in latosol in an advanced secondary forest, (2 ° 19 ' 17.40 " S 44 ° 28 ' 28.20 " W), May 2015, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000206 — 77 juveniles and LB _ ANL 000207, 9 adults, in latosol in a young secondary forest, (2 ° 21 ' 21.90 " S 44 ° 28 ' 44.60 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000208, 7 juveniles, in latosol in a young secondary forest, (2 ° 21 ' 51.10 " S 44 ° 27 ' 16.40 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000209, 1 adult, in latosol in a logged forest, (2 ° 21 ' 56.99 " S 44 ° 27 ' 15.98 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000210 — 21 juveniles and LB _ ANL 000211, 6 adults, in latosol in an advanced secondary forest, (2 ° 21 ' 2.90 " S 44 ° 25 ' 48.60 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000212 — 6 juveniles and LB _ ANL 000213, 7 adults, in latosol in an advanced secondary forest, (2 ° 20 ' 57.70 " S 44 ° 25 ' 42.00 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000214, 6 juveniles, in latosol in a logged forest, (2 ° 24 ' 5.80 " S 44 ° 27 ' 44.40 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls. LB _ ANL 000215, 1 juvenile, in latosol in a young secondary forest, (2 ° 22 ' 3.10 " S 44 ° 26 ' 42.10 " W), June 2016, J. E. B. Guerrero, L. M. Hernández-García, colls.; Caxias: LB _ ANL 000011, 7 juveniles, in neosol in an old-growth riparian forest, (4 ° 53 ' 27.17 " S 43 ° 25 ' 13.87 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000012 — 137 juveniles and LB _ ANL 000013, 3 adults, in neosol in an old-growth riparian forest, (4 ° 53 ' 26.45 " S 43 ° 25 ' 20.82 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000014, 16 juveniles, in neosol in a logged riparian forest, (4 ° 53 ' 33.58 " S 43 ° 25 ' 36.37 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000015 — 45 juveniles and LB _ ANL 000016, 2 adults, in neosol in a logged riparian forest, (4 ° 53 ' 27.92 " S 43 ° 25 ' 32.98 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000017 — 48 juveniles and LB _ ANL 000018, 3 adults, in neosol in a logged riparian forest, (4 ° 53 ' 21.88 " S 43 ° 25 ' 29.14 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000019, 23 juveniles, in neosol in a logged riparian forest, (4 ° 53 ' 14.71 " S 43 ° 25 ' 31.15 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000024 — 28 juveniles and LB _ ANL 000025, 8 adults, in neosol in an old-growth forest, (4 ° 53 ' 59.03 " S 43 ° 28 ' 15.53 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000028, 2 juveniles, in neosol in an old-growth forest, (4 ° 54 ' 24.84 " S 43 ° 27 ' 58.54 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000029 — 17 juveniles and LB _ ANL 000030, 11 adults, in neosol in a logged forest, (4 ° 53 ' 51.28 " S 43 ° 28 ' 15.06 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000031, 28 juveniles, in plinthosol in a pasture, (4 ° 49 ' 16.72 " S 43 ° 20 ' 28.81 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000032 — 45 juveniles and LB _ ANL 000033, 17 adults, in plinthosol in a logged riparian forest, (4 ° 49 ' 9.08 " S 43 ° 20 ' 31.90 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000034 — 40 juveniles and LB _ ANL 000035, 10 adults, in plinthosol in a logged riparian forest, (4 ° 49 ' 5.42 " S 43 ° 20 ' 33.6 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls. LB _ ANL 000037 — 9 adults and LB _ ANL 000038, 34 juveniles, in plinthosol in a logged riparian forest, Caxias (4 ° 49 ' 5.42 " S 43 ° 20 ' 33.60 " W), February 2019, A. M. Sousa, L. M. Hernández-Garcia, colls.; Centro Novo do Maranhão: LB _ ANL 000154, 1 juvenile, in plinthosol in an old-growth forest, (3 ° 41 ' 11.71 " S 46 ° 45 ' 34.91 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000155, 4 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 11.71 " S 46 ° 45 ' 34.91 " W), March 2015, L. M. Hernández-García. LB _ ANL 000156, 8 juveniles, in plinthosol in an old-growth forest, (3 ° 41 ' 16.35 " S 46 ° 45 ' 34.74 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000157, 1 adult, in plinthosol in an old-growth forest, (3 ° 41 ' 14.03 " S 46 ° 45 ' 34.35 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000158 — 4 juveniles and LB _ ANL 000159, 2 adults, in plinthosol in a logged forest, (3 ° 40 ' 50.06 " S 46 ° 46 ' 12.95 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000160 — 1 juvenile and LB _ ANL 000161, 1 adult, in plinthosol in a logged forest, (3 ° 40 ' 50.06 " S 46 ° 46 ' 12.95 " W), March 2015, L. M. Hernández-García. LB _ ANL 000162, 4 juveniles, in plinthosol in a logged forest, (3 ° 40 ' 56.10 " S 46 ° 46 ' 8.78 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000176, 2 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 24.87 " S 46 ° 45 ' 13.87 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000177 — 5 adults and LB _ ANL 000178, 5 juveniles, in plinthosol in a logged forest, (3 ° 41 ' 10.40 " S 46 ° 45 ' 56.76 " W), March 2015, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000193, 1 adult, in plinthosol in a pasture, (3 ° 40 ' 25.30 " S 46 ° 47 ' 20.50 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000194, 3 adults, in plinthosol in a pasture, (3 ° 40 ' 22.00 " S 46 ° 47 ' 46.60 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000195 — 6 adults and LB _ ANL 000196, 8 juveniles, in plinthosol in a pasture, (3 ° 40 ' 8.20 " S 46 ° 47 ' 54.20 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000197 — 156 juveniles and LB _ ANL 000198, 84 adults, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 9.80 " S 46 ° 45 ' 51.10 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000199 — 26 adults and LB _ ANL 000200, 46 juveniles, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 7.90 " S 46 ° 45 ' 42.60 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000201, 3 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 26.10 " S 46 ° 46 ' 14.20 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000202, 3 adults, in plinthosol in an old-growth forest, (3 ° 41 ' 27.74 " S 46 ° 46 ' 2.68 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000203 — 51 adults and LB _ ANL 000204, 58 juveniles, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 18.22 " S 46 ° 45 ' 47.60 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000205, 2 adults, in plinthosol in an intermediate secondary forest, (3 ° 42 ' 7.3 " S 46 ° 45 ' 37.1 " W), April 2016, L. M. Hernández-García, G. X. Rousseau, colls.; Itinga do Maranhão: LB _ ANL 000036, 8 adults, in latosol in a logged forest, Itinga do Maranhão (4 ° 03 ' 20.92 " S 46 ° 55 ' 39.29 " W), June 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000216 — 6 juveniles and LB _ ANL 000217, 3 adults, in latosol in an old-growth forest, (4 ° 2 ' 3.80 " S 46 ° 54 ' 20.90 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000218 — 59 adults and LB _ ANL 000219, 88 juveniles, in latosol in an old-growth forest, (4 ° 2 ' 12.20 " S 46 ° 54 ' 23.70 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000220, 3 adults, in argisol in an old-growth forest, (4 ° 6 ' 4.00 " S 46 ° 54 ' 59.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000221, 8 adults, in argisol in a pasture, (4 ° 4 ' 20.00 " S 46 ° 4 ' 20.78 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000222 — 12 juveniles and LB _ ANL 000223, 7 adults, in argisol in a pasture, (4 ° 5 ' 43.00 " S 46 ° 52 ' 21.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000224 — 309 juveniles and LB _ ANL 000225, 106 adults, in argisol in an intermediate secondary forest, (4 ° 5 ' 7.40 " S 46 ° 52 ' 9.40 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000226 — 6 adults and LB _ ANL 000227, 3 juveniles, in argisol in an intermediate secondary forest, (4 ° 4 ' 50.40 " S 46 ° 4 ' 50.30 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls. LB _ ANL 000228 — 9 adults and LB _ ANL 000229, 5 juveniles, in argisol in an intermediate secondary forest, (4 ° 4 ' 32.00 " S 46 ° 5 ' 31.00 " W), April 2017, L. M. Hernández-García, G. X. Rousseau, colls.; Rosário: LB _ ANL 000128 — 8 adults and LB _ ANL 000129, 82 juveniles, in argisol in an advanced secondary forest, (2 ° 52 ' 21.93 " S 44 ° 7 ' 38.72 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000130 — 15 adults and LB _ ANL 000131, 43 juveniles, in an argisol in a logged advanced secondary riparian forest, (2 ° 49 ' 55.61 " S 44 ° 7 ' 56.01 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000132 — 15 adults and LB _ ANL 000133, 64 juveniles, in argisol in an advanced secondary riparian forest, (2 ° 51 ' 11.76 " S 44 ° 9 ' 2.63 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000134 — 7 adults and LB _ ANL 000135, 35 juveniles, in argisol in an intermediate secondary forest, (2 ° 51 ' 40.42 " S 44 ° 8 ' 0.4 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000136 — 11 adults and LB _ ANL 000137, 25 juveniles, in argisol in an intermediate secondary forest, (2 ° 51 ' 34.83 " S 44 ° 8 ' 25.15 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000138 — 10 adults and LB _ ANL 000139, 54 juveniles, in argisol in an advanced secondary forest, (2 ° 49 ' 5.28 " S 44 ° 8 ' 20.39 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000140 — 8 adults and LB _ ANL 000141, 21 juveniles, in argisol in a young secondary forest, (2 ° 51 ' 46.88 " S 44 ° 8 ' 32.26 " W), May 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000142 — 11 adults and LB _ ANL 000143, 33 juveniles, in argisol in an intermediate secondary forest, (2 ° 51 ' 10.96 " S 44 ° 9 ' 30.14 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000144 — 4 adults and LB _ ANL 000145, 31 juveniles, in argisol in an intermediate secondary forest, (2 ° 51 ' 17.18 " S 44 ° 9 ' 16.68 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000146 — 3 adults and LB _ ANL 000147, 12 juveniles, in argisol in a young secondary forest, (2 ° 51 ' 2.67 " S 44 ° 8 ' 57.33 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000148 — 42 juveniles and LB _ ANL 000149, 4 adults, in argisol in an advanced AFS, (2 ° 51 ' 4.04 " S 44 ° 8 ' 52.01 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000150 — 13 juveniles and LB _ ANL 000151, 6 adults, in argisol in a young AFS, (2 ° 51 ' 25.47 " S 44 ° 8 ' 3.19 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000152 — 58 juveniles and LB _ ANL 000153, 11 adults, in argisol in an advanced AFS, (2 ° 51 ' 25.47 " S 44 ° 8 ' 3.19 " W), June 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls.; São Luís: LB _ ANL 000020 — 8 adults and LB _ ANL 000021, 25 juveniles, in latosol in an intermediate secondary forest, (2 ° 35 ' 15.00 " S 44 ° 12 ' 43.00 " W), April 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000022 — 1 adult and LB _ ANL 000023, 2 juveniles, in latosol in an intermediate secondary forest, (2 ° 35 ' 15.00 " S 44 ° 12 ' 32.00 " W), April 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000026 — 5 adults and LB _ ANL 000027, 17 juveniles, in latosol in an intermediate secondary forest, (2 ° 35 ' 30.99 " S 44 ° 12 ' 33.00 " W), April 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000058, 1 adult, in latosol in a young secondary forest, São Luís (2 ° 35 ' 25.89 " S 44 ° 12 ' 37.10 " W), Jule 2013, J. A. M. Gutiérrez. LB _ ANL 000071 — 4 adultes and LB _ ANL 000072, 7 juveniles, in latosol in an intermediate secondary forest, (2 ° 35 ' 11.19 " S 44 ° 12 ' 31.36 " W), January 2014, J. A. M. Gutiérrez.; Saõ José de Ribamar: LB _ ANL 000073 — 7 adults and LB _ ANL 000074, 10 juveniles, in argisol in an old-growth forest, (2 ° 38 ' 38.96 " S 44 ° 8 ' 17.66 " W), January 2014, J. A. M. Gutiérrez. LB _ ANL 000075 — 4 adults and LB _ ANL 000076, 12 juveniles, in argisol in an old-growth forest, (2 ° 38 ' 59.24 " S 44 ° 8 ' 4.10 " W), January 2014, J. A. M. Gutiérrez. LB _ ANL 000192, 13 juveniles, in argisol in an old-growth riparian forest, (2 ° 38 ' 42.76 " S 44 ° 8 ' 19.99 " W), January 2016, G. X. Rousseau.; PARÁ: Tomé Açú: LB _ ANL 000041 — 12 adults and LB _ ANL 000042, 125 juveniles, in latosol in an old-growth forest, (2 ° 23 ' 39.00 " S 48 ° 9 ' 52.00 " W), July 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000043 — 3 adults and LB _ ANL 000044, 20 juveniles, in latosol in a logged forest, (2 ° 30 ' 06.00 " S 48 ° 17 ' 26.00 " W), July 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000045 — 1 adult and LB _ ANL 000046, 24 juveniles, in latosol in an intermediate secondary forest, (2 ° 24 ' 46.00 " S 48 ° 11 ' 51.00 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000047 — 1 adult and LB _ ANL 000048, 65 juveniles, in latosol in an intermediate secondary forest, (2 ° 24 ' 7.00 " S 48 ° 11 ' 48.0 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000049, 55 juveniles, in latosol in an intermediate secondary forest, (2 ° 24 ' 6.9984 " S 48 ° 11 ' 48.0012 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000050 — 7 adults and LB _ ANL 000051, 71 juveniles, in latosol in an intermediate secondary forest, Tomé Açu (2 ° 24 ' 29.04 " S 48 ° 11 ' 52.32 " W), Jule 2011, P. R. S. Silva, K. H. M. Muchavisoy, colls. LB _ ANL 000052 — 14 adults and LB _ ANL 000053, 70 juveniles, in latosol in an intermediate secondary forest, (2 ° 31 ' 7.6 " S 48 ° 16 ' 32.7 " W), August 2014, J. A. M. Gutiérrez, G. X. Rousseau, colls. LB _ ANL 000054 — 11 adults and LB _ ANL 000055, 59 juveniles, in plinthosol in an intermediate AFS, (2 ° 11 ' 55.94 " S 48 ° 9 ' 17.15 " W), August 2014, J. A. M. Gutiérrez, G. X. Rousseau, colls. LB _ ANL 000056 — 4 adults and LB _ ANL 000057, 47 juveniles, in latosol in an intermediate AFS, (2 ° 32 ' 38.37 " S 48 ° 16 ' 26.67 W), August 2014, J. A. M. Gutiérrez, G. X. Rousseau, colls. LB _ ANL 000059 — 6 adults and LB _ ANL 000060, 37 juveniles, in latosol in a young AFS, (2 ° 20 ' 58.38 " S 48 ° 15 ' 45.83 " W), Jule 2011, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000061 — 16 adults and LB _ ANL 000062, 100 juveniles, in latosol in a young AFS, (2 ° 20 ' 56.76 " S 48 ° 15 ' 42.4 " W), Jule 2011, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000063 — 9 adults and LB _ ANL 000064, 86 juveniles, in latosol in a young AFS, (2 ° 20 ' 55.46 " S 48 ° 15 ' 40.5 " W), Jule 2011, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000065 — 10 adults and LB _ ANL 000066, 83 juveniles, in latosol in a young AFS, (2 ° 24 ' 12 " S 48 ° 14 ' 49 " W), Jule 2011, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000067 — 8 adults and LB _ ANL 000068, 101 juveniles, in latosol in a young AFS, (2 ° 24 ' 15.00 " S 48 ° 14 ' 51.00 " W), Jule 2011, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000069 — 17 adults and LB _ ANL 000070, 109 juveniles, in latosol in a young AFS, (2 ° 24 ' 12.00 " S 48 ° 14 ' 55.20 " W), Jule 2011, P. R. S. Silva, G. X. Rousseau, colls. LB _ ANL 000163 — 21 juveniles and LB _ ANL 000164, 5 adults, in latosol in a logged forest, (2 ° 24 ' 40.82 " S 48 ° 11 ' 34.12 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000165 — 53 juveniles and LB _ ANL 000166, 3 adults, in latosol in an intermediate secondary forest, (2 ° 24 ' 48.65 " S 48 ° 11 ' 34.38 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000167, 45 juveniles, in plinthosol in a young secondary forest, (2 ° 12 ' 33.87 " S 48 ° 9 ' 28.66 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000168 — 45 juveniles and LB _ ANL 000169, 3 adults, in plinthosol in a young secondary forest, (2 ° 12 ' 34.27 " S 48 ° 9 ' 23.68 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000170 — 70 juveniles and LB _ ANL 000171, 7 adults, in plinthosol in an intermediate secondary forest, (2 ° 11 ' 59.71 " S 48 ° 8 ' 45.96 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000172 — 75 juveniles and LB _ ANL 000173, 20 adults, in plinthosol in an intermediate secondary forest, (2 ° 11 ' 59.59 " S 48 ° 8 ' 58.62 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. LB _ ANL 000174 — 36 juveniles and LB _ ANL 000175, 8 adults, in plinthosol in an intermediate secondary forest, (2 ° 11 ' 55.68 " S 48 ° 9 ' 3.06 " W), August 2014, J. A. M. Gutiérrez, E. G. Cardozo, colls. (Figure 1)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE6FF9957C5FB7548239312.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: Caxias (Sousa et al. 2020 b). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE6FF9957C5FA554F2190DE.taxon	distribution	Distribution type: Native. Distribution in Brazil: Present in 10 Brazilian states (Brown & Fragoso, 2007, Sousa et al. 2020 b) Maranhão: Caxias (Sousa et al. 2020 b). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
90278E13FFE6FF9957C5F8E94D8291D2.taxon	distribution	Distribution type: Native. Distribution in Brazil: Maranhão: São Luís (Sousa et al. 2020 b). Pará: Bagagem, near the Tocantins River (Righi, 1985). (Figure 2)	en	Sousa, Sandriel Costa, Hernández-García, Luis Manuel, Beckman, Ruana Aretha Farias Santiago, Rousseau, Guillaume Xavier (2026): Earthworm species (Crassilitellata) from Eastern Amazonia and Northern Cerrado, Brazil. Zootaxa 5777 (1): 119-138, DOI: 10.11646/zootaxa.5777.1.5, URL: https://doi.org/10.11646/zootaxa.5777.1.5
